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<document id="CC1B0D4092786CCDF8D7B58D37C03738" ID-CLB-Dataset="21166" ID-DOI="10.5281/zenodo.3985687" ID-GBIF-Dataset="6921ca7f-2057-41e1-86ee-b9ca833e63aa" ID-ISSN="0253-116X" ID-Zenodo-Dep="3985687" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597414043187" checkinUser="valdenar" docAuthor="Kratochwil, Ockerell Anselm" docDate="2018" docId="03BF87E95779FFDA549172D8FD72FD9B" docLanguage="en" docName="Linzer.biol.Beitr.50.1.0337-0351.pdf.imf" docOrigin="Linzer biologische Beiträge 50 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Andrena wollastoni COCKERELL 1922" docType="treatment" docVersion="8" lastPageNumber="346" masterDocId="FF86FF91577EFFD354237370FFE0FFBE" masterDocTitle="Type specimens of Andrena wollastoni C, 1922 (Hymenoptera, Anthophila): deposition, evaluation and designation of a lectotype" masterLastPageNumber="351" masterPageNumber="337" pageNumber="344" updateTime="1698844652156" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3DC862A03011B9BFEBB14A56021148FD">Type specimens of Andrena wollastoni C, 1922 (Hymenoptera, Anthophila): deposition, evaluation and designation of a lectotype</mods:title>
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<mods:namePart id="0D06934BE70B56DB0EC28ADD493A30E5">Kratochwil, Ockerell Anselm</mods:namePart>
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<mods:date id="9622AE10066D3A4ECBF9B6C6AA1F3E87">2018</mods:date>
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<treatment id="03BF87E95779FFDA549172D8FD72FD9B" ID-DOI="http://doi.org/10.5281/zenodo.4324171" ID-GBIF-Taxon="167293433" ID-Zenodo-Dep="4324171" LSID="urn:lsid:plazi:treatment:03BF87E95779FFDA549172D8FD72FD9B" httpUri="http://treatment.plazi.org/id/03BF87E95779FFDA549172D8FD72FD9B" lastPageId="9" lastPageNumber="346" pageId="7" pageNumber="344">
<subSubSection id="C30C65745779FFD4549172D8FBD6FE00" box="[178,1078,424,446]" pageId="7" pageNumber="344" type="nomenclature">
<paragraph id="8BA936FF5779FFD4549172D8FBD6FE00" blockId="7.[178,1078,424,473]" box="[178,1078,424,446]" pageId="7" pageNumber="344">
<heading id="D0E181935779FFD4549172D8FBD6FE00" box="[178,1078,424,446]" centered="true" fontSize="8" level="2" pageId="7" pageNumber="344" reason="2">
<emphasis id="B962EAED5779FFD4549172D8FBD6FE00" bold="true" box="[178,1078,424,446]" pageId="7" pageNumber="344">
<emphasis id="B962EAED5779FFD4549172D8FE7CFE00" bold="true" box="[178,412,424,446]" italics="true" pageId="7" pageNumber="344">
2.
<taxonomicName id="4C164D7C5779FFD454ED72D8FE7CFE00" ID-CoL="DTBR" authorityName="COCKERELL" authorityYear="1922" box="[206,412,424,446]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="344" phylum="Arthropoda" rank="species" species="wollastoni">Andrena wollastoni</taxonomicName>
</emphasis>
specimens collected by T. V. Wollaston and deposited in the
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C65745779FFDA54F372B3FD72FD9B" lastPageId="9" lastPageNumber="346" pageId="7" pageNumber="344" type="description">
<paragraph id="8BA936FF5779FFD454F372B3FED4FE67" blockId="7.[178,1078,424,473]" box="[208,308,451,473]" pageId="7" pageNumber="344">
<heading id="D0E181935779FFD454F372B3FED4FE67" bold="true" box="[208,308,451,473]" fontSize="9" level="2" pageId="7" pageNumber="344" reason="3">
<emphasis id="B962EAED5779FFD454F372B3FED4FE67" bold="true" box="[208,308,451,473]" pageId="7" pageNumber="344">OUMNH</emphasis>
</heading>
</paragraph>
<paragraph id="8BA936FF5779FFD454917298FE9FFD75" blockId="7.[178,1085,488,715]" pageId="7" pageNumber="344">
One male and two females from
<collectingRegion id="49D2F81D5779FFD456247298FDBBFE40" box="[519,603,488,510]" country="Portugal" name="Madeira" pageId="7" pageNumber="344">Madeira</collectingRegion>
Archipelago, collected by Wollaston (
<figureCitation id="132D2A7A5779FFD457C97298FBD8FE43" box="[1002,1080,488,510]" captionStart="Fig" captionStartId="8.[178,211,906,928]" captionTargetBox="[178,1085,437,895]" captionTargetId="figure@8.[178,1085,435,895]" captionTargetPageId="8" captionText="Fig. 4: Andrena dourada KRATOCHWIL &amp; SCHEUCHL, 2013, male (OUMNH), collected by Wollaston from Porto Santo; (a) card-mounted specimen, (b) labels. Fig. 5: A. dourada, female (OUMNH), collected by Wollaston from Porto Santo; label identical to Fig. 4b. Fig. 6: A. wollastoni COCKERELL, 1922, female (OUMNH), collected by Wollaston from Madeira Island; (a) specimen, (b) labels. Fig. 7: Three labels below the specimens collected by Wollaston. All photos by J. Hogan (OUMNH)." figureDoi="http://doi.org/10.5281/zenodo.3985693" httpUri="https://zenodo.org/record/3985693/files/figure.png" pageId="7" pageNumber="344">Figs 4a</figureCitation>
, 5a, 6a), are deposited in the OUMNH (
<bibRefCitation id="EF874B0E5779FFD4566A7176FD15FDA2" author="SAUNDERS E" box="[585,757,518,540]" pageId="7" pageNumber="344" pagination="207 - 218" refId="ref7952" refString="SAUNDERS E. (1903): Hymenoptera Aculeata collected by Rev. Alfred Eaton M. A., in Madeira and Tenerife in the spring 1902, including notes on the species taken by the late T. Vernon Wollaston and F. A. Bellany. - Trans. Entomol. Soc. London 1903: 207 - 218." type="journal article" year="1903">SAUNDERS 1903</bibRefCitation>
).
<bibRefCitation id="EF874B0E5779FFD457247176FC5EFDA2" author="SAUNDERS E" box="[775,958,518,540]" pageId="7" pageNumber="344" pagination="207 - 218" refId="ref7952" refString="SAUNDERS E. (1903): Hymenoptera Aculeata collected by Rev. Alfred Eaton M. A., in Madeira and Tenerife in the spring 1902, including notes on the species taken by the late T. Vernon Wollaston and F. A. Bellany. - Trans. Entomol. Soc. London 1903: 207 - 218." type="journal article" year="1903">SAUNDERS (1903)</bibRefCitation>
states, ‘The specimens are, of course, old and more or less faded; the smaller ones are often gummed on cards, which renders determination doubtful, and as it would be dangerous to remove specimens of this age, I have not attempted to found any new species on them... In spite of the poor condition of Wollaston’s specimens, it is important to record, as far as possible, the forms which existed over half a century ago in an island so liable to accidential immigration as
<collectingRegion id="49D2F81D5779FFD4553871C5FE97FD75" box="[283,375,693,715]" country="Portugal" name="Madeira" pageId="7" pageNumber="344">Madeira</collectingRegion>
.’
</paragraph>
<paragraph id="8BA936FF5779FFD4549171ABFD7AFCD8" blockId="7.[178,1086,731,870]" pageId="7" pageNumber="344">
The specimens were determined as
<taxonomicName id="4C164D7C5779FFD4563A71ABFD6FFD4E" box="[537,655,731,752]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="344" phylum="Arthropoda" rank="species" species="minutula">
<emphasis id="B962EAED5779FFD4563A71ABFD6FFD4E" box="[537,655,731,752]" italics="true" pageId="7" pageNumber="344">A. minutula</emphasis>
</taxonomicName>
by Saunders (
<figureCitation id="132D2A7A5779FFD4573C71ABFC8EFD4F" box="[799,878,731,753]" captionStart="Fig" captionStartId="8.[178,211,906,928]" captionTargetBox="[178,1085,437,895]" captionTargetId="figure@8.[178,1085,435,895]" captionTargetPageId="8" captionText="Fig. 4: Andrena dourada KRATOCHWIL &amp; SCHEUCHL, 2013, male (OUMNH), collected by Wollaston from Porto Santo; (a) card-mounted specimen, (b) labels. Fig. 5: A. dourada, female (OUMNH), collected by Wollaston from Porto Santo; label identical to Fig. 4b. Fig. 6: A. wollastoni COCKERELL, 1922, female (OUMNH), collected by Wollaston from Madeira Island; (a) specimen, (b) labels. Fig. 7: Three labels below the specimens collected by Wollaston. All photos by J. Hogan (OUMNH)." figureDoi="http://doi.org/10.5281/zenodo.3985693" httpUri="https://zenodo.org/record/3985693/files/figure.png" pageId="7" pageNumber="344">Figs 4b</figureCitation>
, 6b). Not mentioned by
<bibRefCitation id="EF874B0E5779FFD454F07188FE69FCB0" author="SAUNDERS E" box="[211,393,760,782]" pageId="7" pageNumber="344" pagination="207 - 218" refId="ref7952" refString="SAUNDERS E. (1903): Hymenoptera Aculeata collected by Rev. Alfred Eaton M. A., in Madeira and Tenerife in the spring 1902, including notes on the species taken by the late T. Vernon Wollaston and F. A. Bellany. - Trans. Entomol. Soc. London 1903: 207 - 218." type="journal article" year="1903">SAUNDERS (1903)</bibRefCitation>
is the fact that one male (
<figureCitation id="132D2A7A5779FFD456B77188FCE7FCB0" box="[660,775,760,782]" captionStart="Fig" captionStartId="8.[178,211,906,928]" captionTargetBox="[178,1085,437,895]" captionTargetId="figure@8.[178,1085,435,895]" captionTargetPageId="8" captionText="Fig. 4: Andrena dourada KRATOCHWIL &amp; SCHEUCHL, 2013, male (OUMNH), collected by Wollaston from Porto Santo; (a) card-mounted specimen, (b) labels. Fig. 5: A. dourada, female (OUMNH), collected by Wollaston from Porto Santo; label identical to Fig. 4b. Fig. 6: A. wollastoni COCKERELL, 1922, female (OUMNH), collected by Wollaston from Madeira Island; (a) specimen, (b) labels. Fig. 7: Three labels below the specimens collected by Wollaston. All photos by J. Hogan (OUMNH)." figureDoi="http://doi.org/10.5281/zenodo.3985693" httpUri="https://zenodo.org/record/3985693/files/figure.png" pageId="7" pageNumber="344">Figs 4a, 4b</figureCitation>
) and one female (Fig. 5; label identical to
<figureCitation id="132D2A7A5779FFD455087065FE94FC95" box="[299,372,789,811]" captionStart="Fig" captionStartId="8.[178,211,906,928]" captionTargetBox="[178,1085,437,895]" captionTargetId="figure@8.[178,1085,435,895]" captionTargetPageId="8" captionText="Fig. 4: Andrena dourada KRATOCHWIL &amp; SCHEUCHL, 2013, male (OUMNH), collected by Wollaston from Porto Santo; (a) card-mounted specimen, (b) labels. Fig. 5: A. dourada, female (OUMNH), collected by Wollaston from Porto Santo; label identical to Fig. 4b. Fig. 6: A. wollastoni COCKERELL, 1922, female (OUMNH), collected by Wollaston from Madeira Island; (a) specimen, (b) labels. Fig. 7: Three labels below the specimens collected by Wollaston. All photos by J. Hogan (OUMNH)." figureDoi="http://doi.org/10.5281/zenodo.3985693" httpUri="https://zenodo.org/record/3985693/files/figure.png" pageId="7" pageNumber="344">Fig. 4b</figureCitation>
) were collected from
<collectingRegion id="49D2F81D5779FFD456767066FD6CFC95" box="[597,652,790,811]" country="Portugal" name="Porto" pageId="7" pageNumber="344">Porto</collectingRegion>
Santo; one other female (Figs 6a, 6b) was collected from
<collectingRegion id="49D2F81D5779FFD455717043FE46FCF7" box="[338,422,819,841]" country="Portugal" name="Madeira" pageId="7" pageNumber="344">Madeira</collectingRegion>
Island. Both
<collectingRegion id="49D2F81D5779FFD4561A7043FD90FCF6" box="[569,624,819,840]" country="Portugal" name="Porto" pageId="7" pageNumber="344">Porto</collectingRegion>
Santo specimens have to be assigned to
<taxonomicName id="4C164D7C5779FFD450047043FEE8FCDB" authorityName="Kratochwil &amp; Scheuchl" authorityYear="2013" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="344" phylum="Arthropoda" rank="species" species="dourada">
<emphasis id="B962EAED5779FFD450047043FEE8FCDB" italics="true" pageId="7" pageNumber="344">A. dourada</emphasis>
</taxonomicName>
(see KRATOCHWIL &amp; SCHEUCHL 2013).
</paragraph>
<paragraph id="8BA936FF5779FFD454917005FEB1FB85" blockId="7.[178,1085,885,1083]" pageId="7" pageNumber="344">
Cockerell had knowledge of the publication of
<bibRefCitation id="EF874B0E5779FFD456A47006FCDBFC32" author="SAUNDERS E" box="[647,827,886,908]" pageId="7" pageNumber="344" pagination="207 - 218" refId="ref7952" refString="SAUNDERS E. (1903): Hymenoptera Aculeata collected by Rev. Alfred Eaton M. A., in Madeira and Tenerife in the spring 1902, including notes on the species taken by the late T. Vernon Wollaston and F. A. Bellany. - Trans. Entomol. Soc. London 1903: 207 - 218." type="journal article" year="1903">SAUNDERS (1903)</bibRefCitation>
and he probably had seen the
<taxonomicName id="4C164D7C5779FFD454FA70E3FED0FC16" authorityName="Fabricius" authorityYear="1775" box="[217,304,915,936]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="B962EAED5779FFD454FA70E3FED0FC16" box="[217,304,915,936]" italics="true" pageId="7" pageNumber="344">Andrena</emphasis>
</taxonomicName>
specimens of the OUMNH collected by Wollaston because he described in 1921
<emphasis id="B962EAED5779FFD454D170C1FE2DFC7B" box="[242,461,944,966]" italics="true" pageId="7" pageNumber="344">Prosopis maderensis</emphasis>
<taxonomicNameLabel id="A25157965779FFD455F870C0FDEDFC7B" box="[475,525,944,965]" pageId="7" pageNumber="344" rank="species">sp.n.</taxonomicNameLabel>
, today
<emphasis id="B962EAED5779FFD4564670C0FD58FC7B" box="[613,696,944,965]" italics="true" pageId="7" pageNumber="344">Hylaeus</emphasis>
<taxonomicName id="4C164D7C5779FFD456E570C0FC01FC7B" authorityName="Cockerell" authorityYear="1922" box="[710,993,944,966]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="344" phylum="Arthropoda" rank="species" species="maderensis" subGenus="Paraprosopis">
(
<emphasis id="B962EAED5779FFD456ED70C1FCB8FC78" box="[718,856,945,966]" italics="true" pageId="7" pageNumber="344">Paraprosopis</emphasis>
)
<emphasis id="B962EAED5779FFD4574D70C0FC01FC7B" box="[878,993,944,965]" italics="true" pageId="7" pageNumber="344">maderensis</emphasis>
</taxonomicName>
(female specimen), located near the
<taxonomicName id="4C164D7C5779FFD455ED70BDFDC5FC5C" authorityName="Fabricius" authorityYear="1775" box="[462,549,973,994]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="B962EAED5779FFD455ED70BDFDC5FC5C" box="[462,549,973,994]" italics="true" pageId="7" pageNumber="344">Andrena</emphasis>
</taxonomicName>
specimens in the Wollaston collection (
<bibRefCitation id="EF874B0E5779FFD4579F70BEFF08FBBF" author="COCKERELL T" pageId="7" pageNumber="344" pagination="359 - 368" refId="ref7499" refString="COCKERELL T. D. A. (1921): Descriptions and records of bees. XCI. - Journal of Natural History XCI: 359 - 368." type="book chapter" year="1921">COCKERELL, 1921</bibRefCitation>
). This female was labelled by
<bibRefCitation id="EF874B0E5779FFD4561B709BFD13FBBF" author="SAUNDERS E" box="[568,755,1003,1025]" pageId="7" pageNumber="344" pagination="207 - 218" refId="ref7952" refString="SAUNDERS E. (1903): Hymenoptera Aculeata collected by Rev. Alfred Eaton M. A., in Madeira and Tenerife in the spring 1902, including notes on the species taken by the late T. Vernon Wollaston and F. A. Bellany. - Trans. Entomol. Soc. London 1903: 207 - 218." type="journal article" year="1903">SAUNDERS (1903)</bibRefCitation>
as ‘
<emphasis id="B962EAED5779FFD45704709BFC55FBBE" box="[807,949,1003,1024]" italics="true" pageId="7" pageNumber="344">Prosopis ater</emphasis>
’ (marked as ‘
<emphasis id="B962EAED5779FFD454987779FEF5FBA0" box="[187,277,1033,1054]" italics="true" pageId="7" pageNumber="344">Prosopis</emphasis>
n.sp. allied to
<emphasis id="B962EAED5779FFD4559B7779FDE3FBA0" box="[440,515,1033,1054]" italics="true" pageId="7" pageNumber="344">signata</emphasis>
’).
<emphasis id="B962EAED5779FFD456077778FD49FBA0" box="[548,681,1032,1054]" italics="true" pageId="7" pageNumber="344">Hylaeus ater</emphasis>
(
<bibRefCitation id="EF874B0E5779FFD456987778FC8CFBA0" author="SAUNDERS E" box="[699,876,1032,1054]" pageId="7" pageNumber="344" pagination="207 - 218" refId="ref7952" refString="SAUNDERS E. (1903): Hymenoptera Aculeata collected by Rev. Alfred Eaton M. A., in Madeira and Tenerife in the spring 1902, including notes on the species taken by the late T. Vernon Wollaston and F. A. Bellany. - Trans. Entomol. Soc. London 1903: 207 - 218." type="journal article" year="1903">SAUNDERS, 1903</bibRefCitation>
) is restricted to the
<collectingRegion id="49D2F81D5779FFD454917756FEACFB85" box="[178,332,1061,1083]" country="Spain" name="Canarias" pageId="7" pageNumber="344">Canary Islands</collectingRegion>
.
</paragraph>
<paragraph id="8BA936FF5779FFD45491773BFDBDFBC0" blockId="7.[178,1085,1099,1150]" pageId="7" pageNumber="344">
The male and two females of
<taxonomicName id="4C164D7C5779FFD455F8773BFDBFFBDE" authorityName="COCKERELL" authorityYear="1922" box="[475,607,1099,1120]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="344" phylum="Arthropoda" rank="species" species="wollastoni">
<emphasis id="B962EAED5779FFD455F8773BFDBFFBDE" box="[475,607,1099,1120]" italics="true" pageId="7" pageNumber="344">A. wollastoni</emphasis>
</taxonomicName>
are not mentioned in
<bibRefCitation id="EF874B0E5779FFD45719773BFC1CFBDF" author="COCKERELL T" box="[826,1020,1099,1121]" pageId="7" pageNumber="344" pagination="31 - 32" refId="ref7529" refString="COCKERELL T. D. A. (1922): New bees from Madeira islands. - Proc. Ent. Soc. Wash. 24 (1): 31 - 32." type="journal article" year="1922">COCKERELL (1922)</bibRefCitation>
therefore, they do not belong to the
<typeStatus id="54AD885D5779FFD455C97718FDF6FBC3" box="[490,534,1128,1149]" pageId="7" pageNumber="344">type</typeStatus>
series.
</paragraph>
<paragraph id="8BA936FF5779FFD4549177FEFCE5FAED" blockId="7.[178,1085,1165,1363]" pageId="7" pageNumber="344">
a) Male: The whole specimen (
<figureCitation id="132D2A7A5779FFD455CD77FDFDD7FB1A" box="[494,567,1165,1188]" captionStart="Fig" captionStartId="8.[178,211,906,928]" captionTargetBox="[178,1085,437,895]" captionTargetId="figure@8.[178,1085,435,895]" captionTargetPageId="8" captionText="Fig. 4: Andrena dourada KRATOCHWIL &amp; SCHEUCHL, 2013, male (OUMNH), collected by Wollaston from Porto Santo; (a) card-mounted specimen, (b) labels. Fig. 5: A. dourada, female (OUMNH), collected by Wollaston from Porto Santo; label identical to Fig. 4b. Fig. 6: A. wollastoni COCKERELL, 1922, female (OUMNH), collected by Wollaston from Madeira Island; (a) specimen, (b) labels. Fig. 7: Three labels below the specimens collected by Wollaston. All photos by J. Hogan (OUMNH)." figureDoi="http://doi.org/10.5281/zenodo.3985693" httpUri="https://zenodo.org/record/3985693/files/figure.png" pageId="7" pageNumber="344">Fig. 4a</figureCitation>
) is mounted on a squared cardboard (with the time yellowed), wings spread out, in good condition; the cardboard is colour-coded with blue indicating the female specimen comes from
<collectingRegion id="49D2F81D5779FFD456A477B8FD5EFB63" box="[647,702,1224,1245]" country="Portugal" name="Porto" pageId="7" pageNumber="344">Porto</collectingRegion>
Santo (
<bibRefCitation id="EF874B0E5779FFD4573177B8FC20FB60" author="MACHADO A" box="[786,960,1224,1246]" pageId="7" pageNumber="344" refId="ref7871" refString="MACHADO A. C. (2006): T. Vernon Wollaston (1822 - 1878): un entomologo en la Macaronesia. - Fundacion Cesar M anrique. 169 pp." type="book" year="2006">MACHADO 2006</bibRefCitation>
); this information was not recorded by
<bibRefCitation id="EF874B0E5779FFD455F77796FD6EFB42" author="SAUNDERS E" box="[468,654,1254,1276]" pageId="7" pageNumber="344" pagination="207 - 218" refId="ref7952" refString="SAUNDERS E. (1903): Hymenoptera Aculeata collected by Rev. Alfred Eaton M. A., in Madeira and Tenerife in the spring 1902, including notes on the species taken by the late T. Vernon Wollaston and F. A. Bellany. - Trans. Entomol. Soc. London 1903: 207 - 218." type="journal article" year="1903">SAUNDERS (1903)</bibRefCitation>
. There are no other labels from Wollaston with locality data or numbers. A later label (
<figureCitation id="132D2A7A5779FFD456537673FD59FAA7" box="[624,697,1283,1305]" captionStart="Fig" captionStartId="8.[178,211,906,928]" captionTargetBox="[178,1085,437,895]" captionTargetId="figure@8.[178,1085,435,895]" captionTargetPageId="8" captionText="Fig. 4: Andrena dourada KRATOCHWIL &amp; SCHEUCHL, 2013, male (OUMNH), collected by Wollaston from Porto Santo; (a) card-mounted specimen, (b) labels. Fig. 5: A. dourada, female (OUMNH), collected by Wollaston from Porto Santo; label identical to Fig. 4b. Fig. 6: A. wollastoni COCKERELL, 1922, female (OUMNH), collected by Wollaston from Madeira Island; (a) specimen, (b) labels. Fig. 7: Three labels below the specimens collected by Wollaston. All photos by J. Hogan (OUMNH)." figureDoi="http://doi.org/10.5281/zenodo.3985693" httpUri="https://zenodo.org/record/3985693/files/figure.png" pageId="7" pageNumber="344">Fig. 4b</figureCitation>
) was added to each specimen after the 1903 publication by Saunders, ‘
<taxonomicName id="4C164D7C5779FFD455DF7650FD55FA8B" box="[508,693,1312,1333]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="344" phylum="Arthropoda" rank="species" species="minutula">
<emphasis id="B962EAED5779FFD455DF7650FD55FA8B" box="[508,693,1312,1333]" italics="true" pageId="7" pageNumber="344">Andrena minutula</emphasis>
</taxonomicName>
Kirby’ handwritten, ‘Named 1903 by Ed. Saunders, Trans. Ent. Soc., 1903, p. 207-218.’ printed.
</paragraph>
<paragraph id="8BA936FF5779FFD454917613FBEAF997" blockId="7.[178,1085,1379,1577]" pageId="7" pageNumber="344">
b) Female: The whole specimen (Fig. 5) is mounted on a square cardboard (yellowed with the time), wings spread out, in good condition; the cardboard is colour-coded blue, indicating that the female specimen was collected from
<collectingRegion id="49D2F81D5779FFD456E876EEFCE2FA0D" box="[715,770,1438,1459]" country="Portugal" name="Porto" pageId="7" pageNumber="344">Porto</collectingRegion>
Santo (
<bibRefCitation id="EF874B0E5779FFD4577176EEFC1EFA0D" author="MACHADO A" box="[850,1022,1437,1459]" pageId="7" pageNumber="344" refId="ref7871" refString="MACHADO A. C. (2006): T. Vernon Wollaston (1822 - 1878): un entomologo en la Macaronesia. - Fundacion Cesar M anrique. 169 pp." type="book" year="2006">MACHADO 2006</bibRefCitation>
). This information was not recorded by
<bibRefCitation id="EF874B0E5779FFD4563C76CBFD3EFA6F" author="SAUNDERS E" box="[543,734,1467,1489]" pageId="7" pageNumber="344" pagination="207 - 218" refId="ref7952" refString="SAUNDERS E. (1903): Hymenoptera Aculeata collected by Rev. Alfred Eaton M. A., in Madeira and Tenerife in the spring 1902, including notes on the species taken by the late T. Vernon Wollaston and F. A. Bellany. - Trans. Entomol. Soc. London 1903: 207 - 218." type="journal article" year="1903">SAUNDERS (1903)</bibRefCitation>
. There are no other labels from Wollaston with locality data or numbers. A later label (identical to
<figureCitation id="132D2A7A5779FFD4574376A8FC49FA50" box="[864,937,1496,1518]" captionStart="Fig" captionStartId="8.[178,211,906,928]" captionTargetBox="[178,1085,437,895]" captionTargetId="figure@8.[178,1085,435,895]" captionTargetPageId="8" captionText="Fig. 4: Andrena dourada KRATOCHWIL &amp; SCHEUCHL, 2013, male (OUMNH), collected by Wollaston from Porto Santo; (a) card-mounted specimen, (b) labels. Fig. 5: A. dourada, female (OUMNH), collected by Wollaston from Porto Santo; label identical to Fig. 4b. Fig. 6: A. wollastoni COCKERELL, 1922, female (OUMNH), collected by Wollaston from Madeira Island; (a) specimen, (b) labels. Fig. 7: Three labels below the specimens collected by Wollaston. All photos by J. Hogan (OUMNH)." figureDoi="http://doi.org/10.5281/zenodo.3985693" httpUri="https://zenodo.org/record/3985693/files/figure.png" pageId="7" pageNumber="344">Fig. 4b</figureCitation>
) was added to each specimen after the 1903 publication by Saunders: ‘
<taxonomicName id="4C164D7C5779FFD456D47685FC50F9B4" box="[759,944,1525,1546]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="344" phylum="Arthropoda" rank="species" species="minutula">
<emphasis id="B962EAED5779FFD456D47685FC50F9B4" box="[759,944,1525,1546]" italics="true" pageId="7" pageNumber="344">Andrena minutula</emphasis>
</taxonomicName>
Kirby’ handwritten, ‘Named 1903 by Ed. Saunders, Trans. Ent. Soc., 1903, p. 207-218.’ printed.
</paragraph>
<paragraph id="8BA936FF5776FFDB549173ABFCBEFEF7" blockId="8.[178,1085,219,329]" pageId="8" pageNumber="345">
c) Female, pinned, in good condition (Fig. 6a);
<materialsCitation id="3B7E3CA25776FFDB56B773ABFCBAFEF7" ID-GBIF-Occurrence="2848388301" collectorName="Island. A &amp; Ed. Saunders" location="Madeira" pageId="8" pageNumber="345" specimenCount="1" stateProvince="Madeira">
the lacking coloured label indicates that it was collected from
<collectingRegion id="49D2F81D5776FFDB555C7388FE33FEB0" box="[383,467,248,270]" country="Portugal" name="Madeira" pageId="8" pageNumber="345">Madeira</collectingRegion>
<collectorName id="26E353295776FFDB55FF7388FDDBFEB3" box="[476,571,248,270]" pageId="8" pageNumber="345">Island. A</collectorName>
later label (Fig. 6b) was added to each specimen after the 1903 publication by Saunders, ‘
<taxonomicName id="4C164D7C5776FFDB567B7265FCF1FE94" box="[600,785,277,298]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="minutula">
<emphasis id="B962EAED5776FFDB567B7265FCF1FE94" box="[600,785,277,298]" italics="true" pageId="8" pageNumber="345">Andrena minutula</emphasis>
</taxonomicName>
Kirby’ handwritten, ‘Named 1903 by
<collectorName id="26E353295776FFDB55287243FE73FEF7" box="[267,403,307,329]" pageId="8" pageNumber="345">Ed. Saunders</collectorName>
, Trans. Ent. Soc., 1903, p. 207-218.’ printed
</materialsCitation>
.
</paragraph>
<paragraph id="8BA936FF5776FFDB54917228FE5CFE17" blockId="8.[178,1085,344,425]" pageId="8" pageNumber="345">
The two specimens from
<collectingRegion id="49D2F81D5776FFDB559B7228FE0FFED3" box="[440,495,344,365]" country="Portugal" name="Porto" pageId="8" pageNumber="345">Porto</collectingRegion>
Santo were also card-mounted in the same way as specimens of the NHMUK. It is remarkable that Saunders did not publish the locality where the specimens were collected.
</paragraph>
<caption id="DF6966775776FFDB549170FAFEFEFBA7" ID-DOI="http://doi.org/10.5281/zenodo.3985693" ID-Zenodo-Dep="3985693" httpUri="https://zenodo.org/record/3985693/files/figure.png" pageId="8" pageNumber="345" startId="8.[178,211,906,928]" targetBox="[178,1085,437,895]" targetPageId="8">
<paragraph id="8BA936FF5776FFDB549170FAFEFEFBA7" blockId="8.[178,1086,906,1049]" pageId="8" pageNumber="345">
<emphasis id="B962EAED5776FFDB549170FAFF10FC21" bold="true" box="[178,240,906,928]" pageId="8" pageNumber="345">Fig. 4</emphasis>
:
<taxonomicName id="4C164D7C5776FFDB552070FBFE4CFC1E" authorityName="Kratochwil &amp; Scheuchl" authorityYear="2013" box="[259,428,907,928]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="dourada">
<emphasis id="B962EAED5776FFDB552070FBFE4CFC1E" box="[259,428,907,928]" italics="true" pageId="8" pageNumber="345">Andrena dourada</emphasis>
</taxonomicName>
KRATOCHWIL &amp; SCHEUCHL, 2013, male (OUMNH), collected by Wollaston from Porto Santo; (
<emphasis id="B962EAED5776FFDB55F270D3FE3CFC06" bold="true" box="[465,476,931,952]" pageId="8" pageNumber="345">a</emphasis>
) card-mounted specimen, (
<emphasis id="B962EAED5776FFDB56FC70D3FD0CFC06" bold="true" box="[735,748,931,952]" pageId="8" pageNumber="345">b</emphasis>
) labels.
<emphasis id="B962EAED5776FFDB571C70D2FC99FC06" bold="true" box="[831,889,930,952]" pageId="8" pageNumber="345">Fig. 5</emphasis>
:
<taxonomicName id="4C164D7C5776FFDB57AB70D3FC11FC06" authorityName="Kratochwil &amp; Scheuchl" authorityYear="2013" box="[904,1009,931,952]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="dourada">
<emphasis id="B962EAED5776FFDB57AB70D3FC11FC06" box="[904,1009,931,952]" italics="true" pageId="8" pageNumber="345">A. dourada</emphasis>
</taxonomicName>
, female (OUMNH), collected by Wollaston from Porto Santo; label identical to Fig. 4b.
<emphasis id="B962EAED5776FFDB57A770CAFC5AFC71" bold="true" box="[900,954,954,976]" pageId="8" pageNumber="345">Fig. 6</emphasis>
:
<taxonomicName id="4C164D7C5776FFDB57E570CBFBDCFC6E" authorityName="COCKERELL" authorityYear="1922" box="[966,1084,955,976]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="wollastoni">
<emphasis id="B962EAED5776FFDB57E570CBFBDCFC6E" box="[966,1084,955,976]" italics="true" pageId="8" pageNumber="345">A. wollastoni</emphasis>
</taxonomicName>
<bibRefCitation id="EF874B0E5776FFDB549170A3FEBEFC56" author="COCKERELL T" box="[178,350,979,1000]" pageId="8" pageNumber="345" pagination="31 - 32" refId="ref7529" refString="COCKERELL T. D. A. (1922): New bees from Madeira islands. - Proc. Ent. Soc. Wash. 24 (1): 31 - 32." type="journal article" year="1922">COCKERELL, 1922</bibRefCitation>
, female (OUMNH), collected by Wollaston from Madeira Island; (
<emphasis id="B962EAED5776FFDB57EB70A3FC33FC56" bold="true" box="[968,979,979,1000]" pageId="8" pageNumber="345">a</emphasis>
) specimen, (
<emphasis id="B962EAED5776FFDB5499709BFF27FBBE" bold="true" box="[186,199,1003,1024]" pageId="8" pageNumber="345">b</emphasis>
) labels.
<emphasis id="B962EAED5776FFDB5537709AFEAAFBBE" bold="true" box="[276,330,1002,1024]" pageId="8" pageNumber="345">Fig. 7</emphasis>
: Three labels below the specimens collected by Wollaston. All photos by J. Hogan (OUMNH).
</paragraph>
</caption>
<paragraph id="8BA936FF5776FFDB5491775FFC48FAAC" blockId="8.[178,1085,1071,1298]" pageId="8" pageNumber="345">
According to our taxonomical study the former
<taxonomicName id="4C164D7C5776FFDB56B7775FFCF9FBFA" authorityName="COCKERELL" authorityYear="1922" box="[660,793,1071,1092]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="wollastoni">
<emphasis id="B962EAED5776FFDB56B7775FFCF9FBFA" box="[660,793,1071,1092]" italics="true" pageId="8" pageNumber="345">A. wollastoni</emphasis>
</taxonomicName>
specimens from
<collectingRegion id="49D2F81D5776FFDB57E5775FFC1DFBFA" box="[966,1021,1071,1092]" country="Portugal" name="Porto" pageId="8" pageNumber="345">Porto</collectingRegion>
Santo belong to a species of its own:
<taxonomicName id="4C164D7C5776FFDB55C2773DFDB2FBDF" authorityName="Kratochwil &amp; Scheuchl" authorityYear="2013" box="[481,594,1100,1122]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="dourada">
<emphasis id="B962EAED5776FFDB55C2773DFDB2FBDF" box="[481,594,1100,1122]" italics="true" pageId="8" pageNumber="345">A. dourada</emphasis>
</taxonomicName>
KRATOCHWIL &amp; SCHEUCHL, 2013. The OUMNH specimens can be depicted as a male and a female of
<taxonomicName id="4C164D7C5776FFDB56EA771AFCDBFBC1" authorityName="Kratochwil &amp; Scheuchl" authorityYear="2013" box="[713,827,1130,1151]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="dourada">
<emphasis id="B962EAED5776FFDB56EA771AFCDBFBC1" box="[713,827,1130,1151]" italics="true" pageId="8" pageNumber="345">A. dourada</emphasis>
</taxonomicName>
. In the paraocular area of
<taxonomicName id="4C164D7C5776FFDB549177F7FEC9FB22" authorityName="Kratochwil &amp; Scheuchl" authorityYear="2013" box="[178,297,1159,1180]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="dourada">
<emphasis id="B962EAED5776FFDB549177F7FEC9FB22" box="[178,297,1159,1180]" italics="true" pageId="8" pageNumber="345">A. dourada</emphasis>
</taxonomicName>
there are yellowish-white hairs from the malar area and overtop and no brownish hairs, in contrast to
<taxonomicName id="4C164D7C5776FFDB55DA77D5FD63FB07" authorityName="COCKERELL" authorityYear="1922" box="[505,643,1188,1210]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="wollastoni">
<emphasis id="B962EAED5776FFDB55DA77D5FD63FB07" box="[505,643,1188,1210]" italics="true" pageId="8" pageNumber="345">A. wollastoni</emphasis>
</taxonomicName>
with brownish hairs. The tibial scopa is characterised by yellowish-white hairs, but in the case of
<taxonomicName id="4C164D7C5776FFDB56D777B2FC99FB69" authorityName="COCKERELL" authorityYear="1922" box="[756,889,1218,1239]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="wollastoni">
<emphasis id="B962EAED5776FFDB56D777B2FC99FB69" box="[756,889,1218,1239]" italics="true" pageId="8" pageNumber="345">A. wollastoni</emphasis>
</taxonomicName>
, more than the half of the scopa and the basal area bear brownish hairs (KRATOCHWIL &amp; SCHEUCHL 2013). An analysis of the scopa by macro photos proved the affiliation to
<taxonomicName id="4C164D7C5776FFDB5713778DFC42FAAF" authorityName="Kratochwil &amp; Scheuchl" authorityYear="2013" box="[816,930,1276,1298]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="dourada">
<emphasis id="B962EAED5776FFDB5713778DFC42FAAF" box="[816,930,1276,1298]" italics="true" pageId="8" pageNumber="345">A. dourada</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA936FF5776FFDB54917652FC16FA74" blockId="8.[178,1085,1314,1482]" pageId="8" pageNumber="345">
The specimens are in good condition; this is in contradiction to the description given by
<bibRefCitation id="EF874B0E5776FFDB5491764FFE94FAEB" author="SAUNDERS E" box="[178,372,1343,1365]" pageId="8" pageNumber="345" pagination="207 - 218" refId="ref7952" refString="SAUNDERS E. (1903): Hymenoptera Aculeata collected by Rev. Alfred Eaton M. A., in Madeira and Tenerife in the spring 1902, including notes on the species taken by the late T. Vernon Wollaston and F. A. Bellany. - Trans. Entomol. Soc. London 1903: 207 - 218." type="journal article" year="1903">SAUNDERS (1903)</bibRefCitation>
, also cited by
<bibRefCitation id="EF874B0E5776FFDB5609764FFD09FAEB" author="MACHADO A" box="[554,745,1343,1365]" pageId="8" pageNumber="345" refId="ref7871" refString="MACHADO A. C. (2006): T. Vernon Wollaston (1822 - 1878): un entomologo en la Macaronesia. - Fundacion Cesar M anrique. 169 pp." type="book" year="2006">MACHADO (2006)</bibRefCitation>
(‘poor condition’). With high probability, the specimens were collected during the second visit of the
<collectingRegion id="49D2F81D5776FFDB57A1762CFC36FACC" box="[898,982,1372,1394]" country="Portugal" name="Madeira" pageId="8" pageNumber="345">Madeira</collectingRegion>
Archipelago from
<date id="FFA8103F5776FFDB553D760AFE27FA31" box="[286,455,1402,1424]" pageId="8" pageNumber="345" value="1848-11">November 1848</date>
to
<date id="FFA8103F5776FFDB55D2760AFDBDFA31" box="[497,605,1402,1423]" pageId="8" pageNumber="345" value="1949-06">June 1949</date>
(
<collectingRegion id="49D2F81D5776FFDB564C760AFD28FA2E" box="[623,712,1402,1424]" country="Portugal" name="Madeira" pageId="8" pageNumber="345">Madeira</collectingRegion>
,
<collectingRegion id="49D2F81D5776FFDB56F4760AFCEEFA31" box="[727,782,1402,1423]" country="Portugal" name="Porto" pageId="8" pageNumber="345">Porto</collectingRegion>
Santo, Desertas). Wollaston visited in
<date id="FFA8103F5776FFDB553A76E7FE5DFA12" box="[281,445,1431,1453]" pageId="8" pageNumber="345" value="1848-12">December 1848</date>
and in
<date id="FFA8103F5776FFDB562E76E7FD9EFA12" box="[525,638,1431,1453]" pageId="8" pageNumber="345" value="1849-04">April 1849</date>
<collectingRegion id="49D2F81D5776FFDB56A676E7FD5CFA12" box="[645,700,1431,1452]" country="Portugal" name="Porto" pageId="8" pageNumber="345">Porto</collectingRegion>
Santo (together, more than a month) (
<bibRefCitation id="EF874B0E5776FFDB549A76C4FE84FA74" author="MACHADO A" box="[185,356,1460,1482]" pageId="8" pageNumber="345" refId="ref7871" refString="MACHADO A. C. (2006): T. Vernon Wollaston (1822 - 1878): un entomologo en la Macaronesia. - Fundacion Cesar M anrique. 169 pp." type="book" year="2006">MACHADO 2006</bibRefCitation>
). These are probably the first collected specimens of
<taxonomicName id="4C164D7C5776FFDB575D76C5FC11FA77" authorityName="Kratochwil &amp; Scheuchl" authorityYear="2013" box="[894,1009,1460,1482]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="345" phylum="Arthropoda" rank="species" species="dourada">
<emphasis id="B962EAED5776FFDB575D76C5FC11FA77" box="[894,1009,1460,1482]" italics="true" pageId="8" pageNumber="345">A. dourada</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA936FF5776FFDB549176AAFBD3FA51" blockId="8.[178,1075,1498,1520]" box="[178,1075,1498,1520]" pageId="8" pageNumber="345">Below the Wollaston specimens, three labels were mounted with the following contents:</paragraph>
<paragraph id="8BA936FF5776FFDA5491768FFDF7FEDB" blockId="8.[178,1085,1535,1586]" lastBlockId="9.[178,1085,219,358]" lastPageId="9" lastPageNumber="346" pageId="8" pageNumber="345">
a) Blue handwritten label ‘new sp.?’. It was written by Frederick Smith (
<date id="FFA8103F5776FFDB57B9768FFBECF9AB" box="[922,1036,1535,1557]" pageId="8" pageNumber="345" value="1805" valueMax="1879">1805-1879</date>
), the first hymenopterist who examined the collection in the OUMNH and worked on the specimens (Fig. 7). In 1849, he was first assistant, later senior assistent at the British Museum of Natural History. During the 28 years of his period of service, he described 25 genera and 702 species and subspecies (
<bibRefCitation id="EF874B0E5777FFDA56437266FD03FE95" author="HARIS A" box="[608,739,277,299]" pageId="9" pageNumber="346" pagination="1 - 246" refId="ref7644" refString="HARIS A. (2016): Hymenoptera research in the Carpathian Basin (Hymenoptera: Aculeata. - Natura Somogyiensis 29: 1 - 246." type="journal article" year="2016">HARIS 2016</bibRefCitation>
). More than 40 years before the description of Cockerell, Smith realised the existence of a new species, but he did not carry out a description the species status.
</paragraph>
<paragraph id="8BA936FF5777FFDA54917205FDE3FE78" blockId="9.[178,1085,373,454]" pageId="9" pageNumber="346">
b) Label ‘
<emphasis id="B962EAED5777FFDA553B7205FE92FE34" box="[280,370,373,394]" italics="true" pageId="9" pageNumber="346">minutula</emphasis>
K.’, handwritten with ink on paper similar to the square cardboard of the specimens (yellowed with the time) (Fig. 7). A graphological analysis suggested that it is the handwriting of Saunders.
</paragraph>
<paragraph id="8BA936FF5777FFDA549172A6FD72FD9B" blockId="9.[178,1085,469,550]" pageId="9" pageNumber="346">
c) Printed label ‘
<taxonomicName id="4C164D7C5777FFDA554372A5FDB5FE55" authority="" box="[352,597,469,491]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="346" phylum="Arthropoda" rank="subGenus" subGenus="Micrandrena">
<emphasis id="B962EAED5777FFDA554372A5FE57FE54" box="[352,439,469,490]" italics="true" pageId="9" pageNumber="346">Andrena</emphasis>
(
<emphasis id="B962EAED5777FFDA55E472A5FDACFE54" box="[455,588,469,490]" italics="true" pageId="9" pageNumber="346">Micrandrena</emphasis>
)
</taxonomicName>
sp. Not conspecific with
<emphasis id="B962EAED5777FFDA574372A6FC8FFE55" box="[864,879,470,491]" italics="true" pageId="9" pageNumber="346">A</emphasis>
.
<taxonomicName id="4C164D7C5777FFDA575D72A6FBECFE54" box="[894,1036,469,491]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="346" phylum="Arthropoda" rank="species" species="minutula" subGenus="Micrandrena">
(
<emphasis id="B962EAED5777FFDA57A572A6FC7BFE55" box="[902,923,470,491]" italics="true" pageId="9" pageNumber="346">M</emphasis>
.)
<emphasis id="B962EAED5777FFDA579172A5FBECFE54" box="[946,1036,469,490]" italics="true" pageId="9" pageNumber="346">minutula</emphasis>
</taxonomicName>
(K.) both sexes differing in all diagnostic features of surface sculpturing,
<specimenCount id="9D10FD765777FFDA57857283FC59FDB6" box="[934,953,499,520]" pageId="9" pageNumber="346" type="male">♂</specimenCount>
genitalia &amp; accessory sclerites missing. C. O’Toole, 2003’.
</paragraph>
</subSubSection>
</treatment>
</document>
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