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<document id="5D11200A629C9DB7025BF4BFBEDA5969" ID-CLB-Dataset="8501" ID-DOI="10.11646/zootaxa.4920.1.2" ID-GBIF-Dataset="16b7d8d9-c520-44b2-a097-31067e1a9869" ID-ISSN="1175-5326" ID-Zenodo-Dep="4471588" ID-ZooBank="23D36297-E800-4D57-80AF-F80B8399E621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611747055971" checkinUser="plazi" docAuthor="Okayasu, Juriya, Lelej, Arkady S. &amp; Williams, Kevin A." docDate="2021" docId="03BF4A48656DFF9DFF1EFE18FD9EFD1D" docLanguage="en" docName="zootaxa.4920.1.2.pdf" docOrigin="Zootaxa 4920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eotrogaspidia adhabar Okayasu, Lelej &amp; Williams 2021, sp. nov." docType="treatment" docUuid="A5D10C25-921E-498C-9136-1E6F06468504" docUuidSource="ZooBank" docVersion="7" lastPageNumber="62" masterDocId="FF863230656EFF9BFF89FF9FFFE9FFEC" masterDocTitle="Review of Eotrogaspidia Lelej (Hymenoptera: Mutillidae: Trogaspidiini)" masterLastPageNumber="90" masterPageNumber="56" pageNumber="59" updateTime="1698925657998" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="235E223441674A275478DD2CCB65D3A9">Review of Eotrogaspidia Lelej (Hymenoptera: Mutillidae: Trogaspidiini)</mods:title>
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<mods:namePart id="5F4D5E1C288654AB8573E90F06E025B9">Okayasu, Juriya</mods:namePart>
<mods:affiliation id="1D41AFCCB0B316AA4C2994F942EF79CD">Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566 Japan.</mods:affiliation>
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<mods:namePart id="407AB9A9CB8A6F78F07D9C67897AA46E">Lelej, Arkady S.</mods:namePart>
<mods:nameIdentifier id="F5A75DB36DF9F374B25A29CA55EC71FD" type="ORCID">0000-0001-7501-0981</mods:nameIdentifier>
<mods:affiliation id="EC8BF7B257CE749EA45929EB7F76F060">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok, Russia. lelej @ biosoil. ru; https: // orcid. org / 0000 - 0001 - 7501 - 0981</mods:affiliation>
<mods:nameIdentifier id="78670C7A62CADB6C2A7F2F4F8BE5CAD4" type="email">lelej@biosoil.ru</mods:nameIdentifier>
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<mods:namePart id="A0985003AF82F4D9EFFB624347258541">Williams, Kevin A.</mods:namePart>
<mods:nameIdentifier id="62D1BE800136668D01342CB8229D0E56" type="ORCID">0000-0002-6073-1070</mods:nameIdentifier>
<mods:affiliation id="0040B999A8310120E7C9A834A30DC738">Plant Pest Diagnostics Center, California Department of Food &amp; Agriculture, 3294 Meadowview Road, Sacramento, CA 95832, USA. Kevin. Williams @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 6073 - 1070</mods:affiliation>
<mods:nameIdentifier id="A627CB6844A44022C4A0F40C3A60BB62" type="email">illiams@cdfa.ca.gov</mods:nameIdentifier>
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<treatment id="03BF4A48656DFF9DFF1EFE18FD9EFD1D" ID-DOI="http://doi.org/10.5281/zenodo.4471594" ID-GBIF-Taxon="177207642" ID-Zenodo-Dep="4471594" LSID="urn:lsid:zoobank.org:act:A5D10C25-921E-498C-9136-1E6F06468504" httpUri="http://treatment.plazi.org/id/03BF4A48656DFF9DFF1EFE18FD9EFD1D" lastPageId="6" lastPageNumber="62" pageId="3" pageNumber="59">
<subSubSection id="C30CA8D5656DFF98FF1EFE18FC9DFE4E" box="[151,884,391,418]" pageId="3" pageNumber="59" type="nomenclature">
<paragraph id="8BA9FB5E656DFF98FF1EFE18FC9DFE4E" blockId="3.[151,884,391,455]" box="[151,884,391,418]" pageId="3" pageNumber="59">
<heading id="D0E14C32656DFF98FF1EFE18FC9DFE4E" bold="true" box="[151,884,391,418]" fontSize="11" level="1" pageId="3" pageNumber="59" reason="1">
<emphasis id="B962274C656DFF98FF1EFE18FC9DFE4E" bold="true" box="[151,884,391,418]" pageId="3" pageNumber="59">
<taxonomicName id="4C1680DD656DFF98FF1EFE18FCE3FE4E" authority="Okayasu, Lelej &amp; Williams" authorityName="Okayasu, Lelej &amp; Williams" authorityYear="2021" box="[151,778,391,418]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="adhabar" status="sp. nov.">
<emphasis id="B962274C656DFF98FF1EFE18FE44FE4E" bold="true" box="[151,429,391,418]" italics="true" pageId="3" pageNumber="59">Eotrogaspidia adhabar</emphasis>
Okayasu, Lelej &amp; Williams
</taxonomicName>
,
<taxonomicNameLabel id="A2519A37656DFF98FC9EFE17FC9DFE4E" box="[791,884,392,418]" pageId="3" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C30CA8D5656DFF98FF1EFE33FDB2FE2A" box="[151,603,428,455]" pageId="3" pageNumber="59" type="description">
<paragraph id="8BA9FB5E656DFF98FF1EFE33FDB2FE2A" blockId="3.[151,884,391,455]" box="[151,603,428,455]" pageId="3" pageNumber="59">
(
<figureCitation id="132DE7DB656DFF98FF16FE33FEC0FE2A" box="[159,297,428,454]" captionStart="FIGURE 1" captionStartId="4.[151,250,1972,1996]" captionTargetBox="[151,1435,561,1949]" captionTargetId="figure-192@4.[151,1435,561,1949]" captionTargetPageId="4" captionText="FIGURE 1. Eotrogaspidia spp., ♂, habitus. A, E. E. adhabar sp. nov., holotype; B, F. E. amans (André); C, G. E. auroguttata (Smith); D, H. E. buddha sp. nov., paratype; AD. dorsal view; EH. lateral view." figureDoi="http://doi.org/10.5281/zenodo.4471592" httpUri="https://zenodo.org/record/4471592/files/figure.png" pageId="3" pageNumber="59">Figs 1A, 1E</figureCitation>
,
<figureCitation id="132DE7DB656DFF98FEBCFE33FEB3FE2B" box="[309,346,428,455]" captionStart="FIGURE 5" captionStartId="11.[151,250,1498,1522]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-17@11.[151,1435,181,1473]" captionTargetPageId="11" captionText="FIGURE 5. Eotrogaspidia spp., ♂, face view. A. E. adhabar sp. nov., holotype; B. E. amans (André); C. E. auroguttata (Smith); D. E. buddha sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.4471602" httpUri="https://zenodo.org/record/4471602/files/figure.png" pageId="3" pageNumber="59">5A</figureCitation>
,
<figureCitation id="132DE7DB656DFF98FEECFE33FE63FE2A" box="[357,394,428,454]" captionStart="FIGURE 7" captionStartId="16.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1711]" captionTargetId="figure-17@16.[151,1435,181,1714]" captionTargetPageId="16" captionText="FIGURE 7. Eotrogaspidia and Trogaspidia spp., ♂, metacoxa. A. E. adhabar sp. nov., paratype; B. E. amans (André); C. E. auroguttata (Smith); D. E. buddha sp. nov., paratype; E. E. hauseri sp. nov., paratype; F. E. oryzae (Pagden); G. E. saussurei (Lelej); H. T. mackieae (Cockerell)." figureDoi="http://doi.org/10.5281/zenodo.4471608" httpUri="https://zenodo.org/record/4471608/files/figure.png" pageId="3" pageNumber="59">7A</figureCitation>
,
<figureCitation id="132DE7DB656DFF98FE1FFE33FE4BFE2A" box="[406,418,428,454]" captionStart="FIGURE 8" captionStartId="17.[151,250,1956,1980]" captionTargetBox="[151,1435,670,1931]" captionTargetId="figure-231@17.[151,1435,669,1933]" captionTargetPageId="17" captionText="FIGURE 8. Eotrogaspidia spp., ♂, genitalia.AB. E. adhabar sp. nov., holotype; CD. E. amans (André); EF. E. auroguttata (Smith); GH. E. buddha sp. nov., paratype; A, C, E, G. dorsal view; B, D, F, H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.4471610" httpUri="https://zenodo.org/record/4471610/files/figure.png" pageId="3" pageNumber="59">8</figureCitation>
AB, 10A, 10E)
</paragraph>
</subSubSection>
<subSubSection id="C30CA8D5656DFF98FF1EFE6BFAA3FDDE" pageId="3" pageNumber="59" type="diagnosis">
<paragraph id="8BA9FB5E656DFF98FF1EFE6BFAA3FDDE" blockId="3.[151,1437,500,2039]" pageId="3" pageNumber="59">
<emphasis id="B962274C656DFF98FF1EFE6BFEFBFDE2" bold="true" box="[151,274,500,526]" pageId="3" pageNumber="59">Diagnosis.</emphasis>
MALE. Clypeus medially gibbous in pentagonal shape (
<figureCitation id="132DE7DB656DFF98FC16FE6BFC15FDE3" box="[927,1020,500,527]" captionStart="FIGURE 5" captionStartId="11.[151,250,1498,1522]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-17@11.[151,1435,181,1473]" captionTargetPageId="11" captionText="FIGURE 5. Eotrogaspidia spp., ♂, face view. A. E. adhabar sp. nov., holotype; B. E. amans (André); C. E. auroguttata (Smith); D. E. buddha sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.4471602" httpUri="https://zenodo.org/record/4471602/files/figure.png" pageId="3" pageNumber="59">Fig. 5A</figureCitation>
); scape with one strong longitudinal carina ventrally (
<figureCitation id="132DE7DB656DFF98FEDCFD87FE59FDDF" box="[341,432,536,563]" captionStart="FIGURE 5" captionStartId="11.[151,250,1498,1522]" captionTargetBox="[151,1435,181,1473]" captionTargetId="figure-17@11.[151,1435,181,1473]" captionTargetPageId="11" captionText="FIGURE 5. Eotrogaspidia spp., ♂, face view. A. E. adhabar sp. nov., holotype; B. E. amans (André); C. E. auroguttata (Smith); D. E. buddha sp. nov., paratype." figureDoi="http://doi.org/10.5281/zenodo.4471602" httpUri="https://zenodo.org/record/4471602/files/figure.png" pageId="3" pageNumber="59">Fig. 5A</figureCitation>
). Metacoxa evenly convex with long sparse setae (
<figureCitation id="132DE7DB656DFF98FC61FD87FBABFDDE" box="[1000,1090,536,562]" captionStart="FIGURE 7" captionStartId="16.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1711]" captionTargetId="figure-17@16.[151,1435,181,1714]" captionTargetPageId="16" captionText="FIGURE 7. Eotrogaspidia and Trogaspidia spp., ♂, metacoxa. A. E. adhabar sp. nov., paratype; B. E. amans (André); C. E. auroguttata (Smith); D. E. buddha sp. nov., paratype; E. E. hauseri sp. nov., paratype; F. E. oryzae (Pagden); G. E. saussurei (Lelej); H. T. mackieae (Cockerell)." figureDoi="http://doi.org/10.5281/zenodo.4471608" httpUri="https://zenodo.org/record/4471608/files/figure.png" pageId="3" pageNumber="59">Fig. 7A</figureCitation>
). FEMALE. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30CA8D5656DFF9DFF4EFDA3FE62FE19" lastPageId="6" lastPageNumber="62" pageId="3" pageNumber="59" type="description">
<paragraph id="8BA9FB5E656DFF98FF4EFDA3FC10FDBB" blockId="3.[151,1437,500,2039]" box="[199,1017,572,599]" pageId="3" pageNumber="59">
<emphasis id="B962274C656DFF98FF4EFDA3FEB0FDBA" bold="true" box="[199,345,572,598]" pageId="3" pageNumber="59">Description.</emphasis>
MALE. Body length: 13.0 mm. Forewing length:
<quantity id="4CEE56BB656DFF98FC07FDA3FC1CFDBB" box="[910,1013,572,599]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="3" pageNumber="59" unit="mm" value="10.9">10.9 mm</quantity>
.
</paragraph>
<paragraph id="8BA9FB5E656DFF98FF4EFDFFFB3EFAEE" blockId="3.[151,1437,500,2039]" pageId="3" pageNumber="59">
<emphasis id="B962274C656DFF98FF4EFDFFFE9BFD96" box="[199,370,608,634]" italics="true" pageId="3" pageNumber="59">Color and setae</emphasis>
. Black with mandible medially, premetum, stipes, palpi, scape except apex and pedicel dark brown, apex od scape yellow, tegula brownish black, protibial spur pale brown, meso- and metatibial spurs yellowish, T14 and S14 yellowish red except medial carina on S1 black, T57 and S58 brownish black. Frons, vertex and gena entirely covered with sparse long erect silvery setae, dense on antennal scrobe, malar space and clypeus laterally; frons with dense long appressed silvery setae; vertex behind ocellar region with dense short appressed silvery setae; anterior margin of clypeus fringed with long erect golden setae; prementum, stipes and palpi with sparse very short erect pale golden setae; scape with sparse long recumbent and sparse long erect silvery setae; pedicel and F1 with dense short sub-appressed silvery setae; F211 with dense very short erect pale golden setae. Mesosoma mostly with sparse erect silvery setae; pronotal collar posteriorly glabrous; dorsum of pronotum, mesopleuron and dorsum of propodeum between sublateral carinae with dense long appressed silvery setae; mesoscutum with sparse short recumbent and sparser long erect black setae; tegula with sparse short recumbent black setae; mesoscutellum with sparse long erect black setae on anterior half, with sparse long erect yellowish setae on posterior half; axilla and metanotal trough with dense short recumbent silvery setae; lateral face of pronotum, metapleuron and propodeum except dorsum with sparse short recumbent silvery setae. Wings pale brown, veins brown, with short erect black, regularly distributed setae. Legs entirely with sparse long erect silvery setae; procoxa with dense short appressed silvery setae; meso- and metatrochanters, femora and tibiae with sparse short appressed silvery setae; tarsi with dense long recumbent silvery setae, ventrally more or less yellowish. T1, T25 laterally and S18 with sparse long erect white setae; T25 medially with sparse long erect golden setae; T67 with sparse long recumbent black and sparser long erect white setae; T14 with posterior fringe of sparse long golden setae; S24 with posterior fringe of sparse long white setae; S58 with posterior fringe of sparse long pale golden setae; T2 with lateral golden felt line.
</paragraph>
<paragraph id="8BA9FB5E656DFF98FF4EFA93FDA0F8F3" blockId="3.[151,1437,500,2039]" pageId="3" pageNumber="59">
<emphasis id="B962274C656DFF98FF4EFA93FEC1FACA" box="[199,296,1292,1318]" italics="true" pageId="3" pageNumber="59">Structure</emphasis>
. Head 1.79 × wider than long; lateral margins strongly convergent behind eye; vertex rounded posteriorly; POD/OOD = 0.68; OD:MOD:LOD = 1.00:1.38:1.00; ocellar region slightly convex, each ocellus without posterior carina; antennal tubercle and antennal scrobe with weak continuous dorsal carina not reaching inner eye margin; genal carina absent; frons and gena with minute sparse and large dense punctures; ocellar region and vertex laterally to it with large sparse punctures; vertex posteriorly to ocelli with minute dense and large sparse punctures; post genal bridge without medial projection; hypostomal carina weakly developed. Clypeus medially gibbous in pentagonal shape, laterally flattened; anterior margin medially widely protruding anteriorly; medial protruding part depressed ventrally, laterally not defined by carina; surface of clypeus impunctate, polished, except anterior margin and posterior margin of medial convexity with minute dense punctures. Mandible slender, narrowed to apex, apically bidentate, without subbasal denticle; inner denticle smaller than apex; ventral margin excised into large basal tooth; outer face of mandible with minute dense punctures. Scape bending medially, with one strong carina beneath not widened apically; surface of scape with small dense punctures dorsally, impunctate and polished ventrally; surface of pedicel with minute sparse punctures; F1 compressed; ration between length and width of pedicel, F1 and F2 = 1.00:1.18:3.53:1.59:2.29:1.47; F211 almost equal in width; F11 as long as F10, conical; surface of flagellum with minute dense punctures. Prementum flattened, micro-reticulate, with posterior impunctate convex line; stipes with minute sparse punctures.
</paragraph>
<paragraph id="8BA9FB5E656DFF9FFF4EF8B6FBDFFDF5" blockId="3.[151,1437,500,2039]" lastBlockId="4.[151,1437,151,537]" lastPageId="4" lastPageNumber="60" pageId="3" pageNumber="59">Mesosoma broadest at mesonotum including tegulae; ratio between head width, humeral width, pronotal width, mesonotal width = 88:75:84:100; humeral angle rounded, with weak humeral carina ventrally; propodeum abruptly convergent posteriorly; mesoscutal width 1.17 × its length, 0.70 × mesonotal width; tegula short rounded, posteriorly reaching mesoscuto-scutellar articulation, 1.38 × longer than wide; mesoscutum distinctly convex; notaulus evenly wide, reaching anterior 3/8 of mesoscutum, subparallel; parapsidal line length 0.2 × mesoscutal length; parascutal carina forming wide, apically rounded, weakly elevated process on postero-lateral angle of mesoscutum; mesoscutellum gibbous with medial longitudinal carina posteriorly elevated; mesopleuron laterally with distinct transverse mesopleural suture, ventrally with precoxal swelling; precoxal ridge absent; pronotal collar anteriorly with minute dense punctures, posteriorly with small sparse punctures; dorsum of pronotum with minute dense and large dense punctures; mesoscutum and mesoscutellum with large confluent punctures; tegula impunctate polished except anterior and inner margins with small sparse punctures; axilla with minute sparse punctures; metascutellum medially with minute sparse punctures, laterally with small confluent punctures; metanotal trough striate; posterior propodeal face reticulate, with sublateral carina, antero-medially with long triangular cell posteriorly forming small process; dorsal and lateral propodeal faces not separated by carina; lateral face of pronotum transversely striate; anterior part of lateral face of mesopleuron, dorsal half of metapleuron and ventral half of lateral propodeal face with minute dense punctures; medial part of lateral face of mesopleuron and ventral face of mesopleuron with large confluent punctures; posterior part of lateral face of mesopleuron and ventral half of metapleuron with minute dense and small sparse punctures; dorsal half of lateral propodeal face reticulate.</paragraph>
<caption id="DF69ABD6656AFF9FFF1EF82BFC31F81C" ID-DOI="http://doi.org/10.5281/zenodo.4471592" ID-Zenodo-Dep="4471592" httpUri="https://zenodo.org/record/4471592/files/figure.png" pageId="4" pageNumber="60" startId="4.[151,250,1972,1996]" targetBox="[151,1435,561,1949]" targetPageId="4">
<paragraph id="8BA9FB5E656AFF9FFF1EF82BFC31F81C" blockId="4.[151,1437,1972,2032]" pageId="4" pageNumber="60">
<emphasis id="B962274C656AFF9FFF1EF82BFEFAF820" bold="true" box="[151,275,1972,1996]" pageId="4" pageNumber="60">FIGURE 1.</emphasis>
<taxonomicName id="4C1680DD656AFF9FFE93F82BFE43F820" authorityName="Lelej" authorityYear="1996" box="[282,426,1972,1996]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B962274C656AFF9FFE93F82BFE43F820" box="[282,426,1972,1996]" italics="true" pageId="4" pageNumber="60">Eotrogaspidia</emphasis>
</taxonomicName>
spp., ♂, habitus. A, E.
<taxonomicName id="4C1680DD656AFF9FFD1FF82AFCE0F820" authority="Okayasu &amp; Lelej &amp; Williams, 2021" authorityName="Okayasu &amp; Lelej &amp; Williams" authorityYear="2021" box="[662,777,1972,1996]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="adhabar" status="sp. nov.">
<emphasis id="B962274C656AFF9FFD1FF82AFD4CF820" box="[662,677,1973,1996]" italics="true" pageId="4" pageNumber="60">E</emphasis>
.
<emphasis id="B962274C656AFF9FFD3AF82BFCE0F820" box="[691,777,1972,1996]" italics="true" pageId="4" pageNumber="60">adhabar</emphasis>
</taxonomicName>
<emphasis id="B962274C656AFF9FFC86F82BFC89F820" bold="true" box="[783,864,1972,1996]" pageId="4" pageNumber="60">
<taxonomicNameLabel id="A2519A37656AFF9FFC86F82BFC89F820" box="[783,864,1972,1996]" pageId="4" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype; B, F.
<taxonomicName id="4C1680DD656AFF9FFB83F82AFB2BF820" authority="(Andre)" baseAuthorityName="Andre" baseAuthorityYear="1909" box="[1034,1218,1972,1996]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="amans">
<emphasis id="B962274C656AFF9FFB83F82AFBF0F820" box="[1034,1049,1973,1996]" italics="true" pageId="4" pageNumber="60">E</emphasis>
.
<emphasis id="B962274C656AFF9FFBAFF82AFB81F820" box="[1062,1128,1973,1996]" italics="true" pageId="4" pageNumber="60">amans</emphasis>
(André)
</taxonomicName>
; C, G.
<taxonomicName id="4C1680DD656AFF9FFA80F82AFF0EF81C" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1855" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="auroguttata">
<emphasis id="B962274C656AFF9FFA80F82AFAF1F820" box="[1289,1304,1973,1996]" italics="true" pageId="4" pageNumber="60">E</emphasis>
.
<emphasis id="B962274C656AFF9FFAAFF82AFA74F820" box="[1318,1437,1973,1996]" italics="true" pageId="4" pageNumber="60">auroguttata</emphasis>
(Smith)
</taxonomicName>
; D, H.
<taxonomicName id="4C1680DD656AFF9FFEB9F846FE71F81C" authority="Okayasu &amp; Lelej &amp; Williams, 2021" authorityName="Okayasu &amp; Lelej &amp; Williams" authorityYear="2021" box="[304,408,2008,2032]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="buddha" status="sp. nov.">
<emphasis id="B962274C656AFF9FFEB9F846FED6F81C" box="[304,319,2009,2032]" italics="true" pageId="4" pageNumber="60">E</emphasis>
.
<emphasis id="B962274C656AFF9FFEC5F847FE71F81C" box="[332,408,2008,2032]" italics="true" pageId="4" pageNumber="60">buddha</emphasis>
</taxonomicName>
<emphasis id="B962274C656AFF9FFE17F847FE06F81C" bold="true" box="[414,495,2008,2032]" pageId="4" pageNumber="60">
<taxonomicNameLabel id="A2519A37656AFF9FFE17F847FE06F81C" box="[414,495,2008,2032]" pageId="4" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype; AD. dorsal view; EH. lateral view.
</paragraph>
</caption>
<caption id="DF69ABD6656BFF9EFF1EF9A0FBCDF997" ID-DOI="http://doi.org/10.5281/zenodo.4471596" ID-Zenodo-Dep="4471596" httpUri="https://zenodo.org/record/4471596/files/figure.png" pageId="5" pageNumber="61" startId="5.[151,250,1599,1623]" targetBox="[151,1432,181,1574]" targetPageId="5">
<paragraph id="8BA9FB5E656BFF9EFF1EF9A0FBCDF997" blockId="5.[151,1436,1599,1659]" pageId="5" pageNumber="61">
<emphasis id="B962274C656BFF9EFF1EF9A0FEFDF9BB" bold="true" box="[151,276,1599,1623]" pageId="5" pageNumber="61">FIGURE 2.</emphasis>
<taxonomicName id="4C1680DD656BFF9EFE95F9A0FE45F9BB" authorityName="Lelej" authorityYear="1996" box="[284,428,1599,1623]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B962274C656BFF9EFE95F9A0FE45F9BB" box="[284,428,1599,1623]" italics="true" pageId="5" pageNumber="61">Eotrogaspidia</emphasis>
</taxonomicName>
spp., ♀, habitus. A, E.
<taxonomicName id="4C1680DD656BFF9EFD17F9DFFCB0F9BB" authority="(Andre)" baseAuthorityName="Andre" baseAuthorityYear="1909" box="[670,857,1599,1623]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="61" phylum="Arthropoda" rank="species" species="amans">
<emphasis id="B962274C656BFF9EFD17F9DFFD44F9BB" box="[670,685,1600,1623]" italics="true" pageId="5" pageNumber="61">E</emphasis>
.
<emphasis id="B962274C656BFF9EFD35F9DFFD17F9BB" box="[700,766,1600,1623]" italics="true" pageId="5" pageNumber="61">amans</emphasis>
(André)
</taxonomicName>
; B, F.
<taxonomicName id="4C1680DD656BFF9EFC14F9DFFB61F9BB" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1855" box="[925,1160,1599,1623]" class="Insecta" family="Crabronidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="61" phylum="Arthropoda" rank="species" species="auroguttata">
<emphasis id="B962274C656BFF9EFC14F9DFFC45F9BB" box="[925,940,1600,1623]" italics="true" pageId="5" pageNumber="61">E</emphasis>
.
<emphasis id="B962274C656BFF9EFC33F9DFFBD8F9BB" box="[954,1073,1600,1623]" italics="true" pageId="5" pageNumber="61">auroguttata</emphasis>
(Smith)
</taxonomicName>
; C, G.
<taxonomicName id="4C1680DD656BFF9EFB5AF9DFFAD5F9BB" authority="Okayasu &amp; Lelej &amp; Williams, 2021" authorityName="Okayasu &amp; Lelej &amp; Williams" authorityYear="2021" box="[1235,1340,1599,1623]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="61" phylum="Arthropoda" rank="species" species="buddha" status="sp. nov.">
<emphasis id="B962274C656BFF9EFB5AF9DFFB0BF9BB" box="[1235,1250,1600,1623]" italics="true" pageId="5" pageNumber="61">E</emphasis>
.
<emphasis id="B962274C656BFF9EFB79F9A0FAD5F9BB" box="[1264,1340,1599,1623]" italics="true" pageId="5" pageNumber="61">buddha</emphasis>
</taxonomicName>
<emphasis id="B962274C656BFF9EFACDF9A0FA7CF9BB" bold="true" box="[1348,1429,1599,1623]" pageId="5" pageNumber="61">
<taxonomicNameLabel id="A2519A37656BFF9EFACDF9A0FA7CF9BB" box="[1348,1429,1599,1623]" pageId="5" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype; D, H.
<taxonomicName id="4C1680DD656BFF9EFEB3F9FBFE0AF997" authority="Okayasu &amp; Lelej &amp; Williams, 2021" authorityName="Okayasu &amp; Lelej &amp; Williams" authorityYear="2021" box="[314,483,1635,1659]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="61" phylum="Arthropoda" rank="species" species="melanopleura" status="sp. nov.">
<emphasis id="B962274C656BFF9EFEB3F9FBFEA0F997" box="[314,329,1636,1659]" italics="true" pageId="5" pageNumber="61">E</emphasis>
.
<emphasis id="B962274C656BFF9EFEDFF9FCFE0AF997" box="[342,483,1635,1659]" italics="true" pageId="5" pageNumber="61">melanopleura</emphasis>
</taxonomicName>
<emphasis id="B962274C656BFF9EFE60F9FCFDD0F997" bold="true" box="[489,569,1635,1659]" pageId="5" pageNumber="61">
<taxonomicNameLabel id="A2519A37656BFF9EFE60F9FCFDD0F997" box="[489,569,1635,1659]" pageId="5" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype; AD. dorsal view; EH. lateral view.
</paragraph>
</caption>
<paragraph id="8BA9FB5E656BFF9EFF4EF936FD7FF90B" blockId="5.[151,1436,1705,2020]" pageId="5" pageNumber="61">Wings well developed; distance between origin of RS on vein SC and the base of stigmatic cell equal to stigmatic cell length and first abscissa RS length.</paragraph>
<paragraph id="8BA9FB5E656BFF9EFF4EF96DFE49F89B" blockId="5.[151,1436,1705,2020]" pageId="5" pageNumber="61">Procoxa with minute dense punctures; meso- and metacoxae with minute sparse and small sparse punctures, ventro-medially largely impunctate polished; femora and tarsi with minute dense and small sparse punctures; tarsi with small sparse punctures; metacoxa with weak inner carina throughout; tibiae without outer spines; claws simple, without inner denticles.</paragraph>
<paragraph id="8BA9FB5E656BFF9DFF4EF81EFEBAFE3D" blockId="5.[151,1436,1705,2020]" lastBlockId="6.[151,1437,151,753]" lastPageId="6" lastPageNumber="62" pageId="5" pageNumber="61">T1 with distinct dorsum and anterior slope; T1 spiracle weakly tuberculate; T2 evenly convex, 1.33 × wider than long; S1 with straight medial longitudinal carina with its outer margin emarginate medially and elevated posteriorly forming sharp tooth; S2 evenly convex without medial swelling or carina; T1, T2 laterally and S12 with large dense punctures; T2 disc with small sparse punctures; T37 and S38 with small dense punctures except T7 medially with narrow longitudinal impunctate carina not forming tubercle posteriorly; S7 with lateral rounded tubercle; S8 with lateral pointed tubercle reaching posterior 1/2 of sternum. Paramere down-curved posteriorly, without inner protuberance; inner margin of paramere dorsally with row of short erect golden setae; ventral lobe posteriorly projecting, pointed; parapenial lobe large, rounded; cuspis long slender, apex pointed, posteriorly up-curved, clearly surpassing apex of penis valve, ventrally with dense long erect golden setae; digitus S-shaped, posteriorly with dense short erect golden setae on outer face; paracuspis tuberculate with dense long erect golden setae; penis valves slender, weakly asymmetrical, right penis valve slightly longer than left, with posterior ventral hook and subapical short erect setae.</paragraph>
<paragraph id="8BA9FB5E6568FF9DFF1EFE44FE62FE19" blockId="6.[151,1437,151,753]" box="[151,395,475,501]" pageId="6" pageNumber="62">FEMALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30CA8D56568FF9DFF4EFE60FBC4FDD1" pageId="6" pageNumber="62" type="materials_examined">
<paragraph id="8BA9FB5E6568FF9DFF4EFE60FBC4FDD1" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="62">
<emphasis id="B962274C6568FF9DFF4EFE60FE91FDF4" bold="true" box="[199,376,510,537]" pageId="6" pageNumber="62">Type material.</emphasis>
<materialsCitation id="3B7EF1036568FF9DFE08FE60FABDFDF5" ID-GBIF-Occurrence="3028885367" box="[385,1364,510,537]" collectingDate="1975-10-21" collectionCode="HUM" collectorName="S. Takagi" country="Nepal" location="Adhabar" pageId="6" pageNumber="62" specimenCount="1" stateProvince="BARA DISTRICT" typeStatus="holotype">
3♂.
<typeStatus id="54AD45FC6568FF9DFE3CFE60FDC9FDF5" box="[437,544,511,537]" pageId="6" pageNumber="62" type="holotype">Holotype</typeStatus>
, ♂,
<emphasis id="B962274C6568FF9DFDDAFE60FD70FDF5" bold="true" box="[595,665,511,537]" pageId="6" pageNumber="62">
<collectingCountry id="F301BBCE6568FF9DFDDAFE60FD70FDF5" box="[595,665,511,537]" name="Nepal" pageId="6" pageNumber="62">Nepal</collectingCountry>
</emphasis>
:
<collectingRegion id="49D235BC6568FF9DFD23FE60FCA7FDFB" box="[682,846,511,537]" country="Nepal" name="BARA DISTRICT" pageId="6" pageNumber="62">BARA DISTRICT</collectingRegion>
:
<location id="8EC9AD856568FF9DFCD7FE60FC29FDF5" LSID="urn:lsid:plazi:treatment:03BF4A48656DFF9DFF1EFE18FD9EFD1D:8EC9AD856568FF9DFCD7FE60FC29FDF5" box="[862,960,511,537]" country="Nepal" name="Adhabar" pageId="6" pageNumber="62" stateProvince="BARA DISTRICT">Adhabar</location>
,
<date id="FFA8DD9E6568FF9DFC47FE61FBAFFDF5" box="[974,1094,510,537]" pageId="6" pageNumber="62" value="1975-10-21">
<collectingDate id="EFEC24766568FF9DFC47FE61FBAFFDF5" box="[974,1094,510,537]" pageId="6" pageNumber="62" value="1975-10-21">21.X.1975</collectingDate>
</date>
,
<collectorName id="26E39E886568FF9DFBDCFE60FB29FDF5" box="[1109,1216,511,537]" pageId="6" pageNumber="62">S. Takagi</collectorName>
leg. [
<collectionCode id="ED07639B6568FF9DFA8AFE61FAA4FDF5" box="[1283,1357,510,537]" country="Germany" httpUri="http://grbio.org/cool/kch5-hv72" name="Humboldt University Zoologisches Museum" pageId="6" pageNumber="62">HUM</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="3B7EF1036568FF9DFAE8FE60FBC0FDD1" ID-GBIF-Occurrence="3028885464" collectingDate="1959-10" collectionCode="CASC" collectorName="P. S. Nathan" country="India" elevation="213" location="Walayar Forests" pageId="6" pageNumber="62" specimenCount="1" stateProvince="Kerala" typeStatus="paratype">
<typeStatus id="54AD45FC6568FF9DFAE8FE60FF3FFDD1" pageId="6" pageNumber="62" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B962274C6568FF9DFF68FDBDFEC3FDD0" bold="true" box="[225,298,546,572]" pageId="6" pageNumber="62">
<collectingCountry id="F301BBCE6568FF9DFF68FDBDFECCFDD0" box="[225,293,546,572]" name="India" pageId="6" pageNumber="62">India</collectingCountry>
:
</emphasis>
<collectingRegion id="49D235BC6568FF9DFEB8FDBCFE6FFDD7" box="[305,390,547,573]" country="India" name="Kerala" pageId="6" pageNumber="62">KERALA</collectingRegion>
: 2♂,
<location id="8EC9AD856568FF9DFE4FFDBCFD97FDD1" LSID="urn:lsid:plazi:treatment:03BF4A48656DFF9DFF1EFE18FD9EFD1D:8EC9AD856568FF9DFE4FFDBCFD97FDD1" box="[454,638,547,573]" country="India" name="Walayar Forests" pageId="6" pageNumber="62" stateProvince="Kerala">Walayar Forests</location>
,
<quantity id="4CEE56BB6568FF9DFD00FDBCFD3DFDD1" box="[649,724,547,573]" metricMagnitude="2" metricUnit="m" metricValue="2.1336" pageId="6" pageNumber="62" unit="ft" value="700.0">
<elevation id="003B1C6D6568FF9DFD00FDBCFD3DFDD1" box="[649,724,547,573]" metricMagnitude="2" metricUnit="m" metricValue="2.1336" pageId="6" pageNumber="62" unit="ft" value="700.0">700 ft.</elevation>
</quantity>
,
<date id="FFA8DD9E6568FF9DFD69FDBDFCDFFDD1" box="[736,822,546,573]" pageId="6" pageNumber="62" value="1959-10">
<collectingDate id="EFEC24766568FF9DFD69FDBDFCDFFDD1" box="[736,822,546,573]" pageId="6" pageNumber="62" value="1959-10">X.1959</collectingDate>
</date>
,
<collectorName id="26E39E886568FF9DFCC8FDBCFC2AFDD1" box="[833,963,547,573]" pageId="6" pageNumber="62">P.S. Nathan</collectorName>
[
<collectionCode id="ED07639B6568FF9DFC58FDBDFBCAFDD1" box="[977,1059,546,573]" pageId="6" pageNumber="62">CASC</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30CA8D56568FF9DFF4EFDD9FD7FFD8D" box="[199,662,582,609]" pageId="6" pageNumber="62" type="distribution">
<paragraph id="8BA9FB5E6568FF9DFF4EFDD9FD7FFD8D" blockId="6.[151,1437,151,753]" box="[199,662,582,609]" pageId="6" pageNumber="62">
<emphasis id="B962274C6568FF9DFF4EFDD9FE88FD8C" bold="true" box="[199,353,582,608]" pageId="6" pageNumber="62">Distribution.</emphasis>
<collectingCountry id="F301BBCE6568FF9DFEEEFDD8FE4CFD8D" box="[359,421,583,609]" name="India" pageId="6" pageNumber="62">India</collectingCountry>
:
<collectingRegion id="49D235BC6568FF9DFE39FDD8FE17FD8D" box="[432,510,583,609]" country="India" name="Kerala" pageId="6" pageNumber="62">Kerala</collectingRegion>
.
<collectingCountry id="F301BBCE6568FF9DFD81FDD8FDA6FD8D" box="[520,591,583,609]" name="Nepal" pageId="6" pageNumber="62">Nepal</collectingCountry>
: Bara.
</paragraph>
</subSubSection>
<subSubSection id="C30CA8D56568FF9DFF4EFDF4FADDFD69" box="[199,1332,619,645]" pageId="6" pageNumber="62" type="etymology">
<paragraph id="8BA9FB5E6568FF9DFF4EFDF4FADDFD69" blockId="6.[151,1437,151,753]" box="[199,1332,619,645]" pageId="6" pageNumber="62">
<emphasis id="B962274C6568FF9DFF4EFDF4FEA7FD69" bold="true" box="[199,334,619,645]" pageId="6" pageNumber="62">Etymology.</emphasis>
This new species is named after the
<typeStatus id="54AD45FC6568FF9DFD63FDF4FCF3FD69" box="[746,794,619,645]" pageId="6" pageNumber="62">type</typeStatus>
locality in
<collectingCountry id="F301BBCE6568FF9DFC10FDF4FC09FD69" box="[921,992,619,645]" name="Nepal" pageId="6" pageNumber="62">Nepal</collectingCountry>
. Treat as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C30CA8D56568FF9DFF4EFD10FD9EFD1D" pageId="6" pageNumber="62" type="discussion">
<paragraph id="8BA9FB5E6568FF9DFF4EFD10FD9EFD1D" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="62">
<emphasis id="B962274C6568FF9DFF4EFD10FED2FD45" bold="true" box="[199,315,655,681]" pageId="6" pageNumber="62">Remarks.</emphasis>
This new species is readily recognized in
<taxonomicName id="4C1680DD6568FF9DFC82FD10FC43FD45" authorityName="Lelej" authorityYear="1996" box="[779,938,655,681]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B962274C6568FF9DFC82FD10FC43FD45" box="[779,938,655,681]" italics="true" pageId="6" pageNumber="62">Eotrogaspidia</emphasis>
</taxonomicName>
by having the clypeus gibbous and the metacoxa evenly convex with long sparse setae; in these features, it is similar to
<taxonomicName id="4C1680DD6568FF9DFC52FD2CFB88FD21" authorityName="Ashmead" authorityYear="1899" box="[987,1121,691,717]" class="Insecta" family="Mutillidae" genus="Trogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B962274C6568FF9DFC52FD2CFB88FD21" box="[987,1121,691,717]" italics="true" pageId="6" pageNumber="62">Trogaspidia</emphasis>
</taxonomicName>
. The genitalia, however, are consistent with other
<taxonomicName id="4C1680DD6568FF9DFE0EFD48FDCFFD1D" authorityName="Lelej" authorityYear="1996" box="[391,550,727,753]" class="Insecta" family="Mutillidae" genus="Eotrogaspidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B962274C6568FF9DFE0EFD48FDCFFD1D" box="[391,550,727,753]" italics="true" pageId="6" pageNumber="62">Eotrogaspidia</emphasis>
</taxonomicName>
males.
</paragraph>
</subSubSection>
</treatment>
</document>