treatments-xml/data/03/BE/A7/03BEA74211A15D24A2218E195CA4E1A1.xml

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<document ID-CLB-Dataset="89309" ID-DOI="http://dx.doi.org/10.3897/zookeys.1110.82461" ID-GBIF-Dataset="92e148c9-6ca3-489d-8b0b-7bca978ee901" ID-PMC="PMC9848772" ID-Pensoft-Pub="1313-2970-1110-1" ID-Pensoft-UUID="88D66028C7A752528DD9138A2105F895" ID-PubMed="36761453" ID-ZooBank="01302157EE404B9999DD3EC3377D929C" ModsDocID="1313-2970-1110-1" checkinTime="1657006639549" checkinUser="pensoft" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak, Lin, Aung &amp; Panha, Somsak" docDate="2022" docId="03BEA74211A15D24A2218E195CA4E1A1" docLanguage="en" docName="ZooKeys 1110: 1-37" docOrigin="ZooKeys 1110" docPubDate="2022-07-04" docSource="http://dx.doi.org/10.3897/zookeys.1110.82461" docTitle="Burmochlamys fasciola Pholyotha &amp; Panha 2022, sp. nov." docType="treatment" docUuid="BDE9EEDE-1DAC-418E-9097-1EDCC579299B" docUuidSource="ZooBank" docVersion="5" id="88D66028C7A752528DD9138A2105F895" lastPageNumber="1" masterDocId="88D66028C7A752528DD9138A2105F895" masterDocTitle="Uncovering local endemism from southeastern Myanmar: description of the new karst-associated terrestrial snail genus Burmochlamys (Eupulmonata, Helicarionidae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="1" updateTime="1732937909010" updateUser="ExternalLinkService">
<mods:mods id="0E2A7558DC12080C8B4B9D44D20B9C30" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="41B388F11F5ACF101CB480CBE4000B6C">
<mods:title id="363D4A1D1C40BFB6D2BED4639512C1A1">Uncovering local endemism from southeastern Myanmar: description of the new karst-associated terrestrial snail genus Burmochlamys (Eupulmonata, Helicarionidae)</mods:title>
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<mods:name id="CB9FDAAC86EC3A909DF993B13E180249" type="personal">
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<mods:roleTerm id="B4F2BC68A3F72E0042ABCF3987F24827">Author</mods:roleTerm>
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<mods:namePart id="23233B08777EB57B96B57694F03401F1">Pholyotha, Arthit</mods:namePart>
<mods:affiliation id="C9D0AF94EE611BD7503C98496E8C4D8F">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
</mods:name>
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<mods:namePart id="A5C255F74FBEEF9DF18F164F901451F1">Sutcharit, Chirasak</mods:namePart>
<mods:affiliation id="C5159821F136C99E8891DB6FF0CFCD8D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
</mods:name>
<mods:name id="809BAC486D7E695CECF09118055EE26D" type="personal">
<mods:role id="4D02EBCD68AEE6C39137B3DCEC4D681F">
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</mods:role>
<mods:namePart id="3105F1F3E1AFE57BEF20CC854B572AE2">Lin, Aung</mods:namePart>
<mods:affiliation id="8FDC848E10027A54066EA5F7BAD634CF">Fauna and Flora International, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
</mods:name>
<mods:name id="C53680B55781FBDCF4C3B9D7B9949C4C" type="personal">
<mods:role id="EE79287987C951EB5F2CE6DF2557E652">
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</mods:role>
<mods:namePart id="86BE4BA2817686A64E44D7848CD4E9EC">Panha, Somsak</mods:namePart>
<mods:affiliation id="C2B06AE00AB2B08E224027AE382A23FC">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok, Thailand</mods:affiliation>
<mods:nameIdentifier id="76EAC662F79316FECF0522E99B96428F" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0845BE1245D3B0C8383B74F9B39FED43">text</mods:typeOfResource>
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<mods:title id="548A59DFA3B5D9CC225EF026675FCCC3">ZooKeys</mods:title>
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<mods:part id="3D947A72BBEAEF92B97452A4BAFBD596">
<mods:date id="3867D2BBBD4DECCC310583E25867EE47">2022</mods:date>
<mods:detail id="51F5C5578B7CD1374928C1DD8105DAC6" type="pubDate">
<mods:number id="3C04E5DF4744D4188866DF35A47CC8C3">2022-07-04</mods:number>
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<mods:detail id="50EF8F8946304AE3F595F64461E0B318" type="volume">
<mods:number id="F2A58455EB014A8AF0184628929EA75A">1110</mods:number>
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<mods:start id="B24E31528E6EF8DB83F42E0AD0BC7022">1</mods:start>
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<mods:url id="9B212F5ABAF6EDB831054C73B465FA1D">http://dx.doi.org/10.3897/zookeys.1110.82461</mods:url>
</mods:location>
<mods:classification id="1A3686E308152EDC07B281D2FD2C03E0">journal article</mods:classification>
<mods:identifier id="3E66C6D3CBF0A10930C3522B8E96FA57" type="DOI">http://dx.doi.org/10.3897/zookeys.1110.82461</mods:identifier>
<mods:identifier id="87E2183A7F2B19E25663BC1B91741B8F" type="Pensoft-Pub">1313-2970-1110-1</mods:identifier>
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<treatment id="03BEA74211A15D24A2218E195CA4E1A1" ID-GBIF-Taxon="196387962" LSID="urn:lsid:zoobank.org:act:BDE9EEDE-1DAC-418E-9097-1EDCC579299B" httpUri="http://treatment.plazi.org/id/03BEA74211A15D24A2218E195CA4E1A1" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Helicarionidae" scope_subFamily="Durgellinae" scope_superOrder="Eupulmonata">
<subSubSection id="BA34EFDE7E3C492EC42DE7B6D0ED63A2" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="BCA5FC98C574F0A83ACA5E5073356F8D" pageId="0" pageNumber="1">
<taxonomicName id="BBD4D26C756F9DF097251F7C203C1A8A" ID-CoL="7P2B6" LSID="https://zoobank.org/BDE9EEDE-1DAC-418E-9097-1EDCC579299B" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Burmochlamys fasciola" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="fasciola" status="sp. nov.">Burmochlamys fasciola Pholyotha &amp; Panha</taxonomicName>
<taxonomicNameLabel id="C51B6909085710B2821A99572DB6D8EB" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C2E6995E3A009EB5D83326C95D864B75" pageId="0" pageNumber="1" type="description">
<paragraph id="D14FC8C5A39AC3D226032EC5429A7C14" pageId="0" pageNumber="1">
<figureCitation id="94658BF2DF81A35302C422A8E3434ECC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the Salween River basin (known as Thanlwin River) in the southeastern Myanmar showing the sampling sites. Coloured triangles indicate the localities recorded for each Burmochlamys species. White circles indicate the localities recorded for Sophina species. White squares indicate the sampling localities with no records of both Sophina and Burmochlamys species." figureDoi="10.3897/zookeys.1110.82461.figure1" httpUri="https://binary.pensoft.net/fig/709567" pageId="0" pageNumber="1">Figs 1</figureCitation>
<figureCitation id="FD5578567DF8AA083D46B5A36CAAE129" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitat of some species of Burmochlamys gen. nov. in the karst basin of Hpa-An, Kayin State, Myanmar. A type locality of B. fasciola sp. nov. at Bardai Mountain and microhabitat structure of the karst wall B newly discovered locality of B. poongee at Kaw Ka Thaung Cave C type locality of B. albida sp. nov. at Waiponla Monastery D type locality of B. moulmeinica sp. nov. at Lun Nya Pagoda E type locality of B. versicolor sp. nov. at Bayin Nyi Cave. Photos by Ruttapon Srisonchai." figureDoi="10.3897/zookeys.1110.82461.figure2" httpUri="https://binary.pensoft.net/fig/709568" pageId="0" pageNumber="1">, 2A</figureCitation>
<figureCitation id="BA56CDAD7AD74644B8DB62DCB830314D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Synoptic illustration of mantle extensions, body terminology and radular morphology of Burmochlamys gen. nov. A right and left shell lobes of B. fasciola sp. nov. B mantle extensions (shell lobes and dorsal lobes) and posterior body of B. poongee C dorsal view of mantle extensions (shell lobes and dorsal lobes) of B. cassidula D, E representative SEM images of radula showing monocuspid central tooth (yellow) and lateromarginal teeth (blue) attached by the smaller triangular-shaped teeth (red). White arrow indicates pneumostome or breathing pore. Yellow arrow indicates a tiny cusp on outermost lateromarginal teeth." figureDoi="10.3897/zookeys.1110.82461.figure3" httpUri="https://binary.pensoft.net/fig/709569" pageId="0" pageNumber="1">, 3A</figureCitation>
<figureCitation id="CA2E6A4D07A286F66A5FE0282F3118A8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Living snails of Burmochlamys species A B. fasciola sp. nov. paratype CUMZ 14214 showing left and right lateral views B B. cassidula specimen CUMZ 14209 C B. poongee specimen CUMZ 14210 D B. albida sp. nov. paratype CUMZ 14212 E B. moulmeinica sp. nov. paratype CUMZ 14217 F B. versicolor sp. nov. paratype CUMZ 14219 G B. whitteni sp. nov. paratype CUMZ 14222." figureDoi="10.3897/zookeys.1110.82461.figure5" httpUri="https://binary.pensoft.net/fig/709571" pageId="0" pageNumber="1">, 5A</figureCitation>
<figureCitation id="5ECD680337A100AEB57A6239B880716E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells of Burmochlamys species A, B B. albida sp. nov. A holotype CUMZ 14211, and D paratype CUMZ 14212 C, D B. fasciola sp. nov. C holotype CUMZ 14213 D paratype CUMZ 14214 E, F B. moulmeinica sp. nov. E holotype CUMZ 14216 F paratype CUMZ 14217." figureDoi="10.3897/zookeys.1110.82461.figure7" httpUri="https://binary.pensoft.net/fig/709573" pageId="0" pageNumber="1">, 7C, D</figureCitation>
<figureCitation id="5B0ACAE14166E0B7E7C06E2DD0393781" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Representative SEM images of the shell of Burmochlamys species A-C B. cassidula specimen CUMZ 14209 A protoconch B body whorl C zoom-in view of body whorl D-F B. poongee specimen CUMZ 14210 D protoconch E body whorl F zoom-in view of body whorl G-I B. albida sp. nov. paratype CUMZ 14212 G protoconch H body whorl I zoom-in view of body whorl J-L B. fasciola sp. nov. paratype CUMZ 14214 J protoconch K body whorl L zoom-in view of body whorl." figureDoi="10.3897/zookeys.1110.82461.figure8" httpUri="https://binary.pensoft.net/fig/709574" pageId="0" pageNumber="1">, 8J-L</figureCitation>
<figureCitation id="206281A6961C9ED6F94F3749DE157AA1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. General view of the genital system of Burmochlamys species A B. fasciola sp. nov. paratype CUMZ 14214 B B. moulmeinica sp. nov. paratype CUMZ 14217 C B. versicolor sp. nov. paratype CUMZ 14219. White arrow indicates the end of the penis. Yellow arrow indicates the solid at the tip of dart apparatus. Red-dotted line indicates vas deferens missing." figureDoi="10.3897/zookeys.1110.82461.figure10" httpUri="https://binary.pensoft.net/fig/709576" pageId="0" pageNumber="1">, 10A</figureCitation>
<figureCitation id="D640301DE9162E9C928BECC5F7AAE748" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Representative SEM images of the radula of Burmochlamys species A B. cassidula specimen CUMZ 14209 B B. poongee specimen CUMZ 14210 C B. albida sp. nov. paratype CUMZ 14212 D B. fasciola sp. nov. paratype CUMZ 14214. Central tooth indicated by ' C'; yellow colour indicates central tooth row." figureDoi="10.3897/zookeys.1110.82461.figure11" httpUri="https://binary.pensoft.net/fig/709577" pageId="0" pageNumber="1">, 11D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="42812D127A0F046FE8D120F8B847131D" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="C141F6A8F504DDB99E4EA04C9C6AC8E9" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="E836214202ABFEE37356A3C55AF34772" pageId="0" pageNumber="1">
<emphasis id="E80C6E4034E2850A41E248030A92C5B2" bold="true" pageId="0" pageNumber="1">
Type material.
<emphasis id="CC95493B52A87C936E0E970A7E199646" italics="true" pageId="0" pageNumber="1">
<typeStatus id="45B8AC86AA18208FDACCBBB438E89264">Holotype</typeStatus>
</emphasis>
</emphasis>
: CUMZ 14213 (Fig.
<figureCitation id="52C863922767A90DB0C0403D26ABF30E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells of Burmochlamys species A, B B. albida sp. nov. A holotype CUMZ 14211, and D paratype CUMZ 14212 C, D B. fasciola sp. nov. C holotype CUMZ 14213 D paratype CUMZ 14214 E, F B. moulmeinica sp. nov. E holotype CUMZ 14216 F paratype CUMZ 14217." figureDoi="10.3897/zookeys.1110.82461.figure7" httpUri="https://binary.pensoft.net/fig/709573" pageId="0" pageNumber="1">7C</figureCitation>
; width
<quantity id="F154A63876AE8A97F053A1F169AF4EEB" metricMagnitude="-3" metricUnit="m" metricValue="5.6" unit="mm" value="5.6">5.6 mm</quantity>
, height
<quantity id="F533715C6C857CA98E9A1DF50A46F93F" metricMagnitude="-3" metricUnit="m" metricValue="5.1" unit="mm" value="5.1">5.1 mm</quantity>
).
<emphasis id="29C98765E1AF4E07E3E14F52F6F8D153" bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus id="8C3BE21837C79C5473E849969685F9A4">Paratypes</typeStatus>
</emphasis>
: Same locality as holotype: CUMZ 14214 (Fig.
<figureCitation id="BFC149C02FFBD6115B3552CCF3B26887" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells of Burmochlamys species A, B B. albida sp. nov. A holotype CUMZ 14211, and D paratype CUMZ 14212 C, D B. fasciola sp. nov. C holotype CUMZ 14213 D paratype CUMZ 14214 E, F B. moulmeinica sp. nov. E holotype CUMZ 14216 F paratype CUMZ 14217." figureDoi="10.3897/zookeys.1110.82461.figure7" httpUri="https://binary.pensoft.net/fig/709573" pageId="0" pageNumber="1">7D</figureCitation>
; width
<quantity id="D077F6A7729E825D5A577BEB88784BEA" metricMagnitude="-3" metricUnit="m" metricValue="5.8" unit="mm" value="5.8">5.8 mm</quantity>
, height
<quantity id="C17C72CE930D03B92FD2C6ED7BEB6713" metricMagnitude="-3" metricUnit="m" metricValue="4.9" unit="mm" value="4.9">4.9 mm</quantity>
), NHMUK (two shells).
</paragraph>
</subSubSection>
<subSubSection id="2E325AE8056F064EE3D9B01DBB8E5319" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="2B90CBE4F54E4DEA529344C7A26C146A" pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph id="587BDDE2F2DCC8AB434415941EEBF8A7" pageId="0" pageNumber="1">
<materialsCitation id="7D25131E6418DC8505112B59AE52E98F" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Myanmar" latitude="17.016527" location="Kyankaw Mountain" longLatPrecision="1" longitude="97.70344" specimenCount="1" stateProvince="Kayin State">
<location id="C92079C8F580B14F26E0016E7F8C0FB0" LSID="urn:lsid:plazi:treatment:03BEA74211A15D24A2218E195CA4E1A1:C92079C8F580B14F26E0016E7F8C0FB0" country="Myanmar" latitude="17.016527" longLatPrecision="1" longitude="97.70344" name="Kyankaw Mountain" stateProvince="Kayin State">Kyankaw Mountain</location>
, Hpa-An,
<collectingRegion id="FB56806139EB600DE3AF13B8DA46BFE8" country="Myanmar" name="Kayin">Kayin State</collectingRegion>
,
<collectingCountry id="76141EA3D39BD61F9C0ED78E83CFAD02" name="Myanmar">Myanmar</collectingCountry>
(
<geoCoordinate id="0A3E8DDB299135B3C6E929543CC42C8F" degrees="17" direction="north" minutes="00" orientation="latitude" precision="1" seconds="59.5" value="17.016527">17°00'59.5&quot;N</geoCoordinate>
,
<geoCoordinate id="D538F8BC3BB7F0B2A1C25FB677A1D95A" degrees="97" direction="east" minutes="42" orientation="longitude" precision="1" seconds="12.4" value="97.70344">97°42'12.4&quot;E</geoCoordinate>
): CUMZ 14215
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1820D170C35F5D563DB0D19F46F64EF1" pageId="0" pageNumber="1" type="type locality">
<paragraph id="530776DB59FA2E294866C1C9321E77FC" pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph id="1B2E76B0B210DD0188D64EFE50726C98" pageId="0" pageNumber="1">
Bardai Mountain, Hpa-An, Kayin State, Myanmar (
<geoCoordinate id="F9E1B8C7BE341B675B246A014D0240BC" degrees="17" direction="north" minutes="00" orientation="latitude" precision="1" seconds="00.5" value="17.00014">17°00'00.5&quot;N</geoCoordinate>
,
<geoCoordinate id="8886D61E89DC3E4D1A736520825BA41D" degrees="97" direction="east" minutes="41" orientation="longitude" precision="1" seconds="41.6" value="97.69489">97°41'41.6&quot;E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="6E7D4682F6E93FEFAB127CCAFD4D0B8C" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="7E2E690B50C35D42FF4D0AC63CD783F3" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="B23FB4A449CD99B8A765F2AC9EF5BBB0" pageId="0" pageNumber="1">Shell globose and milky white with narrow yellowish brown band. Animal pale freshy-grey with five mantle extensions. Genitalia with rather short epiphallic caecum attached by thin penial retractor muscle, very short vagina, and solid at the tip of dart apparatus.</paragraph>
</subSubSection>
<subSubSection id="CA7FA9E4467A7954B99C1E1CDC003F66" pageId="0" pageNumber="1" type="description">
<paragraph id="E4C4382862440C7BFDE8DBB285B65037" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="460DF9329AA9167B0AD3C3F099CC2369" pageId="0" pageNumber="1">
<emphasis id="945896372D03388DE19A74AD3E159CB4" bold="true" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
(Figs
<figureCitation id="8943AC32B03B2D9AC4952B2F253BD6DF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Shells of Burmochlamys species A, B B. albida sp. nov. A holotype CUMZ 14211, and D paratype CUMZ 14212 C, D B. fasciola sp. nov. C holotype CUMZ 14213 D paratype CUMZ 14214 E, F B. moulmeinica sp. nov. E holotype CUMZ 14216 F paratype CUMZ 14217." figureDoi="10.3897/zookeys.1110.82461.figure7" httpUri="https://binary.pensoft.net/fig/709573" pageId="0" pageNumber="1">7C, D</figureCitation>
,
<figureCitation id="AA33C668E4EE2CCFF9BD4F350011F436" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Representative SEM images of the shell of Burmochlamys species A-C B. cassidula specimen CUMZ 14209 A protoconch B body whorl C zoom-in view of body whorl D-F B. poongee specimen CUMZ 14210 D protoconch E body whorl F zoom-in view of body whorl G-I B. albida sp. nov. paratype CUMZ 14212 G protoconch H body whorl I zoom-in view of body whorl J-L B. fasciola sp. nov. paratype CUMZ 14214 J protoconch K body whorl L zoom-in view of body whorl." figureDoi="10.3897/zookeys.1110.82461.figure8" httpUri="https://binary.pensoft.net/fig/709574" pageId="0" pageNumber="1">8J-L</figureCitation>
). Shell globose, small (width up to 6.1 mm, height up to 6.0 mm), thin, semi-translucent. Colour milky-white with a narrow yellowish brown band above periphery. Protoconch and teleoconch surface with similar sculpture. Surface of body whorl with distinct spiral furrows at regular intervals, cut by distinctly undulating radial lines (Fig.
<figureCitation id="38F73619D79F22339BA9AA131D54320C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Representative SEM images of the shell of Burmochlamys species A-C B. cassidula specimen CUMZ 14209 A protoconch B body whorl C zoom-in view of body whorl D-F B. poongee specimen CUMZ 14210 D protoconch E body whorl F zoom-in view of body whorl G-I B. albida sp. nov. paratype CUMZ 14212 G protoconch H body whorl I zoom-in view of body whorl J-L B. fasciola sp. nov. paratype CUMZ 14214 J protoconch K body whorl L zoom-in view of body whorl." figureDoi="10.3897/zookeys.1110.82461.figure8" httpUri="https://binary.pensoft.net/fig/709574" pageId="0" pageNumber="1">8J-L</figureCitation>
). Whorls 6-
<normalizedToken id="0D2F8F7101E50D9A5AFC2EAF7935F71D" originalValue="6½">61/2</normalizedToken>
, increasing regularly; suture shallowly impressed; spire much elevated; last whorl well-rounded. Aperture obliquely crescent-shaped; peristome simple. Columellar margin simple, slightly reflected near umbilicus. Umbilicus open, narrow, and deep.
</paragraph>
<caption id="93594E56A14176D97A58609C6B03895A" doi="10.3897/zookeys.1110.82461.figure8" httpUri="https://binary.pensoft.net/fig/709574" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
<paragraph id="8DBDC04F5EC114CED48B84EFFB1E55E0" pageId="0" pageNumber="1">
<emphasis id="A53DBF72C66892D670B1AF3A470B2373" bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
Representative SEM images of the shell of
<taxonomicName id="027AF2248637DC97550726B6D86E3967" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="FB2EA3CB641FC97A6836512D04CB2D90" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
species
<emphasis id="2A56DF25CDD31FB03DD36F53C7DE570D" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
<taxonomicName id="AF7DCA69A692B02C3E0B2EDB41CE7595" lsidName="B. cassidula" pageId="0" pageNumber="1" rank="species" species="cassidula">
<emphasis id="85641135522263812E343EC3F2D7A152" italics="true" pageId="0" pageNumber="1">B. cassidula</emphasis>
</taxonomicName>
specimen CUMZ 14209
<emphasis id="4618D548FDFEFCD7E7C9B04D3AD2FD66" bold="true" pageId="0" pageNumber="1">A</emphasis>
protoconch
<emphasis id="A8C8721C75713B226C7A02A254EE9EDE" bold="true" pageId="0" pageNumber="1">B</emphasis>
body whorl
<emphasis id="8C5B7ADA1E7C72979BA56DC84D378BF5" bold="true" pageId="0" pageNumber="1">C</emphasis>
zoom-in view of body whorl
<emphasis id="4A2417252DD6C982277C7B6E94269342" bold="true" pageId="0" pageNumber="1">D-F</emphasis>
<taxonomicName id="480E3506B685ECBAC12DB1D2DD3CCB3D" lsidName="B. poongee" pageId="0" pageNumber="1" rank="species" species="poongee">
<emphasis id="6DA3C38E494E6BAC7A2E432EE4BBBAAE" italics="true" pageId="0" pageNumber="1">B. poongee</emphasis>
</taxonomicName>
specimen CUMZ 14210
<emphasis id="F4FA58CCF4869EF6269410EFCDF432A5" bold="true" pageId="0" pageNumber="1">D</emphasis>
protoconch
<emphasis id="89B8E798756642530C13A02A18D8D940" bold="true" pageId="0" pageNumber="1">E</emphasis>
body whorl
<emphasis id="ED3A0D2100B5414179E64EACEE783E14" bold="true" pageId="0" pageNumber="1">F</emphasis>
zoom-in view of body whorl
<emphasis id="8644B64627FAFB9B96DD261EE73325CD" bold="true" pageId="0" pageNumber="1">G-I</emphasis>
<taxonomicName id="12E441C390608ECE305273E2DE85040F" lsidName="B. albida" pageId="0" pageNumber="1" rank="species" species="albida">
<emphasis id="2D6F3C992984CA9F140170790AE82427" italics="true" pageId="0" pageNumber="1">B. albida</emphasis>
</taxonomicName>
sp. nov. paratype CUMZ 14212
<emphasis id="597697F6B724F0AC7239DC4D09C86539" bold="true" pageId="0" pageNumber="1">G</emphasis>
protoconch
<emphasis id="6069AED3144BBEFE016DC922A49C4479" bold="true" pageId="0" pageNumber="1">H</emphasis>
body whorl
<emphasis id="8A31529C29D83D22E4597928A4CD4647" bold="true" pageId="0" pageNumber="1">I</emphasis>
zoom-in view of body whorl
<emphasis id="2CAEA39E1A26124A5496F38C63F50316" bold="true" pageId="0" pageNumber="1">J-L</emphasis>
<taxonomicName id="9ED91247E9599B0B59622A7AE8FCFEFF" lsidName="B. fasciola" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis id="CE62C0EE7DCE476AB5987DC0F63441D7" italics="true" pageId="0" pageNumber="1">B. fasciola</emphasis>
</taxonomicName>
sp. nov. paratype CUMZ 14214
<emphasis id="1F2DBB278D0CFC946BFDDD35DDF107BF" bold="true" pageId="0" pageNumber="1">J</emphasis>
protoconch
<emphasis id="AB8101AF50E904E3E7CFCE5E79FF7ED8" bold="true" pageId="0" pageNumber="1">K</emphasis>
body whorl
<emphasis id="262C355E4DE6DA171496809313549FD0" bold="true" pageId="0" pageNumber="1">L</emphasis>
zoom-in view of body whorl.
</paragraph>
</caption>
<paragraph id="3CF560BC175E821FB601A8ACAC40B834" pageId="0" pageNumber="1">
<emphasis id="D0350D86721CD5802AA23754169E3F34" bold="true" italics="true" pageId="0" pageNumber="1">Genital organs</emphasis>
(Fig.
<figureCitation id="CE23E419A5876AD3CDB32E668AB48AEB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. General view of the genital system of Burmochlamys species A B. fasciola sp. nov. paratype CUMZ 14214 B B. moulmeinica sp. nov. paratype CUMZ 14217 C B. versicolor sp. nov. paratype CUMZ 14219. White arrow indicates the end of the penis. Yellow arrow indicates the solid at the tip of dart apparatus. Red-dotted line indicates vas deferens missing." figureDoi="10.3897/zookeys.1110.82461.figure10" httpUri="https://binary.pensoft.net/fig/709576" pageId="0" pageNumber="1">10A</figureCitation>
). Atrium (at) very short or indistinct. Penis (p) moderate, cylindrical and with a thin penial sheath. Epiphallus (e1+e2) ca. three-fifths of penis length; e1 cylindrical and smaller diameter than penis and e2; e2 bulbous and ca. half of e1 length. Epiphallic caecum (ec) rather short, cylindrical and with a thin penial retractor muscle (prm) attached at tip. Vas deferens (vd) thin tube. Dart apparatus large, long cylindrical, with solid at the tip (yellow arrow in Fig.
<figureCitation id="1B565BF3FA1C73F5CB9D90F356208768" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. General view of the genital system of Burmochlamys species A B. fasciola sp. nov. paratype CUMZ 14214 B B. moulmeinica sp. nov. paratype CUMZ 14217 C B. versicolor sp. nov. paratype CUMZ 14219. White arrow indicates the end of the penis. Yellow arrow indicates the solid at the tip of dart apparatus. Red-dotted line indicates vas deferens missing." figureDoi="10.3897/zookeys.1110.82461.figure10" httpUri="https://binary.pensoft.net/fig/709576" pageId="0" pageNumber="1">10A</figureCitation>
), and located on atrium at vagina and penis junction. Vagina (v) very short to indistinguishable. Gametolytic sac (gs) bulbous; gametolytic duct (gd) cylindrical, moderate, slightly shorter than penis. Free oviduct (fo) ca. half of penis length, cylindrical, and encircled with thick tissue near vagina.
</paragraph>
<paragraph id="13FF6CC0377D2C481508F749645F10CF" pageId="0" pageNumber="1">
<taxonomicName id="887E44E7A206A51F8174FA7F7F0DCC2D" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" lsidName="Radula" order="Porellales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
<emphasis id="3D6AE2F844D5EBED7C67E6EF2A5CFA49" bold="true" italics="true" pageId="0" pageNumber="1">Radula</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="3806E8E9EC33B581741993415747974F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Representative SEM images of the radula of Burmochlamys species A B. cassidula specimen CUMZ 14209 B B. poongee specimen CUMZ 14210 C B. albida sp. nov. paratype CUMZ 14212 D B. fasciola sp. nov. paratype CUMZ 14214. Central tooth indicated by ' C'; yellow colour indicates central tooth row." figureDoi="10.3897/zookeys.1110.82461.figure11" httpUri="https://binary.pensoft.net/fig/709577" pageId="0" pageNumber="1">11D</figureCitation>
). Teeth arranged in wide U-shaped rows with each row consisting of ~ 50 teeth. Central tooth monocuspid, large and oblong spatulate plate, and attached by two smaller triangular-shaped teeth at its base. Laterals and marginals not differentiated and monocuspid; each tooth with large and oblong spatulate plate attached by only a smaller triangular-shaped tooth at base on outer side. From inner to outer, lateromarginal teeth gradually narrower, smaller, and rather more pointed cusp. Some outermost teeth with a small and pointed cusp at inner side.
</paragraph>
<caption id="4F2DE01F73DE28E52BEB34B090EB98E4" doi="10.3897/zookeys.1110.82461.figure9" httpUri="https://binary.pensoft.net/fig/709575" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
<paragraph id="1B477034918366E62E65FE1B780B730E" pageId="0" pageNumber="1">
<emphasis id="304ADC64D3FD1DAA1AA31E44A78BE438" bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
General view of the genital system of
<taxonomicName id="E69BE65588022B8819C49602290A1E84" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="0C9649A30EB80D743291CF61A4DECD98" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
species
<emphasis id="F570E0CADC9550C66012962F383062F2" bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName id="D3DA0D74ACF302591ACCAE42B335D49A" lsidName="B. cassidula" pageId="0" pageNumber="1" rank="species" species="cassidula">
<emphasis id="B63377C39DF295B60184365C7D05A696" italics="true" pageId="0" pageNumber="1">B. cassidula</emphasis>
</taxonomicName>
specimen CUMZ 14209
<emphasis id="43BE8C11FC8941770811A900C4C7C5AF" bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName id="B1A62FAAB39E77AEBE25D408C74115F7" lsidName="B. poongee" pageId="0" pageNumber="1" rank="species" species="poongee">
<emphasis id="196C2DC54E0B5C5EA3A102EB5CC59093" italics="true" pageId="0" pageNumber="1">B. poongee</emphasis>
</taxonomicName>
specimen CUMZ 14210
<emphasis id="C2F13EFFA7C6462561660C2B012D3A3E" bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName id="86B5221E342681C0D6DA3DB5A64ED2E1" lsidName="B. albida" pageId="0" pageNumber="1" rank="species" species="albida">
<emphasis id="0282E4BDAF6AA36B2C5B8F86D47F37F4" italics="true" pageId="0" pageNumber="1">B. albida</emphasis>
</taxonomicName>
sp. nov. paratype CUMZ 14212. White arrow indicates the end of the penis. Red-dotted line indicates vas deferens missing.
</paragraph>
</caption>
<paragraph id="ED82A6B178790DF4772F1F45DC8F26C1" pageId="0" pageNumber="1">
<emphasis id="8EBE014570207747466FCB374C9D9D4D" bold="true" italics="true" pageId="0" pageNumber="1">External appearance</emphasis>
(Figs
<figureCitation id="4ECF282DA662DBF44886CF2B82A85F54" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Synoptic illustration of mantle extensions, body terminology and radular morphology of Burmochlamys gen. nov. A right and left shell lobes of B. fasciola sp. nov. B mantle extensions (shell lobes and dorsal lobes) and posterior body of B. poongee C dorsal view of mantle extensions (shell lobes and dorsal lobes) of B. cassidula D, E representative SEM images of radula showing monocuspid central tooth (yellow) and lateromarginal teeth (blue) attached by the smaller triangular-shaped teeth (red). White arrow indicates pneumostome or breathing pore. Yellow arrow indicates a tiny cusp on outermost lateromarginal teeth." figureDoi="10.3897/zookeys.1110.82461.figure3" httpUri="https://binary.pensoft.net/fig/709569" pageId="0" pageNumber="1">3A</figureCitation>
,
<figureCitation id="5AE56BE87BD0D045EB1333C27CA0D681" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Living snails of Burmochlamys species A B. fasciola sp. nov. paratype CUMZ 14214 showing left and right lateral views B B. cassidula specimen CUMZ 14209 C B. poongee specimen CUMZ 14210 D B. albida sp. nov. paratype CUMZ 14212 E B. moulmeinica sp. nov. paratype CUMZ 14217 F B. versicolor sp. nov. paratype CUMZ 14219 G B. whitteni sp. nov. paratype CUMZ 14222." figureDoi="10.3897/zookeys.1110.82461.figure5" httpUri="https://binary.pensoft.net/fig/709571" pageId="0" pageNumber="1">5A</figureCitation>
). Living animal with reticulated skin and pale freshy-grey body. Five mantle extensions present, same colour as body. Sole divided into three parts longitudinally. Caudal foss (cf) and caudal horn (ch) present with similar colour to body.
</paragraph>
</subSubSection>
<subSubSection id="50488F00748534C572F59F47A1D88B11" pageId="0" pageNumber="1" type="etymology">
<paragraph id="BBC11E3925E229D675B8A82413BD0DFF" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="C0B46208C1C2D2CD94894F74AC90B6A9" pageId="0" pageNumber="1">
The specific epithet is the Latin word
<emphasis id="219C5C247B1C7F57B25AACC0B8A9CED8" italics="true" pageId="0" pageNumber="1">fasciola</emphasis>
meaning band or stripe. It refers to the presence of a brownish peripheral band, which characterises this species.
</paragraph>
</subSubSection>
<subSubSection id="985AE28F34EB409B52F99CCCFF3ECD36" pageId="0" pageNumber="1" type="distribution">
<paragraph id="09283896F8E58D5B57F803B69F6B0A04" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="F1263FC913E513624A8F3FDC6A5C491A" pageId="0" pageNumber="1">
<taxonomicName id="30B8AE0524DC94958F1D8E752689D9FD" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys fasciola" order="Eupulmonata" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis id="CFCBFBE1DE08780863DDE041CFD50B7A" italics="true" pageId="0" pageNumber="1">Burmochlamys fasciola</emphasis>
</taxonomicName>
sp. nov. is known from two limestone areas in the south of the Salween River basin (Fig.
<figureCitation id="DC4B0CAC148D68078A9DE67E31FDE9AF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of the Salween River basin (known as Thanlwin River) in the southeastern Myanmar showing the sampling sites. Coloured triangles indicate the localities recorded for each Burmochlamys species. White circles indicate the localities recorded for Sophina species. White squares indicate the sampling localities with no records of both Sophina and Burmochlamys species." figureDoi="10.3897/zookeys.1110.82461.figure1" httpUri="https://binary.pensoft.net/fig/709567" pageId="0" pageNumber="1">1</figureCitation>
). The Bardai Mountain (Fig.
<figureCitation id="FFA20FECD2709A0E9BFB37EFB3A5C1A7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitat of some species of Burmochlamys gen. nov. in the karst basin of Hpa-An, Kayin State, Myanmar. A type locality of B. fasciola sp. nov. at Bardai Mountain and microhabitat structure of the karst wall B newly discovered locality of B. poongee at Kaw Ka Thaung Cave C type locality of B. albida sp. nov. at Waiponla Monastery D type locality of B. moulmeinica sp. nov. at Lun Nya Pagoda E type locality of B. versicolor sp. nov. at Bayin Nyi Cave. Photos by Ruttapon Srisonchai." figureDoi="10.3897/zookeys.1110.82461.figure2" httpUri="https://binary.pensoft.net/fig/709568" pageId="0" pageNumber="1">2A</figureCitation>
) and Kyankaw Mountain are surrounded by paddy fields that are temporarily flooded during the monsoon season. In Kyankaw Mountain, this new species is sympatric with the limestone karst-restricted land snail species,
<taxonomicName id="EF53434CB467274212381F5DF3379FF8" authorityName="Sutcharit &amp; Panha" authorityYear="2020" class="Gastropoda" family="Helicarionidae" genus="Sophina" kingdom="Animalia" lsidName="Sophina salweenica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="salweenica">
<emphasis id="EEDD4F41E70399A681C42EF430DAEFEA" italics="true" pageId="0" pageNumber="1">Sophina salweenica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3A2E6C1FE863A1F7C54EB0F94060C483" pageId="0" pageNumber="1" type="remarks">
<paragraph id="9E81D11C18516249FBBBD1139AC85E33" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="7F3AEE27AAB2592F256A8210319ED427" pageId="0" pageNumber="1">
The distinguishing characters of
<taxonomicName id="E4E47E3387BD75C23231493BA3672481" lsidName="B. fasciola" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis id="E6216BFD19FA80C94071886723091C78" italics="true" pageId="0" pageNumber="1">B. fasciola</emphasis>
</taxonomicName>
sp. nov. is a milky-white shell with a narrow yellowish brown peripheral band and rather elongated radular teeth. Whereas the other
<taxonomicName id="1124E7C5A0D22B708C07853C206D940F" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="F47FC5DC71F55618040834C60D512343" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
species have a pale to dark brownish shell without any band and the radular teeth have a broad spatulate shape (see Table
<tableCitation id="1FAEAAEE2A93663656EEEFD591808DD1" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of shell, radula, genital system, and distribution of all members of Burmochlamys gen. nov. in Myanmar." httpUri="http://table.plazi.org/id/A3EF01A52C7ECD353E45707D9F7BAEB8" pageId="0" pageNumber="1" tableUuid="A3EF01A52C7ECD353E45707D9F7BAEB8">2</tableCitation>
).
</paragraph>
<caption id="A3EF01A52C7ECD353E45707D9F7BAEB8" ID-Table-UUID="A3EF01A52C7ECD353E45707D9F7BAEB8" httpUri="http://table.plazi.org/id/A3EF01A52C7ECD353E45707D9F7BAEB8" pageId="0" pageNumber="1" start="Table 2" startId="T2">
<paragraph id="25816BC68FBDC858D31C492D17CD90D4" pageId="0" pageNumber="1">
<emphasis id="2097D8EB891CC71A385CB56A82CE3DDA" bold="true" pageId="0" pageNumber="1">Table 2.</emphasis>
Comparison of shell, radula, genital system, and distribution of all members of
<taxonomicName id="853B818447AD6C350D5CF7AEADBE164A" authorityName="Pholyotha &amp; Panha" authorityYear="2022" family="Helicarionidae" genus="Burmochlamys" kingdom="Animalia" lsidName="Burmochlamys" order="Eupulmonata" pageId="0" pageNumber="1" rank="genus">
<emphasis id="02D910A975B4DA4DF2A5494DFAA7BC91" italics="true" pageId="0" pageNumber="1">Burmochlamys</emphasis>
</taxonomicName>
gen. nov. in Myanmar.
</paragraph>
</caption>
<paragraph id="A5FF902382F2B4B2083DC4AE558B3963" pageId="0" pageNumber="1">
<table id="A6BE9D16C296F1FF864F0A383ABF1A2E" pageId="0" pageNumber="1">
<tr id="5A962659809C7053C10ECA8CEE3273BD" pageId="0" pageNumber="1">
<th id="D31354B326974FF33C20EA0F50D7A8B0" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</th>
<th id="BD65AB25104A8E41D3BB8DB7C2BFEE2A" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="5EE08F230FC905CE271E72980D28587E" lsidName="B. cassidula" pageId="0" pageNumber="1" rank="species" species="cassidula">
<emphasis id="581D3B7B83CB026F505D0E3CA07E23E1" italics="true" pageId="0" pageNumber="1">B. cassidula</emphasis>
</taxonomicName>
</th>
<th id="5FA799F262C477979A5D5F1B365FE11C" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="10B2363AAE1EF98208921836FB203AC1" lsidName="B. cauisa" pageId="0" pageNumber="1" rank="species" species="cauisa">
<emphasis id="9D9814DC93A33179CFE7C455E7A0A83C" italics="true" pageId="0" pageNumber="1">B. cauisa</emphasis>
</taxonomicName>
*
</th>
<th id="4B35CEC5B7F402BF2340855E3588E30C" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="B1CA1BEC59002E4473A79C4BCEF0D42F" lsidName="B. perpaula" pageId="0" pageNumber="1" rank="species" species="perpaula">
<emphasis id="CF3DA1511C637920D835917F88BB9EDA" italics="true" pageId="0" pageNumber="1">B. perpaula</emphasis>
</taxonomicName>
*
</th>
<th id="A4F99D3272626557C1C5A85B1386C6F8" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="A985D4CCA73EB9F4710199C14EACC474" lsidName="B. poongee" pageId="0" pageNumber="1" rank="species" species="poongee">
<emphasis id="B2C9A1EEA9DFB77D7A00A6AF737606DA" italics="true" pageId="0" pageNumber="1">B. poongee</emphasis>
</taxonomicName>
</th>
<th id="CD66609E0758E92D650E8454680B07BE" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="7BE8BFED2701510FC41DE9E65B26275E" lsidName="B. albida" pageId="0" pageNumber="1" rank="species" species="albida">
<emphasis id="9740AD9F7CC42B72646BB537A768C63D" italics="true" pageId="0" pageNumber="1">B. albida</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="FB620B07CBCDABC15246A5270ED2D3D6" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="1F2F5BED30E60BDA00F0B11225FFB1B3" lsidName="B. fasciola" pageId="0" pageNumber="1" rank="species" species="fasciola">
<emphasis id="CD3C09C5CD1AAAC6E2B4155306A74767" italics="true" pageId="0" pageNumber="1">B. fasciola</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="31B3BDC1979BA5B38DE10C5CB068E418" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="7C43388DF26A5A0E432CFE03DE38B709" lsidName="B. moulmeinica" pageId="0" pageNumber="1" rank="species" species="moulmeinica">
<emphasis id="49C4506CEE7B04DC080DEBDAD1091809" italics="true" pageId="0" pageNumber="1">B. moulmeinica</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="51B63EE6AE64A61FCA4AAFFC903829DB" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="475FB011CD811DC2F01A4A53A5B8B966" lsidName="B. versicolor" pageId="0" pageNumber="1" rank="species" species="versicolor">
<emphasis id="F7C77F258892E055756049493B611BF0" italics="true" pageId="0" pageNumber="1">B. versicolor</emphasis>
</taxonomicName>
sp. nov.
</th>
<th id="E487D7A3BDD36954ACC5CD6A801C4FDA" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="448C1D061B95B151F64507C256A362A2" lsidName="B. whitteni" pageId="0" pageNumber="1" rank="species" species="whitteni">
<emphasis id="9A9AAADB3EDE055126EDB18F02437D84" italics="true" pageId="0" pageNumber="1">B. whitteni</emphasis>
</taxonomicName>
sp. nov.
</th>
</tr>
<tr id="B1F3689B092335945B7920A465F640B3" pageId="0" pageNumber="1">
<td id="2A09BE3632DB7126E207F952AC1464F0" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="99C452433A492F8A30724B43BB7C4ECD" bold="true" pageId="0" pageNumber="1">shell colour</emphasis>
</td>
<td id="7AD6AC4990EB16088A4EACE3B23FF1DB" colspan="1" pageId="0" pageNumber="1" rowspan="1">whitish with a yellow tinge</td>
<td id="6500BE9B7D0B30ECBD7F593F324FB940" colspan="1" pageId="0" pageNumber="1" rowspan="1">pale yellowish brown</td>
<td id="62C34B030A1673A991D92DA8B42564FC" colspan="1" pageId="0" pageNumber="1" rowspan="1">dark brown</td>
<td id="3EE1595BC3FCC3A82D65EAA66A7FA640" colspan="1" pageId="0" pageNumber="1" rowspan="1">pale to dark brown</td>
<td id="06DAB42988B6B05DEB9E93F76167DE25" colspan="1" pageId="0" pageNumber="1" rowspan="1">whitish with a yellow tinge</td>
<td id="84FA08A14B43E695D05FC895D6E5C13D" colspan="1" pageId="0" pageNumber="1" rowspan="1">whitish</td>
<td id="82B5D896FBA6802B714D50F16821A9E1" colspan="1" pageId="0" pageNumber="1" rowspan="1">pale to dark brown</td>
<td id="3A4BE647E7C28C84EAECEEEF91DF1226" colspan="1" pageId="0" pageNumber="1" rowspan="1">whitish yellow / yellowish brown</td>
<td id="7D082316D123918038BAB0552F8305E8" colspan="1" pageId="0" pageNumber="1" rowspan="1">pale to dark brown</td>
</tr>
<tr id="B9867386A3FB4FD111FDA19B18145C52" pageId="0" pageNumber="1">
<td id="3BD7CC1B33F9C8D33F18C472B4F42FC9" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="B77143F4A3B85F5F35909A5C72315A2C" bold="true" pageId="0" pageNumber="1">peripheral band</emphasis>
</td>
<td id="6949BB3E23E9F03CAB25692934B7B2C9" colspan="1" pageId="0" pageNumber="1" rowspan="1">wide</td>
<td id="E6709CD1877CEA0EBAA19C4196B66A83" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="FA24139BC262C00727334B02871529F5" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="CC2EC5ABC8B78DA0458FF9A10CBE84A0" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="E6EAC7FB6C6C652F14181F535B0DEB69" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent or rarely indistinct</td>
<td id="9A700753C7B78E1515D7D9AF18D087FC" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow</td>
<td id="A89CD3D77AE8EAD995FF7A83B78151C9" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="1C089629846ADA7A143E4AA3FCB43CBE" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="AA1FF62AC20A2BB8BC23F1586BEE98F1" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
</tr>
<tr id="06ECE4BC3AD6242EF4556D78134799DB" pageId="0" pageNumber="1">
<td id="1F460B1D291A30704465725D59A482D5" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="44751BBA5C608D970A6B4836D15A25F8" bold="true" pageId="0" pageNumber="1">shell shape</emphasis>
</td>
<td id="F2826184F3761FE0BBCF0460BE893999" colspan="1" pageId="0" pageNumber="1" rowspan="1">globose</td>
<td id="00BBB4399ED1B54FB6DB0BDCF49087F8" colspan="1" pageId="0" pageNumber="1" rowspan="1">subglobose</td>
<td id="BA30A92A93DF2671FCF756FAEA5323DC" colspan="1" pageId="0" pageNumber="1" rowspan="1">subglobose</td>
<td id="921D71531A1F22C287C1DAD269419C40" colspan="1" pageId="0" pageNumber="1" rowspan="1">globose</td>
<td id="E69656FB2302F9444D43D2320751A036" colspan="1" pageId="0" pageNumber="1" rowspan="1">globose</td>
<td id="89E63610ADFE594372145CFCB062A3B1" colspan="1" pageId="0" pageNumber="1" rowspan="1">globose</td>
<td id="99739BFBD24ADA6A9A171239DFB6DCE6" colspan="1" pageId="0" pageNumber="1" rowspan="1">globose</td>
<td id="7F1A74AE28311692C325AE52CE736153" colspan="1" pageId="0" pageNumber="1" rowspan="1">subglobose</td>
<td id="D8494C13F3F542FD96E2FC2BC86540C0" colspan="1" pageId="0" pageNumber="1" rowspan="1">globose</td>
</tr>
<tr id="64673433E4A381D8AB7B75190C6B3BB5" pageId="0" pageNumber="1">
<td id="0A3551AF716B983CBD13D8F21A431A40" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="7A3AE95FE03E5014841D2CA913ACB703" bold="true" pageId="0" pageNumber="1">microscopic sculpture</emphasis>
</td>
<td id="F27A8B3AC8DB908D6EEF2337E30A89DF" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="2DF29FE1778A23022F3AF1C92DFDDCCB" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="98A92F5DEB1F2D83BE4BF567B6A7DC8F" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="774330F34A15A77135315D9EB284667E" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="CF72152592D96FC8982D44CFCA15CB74" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="84B0843BFB90380D7F0C3C33E8C5BF9D" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="4B137CEEEDE62A05565056C5C0BF55D2" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
<td id="C8341809CBC509A0CF4B2B8AEAA7349E" colspan="1" pageId="0" pageNumber="1" rowspan="1">present but indistinct</td>
<td id="786F3728B1FF97BDF7861CF0D5652A24" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
</tr>
<tr id="B5054DE23E3A25C25A23704C4E79E384" pageId="0" pageNumber="1">
<td id="85CA45F6F5B39D16800A65994786D374" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="BF8538684AACB9061E198D09A203432C" bold="true" pageId="0" pageNumber="1">umbilicus</emphasis>
</td>
<td id="4E2356916696D4A042A70D16364F1FDE" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow</td>
<td id="7050E78DCF437990DE04690BFF8E564E" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow</td>
<td id="1839AF92B5BDD00613D59D4B520A754F" colspan="1" pageId="0" pageNumber="1" rowspan="1">very narrow</td>
<td id="9B6E701F3D8B2B29DD7213BE45FEDF30" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow</td>
<td id="5A317101537830B7A7A608360D14096C" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow</td>
<td id="0CAF2BA6BB3385BCC06F71AABCE6290E" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow</td>
<td id="F2E14340D3537D889BA86A9A355931BA" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow</td>
<td id="3A24ED9A2F9303542FB660062DA53C38" colspan="1" pageId="0" pageNumber="1" rowspan="1">moderate</td>
<td id="89B9D1C6915DDA40EDA103B71F5040F4" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow</td>
</tr>
<tr id="CDBF2AE5F6EE5A84223D1BD9BA8491E0" pageId="0" pageNumber="1">
<td id="065CAEB571AD061FE034CF16CB7AA692" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="171C50C0347FAA46ED2EF9B4FCDD9CA1" bold="true" pageId="0" pageNumber="1">central tooth shape</emphasis>
</td>
<td id="50D07C38D331465B8E646D6DAF6D458B" colspan="1" pageId="0" pageNumber="1" rowspan="1">broad spatulate</td>
<td id="B661C7E5FC469C4F7B41BAFC333EBAB6" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="CB5E78BF0EBD2188AF0549960B593745" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="1150610ACD9D02D64C740DCD889966E4" colspan="1" pageId="0" pageNumber="1" rowspan="1">broad spatulate</td>
<td id="3709CB0D1B6210FB49062695F5166FBA" colspan="1" pageId="0" pageNumber="1" rowspan="1">broad spatulate</td>
<td id="637627C2178DB6D01F300467BD6FBD20" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow spatulate</td>
<td id="F3B72B1731038D2059A883B881BBF7D8" colspan="1" pageId="0" pageNumber="1" rowspan="1">broad spatulate</td>
<td id="7A7ED89EBFE9BBA163CE913FBC09A7DE" colspan="1" pageId="0" pageNumber="1" rowspan="1">narrow spatulate</td>
<td id="CCF3C4BA823519C4AD9183C9B6A9B309" colspan="1" pageId="0" pageNumber="1" rowspan="1">broad spatulate</td>
</tr>
<tr id="6FA877B3E629ED3ADB7D5492E0A94B1A" pageId="0" pageNumber="1">
<td id="C4577142665D37526DC944552B31E4F7" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="EF6AC4695B16BDB50393489CFECC8219" bold="true" pageId="0" pageNumber="1">penis</emphasis>
</td>
<td id="8A9CA4C88D165E02219CD20183A7EC86" colspan="1" pageId="0" pageNumber="1" rowspan="1">rather short</td>
<td id="E02987A38624C75F2E3C19586C6584D3" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="984CAC6313DB6801265826FF2F0BE8FB" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="7922F93AB1A73837698F34D79C230A1B" colspan="1" pageId="0" pageNumber="1" rowspan="1">moderate</td>
<td id="17161A2E96EE0F88DDF6E08910283E57" colspan="1" pageId="0" pageNumber="1" rowspan="1">moderate</td>
<td id="1A72B9CBF6BD5BE19608E12E0FB87B76" colspan="1" pageId="0" pageNumber="1" rowspan="1">moderate</td>
<td id="0CC97C1E969065EE15A8987BFB452DF3" colspan="1" pageId="0" pageNumber="1" rowspan="1">moderate</td>
<td id="773F1DDA4200C9104D63E4F7AB250FA8" colspan="1" pageId="0" pageNumber="1" rowspan="1">moderate</td>
<td id="78F3BAB1C96E68A165420FFB314BF83C" colspan="1" pageId="0" pageNumber="1" rowspan="1">rather long</td>
</tr>
<tr id="E30BC74A29FDF75D7AE08F61B206E0F5" pageId="0" pageNumber="1">
<td id="5623BC6C3563192688949838417E3CB3" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="D326CA2890724298C5E52CDAF7EB5414" bold="true" pageId="0" pageNumber="1">penial retractor muscle</emphasis>
</td>
<td id="AFCD2B342769BCF8F39A663AFAAB8D1B" colspan="1" pageId="0" pageNumber="1" rowspan="1">thin</td>
<td id="E549D9C54748A08D300B354B0CAD64D7" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="3DD7787563E32A6C229FBECB54E868DA" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="FC4C3B3032578E3A8787C0E76DC9AD88" colspan="1" pageId="0" pageNumber="1" rowspan="1">thin</td>
<td id="36C47D1D281E47806733276D7CCE158E" colspan="1" pageId="0" pageNumber="1" rowspan="1">thick</td>
<td id="440F79F99EEEF97B8EFD108F05C03DFB" colspan="1" pageId="0" pageNumber="1" rowspan="1">thin</td>
<td id="498A526ED6181FA03316096A693B09DB" colspan="1" pageId="0" pageNumber="1" rowspan="1">thin</td>
<td id="2D761C4BCA1B2C27FBB536CC827B974E" colspan="1" pageId="0" pageNumber="1" rowspan="1">thin</td>
<td id="F00240A56390356021FB73F6A499DF67" colspan="1" pageId="0" pageNumber="1" rowspan="1">thick</td>
</tr>
<tr id="B395D35CCEBCFDA6AB332E564BB36166" pageId="0" pageNumber="1">
<td id="8B0AE070E554F52B4335EA6432A6010C" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="7AFBE5307451CFF8B136EB38225AAF19" bold="true" pageId="0" pageNumber="1">vagina</emphasis>
</td>
<td id="F93CCC3ACFD76DDF64426B70BBC18A3F" colspan="1" pageId="0" pageNumber="1" rowspan="1">very short</td>
<td id="9C24CCB297921D2E1F183383B20EE60E" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="99547E51C608B4685D441E9C66BF0CAB" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="2A0A4D1925C89A5CA5934C949D8A9F52" colspan="1" pageId="0" pageNumber="1" rowspan="1">short</td>
<td id="E93538D30F709749A05354A50A3D30FD" colspan="1" pageId="0" pageNumber="1" rowspan="1">short</td>
<td id="C5DE75ABE3F034FC71EAC9F2CF2115A1" colspan="1" pageId="0" pageNumber="1" rowspan="1">very short</td>
<td id="329214800E9287128CBDC8049E7B67EA" colspan="1" pageId="0" pageNumber="1" rowspan="1">short</td>
<td id="D0E552D17A996FC2E7760ABE1633F661" colspan="1" pageId="0" pageNumber="1" rowspan="1">short</td>
<td id="F63E934F508FD8DC471E4A8B18B92061" colspan="1" pageId="0" pageNumber="1" rowspan="1">very short</td>
</tr>
<tr id="0D79497A7F8DE3ED169732B25AB4AA52" pageId="0" pageNumber="1">
<td id="071E53E20F11C7A7176C4D12191A37F9" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="C02A59D9763BEC01120BABC0EFAC6AF4" bold="true" pageId="0" pageNumber="1">the tissue surrounding gametolytic part and free oviduct</emphasis>
</td>
<td id="85BBC2CA3D74D9B48DD5EE3B14079A8E" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="A209C671FFE5BCFE83A27D3ACDBCB99E" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="2D84E6A37A3A8ED6EF927D43E01B4612" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="3511DC7AB494730421620590BEAF0D4D" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="A4A3A3FC7DBA7CB0AC909DF0D1902795" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="4BBBDA985DCDD672E403E3A9C307CC1E" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="BF86161ADF223DECDF087A8DCC30867E" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="D4AE32E38C0DDE731FD51CC43A8FE58B" colspan="1" pageId="0" pageNumber="1" rowspan="1">absent</td>
<td id="E902D98E48AAD96115EFB0FBF4C9F2FB" colspan="1" pageId="0" pageNumber="1" rowspan="1">present</td>
</tr>
<tr id="3B9AA54BC0043600E10BCBFB98F4A24A" pageId="0" pageNumber="1">
<td id="201C659C741A8636EB5CE7DBEB2CECA3" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="F034EF44A09F485C2C66E84EAF637EF2" bold="true" pageId="0" pageNumber="1">dart apparatus near atrium</emphasis>
</td>
<td id="3C7BD17A888DF1EF27BC38C1B5A3B64A" colspan="1" pageId="0" pageNumber="1" rowspan="1">rather long</td>
<td id="E4530DD1F6C799FB0546EE47D4242645" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="6A75A25BB2796D7472FBDC89C6F90C0E" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="12384EE267EE72E25A217A36713FE038" colspan="1" pageId="0" pageNumber="1" rowspan="1">very long</td>
<td id="0FCB7F5E0EFDA609D48AC6B4C7FCB094" colspan="1" pageId="0" pageNumber="1" rowspan="1">rather long</td>
<td id="5BF3AEF9285F1F7B385A1FF8C3270AA8" colspan="1" pageId="0" pageNumber="1" rowspan="1">rather short</td>
<td id="89FDE83B4372CF3C834B1CDE0D88067B" colspan="1" pageId="0" pageNumber="1" rowspan="1">rather short</td>
<td id="54F140381E504CDEE8800D58D06170CC" colspan="1" pageId="0" pageNumber="1" rowspan="1">rather short</td>
<td id="774B4BDB5A3B62C8C65DB74BE4142B01" colspan="1" pageId="0" pageNumber="1" rowspan="1">rather long</td>
</tr>
<tr id="C6AB97B02EB74589D3ED4404E336964D" pageId="0" pageNumber="1">
<td id="7B581BB1BDE4DB68E7D3178319C346DF" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis id="017C7E0EACE7139254CD78150E654B23" bold="true" pageId="0" pageNumber="1">at the tip of dart apparatus</emphasis>
</td>
<td id="B070B8F06B7B0FA07A4782016FFC0A58" colspan="1" pageId="0" pageNumber="1" rowspan="1">soft</td>
<td id="4A94609C25EDCD48D185E02A85DE1D15" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="366E5897B52F117D725927FFDD7F8976" colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="F81A3F9200B3914EC4828417D4777BFE" colspan="1" pageId="0" pageNumber="1" rowspan="1">soft</td>
<td id="904F3B4D2E3AB845D50C8E4ACA09ED88" colspan="1" pageId="0" pageNumber="1" rowspan="1">soft</td>
<td id="0C6B16BCABE4F7209168D72A37B4A5B6" colspan="1" pageId="0" pageNumber="1" rowspan="1">solid</td>
<td id="C162D60E9903391AFC089E32AC6FB0C6" colspan="1" pageId="0" pageNumber="1" rowspan="1">soft</td>
<td id="617AFD74529CF91B48ABAB04AA85AEBA" colspan="1" pageId="0" pageNumber="1" rowspan="1">soft</td>
<td id="5B1FC8B5F0907397B0C30D8EB4485934" colspan="1" pageId="0" pageNumber="1" rowspan="1">soft</td>
</tr>
</table>
</paragraph>
<paragraph id="B55B7AF750D29CE7BA83C800680C3C83" pageId="0" pageNumber="1">
<tableNote id="EC9EBF51E632D75B9F8D0ACD6814E7B0" pageId="0" pageNumber="1">
*Information from
<bibRefCitation id="28F0CBFFE08EDF657A4DB839EF6FC694" DOI="https://doi.org/10.5962/bhl.title.13103" author="Blanford, WT" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B3" refString="Blanford, WT, Godwin-Austen, HH, 1908. The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae. Taylor and Francis, London, DOI: https://doi.org/10.5962/bhl.title.13103" title="The Fauna of British India, including Ceylon and Burma. Mollusca. Testacellidae and Zonitidae." url="https://doi.org/10.5962/bhl.title.13103" year="1908">Blanford and Godwin-Austen (1908)</bibRefCitation>
.
</tableNote>
</paragraph>
</subSubSection>
</treatment>
</document>