144 lines
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144 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.531.6097" ID-GBIF-Dataset="70702787-a6a8-4af7-9702-738f344a5cbe" ID-PMC="PMC4668911" ID-Pensoft-Pub="1313-2970-531-1" ID-PubMed="26692803" ID-ZBK="C845838EC9124BD8AB4E07980F91959E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-531-1" ModsDocOrigin="ZooKeys 531" ModsDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" checkinTime="1451243834596" checkinUser="pensoft" docAuthor="Vermeulen, Jaap J., Liew, Thor-Seng & Schilthuizen, Menno" docDate="2015" docId="AB8CEA057B70A37BB22002F52E065A1C" docLanguage="en" docName="ZooKeys 531: 1-139" docOrigin="ZooKeys 531" docSource="http://dx.doi.org/10.3897/zookeys.531.6097" docTitle="Kaliella calculosa Gould 1852" docType="treatment" docVersion="9" lastPageNumber="78" masterDocId="1527DF5A6645C540FFF1FF85AF4AE358" masterDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" masterLastPageNumber="139" masterPageNumber="1" pageNumber="76" updateTime="1668160884494" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liew, Thor-Seng</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>531</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>139</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.531.6097</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.531.6097</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-531-1</mods:identifier>
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<mods:identifier type="ZBK">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
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<mods:identifier type="ZooBank">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="144057554" LSID="urn:lsid:plazi:treatment:AB8CEA057B70A37BB22002F52E065A1C" httpUri="http://treatment.plazi.org/id/AB8CEA057B70A37BB22002F52E065A1C" lastPageId="77" lastPageNumber="78" pageId="75" pageNumber="76">
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<subSubSection pageId="75" pageNumber="76" type="multiple">
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<paragraph pageId="75" pageNumber="76">Taxon classification Animalia Stylommatophora Euconulidae</paragraph>
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</subSubSection>
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<subSubSection pageId="75" pageNumber="76" type="nomenclature">
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<paragraph pageId="75" pageNumber="76">
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<taxonomicName authority="Gould, 1852" authorityName="Gould" authorityYear="1852" class="Gastropoda" family="Chronidae" genus="Kaliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaliella calculosa" order="Stylommatophora" pageId="75" pageNumber="76" phylum="Mollusca" rank="species" species="calculosa">Kaliella calculosa (Gould, 1852)</taxonomicName>
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Figure 65, 68
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</paragraph>
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</subSubSection>
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<subSubSection pageId="76" pageNumber="77" type="reference_group">
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<paragraph pageId="76" pageNumber="77">
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix calculosa" order="Stylommatophora" pageId="76" pageNumber="77" phylum="Mollusca" rank="species" species="calculosa">
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<pageBreakToken pageId="76" pageNumber="77" start="start">Helix</pageBreakToken>
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calculosa
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</taxonomicName>
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Gould, 1852: 48.
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<taxonomicName class="Gastropoda" family="Euconulidae" genus="Coneuplecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coneuplecta calculosa" order="Stylommatophora" pageId="76" pageNumber="77" phylum="Mollusca" rank="species" species="calculosa">Coneuplecta calculosa</taxonomicName>
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(Gould) H. Burrington Baker, 1941: 234;
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<bibRefCitation author="Solem, A" journalOrPublisher="Invertebrate Taxonomy" pageId="76" pageNumber="77" pagination="455 - 603" title="Non-Camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges." url="10.1071/IT9880455" volume="2" year="1988">Solem 1988</bibRefCitation>
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: 544.
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<taxonomicName class="Gastropoda" family="Chronidae" genus="Kaliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaliella calculosa" order="Stylommatophora" pageId="76" pageNumber="77" phylum="Mollusca" rank="species" species="calculosa">Kaliella calculosa</taxonomicName>
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(Gould) Schilthuizen, 2004: 94;
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<bibRefCitation author="Clements, R" journalOrPublisher="Biological Conservation" pageId="76" pageNumber="77" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
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: 2762;
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<bibRefCitation author="Liew, TS" journalOrPublisher="Journal of Biogeography" pageId="76" pageNumber="77" pagination="1071 - 1078" title="The determinants of land snail diversity along a tropical elevational gradient: insularity, geometry and niches." url="10.1111/j.1365-2699.2009.02243.x" volume="37" year="2010">Liew et al. 2010</bibRefCitation>
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: Appendix S1 in online Supporting Information;
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<bibRefCitation author="Schilthuizen, M" journalOrPublisher="Journal of the Geological Society, London" pageId="76" pageNumber="77" pagination="539 - 545" title="Species diversity patterns in insular land snail communities of Borneo." url="10.1144/jgs2012-014" volume="170" year="2013">Schilthuizen et al. 2013</bibRefCitation>
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: online supplementary data. Type from Society Islands, Tahiti.
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</paragraph>
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<paragraph pageId="76" pageNumber="77">
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<taxonomicName class="Gastropoda" family="Helicarionidae" genus="Sitala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sitala demissa" order="Stylommatophora" pageId="76" pageNumber="77" phylum="Mollusca" rank="species" species="demissa">Sitala demissa</taxonomicName>
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E.A.
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<bibRefCitation author="Smith, EA" journalOrPublisher="London" pageId="76" pageNumber="77" pagination="97 - 127" title="On a collection of land-shells from Sarawak, British North Borneo, Palawan, and other neighbouring islands." volumeTitle="Proceedings of the Zoological Society" year="1895">Smith 1895</bibRefCitation>
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: 110. Type from Malaysia, Sarawak, Mount Mulu and Busau.
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</paragraph>
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<paragraph pageId="76" pageNumber="77">
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<taxonomicName class="Gastropoda" family="Euconulidae" genus="Coneuplecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coneuplecta sitaliformis" order="Stylommatophora" pageId="76" pageNumber="77" phylum="Mollusca" rank="species" species="sitaliformis">Coneuplecta sitaliformis</taxonomicName>
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auct.
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<bibRefCitation author="Schilthuizen, M" journalOrPublisher="Journal of Molluscan Studies" pageId="76" pageNumber="77" pagination="255 - 258" title="Microsnails at microscales in Borneo: distributions of Prosobranchia versus Pulmonata." url="10.1093/mollus/68.3.255" volume="68" year="2002">Schilthuizen et al. 2002</bibRefCitation>
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: 256.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="materials_examined">
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<paragraph pageId="76" pageNumber="77">Examined material from Sabah.</paragraph>
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<paragraph lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
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Interior Province. Upper Padas valley, Long Pasia (leg. T.S. Liew & Meckson, BOR/MOL 4357). Crocker Range N.P., West of the km 10 marker on the road Tambunan-Ranau, Mahua Waterfall (leg. J.J. Vermeulen & M. Schilthuizen, V 9736; leg. M. Schilthuizen, BOR/MOL 814). Gunung Trusmadi slopes, Gua Loloposon (leg. J.J. Vermeulen, V 13234). Pun Batu c. 30 km West of Sepulut (leg. J.J. Vermeulen, V 1282). Sepulut valley, Batu Punggul (leg. J.J. Vermeulen, V 1985); Batu Temurung (leg. J.J. Vermeulen, V 8050); Gua Pungiton (leg. J.J. Vermeulen & M. Schilthuizen, V 7560). Sandakan Province. Kinabatangan valley, Gomantong (leg. A. Van Til, BOR/MOL 3269); Batu Keruak (leg. M. Salverda & H. van Oosten, BOR/MOL 2358); Batu Materis (leg. T.S. Liew & B. Elahan, BOR/MOL 2121); Batu Tomanggong Besar (leg. T.S. Liew & B. Elahan, BOR/MOL 2258; leg. M. Schilthuizen, A. van Til & B. Elahan, BOR/MOL 2972); Batu Tomanggong Kecil (leg. T.S. Liew & B. Elahan, BOR/MOL 2058); Unnamed hill near Sukau Police Station (leg. T.S. Liew & B. Elahan, BOR/MOL 2225). Tawau Province. Batu Baturong c. 50 km W.S.W. of Lahad Datu (leg. J.J. Vermeulen & H. Duistermaat, V 14247); Danum Valley (leg. H.A. Rutjes, BOR/MOL 813, BOR/MOL 815; leg. UMS students, BOR/MOL 818). Segama valley,
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<normalizedToken originalValue="‘Kirk’s" pageId="76" pageNumber="77">'Kirk's</normalizedToken>
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<normalizedToken originalValue="Cave’" pageId="76" pageNumber="77">Cave'</normalizedToken>
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8 km North of Lahad Datu (leg. J.J. Vermeulen, V 14248). West Coast Province. Kinabalu N.P., Poring Hot Springs, Garden near hot springs (leg. J.J. Vermeulen, V 13023); Orchid Garden (leg. J.J. Vermeulen, V 13025); Serinsim (leg. M. Schilthuizen, BOR/MOL 3076); Mesilau
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<pageBreakToken pageId="77" pageNumber="78" start="start">trail</pageBreakToken>
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at 2112 m (leg. T.S. Liew, BOR/MOL 6042). Kota Kinabalu, Kiansom (leg. UMS students, BOR/MOL 816), Papar, Kg. Langsat (leg. T.S. Liew, M. Schilthuizen & F. Disuk, BOR/MOL 3095).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="description">
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<paragraph pageId="77" pageNumber="78">Description.</paragraph>
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<paragraph pageId="77" pageNumber="78">Shell small, very thin, translucent, pale yellowish-green to pale yellowish-brown, conical with about flat sides. Surface with a silky luster, shiny below the periphery. Whorls somewhat convex, last whorl almost rounded to angular at the periphery, moderately convex below the periphery. Protoconch sculpture: distinct, fine, densely placed radial riblets; spiral sculpture slightly subordinate, equally densely placed or slightly wider spaced grooves cutting into the crests of the radial riblets. Teleoconch: Last whorl with or without a thin peripheral spiral thread coinciding with the suture of the penultimate whorl, above and below this with very fine to rather coarse, moderately to widely spaced, continuous, sometimes rather shallow and wide grooves cutting through the radial riblets. Radial sculpture teleoconch subordinate: above the periphery inconspicuous growth lines, next to these very fine, densely and regularly placed riblets, sometimes on the outer whorls locally interrupted by approximately smooth areas; below the periphery with occasional growth lines only. Umbilicus open, very narrow. Dimensions: Height up to 5.0 mm; width up to 4.6 mm; diameters of the first four whorls 0.65-0.75 mm, 1.15-1.25 mm, 1.75-2.10 mm, 2.6-3.2 mm respectively; number of whorls up to c. 5 3/4; height aperture up to 2.5 mm; width aperture up to 2.7 mm.</paragraph>
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<caption pageId="77" pageNumber="78">
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<paragraph pageId="77" pageNumber="78">
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Figure 68-69. 68
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<taxonomicName class="Gastropoda" family="Chronidae" genus="Kaliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaliella calculosa" order="Stylommatophora" pageId="77" pageNumber="78" phylum="Mollusca" rank="species" species="calculosa">Kaliella calculosa</taxonomicName>
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(Gould, 1852). 68A Frontal view, subadult, shell 3,6 mm high 68B Frontal view, shell 4.0 mm high 68C Umbilical view, subadult (68A, 68C Malaysia, Sarawak, Gunung Mulu N.P., Gunung Api, path from camp to pinnacles, incl. pinnacle area, V 11408 68B Malaysia, Sarawak, Upper Penrissen valley, Mount
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<normalizedToken originalValue="Bra’ang" pageId="77" pageNumber="78">Bra'ang</normalizedToken>
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, West flank, about 2 km North of Temurang village, V 13996) 69
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<taxonomicName class="Gastropoda" family="Chronidae" genus="Kaliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaliella microsoma" order="Stylommatophora" pageId="77" pageNumber="78" phylum="Mollusca" rank="species" species="microsoma">Kaliella microsoma</taxonomicName>
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, sp. n. 69A Frontal view, shell 1.6 mm high 69B Umbilical view (Malaysia, West Coast Province. Crocker Range N.P., km 54 marker on the road Kota Kinabalu-Tambunan, Gunung Mas, RMNH.5003926, holotype).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="habitat in sabah and distribution">
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<paragraph pageId="77" pageNumber="78">Habitat in Sabah and distribution.</paragraph>
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<paragraph pageId="77" pageNumber="78">Primary and secondary forest on limestone and sandstone soil. Alt. up to 1000 m. Sabah: Crocker Range; Interior; East coast; scattered localities. Distribution elsewhere: From India, Laos, to Australia; Eastwards to Tahiti.</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="cross diagnosis">
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<paragraph pageId="77" pageNumber="78">Cross diagnosis.</paragraph>
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<paragraph pageId="77" pageNumber="78">
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When fresh, shell thinner and more translucent than the other species in this group,
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<taxonomicName class="Gastropoda" family="Chronidae" genus="Kaliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaliella dendrophila" order="Stylommatophora" pageId="77" pageNumber="78" phylum="Mollusca" rank="species" species="dendrophila">Kaliella dendrophila</taxonomicName>
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excepted. Juveniles can be distinguished from that species by the different apical sculpture.
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</paragraph>
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<paragraph pageId="77" pageNumber="78">
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Elsewhere,
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<taxonomicName class="Gastropoda" family="Chronidae" genus="Kaliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaliella sitaliformis" order="Stylommatophora" pageId="77" pageNumber="78" phylum="Mollusca" rank="species" species="sitaliformis">Kaliella sitaliformis</taxonomicName>
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<bibRefCitation pageId="77" pageNumber="78">Von Moellendorff 1897</bibRefCitation>
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, from Java, Bali, and Sulawesi, differs by having a raised spiral sculpture, visible in between the radial riblets, on the outer whorls.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="remarks">
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<paragraph pageId="77" pageNumber="78">Remarks.</paragraph>
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<paragraph pageId="77" pageNumber="78">
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<taxonomicName class="Gastropoda" family="Chronidae" genus="Kaliella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kaliella calculosa" order="Stylommatophora" pageId="77" pageNumber="78" phylum="Mollusca" rank="species" species="calculosa">Kaliella calculosa</taxonomicName>
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shows considerable variability in the prominence and the density of the spiral sculpture on the teleoconch. SEM-images show that the widest spiral grooves have a distinctly convex bottom; it is as if a thin thread is embedded in the groove.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |