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<document id="4ED1179FB82FC0B5BA712099AABE3EFF" ID-DOI="10.11646/zootaxa.5418.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10726419" ID-ZooBank="928EFF22-3EE9-4C29-92E0-4490C3FF0490" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709201492302" checkinUser="plazi" docAuthor="Heppner, John B. &amp; Bae, Yang-Seop" docDate="2024" docId="AB0F87F37935191AFF6EFF47FC84FA04" docLanguage="en" docName="zootaxa.5418.3.5.pdf" docOrigin="Zootaxa 5418 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5418.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cirrothaumatia pichita Heppner &amp; Bae 2024, new species" docType="treatment" docVersion="2" lastPageNumber="272" masterDocId="5736FF8B7937191EFFF9FFD0FFD6FF99" masterDocTitle="Three new species of Cirrothaumatia Razowski &amp; Becker from Peru, with notes on the genus (Lepidoptera: Tortricidae: Tortricinae: Cochylini)" masterLastPageNumber="278" masterPageNumber="268" pageNumber="270" updateTime="1709201945160" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F6795F3497A71C7D5DEEA6FF72BC86B2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="65E2FBB28B257723A98447F197C4206B">Three new species of Cirrothaumatia Razowski &amp; Becker from Peru, with notes on the genus (Lepidoptera: Tortricidae: Tortricinae: Cochylini)</mods:title>
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<mods:namePart id="85061FA0414597B088AEA8C6EF38F243">Heppner, John B.</mods:namePart>
<mods:affiliation id="6B08F6DEFF719675CA6D6806D7352487">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA.</mods:affiliation>
<mods:nameIdentifier id="281757486FD3A50B4074DB5FBF60C257" type="email">jheppner@flmnh.ufl.edu</mods:nameIdentifier>
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<mods:namePart id="6779378F1A2AD5BF6872356C9C665FE1">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="F1C808CC54689CEBED46389FB5D58F9A" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="1F49FA02239B6C2DC6BE46F6F5FD3DFF">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="CD9F3B2EBC1E9CEB4EFA5C11DCE2FFBA" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<mods:title id="38093413B99EFE1E3C8658777F35D246">Zootaxa</mods:title>
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<mods:date id="355D244E84000192BD4C45E704BBF2BE">2024</mods:date>
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<mods:number id="3CD847176FAE567073AD5600853B4B59">2024-02-29</mods:number>
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<mods:number id="E4FCA8402DA3E92B247489C0F01DE26C">5418</mods:number>
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<mods:number id="4B224CB7B0A14BA68E3D558DA61CFDE9">3</mods:number>
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<treatment id="AB0F87F37935191AFF6EFF47FC84FA04" ID-DOI="http://doi.org/10.5281/zenodo.10726431" ID-Zenodo-Dep="10726431" LSID="urn:lsid:plazi:treatment:AB0F87F37935191AFF6EFF47FC84FA04" httpUri="http://treatment.plazi.org/id/AB0F87F37935191AFF6EFF47FC84FA04" lastPageId="4" lastPageNumber="272" pageId="2" pageNumber="270">
<subSubSection id="6BBC656E7935191CFF6EFF47FCCFFF28" box="[151,793,151,178]" pageId="2" pageNumber="270" type="nomenclature">
<paragraph id="231936E57935191CFF6EFF47FCCFFF28" blockId="2.[151,793,151,213]" box="[151,793,151,178]" pageId="2" pageNumber="270">
<heading id="785181897935191CFF6EFF47FCCFFF28" bold="true" box="[151,793,151,178]" fontSize="11" level="1" pageId="2" pageNumber="270" reason="1">
<emphasis id="11D2EAF77935191CFF6EFF47FCCFFF28" bold="true" box="[151,793,151,178]" pageId="2" pageNumber="270">
<taxonomicName id="E4A64D667935191CFF6EFF47FDAAFF28" authority="Heppner &amp; Bae" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[151,636,151,178]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="270" phylum="Arthropoda" rank="species" species="pichita" status="sp. nov.">
<emphasis id="11D2EAF77935191CFF6EFF47FE79FF2B" bold="true" box="[151,431,151,178]" italics="true" pageId="2" pageNumber="270">Cirrothaumatia pichita</emphasis>
Heppner &amp; Bae
</taxonomicName>
,
<taxonomicNameLabel id="0AE1578C7935191CFD70FF47FCCFFF28" box="[649,793,151,177]" pageId="2" pageNumber="270" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7935191CFF6EFF6BFCB2FE87" pageId="2" pageNumber="270" type="description">
<paragraph id="231936E57935191CFF6EFF6BFEB0FF4C" blockId="2.[151,793,151,213]" box="[151,358,187,213]" pageId="2" pageNumber="270">
(
<figureCitation id="BB9D2A607935191CFF66FF6BFF3BFF4C" box="[159,237,187,213]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="2" pageNumber="270">Figs. 1</figureCitation>
,
<figureCitation id="BB9D2A607935191CFF02FF6BFEDDFF4C" box="[251,267,187,213]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="2" pageNumber="270">4</figureCitation>
,
<figureCitation id="BB9D2A607935191CFEEEFF6BFEE3FF4C" box="[279,309,187,213]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1718,1742]" captionTargetBox="[189,1399,181,1694]" captionTargetId="figure-28@7.[189,1399,181,1694]" captionTargetPageId="7" captionText="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." figureDoi="http://doi.org/10.5281/zenodo.10726433" httpUri="https://zenodo.org/record/10726433/files/figure.png" pageId="2" pageNumber="270">10</figureCitation>
,
<figureCitation id="BB9D2A607935191CFEB8FF6BFE88FF4C" box="[321,350,187,213]" captionStart="FIGURE 18" captionStartId="8.[151,250,1873,1897]" captionTargetBox="[218,1370,665,1850]" captionTargetId="figure-209@8.[218,1370,665,1850]" captionTargetPageId="8" captionText="FIGURE 18. Distribution map of known localities for Cirrothaumatia species (NGS base map): C. huemboana n. sp., yellow square (Peru:Amazonas). C. moyobamba n. sp., black-bordered white square (Peru: San Martín). C. pichita n. sp., white square (Peru: Junín). C. tornocarpa (Meyrick), red square (Panama: Chiriquí). C. tornosema (Clarke), black squares (Mexico:Veracruz; Guatemala: Quetzaltenango; Costa Rica: Cartago, Heredia, and Puntarenas; and Ecuador: Pichincha). C. vesta (Clarke), blue square (Venezuela: Aragua)." figureDoi="http://doi.org/10.5281/zenodo.10726435" httpUri="https://zenodo.org/record/10726435/files/figure.png" pageId="2" pageNumber="270">18</figureCitation>
)
</paragraph>
<paragraph id="231936E57935191CFF6EFED3FCB2FE87" blockId="2.[151,1437,259,465]" box="[151,868,259,286]" pageId="2" pageNumber="270">
<emphasis id="11D2EAF77935191CFF6EFED3FEEAFE84" bold="true" box="[151,316,259,285]" pageId="2" pageNumber="270">Type-locality:</emphasis>
Cerro Pichita Res. Sta., San Ramon,
<collectingRegion id="E162F8077935191CFD26FED3FCC8FE84" box="[735,798,259,285]" country="Peru" name="Junin" pageId="2" pageNumber="270">Junín</collectingRegion>
,
<collectingCountry id="5BB176757935191CFCD0FED4FCB6FE87" box="[809,864,260,286]" name="Peru" pageId="2" pageNumber="270">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7935191CFF3EFEF7FF2FFE48" pageId="2" pageNumber="270" type="diagnosis">
<paragraph id="231936E57935191CFF3EFEF7FF2FFE48" blockId="2.[151,1437,259,465]" pageId="2" pageNumber="270">
<emphasis id="11D2EAF77935191CFF3EFEF7FEECFED8" bold="true" box="[199,314,295,321]" pageId="2" pageNumber="270">Diagnosis</emphasis>
. This species can be distinguished by its larger size (16.2 mm vs. 11.8–13.7 mm) and distinctive female genitalia (
<figureCitation id="BB9D2A607935191CFE98FE9BFE71FEFC" box="[353,423,331,357]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="2" pageNumber="270">Fig. 4</figureCitation>
) with a thin carina-like cestum from the caudal venter of the bursa extending caudally along the lateral wall of the ductus bursae to the colliculum-like band (not as well-developed in the other species of the genus), and the stentor-like, uniformly wide ostial funnel (
<figureCitation id="BB9D2A607935191CFCA0FE43FC7CFE34" box="[857,938,403,429]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="2" pageNumber="270">Fig. 4a</figureCitation>
) (narrowing or bulging in most of the other species).
</paragraph>
</subSubSection>
<caption id="77D9666D7935191CFF6EF8D2FCDEF833" ID-DOI="http://doi.org/10.5281/zenodo.10726421" ID-Zenodo-Dep="10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="2" pageNumber="270" startId="2.[151,264,1794,1818]" targetBox="[152,1436,487,1768]" targetPageId="2" targetType="figure">
<paragraph id="231936E57935191CFF6EF8D2FCDEF833" blockId="2.[151,1436,1794,1962]" pageId="2" pageNumber="270">
<emphasis id="11D2EAF77935191CFF6EF8D2FEE3F883" bold="true" box="[151,309,1794,1818]" pageId="2" pageNumber="270">FIGURES 1–3</emphasis>
. Adult features of
<taxonomicName id="E4A64D667935191CFE0EF8D2FD3EF883" box="[503,744,1794,1818]" pageId="2" pageNumber="270">
<emphasis id="11D2EAF77935191CFE0EF8D2FD43F883" box="[503,661,1794,1818]" italics="true" pageId="2" pageNumber="270">Cirrothaumatia</emphasis>
species
</taxonomicName>
, with details of the head laterally (a) and dorsally (b):
<emphasis id="11D2EAF77935191CFAE2F8D2FAE6F883" bold="true" box="[1307,1328,1794,1818]" pageId="2" pageNumber="270">1)</emphasis>
<taxonomicName id="E4A64D667935191CFACEF8D3FA4AF883" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[1335,1436,1794,1818]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="270" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77935191CFACEF8D3FA4AF883" box="[1335,1436,1794,1818]" italics="true" pageId="2" pageNumber="270">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77935191CFF6EF8F6FF05F8A7" bold="true" box="[151,211,1830,1854]" pageId="2" pageNumber="270">
<taxonomicNameLabel id="0AE1578C7935191CFF6EF8F6FF05F8A7" box="[151,211,1830,1854]" pageId="2" pageNumber="270" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL);
<emphasis id="11D2EAF77935191CFEF8F89AFEC0F8FB" bold="true" box="[257,278,1866,1890]" pageId="2" pageNumber="270">2)</emphasis>
<taxonomicName id="E4A64D667935191CFEE7F89BFE62F8FB" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[286,436,1866,1890]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="270" phylum="Arthropoda" rank="species" species="huemboana" status="n. sp.">
<emphasis id="11D2EAF77935191CFEE7F89BFE62F8FB" box="[286,436,1866,1890]" italics="true" pageId="2" pageNumber="270">C. huemboana</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77935191CFE45F89AFE2EF8FB" bold="true" box="[444,504,1866,1890]" pageId="2" pageNumber="270">
<taxonomicNameLabel id="0AE1578C7935191CFE45F89AFE2EF8FB" box="[444,504,1866,1890]" pageId="2" pageNumber="270" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL);
<emphasis id="11D2EAF77935191CFDF0F8BEFDC9F81F" bold="true" box="[521,543,1902,1926]" pageId="2" pageNumber="270">3)</emphasis>
<taxonomicName id="E4A64D667935191CFDDFF8BFFD17F81F" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[550,705,1902,1926]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="270" phylum="Arthropoda" rank="species" species="moyobamba" status="n. sp.">
<emphasis id="11D2EAF77935191CFDDFF8BFFD17F81F" box="[550,705,1902,1926]" italics="true" pageId="2" pageNumber="270">C. moyobamba</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77935191CFD31F8BEFCD5F81F" bold="true" box="[712,771,1902,1926]" pageId="2" pageNumber="270">
<taxonomicNameLabel id="0AE1578C7935191CFD31F8BEFCD5F81F" box="[712,771,1902,1926]" pageId="2" pageNumber="270" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL).
</paragraph>
</caption>
<caption id="77D9666D7934191DFF6EFA98FE70FA69" ID-DOI="http://doi.org/10.5281/zenodo.10726425" ID-Zenodo-Dep="10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="3" pageNumber="271" startId="3.[151,264,1352,1376]" targetBox="[151,1436,182,1328]" targetPageId="3" targetType="figure">
<paragraph id="231936E57934191DFF6EFA98FE70FA69" blockId="3.[151,1437,1352,1520]" pageId="3" pageNumber="271">
<emphasis id="11D2EAF77934191DFF6EFA98FEE0FAF9" bold="true" box="[151,310,1352,1376]" pageId="3" pageNumber="271">FIGURES 4–6</emphasis>
. Female genitalia of
<taxonomicName id="E4A64D667934191DFDEEFA98FCDCFAF9" box="[535,778,1352,1376]" pageId="3" pageNumber="271">
<emphasis id="11D2EAF77934191DFDEEFA98FD63FAF9" box="[535,693,1352,1376]" italics="true" pageId="3" pageNumber="271">Cirrothaumatia</emphasis>
species
</taxonomicName>
, with details of the ostium, sterigma, and caudal ductus bursae region (a):
<emphasis id="11D2EAF77934191DFEF2FABCFEF6FA1D" bold="true" box="[267,288,1388,1412]" pageId="3" pageNumber="271">4)</emphasis>
<taxonomicName id="E4A64D667934191DFED1FABDFE5BFA1D" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[296,397,1388,1412]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="271" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77934191DFED1FABDFE5BFA1D" box="[296,397,1388,1412]" italics="true" pageId="3" pageNumber="271">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77934191DFE6CFABCFE06FA1D" bold="true" box="[405,464,1388,1412]" pageId="3" pageNumber="271">
<taxonomicNameLabel id="0AE1578C7934191DFE6CFABCFE06FA1D" box="[405,464,1388,1412]" pageId="3" pageNumber="271" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL);
<emphasis id="11D2EAF77934191DFEB9FA40FE80FA31" bold="true" box="[320,342,1424,1448]" pageId="3" pageNumber="271">5)</emphasis>
<taxonomicName id="E4A64D667934191DFEA5FA41FE39FA31" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[348,495,1424,1448]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="271" phylum="Arthropoda" rank="species" species="huemboana" status="n. sp.">
<emphasis id="11D2EAF77934191DFEA5FA41FE39FA31" box="[348,495,1424,1448]" italics="true" pageId="3" pageNumber="271">C. huemboana</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77934191DFE0FFA40FDF9FA31" bold="true" box="[502,559,1424,1448]" pageId="3" pageNumber="271">
<taxonomicNameLabel id="0AE1578C7934191DFE0FFA40FDF9FA31" box="[502,559,1424,1448]" pageId="3" pageNumber="271" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL);
<emphasis id="11D2EAF77934191DFE8DFA64FE5FFA55" bold="true" box="[372,393,1460,1484]" pageId="3" pageNumber="271">6)</emphasis>
<taxonomicName id="E4A64D667934191DFE77FA65FDF1FA55" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[398,551,1460,1484]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="271" phylum="Arthropoda" rank="species" species="moyobamba" status="n. sp.">
<emphasis id="11D2EAF77934191DFE77FA65FDF1FA55" box="[398,551,1460,1484]" italics="true" pageId="3" pageNumber="271">C. moyobamba</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77934191DFDD5FA64FDB2FA55" bold="true" box="[556,612,1460,1484]" pageId="3" pageNumber="271">
<taxonomicNameLabel id="0AE1578C7934191DFDD5FA64FDB2FA55" box="[556,612,1460,1484]" pageId="3" pageNumber="271" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL).
</paragraph>
</caption>
<subSubSection id="6BBC656E7934191AFF3EF992FDC8FC8C" lastPageId="4" lastPageNumber="272" pageId="3" pageNumber="271" type="description">
<paragraph id="231936E57934191AFF3EF992FCF6FE34" blockId="3.[151,1437,1602,2025]" lastBlockId="4.[151,1437,151,1437]" lastPageId="4" lastPageNumber="272" pageId="3" pageNumber="271">
<emphasis id="11D2EAF77934191DFF3EF992FE87F9C5" bold="true" box="[199,337,1602,1628]" pageId="3" pageNumber="271">Description</emphasis>
. Wing expanse 16.2 mm (n = 1). Female (
<figureCitation id="BB9D2A607934191DFCC3F992FCABF9C4" box="[826,893,1602,1629]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="3" pageNumber="271">Fig. 1</figureCitation>
).
<emphasis id="11D2EAF77934191DFC6EF992FC02F9C5" box="[919,980,1602,1628]" italics="true" pageId="3" pageNumber="271">Head</emphasis>
(
<figureCitation id="BB9D2A607934191DFC1DF992FB82F9C5" box="[996,1108,1602,1628]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="3" pageNumber="271">Fig. 1a–b</figureCitation>
): Vertex white; caudo-lateral scale tufts white to slightly tan-gray caudally; labial palpus white with few tan to brown-gray scales laterally and on basal segment; antenna brown, with white scaling dorsally and a few maroon scales; scape white.
<emphasis id="11D2EAF77934191DFB16F95AFA96F93D" box="[1263,1344,1674,1700]" italics="true" pageId="3" pageNumber="271">Thorax</emphasis>
: Brown mottled with white, many scales dark orange-tipped, slightly more orange scaling distally on metathorax; tegula same, with more white antero-laterally; patagia gray-brown with scales tipped pale white; venter white with gray-brown mixed; fore- and midlegs mostly dark brown, with some white scales (coxae mostly white), tibiae with white band at middle, tarsi dark brown with white distal margin each segment; hindleg lustrous gray-tan and mesally dull white; tibial spurs gray-brown.
<emphasis id="11D2EAF77934191DFE0BF8EFFD88F8C1" box="[498,606,1855,1880]" italics="true" pageId="3" pageNumber="271">Forewing</emphasis>
(
<figureCitation id="BB9D2A607934191DFD95F8EEFD7DF8C0" box="[620,683,1854,1881]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="3" pageNumber="271">Fig. 1</figureCitation>
) mottled dark brown and black-brown, with prominent darker spots at base of anal margin, near midwing along costa and on apical one-fourth, and some maroon scaling on brown and dark spots; dark areas separated by large white blotches with diffuse borders near base and end of discal cell to costa, plus narrow irregular triangular-like spots along termen and near apex; prominent white dash at end of cell; subterminal fascia brown-gray oblique from near tornus to near costal margin, tapering toward costa and with median short but broad knob-like extension on distal side; tornus with a prominent quadratic orange patch; scattered slight orange scaling near mid-wing and near wing base and mixed in on dorsal half of wing from near tornus to near wing base; termen with a thin black-brown line; fringe mostly white, interspersed with two dark brown portions on apical half sector of terminal fringe and brown near tornal fringe sector; venter dark gray-brown with some white on outer third, and cubital field pale tan-white.
<emphasis id="11D2EAF77933191AFD73FEF7FD2EFED8" box="[650,760,295,321]" italics="true" pageId="4" pageNumber="272">Hindwing</emphasis>
(
<figureCitation id="BB9D2A607933191AFCF1FEF7FC9FFED8" box="[776,841,295,321]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="4" pageNumber="272">Fig. 1</figureCitation>
) brown, translucent, with darker brown along veins; fringe in three parts from thin white line at termen, middle black-brown portion, and distal pale gray portion; venter gray-brown with dark tan on radial sector from base, with alternating short brown bars on main veins, and with dorsal half of wing gray-brown with veins slightly darker.
</paragraph>
<paragraph id="231936E57933191AFF3EFE67FDC8FC8C" blockId="4.[151,1437,151,1437]" pageId="4" pageNumber="272">
<emphasis id="11D2EAF77933191AFF3EFE67FEE7FE48" box="[199,305,439,465]" italics="true" pageId="4" pageNumber="272">Abdomen</emphasis>
(
<figureCitation id="BB9D2A607933191AFEBDFE67FE5FFE48" box="[324,393,439,465]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="4" pageNumber="272">Fig. 1</figureCitation>
). Dark gray-brown with pale silvery posterior margins per tergite; venter white on S2–3, remaining sternites mottled white and gray-brown, with dark brown anterior of each sternite; genital tuft dark brown with pale tan distally and dull white ventrally.
<emphasis id="11D2EAF77933191AFCFCFE2FFC11FD80" box="[773,967,511,537]" italics="true" pageId="4" pageNumber="272">Female genitalia</emphasis>
(
<figureCitation id="BB9D2A607933191AFC21FE2FFBC8FD80" box="[984,1054,511,537]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="4" pageNumber="272">Fig. 4</figureCitation>
) similar to congeners, with short ovipositor; apophyses with anterior pair slightly longer than posterior pair; sterigma an indistinct, thin bordering margin; ostial funnel (
<figureCitation id="BB9D2A607933191AFE68FD97FE34FDF8" box="[401,482,583,609]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="4" pageNumber="272">Fig. 4a</figureCitation>
) straight and uniformly broad, short stentor-like, to antrum, with diffuse colliculum-like proximal band following a slight constriction before merging into ductus bursae; ductus bursae slightly and gradually widening to merge with bulbous bursa; a narrow linear carina-like cestum, with a slightly more broadened portion on the caudo-ventral end of the bursa; corpus bursa ovate, evenly merged into ductus bursae; large, elongated accessory bursa emergent from caudo-proximal and slightly extruded end of the bursa, with a long duct (ca. as long as bursa is wide).
<emphasis id="11D2EAF77933191AFE98FD2BFE74FC8C" box="[353,418,763,789]" italics="true" pageId="4" pageNumber="272">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7933191AFF3EFCCFFE20FC18" pageId="4" pageNumber="272" type="materials_examined">
<paragraph id="231936E57933191AFF3EFCCFFE20FC18" blockId="4.[151,1437,151,1437]" pageId="4" pageNumber="272">
<emphasis id="11D2EAF77933191AFF3EFCCFFE57FCA1" bold="true" box="[199,385,798,825]" pageId="4" pageNumber="272">Type materials.</emphasis>
<materialsCitation id="93CE3CB87933191AFE71FCCFFE20FC18" collectingDate="2012-11-04" collectingDateMax="2012-11-07" collectingDateMin="2012-11-04" collectionCode="MGCL, UNALM" collectorName="J. B. Heppner &amp; C. Carrera" country="Peru" county="San Ramon" elevation="2165" location="Female" municipality="Cerro Pichita Research Station" pageId="4" pageNumber="272" specimenCount="1" stateProvince="Junin" typeStatus="holotype">
<emphasis id="11D2EAF77933191AFE71FCCFFE23FCA0" bold="true" box="[392,501,799,825]" pageId="4" pageNumber="272">
<typeStatus id="FC1D88477933191AFE71FCCFFE23FCA0" box="[392,501,799,825]" pageId="4" pageNumber="272" type="holotype">Holotype</typeStatus>
</emphasis>
.
<location id="2679603E7933191AFDFDFCCFFD8FFCA0" LSID="urn:lsid:plazi:treatment:AB0F87F37935191AFF6EFF47FC84FA04:2679603E7933191AFDFDFCCFFD8FFCA0" box="[516,601,799,825]" country="Peru" county="San Ramon" municipality="Cerro Pichita Research Station" name="Female" pageId="4" pageNumber="272" stateProvince="Junin">Female</location>
,
<collectingMunicipality id="C37DAC9F7933191AFD9CFCCFFC16FCA0" box="[613,960,799,825]" pageId="4" pageNumber="272">Cerro Pichita Research Station</collectingMunicipality>
(
<quantity id="E45E9B007933191AFC29FCCEFBFDFCA0" box="[976,1067,798,825]" metricMagnitude="3" metricUnit="m" metricValue="2.165" pageId="4" pageNumber="272" unit="m" value="2165.0">
<elevation id="A88BD1D67933191AFC29FCCEFBFDFCA0" box="[976,1067,798,825]" metricMagnitude="3" metricUnit="m" metricValue="2.165" pageId="4" pageNumber="272" unit="m" value="2165.0">2165 m</elevation>
</quantity>
),
<collectingCounty id="CA784E697933191AFBC6FCCFFB13FCA0" box="[1087,1221,799,825]" pageId="4" pageNumber="272">San Ramon</collectingCounty>
, Dept.
<collectingRegion id="E162F8077933191AFAEFFCCFFA83FCA0" box="[1302,1365,799,825]" country="Peru" name="Junin" pageId="4" pageNumber="272">Junín</collectingRegion>
,
<collectingCountry id="5BB176757933191AFA9BFCCFFA4EFCA0" box="[1378,1432,799,825]" name="Peru" pageId="4" pageNumber="272">Peru</collectingCountry>
,
<date id="571810257933191AFF6EFC93FE96FCC4" box="[151,320,835,861]" pageId="4" pageNumber="272" value="2012-11-04" valueMax="2012-11-07" valueMin="2012-11-04">
<collectingDate id="475CE9CD7933191AFF6EFC93FE96FCC4" box="[151,320,835,861]" pageId="4" pageNumber="272" value="2012-11-04" valueMax="2012-11-07" valueMin="2012-11-04">4–7 Nov 2012</collectingDate>
</date>
,
<collectorName id="8E5353337933191AFEB7FC93FE3BFCC4" box="[334,493,835,861]" pageId="4" pageNumber="272">J. B. Heppner</collectorName>
&amp;
<collectorName id="8E5353337933191AFDEEFC93FD58FCC4" box="[535,654,835,861]" pageId="4" pageNumber="272">C. Carrera</collectorName>
(gen. slide JBH-4361; adult photo 15533) (deposited at
<collectionCode id="45B7AE207933191AFAE7FC93FAA4FCC4" box="[1310,1394,835,861]" pageId="4" pageNumber="272">MGCL</collectionCode>
, on indefinite loan from
<collectionCode id="45B7AE207933191AFE87FCB7FE3CFC18" box="[382,490,871,897]" pageId="4" pageNumber="272">UNALM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7933191AFF3EFC5BFA41FC50" pageId="4" pageNumber="272" type="etymology">
<paragraph id="231936E57933191AFF3EFC5BFA41FC50" blockId="4.[151,1437,151,1437]" pageId="4" pageNumber="272">
<emphasis id="11D2EAF77933191AFF3EFC5BFE9EFC3C" bold="true" box="[199,328,907,933]" pageId="4" pageNumber="272">Etymology</emphasis>
. The species is named after the Cerro Pichita Research Station, San Ramon, Dept.
<collectingRegion id="E162F8077933191AFAEDFC5BFA85FC3C" box="[1300,1363,907,933]" country="Peru" name="Junin" pageId="4" pageNumber="272">Junín</collectingRegion>
,
<collectingCountry id="5BB176757933191AFA9BFC5BFA4EFC3C" box="[1378,1432,907,933]" name="Peru" pageId="4" pageNumber="272">Peru</collectingCountry>
, located about two-thirds the way up the eastern slope of Cerro Pichita (apex slightly above timberline, at
<quantity id="E45E9B007933191AFACBFC7FFA5DFC50" box="[1330,1419,943,969]" metricMagnitude="3" metricUnit="m" metricValue="2.902" pageId="4" pageNumber="272" unit="m" value="2902.0">2902 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7933191AFF3EFC02FC76FC74" box="[199,928,978,1005]" pageId="4" pageNumber="272" type="biology_ecology">
<paragraph id="231936E57933191AFF3EFC02FC76FC74" blockId="4.[151,1437,151,1437]" box="[199,928,978,1005]" pageId="4" pageNumber="272">
<emphasis id="11D2EAF77933191AFF3EFC02FEF7FC75" bold="true" box="[199,289,978,1004]" pageId="4" pageNumber="272">Biology</emphasis>
. Unknown. Habitat submontane Andean forest (
<figureCitation id="BB9D2A607933191AFCC6FC03FC45FC74" box="[831,915,979,1005]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1718,1742]" captionTargetBox="[189,1399,181,1694]" captionTargetId="figure-28@7.[189,1399,181,1694]" captionTargetPageId="7" captionText="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." figureDoi="http://doi.org/10.5281/zenodo.10726433" httpUri="https://zenodo.org/record/10726433/files/figure.png" pageId="4" pageNumber="272">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7933191AFF3EFC26FCD5FBC0" pageId="4" pageNumber="272" type="distribution">
<paragraph id="231936E57933191AFF3EFC26FCD5FBC0" blockId="4.[151,1437,151,1437]" pageId="4" pageNumber="272">
<emphasis id="11D2EAF77933191AFF3EFC26FE8FFB89" bold="true" box="[199,345,1014,1040]" pageId="4" pageNumber="272">Distribution</emphasis>
. Known only from the eastern Andean submontane site at Cerro Pichita (
<quantity id="E45E9B007933191AFB57FC26FADFFB88" box="[1198,1289,1014,1041]" metricMagnitude="3" metricUnit="m" metricValue="2.165" pageId="4" pageNumber="272" unit="m" value="2165.0">2165 m</quantity>
) (
<figureCitation id="BB9D2A607933191AFADAFC27FAAEFB88" box="[1315,1400,1015,1041]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1718,1742]" captionTargetBox="[189,1399,181,1694]" captionTargetId="figure-28@7.[189,1399,181,1694]" captionTargetPageId="7" captionText="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." figureDoi="http://doi.org/10.5281/zenodo.10726433" httpUri="https://zenodo.org/record/10726433/files/figure.png" pageId="4" pageNumber="272">Fig. 10</figureCitation>
), a mountain upland area just west of the small town of San Ramon (
<quantity id="E45E9B007933191AFC94FBCAFC6EFBAC" box="[877,952,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="272" unit="m" value="650.0">650 m</quantity>
), Dept.
<collectingRegion id="E162F8077933191AFBF6FBCBFB99FBAC" box="[1039,1103,1051,1077]" country="Peru" name="Junin" pageId="4" pageNumber="272">Junín</collectingRegion>
. This locality is due east from
<collectingRegion id="E162F8077933191AFF6EFBEFFF03FBC0" box="[151,213,1087,1113]" country="Peru" name="Lima" pageId="4" pageNumber="272">Lima</collectingRegion>
, on the eastern side of the Andes Range (
<figureCitation id="BB9D2A607933191AFD5BFBEFFD20FBC0" box="[674,758,1087,1113]" captionStart="FIGURE 18" captionStartId="8.[151,250,1873,1897]" captionTargetBox="[218,1370,665,1850]" captionTargetId="figure-209@8.[218,1370,665,1850]" captionTargetPageId="8" captionText="FIGURE 18. Distribution map of known localities for Cirrothaumatia species (NGS base map): C. huemboana n. sp., yellow square (Peru:Amazonas). C. moyobamba n. sp., black-bordered white square (Peru: San Martín). C. pichita n. sp., white square (Peru: Junín). C. tornocarpa (Meyrick), red square (Panama: Chiriquí). C. tornosema (Clarke), black squares (Mexico:Veracruz; Guatemala: Quetzaltenango; Costa Rica: Cartago, Heredia, and Puntarenas; and Ecuador: Pichincha). C. vesta (Clarke), blue square (Venezuela: Aragua)." figureDoi="http://doi.org/10.5281/zenodo.10726435" httpUri="https://zenodo.org/record/10726435/files/figure.png" pageId="4" pageNumber="272">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7933191AFF3EFBB2FC84FA04" pageId="4" pageNumber="272" type="discussion">
<paragraph id="231936E57933191AFF3EFBB2FC84FA04" blockId="4.[151,1437,151,1437]" pageId="4" pageNumber="272">
<emphasis id="11D2EAF77933191AFF3EFBB2FE93FBE5" bold="true" box="[199,325,1122,1148]" pageId="4" pageNumber="272">Discussion</emphasis>
. The cestum in the female genitalia of this new species is strong and very narrow, extending along most of the ductus bursae (
<figureCitation id="BB9D2A607933191AFE3DFB57FDDFFB38" box="[452,521,1159,1185]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="4" pageNumber="272">Fig. 4</figureCitation>
).
<bibRefCitation id="47374B147933191AFDE4FB57FD6FFB38" author="Clarke, J. F. G." box="[541,697,1159,1185]" pageId="4" pageNumber="272" pagination="1 - 58" refId="ref6024" refString="Clarke, J. F. G. (1968) Neotropical Microlepidoptera. XVII. Notes and new species of Phaloniidae. Proceedings of the United States. National Museum, 125, 1 - 58, 4 pls. https: // doi. org / 10.5479 / si. 00963801.125 - 3660.1" type="journal article" year="1968">Clarke (1968)</bibRefCitation>
and
<bibRefCitation id="47374B147933191AFD16FB57FBE2FB38" author="Razowski, J. &amp; Becker, V. O." box="[751,1076,1159,1185]" pageId="4" pageNumber="272" pagination="441 - 500" refId="ref6401" refString="Razowski, J. &amp; Becker, V. O. (1986) Cochylidii (Lepidoptera: Tortricidae) collected in Central America and Mexico. Acta Zoologica Cracoviensia, 29, 441 - 500." type="journal article" year="1986">Razowski and Becker (1986)</bibRefCitation>
did not mention the presence of a cestum in their species, but their genital figures appear to indicate a similar structure along the ductus bursa and onto the bursa (
<figureCitation id="BB9D2A607933191AFEB5FB1FFE30FB70" box="[332,486,1231,1257]" captionStart="FIGURES 7–9" captionStartId="6.[151,264,1828,1852]" captionTargetBox="[156,1426,490,1796]" captionTargetId="figure-133@6.[151,1436,489,1805]" captionTargetPageId="6" captionText="FIGURES 7–9. Genital features of related species of Cirrothaumatia: 7) C. tornosema (Clarke), female holotype (Guatemala, USNM) and male (a, shown without left valva; valval scale tuft shown intact on left valval base: Costa Rica, VOB); 8) C. tornocarpa (Meyrick), female holotype (Panama, NHMV); 9) C. vesta (Clarke), female paratype and male holotype (a, with aedeagus to right) (Venezuela, USNM) (Fig. 7, 9 and 9a, after Clarke (1968); Fig. 7a, after Razowski (1964); Fig. 8, after Razowski and Becker (1986))." figureDoi="http://doi.org/10.5281/zenodo.10726429" httpUri="https://zenodo.org/record/10726429/files/figure.png" pageId="4" pageNumber="272">Figs. 7 and 9</figureCitation>
), while
<taxonomicName id="E4A64D667933191AFDBCFB1FFC74FB70" authority="Meyrick (1932)" authorityName="Meyrick" authorityYear="1932" box="[581,930,1231,1257]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="tornocarpa">
<emphasis id="11D2EAF77933191AFDBCFB1FFD30FB71" box="[581,742,1231,1257]" italics="true" pageId="4" pageNumber="272">C. tornocarpa</emphasis>
Meyrick (1932)
</taxonomicName>
has a small semicircular signum (
<bibRefCitation id="47374B147933191AFAD5FB1FFF05FA94" author="Razowski, J." pageId="4" pageNumber="272" pagination="451 - 481" refId="ref6079" refString="Razowski, J. (1964) Meyrick types of Tortricoidea (Lepidoptera) in the Vienna Museum. Annales Zoologici, 22, 451 - 481." type="journal article" year="1964">Razowski 1964</bibRefCitation>
). The dissections for these other species were not available for the present study. Based on female genitalia,
<taxonomicName id="E4A64D667933191AFF6EFAC7FED0FAA8" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[151,262,1303,1329]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="pichita">
<emphasis id="11D2EAF77933191AFF6EFAC7FED0FAA8" box="[151,262,1303,1329]" italics="true" pageId="4" pageNumber="272">C. pichita</emphasis>
</taxonomicName>
appears to be closest to
<taxonomicName id="E4A64D667933191AFDEFFAC7FDA6FAA9" baseAuthorityName="Clarke" baseAuthorityYear="1968" box="[534,624,1303,1329]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="vesta">
<emphasis id="11D2EAF77933191AFDEFFAC7FDA6FAA9" box="[534,624,1303,1329]" italics="true" pageId="4" pageNumber="272">C. vesta</emphasis>
</taxonomicName>
from
<collectingCountry id="5BB176757933191AFD4DFAC7FCFCFAA8" box="[692,810,1303,1329]" name="Venezuela" pageId="4" pageNumber="272">Venezuela</collectingCountry>
, with the ostial funnel straight and uniformly wide in
<taxonomicName id="E4A64D667933191AFA7AFAC7FF33FACC" authorityName="Heppner &amp; Bae" authorityYear="2024" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="pichita">
<emphasis id="11D2EAF77933191AFA7AFAC7FF33FACC" italics="true" pageId="4" pageNumber="272">C. pichita</emphasis>
</taxonomicName>
(
<figureCitation id="BB9D2A607933191AFF0DFAEBFE93FACC" box="[244,325,1339,1365]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="4" pageNumber="272">Fig. 4a</figureCitation>
), while tapering and broadened caudally in
<taxonomicName id="E4A64D667933191AFCD0FAEBFC55FACD" baseAuthorityName="Clarke" baseAuthorityYear="1968" box="[809,899,1339,1365]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="vesta">
<emphasis id="11D2EAF77933191AFCD0FAEBFC55FACD" box="[809,899,1339,1365]" italics="true" pageId="4" pageNumber="272">C. vesta</emphasis>
</taxonomicName>
(
<figureCitation id="BB9D2A607933191AFC68FAEBFC03FACC" box="[913,981,1339,1365]" captionStart="FIGURES 7–9" captionStartId="6.[151,264,1828,1852]" captionTargetBox="[156,1426,490,1796]" captionTargetId="figure-133@6.[151,1436,489,1805]" captionTargetPageId="6" captionText="FIGURES 7–9. Genital features of related species of Cirrothaumatia: 7) C. tornosema (Clarke), female holotype (Guatemala, USNM) and male (a, shown without left valva; valval scale tuft shown intact on left valval base: Costa Rica, VOB); 8) C. tornocarpa (Meyrick), female holotype (Panama, NHMV); 9) C. vesta (Clarke), female paratype and male holotype (a, with aedeagus to right) (Venezuela, USNM) (Fig. 7, 9 and 9a, after Clarke (1968); Fig. 7a, after Razowski (1964); Fig. 8, after Razowski and Becker (1986))." figureDoi="http://doi.org/10.5281/zenodo.10726429" httpUri="https://zenodo.org/record/10726429/files/figure.png" pageId="4" pageNumber="272">Fig. 9</figureCitation>
); bulging medially in
<taxonomicName id="E4A64D667933191AFB33FAEBFAB6FACD" authorityName="Clarke" authorityYear="1968" box="[1226,1376,1339,1365]" class="Insecta" family="Tortricidae" genus="Phalonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="tornosema">
<emphasis id="11D2EAF77933191AFB33FAEBFAB6FACD" box="[1226,1376,1339,1365]" italics="true" pageId="4" pageNumber="272">C. tornosema</emphasis>
</taxonomicName>
(
<figureCitation id="BB9D2A607933191AFA96FAEBFF71FAE0" captionStart="FIGURES 7–9" captionStartId="6.[151,264,1828,1852]" captionTargetBox="[156,1426,490,1796]" captionTargetId="figure-133@6.[151,1436,489,1805]" captionTargetPageId="6" captionText="FIGURES 7–9. Genital features of related species of Cirrothaumatia: 7) C. tornosema (Clarke), female holotype (Guatemala, USNM) and male (a, shown without left valva; valval scale tuft shown intact on left valval base: Costa Rica, VOB); 8) C. tornocarpa (Meyrick), female holotype (Panama, NHMV); 9) C. vesta (Clarke), female paratype and male holotype (a, with aedeagus to right) (Venezuela, USNM) (Fig. 7, 9 and 9a, after Clarke (1968); Fig. 7a, after Razowski (1964); Fig. 8, after Razowski and Becker (1986))." figureDoi="http://doi.org/10.5281/zenodo.10726429" httpUri="https://zenodo.org/record/10726429/files/figure.png" pageId="4" pageNumber="272">Fig. 7</figureCitation>
), and constricted prior to merging with the bursa. The maculation of the species is similar, but
<taxonomicName id="E4A64D667933191AFB3AFA8FFAE7FAE0" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[1219,1329,1375,1401]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="pichita">
<emphasis id="11D2EAF77933191AFB3AFA8FFAE7FAE0" box="[1219,1329,1375,1401]" italics="true" pageId="4" pageNumber="272">C. pichita</emphasis>
</taxonomicName>
has more extensive forewing white areas, particularly on the basal third.
</paragraph>
</subSubSection>
</treatment>
</document>
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