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<document id="A26BABE4EF83E4C8DBB00BB29E0E8F57" ID-DOI="10.11646/zootaxa.5418.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10726419" ID-ZooBank="928EFF22-3EE9-4C29-92E0-4490C3FF0490" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709201492302" checkinUser="plazi" docAuthor="Heppner, John B. &amp; Bae, Yang-Seop" docDate="2024" docId="AB0F87F37933191BFF6EFA3AFA5DFD54" docLanguage="en" docName="zootaxa.5418.3.5.pdf" docOrigin="Zootaxa 5418 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5418.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cirrothaumatia huemboana Heppner &amp; Bae 2024, new species" docType="treatment" docVersion="2" lastPageNumber="273" masterDocId="5736FF8B7937191EFFF9FFD0FFD6FF99" masterDocTitle="Three new species of Cirrothaumatia Razowski &amp; Becker from Peru, with notes on the genus (Lepidoptera: Tortricidae: Tortricinae: Cochylini)" masterLastPageNumber="278" masterPageNumber="268" pageNumber="272" updateTime="1709201945160" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7E3A4B6E48DD53E0AC1461C74C1F9ED0">Three new species of Cirrothaumatia Razowski &amp; Becker from Peru, with notes on the genus (Lepidoptera: Tortricidae: Tortricinae: Cochylini)</mods:title>
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<mods:name id="217F24705A5943808FC7F222A3DDB19E" type="personal">
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<mods:namePart id="4433E535861E87C9ACBC8DE97BFBA8A6">Heppner, John B.</mods:namePart>
<mods:affiliation id="A5AE57656B7200284F048DF4087EA85C">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA.</mods:affiliation>
<mods:nameIdentifier id="5FDF2FECDFAF6014B0165B55F038DF46" type="email">jheppner@flmnh.ufl.edu</mods:nameIdentifier>
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<mods:roleTerm id="F059D22E2956193CA1344081785B414F">Author</mods:roleTerm>
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<mods:namePart id="6218B103C977DD4477CA85967E157662">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="ED9D5B8FC0BEBB411334F083319F6D13" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="F631DEFA2AB6898B25BE5F95A34E84DC">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="E608A0E8E0C603738610B97893880C72" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<mods:title id="EDA6297ECAC98B69469F298DA2FA6E46">Zootaxa</mods:title>
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<mods:part id="1A872A37B361C81FFF1C2B7523DA6277">
<mods:date id="9FB98DDA31FDF6C94BB4785BEFE69E5C">2024</mods:date>
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<mods:number id="18C59BC123C5D4499F9C5A3ECEB39F16">2024-02-29</mods:number>
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<mods:number id="09DC99ADD225417F3E312BDBF0E9C27A">5418</mods:number>
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<mods:detail id="F31A3C80364520F05C3A79A549A56131" type="issue">
<mods:number id="94DF7683338BF376C4D22A8254F7174A">3</mods:number>
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<mods:classification id="CDA75845857594AEDDA1B1886E685C09">journal article</mods:classification>
<mods:identifier id="7DFD1EAD6DD47589E22D3F4EB24A0B04" type="DOI">10.11646/zootaxa.5418.3.5</mods:identifier>
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<treatment id="AB0F87F37933191BFF6EFA3AFA5DFD54" ID-DOI="http://doi.org/10.5281/zenodo.10726437" ID-Zenodo-Dep="10726437" LSID="urn:lsid:plazi:treatment:AB0F87F37933191BFF6EFA3AFA5DFD54" httpUri="http://treatment.plazi.org/id/AB0F87F37933191BFF6EFA3AFA5DFD54" lastPageId="5" lastPageNumber="273" pageId="4" pageNumber="272">
<subSubSection id="6BBC656E7933191AFF6EFA3AFC80F99D" box="[151,854,1514,1541]" pageId="4" pageNumber="272" type="nomenclature">
<paragraph id="231936E57933191AFF6EFA3AFC80F99D" blockId="4.[151,854,1514,1577]" box="[151,854,1514,1541]" pageId="4" pageNumber="272">
<heading id="785181897933191AFF6EFA3AFC80F99D" bold="true" box="[151,854,1514,1541]" fontSize="11" level="1" pageId="4" pageNumber="272" reason="1">
<emphasis id="11D2EAF77933191AFF6EFA3AFC80F99D" bold="true" box="[151,854,1514,1541]" pageId="4" pageNumber="272">
<taxonomicName id="E4A64D667933191AFF6EFA3AFD6FF99C" authority="Heppner &amp; Bae" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[151,697,1514,1541]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="huemboana" status="sp. nov.">
<emphasis id="11D2EAF77933191AFF6EFA3AFE3DF99C" bold="true" box="[151,491,1514,1541]" italics="true" pageId="4" pageNumber="272">Cirrothaumatia huemboana</emphasis>
Heppner &amp; Bae
</taxonomicName>
,
<taxonomicNameLabel id="0AE1578C7933191AFD3FFA3BFC80F99D" box="[710,854,1514,1541]" pageId="4" pageNumber="272" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7933191AFF6EF9DFFC85F9E8" pageId="4" pageNumber="272" type="description">
<paragraph id="231936E57933191AFF6EF9DFFEB3F9B0" blockId="4.[151,854,1514,1577]" box="[151,357,1550,1577]" pageId="4" pageNumber="272">
(
<figureCitation id="BB9D2A607933191AFF66F9DFFF39F9B0" box="[159,239,1551,1577]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="4" pageNumber="272">Figs. 2</figureCitation>
,
<figureCitation id="BB9D2A607933191AFF02F9DEFEDDF9B0" box="[251,267,1550,1577]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="4" pageNumber="272">5</figureCitation>
,
<figureCitation id="BB9D2A607933191AFEEEF9DFFEE2F9B0" box="[279,308,1551,1577]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1718,1742]" captionTargetBox="[189,1399,181,1694]" captionTargetId="figure-28@7.[189,1399,181,1694]" captionTargetPageId="7" captionText="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." figureDoi="http://doi.org/10.5281/zenodo.10726433" httpUri="https://zenodo.org/record/10726433/files/figure.png" pageId="4" pageNumber="272">11</figureCitation>
,
<figureCitation id="BB9D2A607933191AFEB9F9DFFE8BF9B0" box="[320,349,1551,1577]" captionStart="FIGURE 18" captionStartId="8.[151,250,1873,1897]" captionTargetBox="[218,1370,665,1850]" captionTargetId="figure-209@8.[218,1370,665,1850]" captionTargetPageId="8" captionText="FIGURE 18. Distribution map of known localities for Cirrothaumatia species (NGS base map): C. huemboana n. sp., yellow square (Peru:Amazonas). C. moyobamba n. sp., black-bordered white square (Peru: San Martín). C. pichita n. sp., white square (Peru: Junín). C. tornocarpa (Meyrick), red square (Panama: Chiriquí). C. tornosema (Clarke), black squares (Mexico:Veracruz; Guatemala: Quetzaltenango; Costa Rica: Cartago, Heredia, and Puntarenas; and Ecuador: Pichincha). C. vesta (Clarke), blue square (Venezuela: Aragua)." figureDoi="http://doi.org/10.5281/zenodo.10726435" httpUri="https://zenodo.org/record/10726435/files/figure.png" pageId="4" pageNumber="272">18</figureCitation>
)
</paragraph>
<paragraph id="231936E57933191AFF6EF986FC85F9E8" blockId="4.[151,1437,1622,2009]" box="[151,851,1622,1649]" pageId="4" pageNumber="272">
<emphasis id="11D2EAF77933191AFF6EF986FEEAF9E9" bold="true" box="[151,316,1622,1648]" pageId="4" pageNumber="272">Type-locality:</emphasis>
Huembo Reserve, La Florida, Amazonas,
<collectingCountry id="5BB176757933191AFCE1F987FC99F9E8" box="[792,847,1623,1649]" name="Peru" pageId="4" pageNumber="272">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7933191AFF3EF9AAFEBCF944" pageId="4" pageNumber="272" type="diagnosis">
<paragraph id="231936E57933191AFF3EF9AAFEBCF944" blockId="4.[151,1437,1622,2009]" pageId="4" pageNumber="272">
<emphasis id="11D2EAF77933191AFF3EF9AAFEECF90D" bold="true" box="[199,314,1658,1684]" pageId="4" pageNumber="272">Diagnosis</emphasis>
. This species can be distinguished by the much smaller accessory bursa of the female genitalia (
<figureCitation id="BB9D2A607933191AFA96F9ABFF73F920" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="4" pageNumber="272">Fig. 5</figureCitation>
) (i.e., one-third the size of that of
<taxonomicName id="E4A64D667933191AFDD1F94FFD41F920" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[552,663,1695,1721]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77933191AFDD1F94FFD41F920" box="[552,663,1695,1721]" italics="true" pageId="4" pageNumber="272">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77933191AFD67F94FFD0BF920" bold="true" box="[670,733,1695,1721]" pageId="4" pageNumber="272">
<taxonomicNameLabel id="0AE1578C7933191AFD67F94FFD0BF920" box="[670,733,1695,1721]" pageId="4" pageNumber="272" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="E4A64D667933191AFCECF94FFCA6F921" baseAuthorityName="Clarke" baseAuthorityYear="1968" box="[789,880,1695,1721]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="vesta">
<emphasis id="11D2EAF77933191AFCECF94FFCA6F921" box="[789,880,1695,1721]" italics="true" pageId="4" pageNumber="272">C. vesta</emphasis>
</taxonomicName>
), and the stentor-like ostial funnel, similar to that of
<taxonomicName id="E4A64D667933191AFF4FF913FEF2F944" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[182,292,1731,1757]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77933191AFF4FF913FEF2F944" box="[182,292,1731,1757]" italics="true" pageId="4" pageNumber="272">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77933191AFED2F913FEBCF944" bold="true" box="[299,362,1731,1757]" pageId="4" pageNumber="272">
<taxonomicNameLabel id="0AE1578C7933191AFED2F913FEBCF944" box="[299,362,1731,1757]" pageId="4" pageNumber="272" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7933191BFF3EF936FA45FF60" lastPageId="5" lastPageNumber="273" pageId="4" pageNumber="272" type="description">
<paragraph id="231936E57933191AFF3EF936FD54F840" blockId="4.[151,1437,1622,2009]" pageId="4" pageNumber="272">
<emphasis id="11D2EAF77933191AFF3EF936FE87F899" bold="true" box="[199,337,1766,1792]" pageId="4" pageNumber="272">Description</emphasis>
. Wing expanse: 13.7 mm female (n = 1). Female (
<figureCitation id="BB9D2A607933191AFC5AF937FC3DF898" box="[931,1003,1767,1793]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="4" pageNumber="272">Fig. 2</figureCitation>
). As in
<taxonomicName id="E4A64D667933191AFBB3F937FB6DF898" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[1098,1211,1767,1793]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77933191AFBB3F937FB6DF898" box="[1098,1211,1767,1793]" italics="true" pageId="4" pageNumber="272">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77933191AFB3CF937FAD0F898" bold="true" box="[1221,1286,1767,1793]" pageId="4" pageNumber="272">
<taxonomicNameLabel id="0AE1578C7933191AFB3CF937FAD0F898" box="[1221,1286,1767,1793]" pageId="4" pageNumber="272" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
but with the following particulars:
<emphasis id="11D2EAF77933191AFE61F8DBFE03F8BC" box="[408,469,1803,1829]" italics="true" pageId="4" pageNumber="272">Head</emphasis>
(
<figureCitation id="BB9D2A607933191AFE11F8DBFD8DF8BC" box="[488,603,1803,1829]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="4" pageNumber="272">Fig. 2a–b</figureCitation>
): caudo-lateral scale tufts white, to slightly brown-gray caudally; labial palpus white, with many tan to brown-gray scales laterally on segment two and more so on basal segment; antenna scape tan.
<emphasis id="11D2EAF77933191AFEF6F883FEB6F8F4" box="[271,352,1875,1901]" italics="true" pageId="4" pageNumber="272">Thorax</emphasis>
: Dark brown mottled with white anteriorly, with many scales dark orange-tipped (especially on metathorax), and some maroon scales mixed in.
<emphasis id="11D2EAF77933191AFD54F8A7FCCFF809" box="[685,793,1911,1936]" italics="true" pageId="4" pageNumber="272">Forewing</emphasis>
(
<figureCitation id="BB9D2A607933191AFCDEF8A7FCBCF808" box="[807,874,1911,1937]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="4" pageNumber="272">Fig. 2</figureCitation>
) darker than in
<taxonomicName id="E4A64D667933191AFBEEF8A7FB53F808" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[1047,1157,1911,1937]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77933191AFBEEF8A7FB53F808" box="[1047,1157,1911,1937]" italics="true" pageId="4" pageNumber="272">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77933191AFB73F8A7FB1EF808" bold="true" box="[1162,1224,1911,1937]" pageId="4" pageNumber="272">
<taxonomicNameLabel id="0AE1578C7933191AFB73F8A7FB1EF808" box="[1162,1224,1911,1937]" pageId="4" pageNumber="272" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
with much smaller areas of white scaling, and subterminal fascia with median knob-like extension very short.
<emphasis id="11D2EAF77933191AFB7EF84BFB23F82C" box="[1159,1269,1947,1973]" italics="true" pageId="4" pageNumber="272">Hindwing</emphasis>
(
<figureCitation id="BB9D2A607933191AFAFDF84BFA90F82C" box="[1284,1350,1947,1973]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1794,1818]" captionTargetBox="[152,1436,487,1768]" captionTargetId="figure-114@2.[151,1436,487,1768]" captionTargetPageId="2" captionText="FIGURES 1–3. Adult features of Cirrothaumatia species, with details of the head laterally (a) and dorsally (b): 1) C. pichita n. sp., female holotype (16.2mm wingspan), Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru (adult photo 15533) (MGCL); 2) C. huemboana n. sp., female holotype (13.7mm wingspan), Huembo Reserve (2085m), La Florida, Amazonas, Peru (adult photo 15857) (MGCL); 3) C. moyobamba n. sp., female holotype (12.4mm wingspan), Rumipata Lodge (970m), Moyobamba, San Martín, Peru (adult photo 15858) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726421" httpUri="https://zenodo.org/record/10726421/files/figure.png" pageId="4" pageNumber="272">Fig. 2</figureCitation>
) brown and less translucent than in
<taxonomicName id="E4A64D667933191AFE34F86FFDEAF840" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[461,572,1983,2009]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="272" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77933191AFE34F86FFDEAF840" box="[461,572,1983,2009]" italics="true" pageId="4" pageNumber="272">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77933191AFDBAF86FFD54F840" bold="true" box="[579,642,1983,2009]" pageId="4" pageNumber="272">
<taxonomicNameLabel id="0AE1578C7933191AFDBAF86FFD54F840" box="[579,642,1983,2009]" pageId="4" pageNumber="272" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="231936E57932191BFF3EFF47FA45FF60" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFF47FEE7FF28" box="[199,305,151,177]" italics="true" pageId="5" pageNumber="273">Abdomen</emphasis>
. Female genitalia (
<figureCitation id="BB9D2A607932191BFDEBFF47FD8EFF28" box="[530,600,150,177]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="5" pageNumber="273">Fig. 5</figureCitation>
) as in
<taxonomicName id="E4A64D667932191BFD54FF47FCC9FF28" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[685,799,151,177]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="pichita" status="n. sp.">
<emphasis id="11D2EAF77932191BFD54FF47FCC9FF28" box="[685,799,151,177]" italics="true" pageId="5" pageNumber="273">C. pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77932191BFCD0FF47FCBDFF28" bold="true" box="[809,875,151,177]" pageId="5" pageNumber="273">
<taxonomicNameLabel id="0AE1578C7932191BFCD0FF47FCBDFF28" box="[809,875,151,177]" pageId="5" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but sterigma band slightly broader and without small ventral triangular caudal extension; ostial funnel (
<figureCitation id="BB9D2A607932191BFCFAFF6BFC85FF4C" box="[771,851,186,213]" captionStart="FIGURES 4–6" captionStartId="3.[151,264,1352,1376]" captionTargetBox="[151,1436,182,1328]" captionTargetId="figure-28@3.[151,1436,181,1328]" captionTargetPageId="3" captionText="FIGURES 4–6. Female genitalia of Cirrothaumatia species, with details of the ostium, sterigma, and caudal ductus bursae region (a): 4) C. pichita n. sp., holotype female (Cerro Pichita Res. Sta. (2165m), San Ramon, Junín, Peru) (gen. slide JBH- 4361) (MGCL); 5) C. huemboana n. sp., holotype female (Huembo Reserve (2085m), La Florida, Amazonas, Peru) (gen. slide JBH-4469) (MGCL); 6) C. moyobamba n. sp., holotype female (Rumipata Lodge (970m), Moyobamba, San Martín, Peru) (gen. slide JBH-4470) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.10726425" httpUri="https://zenodo.org/record/10726425/files/figure.png" pageId="5" pageNumber="273">Fig. 5a</figureCitation>
) straight and uniformly broad, short stentor-like, but ductus bursae with more ridges along its length; accessory bursa very small and from short duct.
<emphasis id="11D2EAF77932191BFB2CFF0FFAC1FF60" box="[1237,1303,223,249]" italics="true" pageId="5" pageNumber="273">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EFED3FE52FEFC" pageId="5" pageNumber="273" type="materials_examined">
<paragraph id="231936E57932191BFF3EFED3FE52FEFC" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFED3FE56FE85" bold="true" box="[199,384,258,285]" pageId="5" pageNumber="273">Type materials.</emphasis>
<materialsCitation id="93CE3CB87932191BFE71FED3FE52FEFC" collectingDate="2012-10-18" collectingDateMax="2012-10-20" collectingDateMin="2012-10-18" collectionCode="MGCL, UNALM" collectorName="La Florida &amp; J. B. Heppner &amp; C. Carrera" country="Peru" county="La Florida" elevation="2085" location="Female" municipality="Huembo Reserve" pageId="5" pageNumber="273" specimenCount="1" typeStatus="holotype">
<emphasis id="11D2EAF77932191BFE71FED3FE23FE84" bold="true" box="[392,501,259,285]" pageId="5" pageNumber="273">
<typeStatus id="FC1D88477932191BFE71FED3FE23FE84" box="[392,501,259,285]" pageId="5" pageNumber="273" type="holotype">Holotype</typeStatus>
</emphasis>
.
<location id="2679603E7932191BFDFAFED3FD8EFE84" LSID="urn:lsid:plazi:treatment:AB0F87F37933191BFF6EFA3AFA5DFD54:2679603E7932191BFDFAFED3FD8EFE84" box="[515,600,259,285]" country="Peru" county="La Florida" municipality="Huembo Reserve" name="Female" pageId="5" pageNumber="273">Female</location>
,
<collectingMunicipality id="C37DAC9F7932191BFD9DFED3FCF0FE84" box="[612,806,259,285]" pageId="5" pageNumber="273">Huembo Reserve</collectingMunicipality>
(
<quantity id="E45E9B007932191BFCCCFED2FC59FE84" box="[821,911,258,285]" metricMagnitude="3" metricUnit="m" metricValue="2.085" pageId="5" pageNumber="273" unit="m" value="2085.0">
<elevation id="A88BD1D67932191BFCCCFED2FC59FE84" box="[821,911,258,285]" metricMagnitude="3" metricUnit="m" metricValue="2.085" pageId="5" pageNumber="273" unit="m" value="2085.0">2085 m</elevation>
</quantity>
), nr.
<collectorName id="8E5353337932191BFC31FED3FB97FE84" box="[968,1089,259,285]" pageId="5" pageNumber="273">
<collectingCounty id="CA784E697932191BFC31FED3FB97FE84" box="[968,1089,259,285]" pageId="5" pageNumber="273">La Florida</collectingCounty>
</collectorName>
, Dept. Amazonas,
<collectingCountry id="5BB176757932191BFAEDFED3FA9CFE84" box="[1300,1354,259,285]" name="Peru" pageId="5" pageNumber="273">Peru</collectingCountry>
,
<date id="571810257932191BFAAFFED3FED7FED8" pageId="5" pageNumber="273" value="2012-10-18" valueMax="2012-10-20" valueMin="2012-10-18">
<collectingDate id="475CE9CD7932191BFAAFFED3FED7FED8" pageId="5" pageNumber="273" value="2012-10-18" valueMax="2012-10-20" valueMin="2012-10-18">18‒20 Oct 2012</collectingDate>
</date>
,
<collectorName id="8E5353337932191BFEF5FEF7FE73FED8" box="[268,421,295,321]" pageId="5" pageNumber="273">J. B. Heppner</collectorName>
&amp;
<collectorName id="8E5353337932191BFE31FEF7FDEAFED8" box="[456,572,295,321]" pageId="5" pageNumber="273">C. Carrera</collectorName>
(gen. slide JBH-4469; adult photo 15857) (deposited at
<collectionCode id="45B7AE207932191BFB49FEF7FAD2FED8" box="[1200,1284,295,321]" pageId="5" pageNumber="273">MGCL</collectionCode>
, on indefinite loan from
<collectionCode id="45B7AE207932191BFEF5FE9BFEAEFEFC" box="[268,376,331,357]" pageId="5" pageNumber="273">UNALM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EFEBFFD35FE34" pageId="5" pageNumber="273" type="etymology">
<paragraph id="231936E57932191BFF3EFEBFFD35FE34" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFEBFFE9EFE10" bold="true" box="[199,328,367,393]" pageId="5" pageNumber="273">Etymology</emphasis>
. The species is named after the Huembo Reserve, near the small town of La Florida (ca.
<quantity id="E45E9B007932191BFAE3FEBFFA9DFE10" box="[1306,1355,367,393]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="273" unit="km" value="7.0">7km</quantity>
distant on the main road eastwards), Dept. Amazonas,
<collectingCountry id="5BB176757932191BFD51FE43FD09FE34" box="[680,735,403,429]" name="Peru" pageId="5" pageNumber="273">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EFE66FC49FE48" box="[199,927,438,465]" pageId="5" pageNumber="273" type="biology_ecology">
<paragraph id="231936E57932191BFF3EFE66FC49FE48" blockId="5.[151,1437,150,717]" box="[199,927,438,465]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFE66FEF7FE49" bold="true" box="[199,289,438,464]" pageId="5" pageNumber="273">Biology</emphasis>
. Unknown. Habitat submontane Andean forest (
<figureCitation id="BB9D2A607932191BFCC6FE67FC47FE48" box="[831,913,439,465]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1718,1742]" captionTargetBox="[189,1399,181,1694]" captionTargetId="figure-28@7.[189,1399,181,1694]" captionTargetPageId="7" captionText="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." figureDoi="http://doi.org/10.5281/zenodo.10726433" httpUri="https://zenodo.org/record/10726433/files/figure.png" pageId="5" pageNumber="273">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EFE0AFF15FDA4" pageId="5" pageNumber="273" type="distribution">
<paragraph id="231936E57932191BFF3EFE0AFF15FDA4" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFE0AFE8FFE6D" bold="true" box="[199,345,474,500]" pageId="5" pageNumber="273">Distribution</emphasis>
. Known only from the eastern Andean submontane site at Huembo Reserve (
<quantity id="E45E9B007932191BFB33FE0AFAF5FE6C" box="[1226,1315,474,501]" metricMagnitude="3" metricUnit="m" metricValue="2.085" pageId="5" pageNumber="273" unit="m" value="2085.0">2085 m</quantity>
) (
<figureCitation id="BB9D2A607932191BFAC2FE0BFA58FE6C" box="[1339,1422,475,501]" captionStart="FIGURES 10–17" captionStartId="7.[151,264,1718,1742]" captionTargetBox="[189,1399,181,1694]" captionTargetId="figure-28@7.[189,1399,181,1694]" captionTargetPageId="7" captionText="FIGURES 10–17. Habitats of Cirrothaumatia species: 10) type-locality of C. pichita n. sp., Cerro Pichita Res. Sta. (2165 m), San Ramon, Junín, Peru (Nov 2012, JBH); 11) type-locality of C. huemboana n. sp., Huembo Reserve (2085 m), La Florida, Amazonas, Peru (Oct 2012, JBH); 12) type-locality of C. moyobamba n. sp., Rumipata Lodge (970 m), Moyobamba, San Martín, Peru (Oct 2012, JBH); 13) type- locality of C. vesta (Clarke), Rancho Grande (1100 m), Aragua, Venezuela (Venezuela Tourism photo); 14) type-locality of C. tornosema, Volcan Santa Maria (3810 m), with forest sites ca. 800–2000 m, Quetzaltenango, Guatemala (Guatemala Tourism photo); 15) site in Mexico for C. tornosema (Clarke), Huatusco (1300 m), Veracruz, Mexico (Volcan Orizaba (5636 m) in background) (Mexico Tourism photo); 16) site in Costa Rica for C. tornosema, Braulio Carrillo Natl. Park (forest ca. 1100 m), Heredia, Costa Rica (Costa Rica Natl. Pks. photo); 17) type-locality of C. tornocarpa (Meyrick), Lino (ca. 800 m), Boquete Dist., Chiriquí, Panama (Panama Tourism photo)." figureDoi="http://doi.org/10.5281/zenodo.10726433" httpUri="https://zenodo.org/record/10726433/files/figure.png" pageId="5" pageNumber="273">Fig. 11</figureCitation>
), a mountain upland area above a deep canyon, near the town of La Florida, Dept. Amazonas, in northern
<collectingCountry id="5BB176757932191BFAD5FE2FFA89FD80" box="[1324,1375,511,537]" name="Peru" pageId="5" pageNumber="273">Peru</collectingCountry>
(
<figureCitation id="BB9D2A607932191BFA96FE2FFF60FDA4" captionStart="FIGURE 18" captionStartId="8.[151,250,1873,1897]" captionTargetBox="[218,1370,665,1850]" captionTargetId="figure-209@8.[218,1370,665,1850]" captionTargetPageId="8" captionText="FIGURE 18. Distribution map of known localities for Cirrothaumatia species (NGS base map): C. huemboana n. sp., yellow square (Peru:Amazonas). C. moyobamba n. sp., black-bordered white square (Peru: San Martín). C. pichita n. sp., white square (Peru: Junín). C. tornocarpa (Meyrick), red square (Panama: Chiriquí). C. tornosema (Clarke), black squares (Mexico:Veracruz; Guatemala: Quetzaltenango; Costa Rica: Cartago, Heredia, and Puntarenas; and Ecuador: Pichincha). C. vesta (Clarke), blue square (Venezuela: Aragua)." figureDoi="http://doi.org/10.5281/zenodo.10726435" httpUri="https://zenodo.org/record/10726435/files/figure.png" pageId="5" pageNumber="273">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6BBC656E7932191BFF3EFD96FA5DFD54" pageId="5" pageNumber="273" type="discussion">
<paragraph id="231936E57932191BFF3EFD96FA5DFD54" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="273">
<emphasis id="11D2EAF77932191BFF3EFD96FE93FDF9" bold="true" box="[199,325,582,608]" pageId="5" pageNumber="273">Discussion</emphasis>
.
<taxonomicName id="E4A64D667932191BFEADFD97FD8FFDF8" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[340,601,583,609]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="pichita" status="sp. nov.">
<emphasis id="11D2EAF77932191BFEADFD97FD8FFDF8" box="[340,601,583,609]" italics="true" pageId="5" pageNumber="273">Cirrothaumatia pichita</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77932191BFD98FD97FD76FDF8" bold="true" box="[609,672,583,609]" pageId="5" pageNumber="273">
<taxonomicNameLabel id="0AE1578C7932191BFD98FD97FD76FDF8" box="[609,672,583,609]" pageId="5" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="E4A64D667932191BFD20FD97FCA8FDF8" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[729,894,583,609]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="huemboana" status="n. sp.">
<emphasis id="11D2EAF77932191BFD20FD97FCA8FDF8" box="[729,894,583,609]" italics="true" pageId="5" pageNumber="273">C. huemboana</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77932191BFC7CFD97FC13FDF8" bold="true" box="[901,965,583,609]" pageId="5" pageNumber="273">
<taxonomicNameLabel id="0AE1578C7932191BFC7CFD97FC13FDF8" box="[901,965,583,609]" pageId="5" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are clearly sister-species, but they can be separated by maculation and genital features. Discovery of the males of the two may provide further distinctions. The seemingly slight maculation differences between these two species can be interpreted as significant when compared to
<taxonomicName id="E4A64D667932191BFED0FD63FE05FD54" authority="Heppner &amp; Bae, 2024" authorityName="Heppner &amp; Bae" authorityYear="2024" box="[297,467,691,717]" class="Insecta" family="Tortricidae" genus="Cirrothaumatia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="273" phylum="Arthropoda" rank="species" species="moyobamba" status="n. sp.">
<emphasis id="11D2EAF77932191BFED0FD63FE05FD54" box="[297,467,691,717]" italics="true" pageId="5" pageNumber="273">C. moyobamba</emphasis>
</taxonomicName>
<emphasis id="11D2EAF77932191BFE23FD63FDCFFD54" bold="true" box="[474,537,691,717]" pageId="5" pageNumber="273">
<taxonomicNameLabel id="0AE1578C7932191BFE23FD63FDCFFD54" box="[474,537,691,717]" pageId="5" pageNumber="273" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, which also has similar maculation and yet has very different female genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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