381 lines
49 KiB
XML
381 lines
49 KiB
XML
<document id="DB319E17D1D0D76D045851C1D101245B" ID-DOI="10.11646/zootaxa.4804.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3927094" ID-ZooBank="8C1C1703-9BBC-4B33-8045-78BDD9738F51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593672611012" checkinUser="plazi" docAuthor="Westrop, Madison Armstrong Stephen R. & Eoff, Jennifer D." docDate="2020" docId="03F76D37FFBBFFDE6BD2E01EFB9892FD" docLanguage="en" docName="zootaxa.4804.1.1.pdf" docOrigin="Zootaxa 4804 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maryvillia arion Walcott 1916" docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="FFCE154FFFAFFFC86B45E438FF8E9206" masterDocTitle="Systematics of a survivor: the Cambrian kingstoniid trilobite Blountia Walcott, 1916 across the Marjuman-Steptoean (Guzhangian-Paibian) extinction interval in Laurentian North America" masterLastPageNumber="79" masterPageNumber="1" pageNumber="21" updateTime="1698776919363" updateUser="plazi">
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<mods:title id="EEF86917E4006D2AE4B5DECA4CABE5BF">Systematics of a survivor: the Cambrian kingstoniid trilobite Blountia Walcott, 1916 across the Marjuman-Steptoean (Guzhangian-Paibian) extinction interval in Laurentian North America</mods:title>
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<mods:namePart id="A46E2F4C8842E6A368B7D98777A3AA8F">Westrop, Madison Armstrong Stephen R.</mods:namePart>
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<mods:namePart id="A9A8DC70433023C564CACD7560741553">Eoff, Jennifer D.</mods:namePart>
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<mods:title id="6D9CD1AC91C289307DBAB4724FA70343">Zootaxa</mods:title>
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<mods:part id="4D00D5487BD539532F5C0B2EA4086A89">
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<mods:date id="0DD35D79ABE9C4331146DB43881C34C5">2020</mods:date>
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<mods:number id="2FF66D351DFF76D8D6C3868061F288B8">2020-06-30</mods:number>
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<mods:number id="9465D344A1C093E2F827C78963F13CEC">4804</mods:number>
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<mods:classification id="EFB094D745CC02BA39ABA0C5A8459768">journal article</mods:classification>
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<mods:identifier id="B770BCB9D2803A42413433E4F1A81394" type="DOI">10.11646/zootaxa.4804.1.1</mods:identifier>
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<mods:identifier id="A931C99F10763C679FF30454E80AC490" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F76D37FFBBFFDE6BD2E01EFB9892FD" LSID="urn:lsid:plazi:treatment:03F76D37FFBBFFDE6BD2E01EFB9892FD" httpUri="http://treatment.plazi.org/id/03F76D37FFBBFFDE6BD2E01EFB9892FD" lastPageId="22" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="C3448FAAFFBBFFDC6BD2E01EFD819646" box="[151,527,1062,1089]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BE1DC21FFBBFFDC6BD2E01EFD819646" blockId="20.[151,527,1062,1124]" box="[151,527,1062,1089]" pageId="20" pageNumber="21">
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<heading id="D0A96B4DFFBBFFDC6BD2E01EFD819646" bold="true" box="[151,527,1062,1089]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<taxonomicName id="4C5EA7A2FFBBFFDC6BD2E01EFD819646" ID-CoL="6R7KL" authority="Walcott, 1916" authorityName="Walcott" authorityYear="1916" box="[151,527,1062,1089]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arion">
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<emphasis id="B92A0033FFBBFFDC6BD2E01EFD819646" bold="true" box="[151,527,1062,1089]" pageId="20" pageNumber="21">
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<emphasis id="B92A0033FFBBFFDC6BD2E01EFED39646" bold="true" box="[151,349,1062,1089]" italics="true" pageId="20" pageNumber="21">Maryvillia arion</emphasis>
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Walcott, 1916
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3448FAAFFBBFFDC6BD2E072FE1E977E" pageId="20" pageNumber="21" type="description">
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<paragraph id="8BE1DC21FFBBFFDC6BD2E072FEA99662" blockId="20.[151,527,1062,1124]" box="[151,295,1098,1124]" pageId="20" pageNumber="21">
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<figureCitation id="1365C0A4FFBBFFDC6BD2E072FE8E9662" box="[151,256,1098,1124]" captionStart="PLATE 22" captionStartId="68.[151,234,1868,1893]" captionText="PLATE 22. Maryvillia arion Walcott, 1916, all from the Nolichucky Formation; 0.8 km (0.5 mile) east of Rogersville, Hawkins County, Tennessee (USNM locality 123b), except figs 9–11 (from Rasetti’s 1965 locality cnm/2, also in the Rogersville area) 1–3. Cranidium, holotype, USNM 62826, dorsal, anterior and lateral views, x8 (illustrated by Walcott 1916, pl. 64, figs 4, 4’, 4b). 4–7. Cranidium, paratype, USNM 62826, anterior, dorsal, lateral, and posterior views, x5 (previously unillustrated). 8–11. Cranidium, USNM 144603, anterior, posterior, dorsal and lateral views, x4.75 (illustrated by Rasetti 1965, pl. 9, figs 22, 23)." pageId="20" pageNumber="21">Plates 22</figureCitation>
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,
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<figureCitation id="1365C0A4FFBBFFDC6A4EE072FEA99662" box="[267,295,1098,1124]" captionStart="PLATE 23" captionStartId="70.[151,234,1751,1776]" captionText="PLATE 23. Maryvillia arion Walcott, 1916, all from the Nolichucky Formation; 0.8 km (0.5 mile) east of Rogersville, Hawkins County, Tennessee (U.S.N.M locality 123b), except figs 5, 8, 9 (Rasetti’s 1965 locality cnm/2), and figs 6, 7 (Rasetti’s 1965 locality cnn/1), both of which are also from the Rogersville area. 1, 2. Pygidium, paratype (also holotype of Blountia arguta Resser, 1942a), USNM 62827, dorsal and posterior views, x10 (illustrated by Walcott 1916, pl. 64, figs 4a, 4a’). 3, 4. Pygidium, USNM 94968 (paratype of Blountia arguta Resser, 1942a), dorsal and posterior views, x4.5. 5. Pygidium, USNM 144603, dorsal view, x3.75 (illustrated previously by Rasetti 1965, pl. 9, fig. 24). 6, 7. Pygidium, USNM 144604, dorsal and posterior views, x7.5 (illustrated by Rasetti 1965, pl. 9, fig. 26). 8, 9. Pygidium, USNM 144603, posterior and dorsal views, x3.75 (illustrated by Rasetti 1965, pl. 9, fig. 25)." pageId="20" pageNumber="21">23</figureCitation>
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</paragraph>
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<paragraph id="8BE1DC21FFBBFFDC6BD2E0B0FD5E96A7" blockId="20.[151,1437,1160,1400]" box="[151,720,1160,1185]" pageId="20" pageNumber="21">
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1916
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<taxonomicName id="4C5EA7A2FFBBFFDC6B95E0B1FE4196A7" authority="Walcott" authorityName="Walcott" authorityYear="1916" box="[208,463,1160,1185]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arion">
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<emphasis id="B92A0033FFBBFFDC6B95E0B1FEF696A7" box="[208,376,1161,1185]" italics="true" pageId="20" pageNumber="21">Maryvillia arion</emphasis>
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Walcott
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</taxonomicName>
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, p. 400, pl. 64, figs 4–4c.
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</paragraph>
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<paragraph id="8BE1DC21FFBBFFDC6BD2E090FDD796C6" blockId="20.[151,1437,1160,1400]" box="[151,601,1191,1216]" pageId="20" pageNumber="21">
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1942a
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<taxonomicName id="4C5EA7A2FFBBFFDC6B9EE090FE5A96C6" authority="Walcott" authorityName="Walcott" box="[219,468,1191,1216]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arguta">
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<emphasis id="B92A0033FFBBFFDC6B9EE090FEF296C6" box="[219,380,1192,1216]" italics="true" pageId="20" pageNumber="21">Blountia arguta</emphasis>
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Walcott
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</taxonomicName>
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; Resser, p. 8.
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</paragraph>
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<paragraph id="8BE1DC21FFBBFFDC6BD2E0FEFC3196D9" blockId="20.[151,1437,1160,1400]" box="[151,959,1222,1247]" pageId="20" pageNumber="21">
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1956
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<taxonomicName id="4C5EA7A2FFBBFFDC6B95E0FEFE5E96D9" authority="Walcott" authorityName="Walcott" authorityYear="1916" box="[208,464,1222,1247]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arion">
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<emphasis id="B92A0033FFBBFFDC6B95E0FEFEF696D8" box="[208,376,1222,1246]" italics="true" pageId="20" pageNumber="21">Maryvillia arion</emphasis>
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Walcott
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</taxonomicName>
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; Rasetti, p. 1267 (includes complete synonymy).
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</paragraph>
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<paragraph id="8BE1DC21FFBBFFDC6BD2E0DCFCBF96FB" blockId="20.[151,1437,1160,1400]" box="[151,817,1252,1277]" pageId="20" pageNumber="21">
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1961
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<taxonomicName id="4C5EA7A2FFBBFFDC6B95E0DDFE5E96FB" authority="Walcott" authorityName="Walcott" authorityYear="1916" box="[208,464,1252,1277]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arion">
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<emphasis id="B92A0033FFBBFFDC6B95E0DDFEF696FB" box="[208,376,1253,1277]" italics="true" pageId="20" pageNumber="21">Maryvillia arion</emphasis>
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Walcott
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</taxonomicName>
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; Rasetti, p. 116, pl. 21, figs 14–15.
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</paragraph>
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<paragraph id="8BE1DC21FFBBFFDC6BD2E13BFC97971A" blockId="20.[151,1437,1160,1400]" box="[151,793,1283,1308]" pageId="20" pageNumber="21">
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1965
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<taxonomicName id="4C5EA7A2FFBBFFDC6B95E13CFE5E971A" authority="Walcott" authorityName="Walcott" authorityYear="1916" box="[208,464,1283,1308]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arion">
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<emphasis id="B92A0033FFBBFFDC6B95E13CFEF6971A" box="[208,376,1284,1308]" italics="true" pageId="20" pageNumber="21">Maryvillia arion</emphasis>
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Walcott
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</taxonomicName>
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; Rasetti, p. 59, pl. 9, figs 22–26.
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</paragraph>
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<paragraph id="8BE1DC21FFBBFFDC6BD2E11BFBB6973D" blockId="20.[151,1437,1160,1400]" box="[151,1080,1314,1339]" pageId="20" pageNumber="21">
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<emphasis id="B92A0033FFBBFFDC6BD2E11BFF33973C" box="[151,189,1315,1338]" italics="true" pageId="20" pageNumber="21">non</emphasis>
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1992 cf.
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<taxonomicName id="4C5EA7A2FFBBFFDC6A58E11AFD94973D" authority="(Walcott)" baseAuthorityName="Walcott" box="[285,538,1314,1339]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arion">
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<emphasis id="B92A0033FFBBFFDC6A58E11AFE3F973C" box="[285,433,1314,1338]" italics="true" pageId="20" pageNumber="21">Blountia arion</emphasis>
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(Walcott)
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</taxonomicName>
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; Pratt, p. 66, pl. 23, figs 22–25 [=
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<taxonomicName id="4C5EA7A2FFBBFFDC6830E11AFC42973C" authorityName="Walcott" authorityYear="1916" box="[885,972,1314,1338]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="B92A0033FFBBFFDC6830E11AFC42973C" box="[885,972,1314,1338]" italics="true" pageId="20" pageNumber="21">Blountia</emphasis>
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</taxonomicName>
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sp. indet].
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</paragraph>
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<paragraph id="8BE1DC21FFBBFFDC6BD2E17AFE1E977E" blockId="20.[151,1437,1160,1400]" pageId="20" pageNumber="21">
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<emphasis id="B92A0033FFBBFFDC6BD2E17AFF33975F" box="[151,189,1346,1369]" italics="true" pageId="20" pageNumber="21">non</emphasis>
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2000
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<taxonomicName id="4C5EA7A2FFBBFFDC6BBBE179FE73975F" authority="(Walcott)" baseAuthorityName="Walcott" box="[254,509,1344,1369]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arion">
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<emphasis id="B92A0033FFBBFFDC6BBBE179FE1A975F" box="[254,404,1345,1369]" italics="true" pageId="20" pageNumber="21">Blountia arion</emphasis>
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(Walcott)
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</taxonomicName>
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; Stitt & Perfetta, p. fig. 11.4, 11.5 [=
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<taxonomicName id="4C5EA7A2FFBBFFDC683BE179FC67975F" authorityName="Walcott" authorityYear="1916" box="[894,1001,1345,1369]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<emphasis id="B92A0033FFBBFFDC683BE179FC67975F" box="[894,1001,1345,1369]" italics="true" pageId="20" pageNumber="21">Maryvillia</emphasis>
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</taxonomicName>
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cf.
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<taxonomicName id="4C5EA7A2FFBBFFDC6F57E17AFE08977E" authority="Lochman, in Lochman and Duncan, 1944" authorityName="Lochman, in Lochman and Duncan" authorityYear="1944" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="triangularis">
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<emphasis id="B92A0033FFBBFFDC6F57E17AFBA9975F" box="[1042,1063,1346,1369]" italics="true" pageId="20" pageNumber="21">M</emphasis>
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.
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<emphasis id="B92A0033FFBBFFDC6F71E179FB23975F" box="[1076,1197,1345,1369]" italics="true" pageId="20" pageNumber="21">triangularis</emphasis>
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Lochman, in
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<bibRefCitation id="EFCFA1D0FFBBFFDC6E78E178FE08977E" author="Lochman, C. & Duncan, D." pageId="20" pageNumber="21" pagination="1 - 172" refId="ref23573" refString="Lochman, C. & Duncan, D. (1944) Early Upper Cambrian faunas of Central Montana. Geological Society of America Special Papers, 54, 1 - 172. https: // doi. org / 10.1130 / SPE 54 - p 1" type="journal article" year="1944">Lochman and Duncan, 1944</bibRefCitation>
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</taxonomicName>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="C3448FAAFFBBFFDC6BD2E198FE059495" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="8BE1DC21FFBBFFDC6BD2E198FE059495" blockId="20.[151,1437,1440,2043]" pageId="20" pageNumber="21">
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<emphasis id="B92A0033FFBBFFDC6BD2E198FE9C97BD" bold="true" box="[151,274,1440,1467]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
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Evenly rounded anterior margin of cranidium. Frontal area occupies one-third (32%; 31–33) cranidial length. Steeply sloping preglabellar field grades into long (occupies two-thirds [66%; 65–67] of frontal area length), concave anterior border across broad, very shallow border furrow. Subtrapezoidal glabella barely elevated above fixigenae. Pygidium semielliptical, length (sag.) equal to 70% (68–74) of maximum width, with smooth external surface, shallow furrows. Axis extends beyond border furrow onto down-sloping border. Pitting on internal mould (corresponding to small projections on underside of exoskeleton) is bimodal, with larger pits scattered among small- er ones (Pl. 22 fig. 5).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3448FAAFFBBFFDD6B82E2A4FD1E92D1" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="materials_examined">
|
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<paragraph id="8BE1DC21FFBBFFDC6B82E2A4FAB095B2" blockId="20.[151,1437,1440,2043]" pageId="20" pageNumber="21">
|
||
<emphasis id="B92A0033FFBBFFDC6B82E2A4FEBA94B1" bold="true" box="[199,308,1692,1719]" pageId="20" pageNumber="21">Material.</emphasis>
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<typeStatus id="54E56283FFBBFFDC6A79E2A5FE2C94B1" box="[316,418,1693,1719]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||
cranidium (
|
||
<collectionCode id="ED4F44E4FFBBFFDC6962E2A5FDF694B1" box="[551,632,1693,1719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
62826; Pl. 22, figs 1–3), one
|
||
<typeStatus id="54E56283FFBBFFDC6881E2A5FBAC94B1" box="[964,1058,1693,1719]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
|
||
cranidium (
|
||
<collectionCode id="ED4F44E4FFBBFFDC6FEDE2A5FB7794B1" box="[1192,1273,1693,1719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
62826, Pl. 22, figs 4–7) and one
|
||
<typeStatus id="54E56283FFBBFFDC6A1FE2F9FE3694DD" box="[346,440,1729,1755]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
|
||
pygidium (
|
||
<collectionCode id="ED4F44E4FFBBFFDC6971E2F9FD0B94DD" box="[564,645,1729,1755]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
62827, Pl. 23, figs 1, 2), all from the Nolichucky Formation; 0.8 km (0.5 mile) east of Rogersville, Hawkins County,
|
||
<collectingRegion id="499A12C3FFBBFFDC6931E2DDFD6A94F9" box="[628,740,1765,1791]" country="United States of America" name="Tennessee" pageId="20" pageNumber="21">Tennessee</collectingRegion>
|
||
(
|
||
<collectionCode id="ED4F44E4FFBBFFDC69B7E2DDFCCD94F9" box="[754,835,1765,1791]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
locality 123b). A pygidium (
|
||
<collectionCode id="ED4F44E4FFBBFFDC6F38E2DDFB4094F9" box="[1149,1230,1765,1791]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
94968, Pl. 23, figs 3, 4) that was designated by
|
||
<bibRefCitation id="EFCFA1D0FFBBFFDC6A9CE331FD0E9525" author="Resser, C. E." box="[473,640,1801,1827]" pageId="20" pageNumber="21" pagination="1 - 58" refId="ref23949" refString="Resser, C. E. (1942 a) Fifth contribution to nomenclature of Cambrian fossils. Smithsonian Miscellaneous Collections, 101, 1 - 58." type="journal article" year="1942">Resser (1942a)</bibRefCitation>
|
||
as a
|
||
<typeStatus id="54E56283FFBBFFDC69F9E331FC949525" box="[700,794,1801,1827]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5EA7A2FFBBFFDC6807E331FC7E9525" box="[834,1008,1801,1827]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arguta">
|
||
<emphasis id="B92A0033FFBBFFDC6807E331FC7E9525" box="[834,1008,1801,1827]" italics="true" pageId="20" pageNumber="21">Blountia arguta</emphasis>
|
||
</taxonomicName>
|
||
is a previously unillustrated specimen from the type lot of
|
||
<taxonomicName id="4C5EA7A2FFBBFFDC6A34E316FE5E9541" box="[369,464,1837,1863]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arion">
|
||
<emphasis id="B92A0033FFBBFFDC6A34E316FE069541" box="[369,392,1838,1863]" italics="true" pageId="20" pageNumber="21">M</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFBBFFDC6AD0E316FE5E9541" box="[405,464,1838,1863]" italics="true" pageId="20" pageNumber="21">arion</emphasis>
|
||
</taxonomicName>
|
||
. Non-type specimens, also from the Nolichucky Formation in the Rogersville region, are a cranidium (
|
||
<collectionCode id="ED4F44E4FFBBFFDC6A63E369FEF9956D" box="[294,375,1873,1899]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
144603, Pl. 22, figs 8–11) and two pygidia (
|
||
<collectionCode id="ED4F44E4FFBBFFDC681CE369FC24956D" box="[857,938,1873,1899]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
144603, Pl. 23, fig. 5;
|
||
<collectionCode id="ED4F44E4FFBBFFDC6FE4E369FB7F956D" box="[1185,1265,1873,1899]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
144603, Pl. 23, figs 8, 9) from
|
||
<bibRefCitation id="EFCFA1D0FFBBFFDC6A7FE34DFE699589" author="Rasetti, F." box="[314,487,1908,1935]" pageId="20" pageNumber="21" pagination="1 - 127" refId="ref23892" refString="Rasetti, F. (1965) Upper Cambrian trilobite faunas of Northeastern Tennessee. Smithsonian Miscellaneous Collections, 148, 1 - 127." type="journal article" year="1965">Rasetti’s (1965)</bibRefCitation>
|
||
locality cnm/2, and a pygidium (
|
||
<collectionCode id="ED4F44E4FFBBFFDC680AE34DFC2E9589" box="[847,928,1909,1935]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
144604, Pl. 23, figs 6.7) from
|
||
<bibRefCitation id="EFCFA1D0FFBBFFDC6FAAE34DFA129589" author="Rasetti, F." box="[1263,1436,1908,1935]" pageId="20" pageNumber="21" pagination="1 - 127" refId="ref23892" refString="Rasetti, F. (1965) Upper Cambrian trilobite faunas of Northeastern Tennessee. Smithsonian Miscellaneous Collections, 148, 1 - 127." type="journal article" year="1965">Rasetti’s (1965)</bibRefCitation>
|
||
locality cnn/1. All from the Nolichucky Formation, Rogersville, Hawkins County,
|
||
<collectingRegion id="499A12C3FFBBFFDC6F59E3A1FB0295B5" box="[1052,1164,1945,1971]" country="United States of America" name="Tennessee" pageId="20" pageNumber="21">Tennessee</collectingRegion>
|
||
(Walcott 1916).
|
||
</paragraph>
|
||
<paragraph id="8BE1DC21FFBBFFDD6B82E384FD1E92D1" blockId="20.[151,1437,1440,2043]" lastBlockId="21.[151,1437,153,2039]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
|
||
<emphasis id="B92A0033FFBBFFDC6B82E384FED495D1" bold="true" box="[199,346,1980,2007]" pageId="20" pageNumber="21">Occurrence.</emphasis>
|
||
<taxonomicName id="4C5EA7A2FFBBFFDC6A1AE385FCC095D1" authority="Zone, Nolichucky Formation" authorityName="Zone, Nolichucky Formation" box="[351,846,1981,2007]" class="Trilobita" family="Crepicephalidae" genus="Crepicephalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92A0033FFBBFFDC6A1AE385FD8F95D1" box="[351,513,1981,2007]" italics="true" pageId="20" pageNumber="21">Crepicephalus</emphasis>
|
||
Zone, Nolichucky Formation
|
||
</taxonomicName>
|
||
,
|
||
<collectingRegion id="499A12C3FFBBFFDC6813E385FC4295D1" box="[854,972,1981,2007]" country="United States of America" name="Tennessee" pageId="20" pageNumber="21">Tennessee</collectingRegion>
|
||
.
|
||
<materialsCitation id="3B36D67CFFBBFFDD6890E385FD0292D1" collectionCode="USNM" county="Hawkins County" lastPageId="21" lastPageNumber="22" location="Nolichucky Formation" municipality="Rogersville" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Tennessee">
|
||
<typeStatus id="54E56283FFBBFFDC6890E385FB8395D1" box="[981,1037,1981,2007]" pageId="20" pageNumber="21">Type</typeStatus>
|
||
locality is
|
||
<collectionCode id="ED4F44E4FFBBFFDC6FC1E385FB5B95D1" box="[1156,1237,1981,2007]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21" type="Museum">USNM</collectionCode>
|
||
locality 123b, 0.5 mile east of
|
||
<collectingMunicipality id="6B85465BFFBBFFDC6A60E3D9FE2795FD" box="[293,425,2017,2043]" pageId="20" pageNumber="21">Rogersville</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="6280A4ADFFBBFFDC6AF0E3D9FDFB95FD" box="[437,629,2017,2043]" pageId="20" pageNumber="21">Hawkins County</collectingCounty>
|
||
.
|
||
<bibRefCitation id="EFCFA1D0FFBBFFDC69C4E3D9FCBB95FD" author="Rasetti, F." box="[641,821,2016,2043]" pageId="20" pageNumber="21" pagination="1 - 127" refId="ref23892" refString="Rasetti, F. (1965) Upper Cambrian trilobite faunas of Northeastern Tennessee. Smithsonian Miscellaneous Collections, 148, 1 - 127." type="journal article" year="1965">Rasetti’s (1965)</bibRefCitation>
|
||
localities (cnm/2, cnn/1, cnn/2, cnn/3, cnn/4, cnn/14),
|
||
<location id="8E818AFAFFBAFFDD6BD2E4A1FE1992B5" LSID="urn:lsid:plazi:treatment:03F76D37FFBBFFDE6BD2E01EFB9892FD:8E818AFAFFBAFFDD6BD2E4A1FE1992B5" box="[151,407,153,179]" county="Hawkins County" municipality="Rogersville" name="Nolichucky Formation" pageId="21" pageNumber="22" stateProvince="Tennessee">Nolichucky Formation</location>
|
||
at the
|
||
<location id="8E818AFAFFBAFFDD6AA0E4A1FDD692B5" LSID="urn:lsid:plazi:treatment:03F76D37FFBBFFDE6BD2E01EFB9892FD:8E818AFAFFBAFFDD6AA0E4A1FDD692B5" box="[485,600,153,179]" county="Hawkins County" municipality="Rogersville" name="Big Creek" pageId="21" pageNumber="22" stateProvince="Tennessee">Big Creek</location>
|
||
and
|
||
<location id="8E818AFAFFBAFFDD69D5E4A1FC9F92B5" LSID="urn:lsid:plazi:treatment:03F76D37FFBBFFDE6BD2E01EFB9892FD:8E818AFAFFBAFFDD69D5E4A1FC9F92B5" box="[656,785,153,179]" county="Hawkins County" municipality="Rogersville" name="Rogersville" pageId="21" pageNumber="22" stateProvince="Tennessee">Rogersville</location>
|
||
sections in
|
||
<location id="8E818AFAFFBAFFDD68DCE4A1FBD792B5" LSID="urn:lsid:plazi:treatment:03F76D37FFBBFFDE6BD2E01EFB9892FD:8E818AFAFFBAFFDD68DCE4A1FBD792B5" box="[921,1113,153,179]" county="Hawkins County" municipality="Rogersville" name="Hawkins County" pageId="21" pageNumber="22" stateProvince="Tennessee">Hawkins County</location>
|
||
, and
|
||
<location id="8E818AFAFFBAFFDD6FD0E4A1FA9C92B5" LSID="urn:lsid:plazi:treatment:03F76D37FFBBFFDE6BD2E01EFB9892FD:8E818AFAFFBAFFDD6FD0E4A1FA9C92B5" box="[1173,1298,153,179]" county="Hawkins County" municipality="Rogersville" name="Lost Creek" pageId="21" pageNumber="22" stateProvince="Tennessee">Lost Creek</location>
|
||
and
|
||
<location id="8E818AFAFFBAFFDD6E0FE4A1FF4892D1" LSID="urn:lsid:plazi:treatment:03F76D37FFBBFFDE6BD2E01EFB9892FD:8E818AFAFFBAFFDD6E0FE4A1FF4892D1" county="Hawkins County" municipality="Rogersville" name="Russell Gap" pageId="21" pageNumber="22" stateProvince="Tennessee">Russell Gap</location>
|
||
sections in
|
||
<location id="8E818AFAFFBAFFDD6A0EE485FD8292D1" LSID="urn:lsid:plazi:treatment:03F76D37FFBBFFDE6BD2E01EFB9892FD:8E818AFAFFBAFFDD6A0EE485FD8292D1" box="[331,524,189,215]" county="Hawkins County" municipality="Rogersville" name="Jefferson County" pageId="21" pageNumber="22" stateProvince="Tennessee">Jefferson County</location>
|
||
,
|
||
<collectingRegion id="499A12C3FFBAFFDD6953E485FD0292D1" box="[534,652,189,215]" country="United States of America" name="Tennessee" pageId="21" pageNumber="22">Tennessee</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3448FAAFFBAFFDD6B82E4D8FD9F9631" pageId="21" pageNumber="22" type="description">
|
||
<paragraph id="8BE1DC21FFBAFFDD6B82E4D8FB3290F5" blockId="21.[151,1437,153,2039]" pageId="21" pageNumber="22">
|
||
<emphasis id="B92A0033FFBAFFDD6B82E4D8FED792FD" bold="true" box="[199,345,224,251]" pageId="21" pageNumber="22">Description.</emphasis>
|
||
Cranidium subtrapezoidal in outline; width at palpebral lobe equal to 97% of length. Anterior margin evenly rounded. Frontal area occupies one-third (32%; 31–33) cranidial length (sag.); two-thirds (66%; 65–67) of frontal area consists of long, concave anterior border. Preglabellar and preocular fields steeply sloping, then gradually merge with border. Broad (sag., exsag.) anterior border furrow is gently impressed on internal mould. Glabella subtrapezoidal in outline with gently rounded front; three pairs of furrows barely perceptible only on internal mould. Glabellar width at palpebral lobe is 85% (83–86) of length; convexity low, only subtly raised above fixed cheeks. LO short, accounting for less than one-tenth (7%; 6–8) of glabellar length; crescent-shaped, tapers (tr.) before reaching edge of glabella. SO and axial furrows well incised on internal mould. Combined width of palpebral areas makes up 45% (45–47) of cranidial width. Palpebral lobes situated in front of glabellar mid-length; moderately curved, short, equal to 13% (13–14) of cranidial length. On internal mould, raised palpebral ridge extends from lobe to anterior corner of glabella.Anterior branches of facial suture nearly parallel, then curve into anterior margin; posterior branches diverge backward as faintly sigmoid curves. Posterolateral projections extend slightly backward, increasing curvature abaxially to a tapered point. Posterior border furrow well defined on internal mould, extending abaxially across threequarters of width (tr.) of posterolateral projection, terminating short of sutural margin. Dense pitting with bimodal size distribution preserved on internal mould; large pits are more than twice diameter of smaller ones.
|
||
</paragraph>
|
||
<paragraph id="8BE1DC21FFBAFFDD6B82E6C5FD9F9631" blockId="21.[151,1437,153,2039]" pageId="21" pageNumber="22">Pygidium semielliptical in outline, length (sag.) equal to 70% (68–74) of maximum width. Very short articulating half-ring; articulating furrow slightly longer and well-incised only on internal mould. Maximum axis width about one-third (36%; 34–38) of pygidial width, tapers posteriorly and extends beyond border furrow. Twelve axial rings plus terminal piece expressed on internal mould. Axial furrows shallow, weakening towards terminal piece. Axis raised only slightly above pleural field. Pleural furrows weakly expressed on internal mould, indistinct on exoskeleton. Border down-sloping; approximately one-tenth maximum pygidial length (sag.); differentiated from pleural field by shallow border furrow which weakens posteriorly, becoming indistinct at axis tip. External surface smooth, except for terrace ridges along posterior margin. Internal mould has dense pitting with bimodal size distribution similar to cranidial pitting.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3448FAAFFBAFFDE6B82E078FB9892FD" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="discussion">
|
||
<paragraph id="8BE1DC21FFBAFFDD6B82E078FCAF97B1" blockId="21.[151,1437,153,2039]" pageId="21" pageNumber="22">
|
||
<emphasis id="B92A0033FFBAFFDD6B82E078FEC2965D" bold="true" box="[199,332,1088,1115]" pageId="21" pageNumber="22">Discussion.</emphasis>
|
||
The
|
||
<typeStatus id="54E56283FFBAFFDD6AC0E079FE4E965D" box="[389,448,1089,1115]" pageId="21" pageNumber="22">types</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6AA0E079FD10965D" authorityName="Walcott" authorityYear="1916" box="[485,670,1089,1115]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arion">
|
||
<emphasis id="B92A0033FFBAFFDD6AA0E079FD10965D" box="[485,670,1089,1115]" italics="true" pageId="21" pageNumber="22">Maryvillia arion</emphasis>
|
||
</taxonomicName>
|
||
are a cranidium and two pygidia (Walcott 1916b, pl. 64, fig. 4) from Nolichucky Formation near Rogersville,
|
||
<collectingRegion id="499A12C3FFBAFFDD6927E05DFD569679" box="[610,728,1125,1151]" country="United States of America" name="Tennessee" pageId="21" pageNumber="22">Tennessee</collectingRegion>
|
||
; the cranidium and one pygidium are reillustrated in this paper, along with a cranidium from the
|
||
<typeStatus id="54E56283FFBAFFDD6957E0B0FDCC96A4" box="[530,578,1160,1186]" pageId="21" pageNumber="22">type</typeStatus>
|
||
lot that has not been figured previously (Pl. 22, figs 1–8, Pl. 23, figs 1–2).
|
||
<bibRefCitation id="EFCFA1D0FFBAFFDD6BD2E093FECF96C2" author="Resser, C. E." box="[151,321,1194,1221]" pageId="21" pageNumber="22" pagination="1 - 58" refId="ref23949" refString="Resser, C. E. (1942 a) Fifth contribution to nomenclature of Cambrian fossils. Smithsonian Miscellaneous Collections, 101, 1 - 58." type="journal article" year="1942">Resser (1942a)</bibRefCitation>
|
||
considered the pygidia to have been associated incorrectly with the cranidia, and assigned them to a new species,
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6A06E0F5FE7B96E1" box="[323,501,1229,1255]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arguta">
|
||
<emphasis id="B92A0033FFBAFFDD6A06E0F5FE7B96E1" box="[323,501,1229,1255]" italics="true" pageId="21" pageNumber="22">Blountia arguta</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFCFA1D0FFBAFFDD6941E0F5FD2B96E1" author="Rasetti, F." box="[516,677,1228,1255]" pageId="21" pageNumber="22" pagination="1266 - 1269" refId="ref23835" refString="Rasetti, F. (1956) Revision of the trilobite genus Maryvillia Walcott. Journal of Paleontology, 30, 1266 - 1269." type="journal article" year="1956">Rasetti (1956)</bibRefCitation>
|
||
later suggested that Resser was incorrect, and returned them to
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6E3BE0F6FF5A970F" authorityName="Walcott" authorityYear="1916" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arion">
|
||
<emphasis id="B92A0033FFBAFFDD6E3BE0F6FF5A970F" italics="true" pageId="21" pageNumber="22">M. arion</emphasis>
|
||
</taxonomicName>
|
||
, which is followed here. The pygidium of
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD69F2E0C8FC94970F" authorityName="Walcott" authorityYear="1916" box="[695,794,1264,1290]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arion">
|
||
<emphasis id="B92A0033FFBAFFDD69F2E0C8FD40970F" box="[695,718,1264,1289]" italics="true" pageId="21" pageNumber="22">M</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFBAFFDD6998E0C8FC94970F" box="[733,794,1264,1289]" italics="true" pageId="21" pageNumber="22">arion</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD68EBE0C8FBAC970C" authorityName="Walcott" authorityYear="1916" box="[942,1058,1264,1290]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mimula">
|
||
<emphasis id="B92A0033FFBAFFDD68EBE0C8FBAC970C" box="[942,1058,1264,1290]" italics="true" pageId="21" pageNumber="22">B. mimula</emphasis>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="54E56283FFBAFFDD6F1FE0C8FB04970C" box="[1114,1162,1264,1290]" pageId="21" pageNumber="22">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6E4CE0C8FAE7970C" authorityName="Walcott" authorityYear="1916" box="[1289,1385,1264,1290]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92A0033FFBAFFDD6E4CE0C8FAE7970C" box="[1289,1385,1264,1290]" italics="true" pageId="21" pageNumber="22">Blountia</emphasis>
|
||
</taxonomicName>
|
||
, but differs by having a more strongly rounded outline, a more clearly defined border furrow, a longer axis that is barely raised above the pleural fields, and 12 rings in front of the terminal piece, versus
|
||
<quantity id="4CA671C4FFBAFFDD6F6DE10DFBFE9749" box="[1064,1136,1333,1359]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="21" pageNumber="22" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7–8 in</quantity>
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6F32E10EFB649749" authorityName="Walcott" authorityYear="1916" box="[1143,1258,1333,1359]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mimula">
|
||
<emphasis id="B92A0033FFBAFFDD6F32E10EFB069749" box="[1143,1160,1334,1359]" italics="true" pageId="21" pageNumber="22">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFBAFFDD6FD3E10DFB649749" box="[1174,1258,1333,1359]" italics="true" pageId="21" pageNumber="22">mimula</emphasis>
|
||
</taxonomicName>
|
||
. The cranidium of
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6BF2E160FEA59774" authorityName="Walcott" authorityYear="1916" box="[183,299,1368,1394]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mimula">
|
||
<emphasis id="B92A0033FFBAFFDD6BF2E160FF469777" box="[183,200,1368,1393]" italics="true" pageId="21" pageNumber="22">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFBAFFDD6B92E160FEA59774" box="[215,299,1368,1394]" italics="true" pageId="21" pageNumber="22">mimula</emphasis>
|
||
</taxonomicName>
|
||
differs by having a far more elevated, convex glabella, anterior branches of facial sutures diverging outwards from palpebral lobes, rather than proceeding straight forward as in
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD68ADE143FBC79792" authorityName="Walcott" authorityYear="1916" box="[1000,1097,1403,1429]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arion">
|
||
<emphasis id="B92A0033FFBAFFDD68ADE143FC719792" box="[1000,1023,1403,1428]" italics="true" pageId="21" pageNumber="22">M</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFBAFFDD6F49E143FBC79792" box="[1036,1097,1403,1428]" italics="true" pageId="21" pageNumber="22">arion</emphasis>
|
||
</taxonomicName>
|
||
, and with the posterior margin of the posterolateral projection curved strongly backward.
|
||
</paragraph>
|
||
<paragraph id="8BE1DC21FFBAFFDD6B82E1F9FD389545" blockId="21.[151,1437,153,2039]" pageId="21" pageNumber="22">
|
||
<bibRefCitation id="EFCFA1D0FFBAFFDD6B82E1F9FEE697DD" author="Rasetti, F." box="[199,360,1472,1499]" pageId="21" pageNumber="22" pagination="1266 - 1269" refId="ref23835" refString="Rasetti, F. (1956) Revision of the trilobite genus Maryvillia Walcott. Journal of Paleontology, 30, 1266 - 1269." type="journal article" year="1956">Rasetti (1956)</bibRefCitation>
|
||
also considered
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD696FE1F9FB7797DD" authority="Lochman, in Lochman & Duncan, 1944" authorityName="Lochman, in Lochman & Duncan" authorityYear="1944" box="[554,1273,1473,1499]" class="Trilobita" family="Asaphiscidae" genus="Blountina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="triangularis">
|
||
<emphasis id="B92A0033FFBAFFDD696FE1F9FCA997DD" box="[554,807,1473,1499]" italics="true" pageId="21" pageNumber="22">Blountina triangularis</emphasis>
|
||
Lochman, in
|
||
<bibRefCitation id="EFCFA1D0FFBAFFDD688DE1F9FB7797DD" author="Lochman, C. & Duncan, D." box="[968,1273,1473,1499]" pageId="21" pageNumber="22" pagination="1 - 172" refId="ref23573" refString="Lochman, C. & Duncan, D. (1944) Early Upper Cambrian faunas of Central Montana. Geological Society of America Special Papers, 54, 1 - 172. https: // doi. org / 10.1130 / SPE 54 - p 1" type="journal article" year="1944">Lochman & Duncan, 1944</bibRefCitation>
|
||
</taxonomicName>
|
||
to be a junior synonym of
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6A63E1DEFE0297F9" authorityName="Walcott" authorityYear="1916" box="[294,396,1509,1535]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arion">
|
||
<emphasis id="B92A0033FFBAFFDD6A63E1DEFEB397F9" box="[294,317,1510,1535]" italics="true" pageId="21" pageNumber="22">M</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFBAFFDD6A09E1DEFE0297F9" box="[332,396,1510,1535]" italics="true" pageId="21" pageNumber="22">arion</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92A0033FFBAFFDD6AC8E1DEFE1F97F9" box="[397,401,1510,1535]" italics="true" pageId="21" pageNumber="22">.</emphasis>
|
||
Restudy of Lochman & Duncan’s
|
||
<typeStatus id="54E56283FFBAFFDD685BE1DDFCD797F9" box="[798,857,1509,1535]" pageId="21" pageNumber="22">types</typeStatus>
|
||
(Pls 24, 25) shows that these species share several characters supporting assignment of the former to
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6980E231FCB59425" authorityName="Walcott" authorityYear="1916" box="[709,827,1545,1571]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92A0033FFBAFFDD6980E231FCB59425" box="[709,827,1545,1571]" italics="true" pageId="21" pageNumber="22">Maryvillia</emphasis>
|
||
</taxonomicName>
|
||
, including a concave anterior border on the cranidium, a glabella that is just slightly raised above the fixed cheeks, and facial sutures that extend nearly straight forward from palpebral lobes. However, pygidial anatomy indicates unequivocally that they are distinct species. The pygidium of
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6A49E24EFE369489" authorityName="Lochman, in Lochman & Duncan" authorityYear="1944" box="[268,440,1653,1679]" class="Trilobita" family="Asaphiscidae" genus="Blountina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="triangularis">
|
||
<emphasis id="B92A0033FFBAFFDD6A49E24EFE369489" box="[268,440,1653,1679]" italics="true" pageId="21" pageNumber="22">M. triangularis</emphasis>
|
||
</taxonomicName>
|
||
(Pl. 24, figs 9, 10; Pl. 25, figs 1–5, 9) is narrower and distinctly subtriangular in outline, which contrasts with the broad, posteriorly rounded outline of
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6813E2A2FC3794B5" authorityName="Walcott" authorityYear="1916" box="[854,953,1689,1715]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arion">
|
||
<emphasis id="B92A0033FFBAFFDD6813E2A2FCE394B5" box="[854,877,1690,1715]" italics="true" pageId="21" pageNumber="22">M</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFBAFFDD6839E2A2FC3794B5" box="[892,953,1690,1715]" italics="true" pageId="21" pageNumber="22">arion</emphasis>
|
||
</taxonomicName>
|
||
(Pl. 23), which also has a relatively longer axis. The cranidium of
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6AE4E286FDC094D1" authorityName="Lochman, in Lochman & Duncan" authorityYear="1944" box="[417,590,1725,1751]" class="Trilobita" family="Asaphiscidae" genus="Blountina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="triangularis">
|
||
<emphasis id="B92A0033FFBAFFDD6AE4E286FE3694D1" box="[417,440,1726,1751]" italics="true" pageId="21" pageNumber="22">M</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFBAFFDD6A8DE285FDC094D1" box="[456,590,1725,1751]" italics="true" pageId="21" pageNumber="22">triangularis</emphasis>
|
||
</taxonomicName>
|
||
differs most obviously by a crudely triangular-shaped outline, with an anterior border that is longest medially (sag.) rather than maintaining a more even length along the anterior cranidial margin. In addition, the margins of posterolateral projections are directed posteriorly (Pl. 24, fig. 5, Pl. 25, fig. 6) rather than nearly transversely (Pl. 22, figs 1, 5).
|
||
</paragraph>
|
||
<paragraph id="8BE1DC21FFBAFFDD6B82E375FE9495D5" blockId="21.[151,1437,153,2039]" pageId="21" pageNumber="22">
|
||
Sclerites from the Rabbitkettle Formation of northern
|
||
<collectingCountry id="F3499CB1FFBAFFDD686CE375FCF59561" box="[809,891,1869,1895]" name="Canada" pageId="21" pageNumber="22">Canada</collectingCountry>
|
||
that
|
||
<bibRefCitation id="EFCFA1D0FFBAFFDD68F2E375FBB89561" author="Pratt, B. R." box="[951,1078,1869,1895]" pageId="21" pageNumber="22" pagination="1 - 179" refId="ref23771" refString="Pratt, B. R. (1992) Trilobites of the Marjuman and Steptoean stages (Upper Cambrian), Rabbitkettle Formation, southern Mackenzie Mountains, northwest Canada. Palaeontographica Canadiana, 9, 1 - 179" type="journal article" year="1992">Pratt (1992</bibRefCitation>
|
||
, pl. 23, figs 23–25) assigned to cf.
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6BFEE349FED0958D" box="[187,350,1905,1931]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arion">
|
||
<emphasis id="B92A0033FFBAFFDD6BFEE349FED0958D" box="[187,350,1905,1931]" italics="true" pageId="21" pageNumber="22">Blountia arion</emphasis>
|
||
</taxonomicName>
|
||
are incomplete and difficult to assess. However, the strong convexity of both the glabella and the pygidial axis separate them immediately from
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD69D6E3ADFC8895A9" authorityName="Walcott" authorityYear="1916" box="[659,774,1941,1967]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92A0033FFBAFFDD69D6E3ADFC8895A9" box="[659,774,1941,1967]" italics="true" pageId="21" pageNumber="22">Maryvillia</emphasis>
|
||
</taxonomicName>
|
||
, and we consider them to represent an indeterminate species of
|
||
<taxonomicName id="4C5EA7A2FFBAFFDD6BF0E381FE9D95D5" authorityName="Walcott" authorityYear="1916" box="[181,275,1977,2003]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92A0033FFBAFFDD6BF0E381FE9D95D5" box="[181,275,1977,2003]" italics="true" pageId="21" pageNumber="22">Blountia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE1DC21FFBAFFDE6B82E3E5FB9892FD" blockId="21.[151,1437,153,2039]" lastBlockId="22.[151,1437,153,251]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
<bibRefCitation id="EFCFA1D0FFBAFFDD6B82E3E5FE3195F1" author="Stitt, J. H. & Perfetta, P. J." box="[199,447,2013,2039]" pageId="21" pageNumber="22" pagination="199 - 223" refId="ref24040" refString="Stitt, J. H. & Perfetta, P. J. (2000) Trilobites, biostratigraphy, and lithostratigraphy of the Crepicephalus and Aphelaspis zones, lower Deadwood Formation (Marjuman and Steptoean stages, Upper Cambrian), Black Hills, South Dakota. Journal of Paleontology, 74, 199 - 223. https: // doi. org / 10.1017 / S 0022336000031437" type="journal article" year="2000">Stitt & Perfetta (2000</bibRefCitation>
|
||
, fig. 11.4, 11.5) illustrated two pygidia from the Deadwood Formation under the name,
|
||
<taxonomicName id="4C5EA7A2FFB9FFDE6BD2E4A1FE2092B2" authority="(Walcott)" baseAuthorityName="Walcott" box="[151,430,153,180]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="arion">
|
||
<emphasis id="B92A0033FFB9FFDE6BD2E4A1FEB592B5" box="[151,315,153,179]" italics="true" pageId="22" pageNumber="23">Blountia arion</emphasis>
|
||
(Walcott)
|
||
</taxonomicName>
|
||
. The subtriangular outlines of these sclerites demonstrate that they are misidentified. They can be compared to
|
||
<taxonomicName id="4C5EA7A2FFB9FFDE6A3CE486FDAA92D1" authorityName="Lochman, in Lochman & Duncan" authorityYear="1944" box="[377,548,189,215]" class="Trilobita" family="Asaphiscidae" genus="Blountina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ptychopariida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="triangularis">
|
||
<emphasis id="B92A0033FFB9FFDE6A3CE486FE1E92D1" box="[377,400,190,215]" italics="true" pageId="22" pageNumber="23">M</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFB9FFDE6ADBE485FDAA92D1" box="[414,548,189,215]" italics="true" pageId="22" pageNumber="23">triangularis</emphasis>
|
||
</taxonomicName>
|
||
, although if the larger of the two specimens is complete, it is shorter and wider than any of Lochman and Duncan’s
|
||
<typeStatus id="54E56283FFB9FFDE6968E4D9FDE692FD" box="[557,616,225,251]" pageId="22" pageNumber="23">types</typeStatus>
|
||
(Pl. 24, figs 9, 10; Pl. 25, figs 1–5, 9).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |