304 lines
40 KiB
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304 lines
40 KiB
XML
<document id="D1D7880DE92125A247F13A6D60D7B639" ID-DOI="10.11646/zootaxa.4804.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3927094" ID-ZooBank="8C1C1703-9BBC-4B33-8045-78BDD9738F51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593672611012" checkinUser="plazi" docAuthor="Westrop, Madison Armstrong Stephen R. & Eoff, Jennifer D." docDate="2020" docId="03F76D37FFA5FFC36BD2E0F7FC4891A1" docLanguage="en" docName="zootaxa.4804.1.1.pdf" docOrigin="Zootaxa 4804 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Blountia bristolensis Resser 1938" docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="FFCE154FFFAFFFC86B45E438FF8E9206" masterDocTitle="Systematics of a survivor: the Cambrian kingstoniid trilobite Blountia Walcott, 1916 across the Marjuman-Steptoean (Guzhangian-Paibian) extinction interval in Laurentian North America" masterLastPageNumber="79" masterPageNumber="1" pageNumber="11" updateTime="1698776919363" updateUser="plazi">
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<mods:title id="81492ADDB31C7916C816957571157A5B">Systematics of a survivor: the Cambrian kingstoniid trilobite Blountia Walcott, 1916 across the Marjuman-Steptoean (Guzhangian-Paibian) extinction interval in Laurentian North America</mods:title>
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<mods:namePart id="B45F626820F9C185200AD36F3262DC2F">Westrop, Madison Armstrong Stephen R.</mods:namePart>
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<mods:namePart id="E5DCE03986B0081970212CB3BD5D37E7">Eoff, Jennifer D.</mods:namePart>
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<mods:typeOfResource id="25F7A3162D0247508DA8060AF592173A">text</mods:typeOfResource>
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<mods:title id="C40B29FF541EB8BEB0C96F4EBE5C04D3">Zootaxa</mods:title>
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<mods:part id="726720C6AF545651860AC0A0548F331E">
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<mods:date id="EF605CD295CEDFC7D5F10B15B49394F6">2020</mods:date>
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<mods:number id="7D3DC7A9620B45D28886EEB2B292F514">2020-06-30</mods:number>
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<mods:number id="FA1ABAB9720C96BC8A4ADBB62E84C0AF">4804</mods:number>
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<mods:classification id="FECA8827EB0DD13B701FE5D0FD5D7920">journal article</mods:classification>
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<mods:identifier id="32E5C10E37F663C56E488E90A75E8D10" type="DOI">10.11646/zootaxa.4804.1.1</mods:identifier>
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<mods:identifier id="3B55ECEB5C735BBFBDB306ED0B26CDCF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C7F49D87EA7D55B96DC20ABD0FC8B244" type="Zenodo-Dep">3927094</mods:identifier>
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<subSubSection id="C3448FAAFFA5FFC26BD2E0F7FDBD96EC" box="[151,563,1231,1258]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BE1DC21FFA5FFC26BD2E0F7FDBD96EC" blockId="10.[151,563,1231,1294]" box="[151,563,1231,1258]" pageId="10" pageNumber="11">
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<heading id="D0A96B4DFFA5FFC26BD2E0F7FDBD96EC" bold="true" box="[151,563,1231,1258]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<taxonomicName id="4C5EA7A2FFA5FFC26BD2E0F7FDBD96EC" authority="Resser, 1938" authorityName="Resser" authorityYear="1938" box="[151,563,1231,1258]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
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<emphasis id="B92A0033FFA5FFC26BD2E0F7FDBD96EC" bold="true" box="[151,563,1231,1258]" pageId="10" pageNumber="11">
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<emphasis id="B92A0033FFA5FFC26BD2E0F7FE1F96EC" bold="true" box="[151,401,1231,1258]" italics="true" pageId="10" pageNumber="11">Blountia bristolensis</emphasis>
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<bibRefCitation id="EFCFA1D0FFA5FFC26ADCE0E8FDBD96EC" author="Resser, C. E." box="[409,563,1232,1258]" pageId="10" pageNumber="11" pagination="1 - 140" refId="ref23917" refString="Resser, C. E. (1938) The Cambrian (Restricted) of the southern Appalachians. Geological Society of America Special Papers, 15, 1 - 140." type="journal article" year="1938">Resser, 1938</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3448FAAFFA5FFC26BD2E0CCFB1297CC" pageId="10" pageNumber="11" type="description">
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<paragraph id="8BE1DC21FFA5FFC26BD2E0CCFE859708" blockId="10.[151,563,1231,1294]" box="[151,267,1267,1294]" pageId="10" pageNumber="11">
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<figureCitation id="1365C0A4FFA5FFC26BD2E0CCFE859708" box="[151,267,1267,1294]" captionStart-0="PLATE 2" captionStart-1="PLATE 3" captionStart-2="PLATE 4" captionStart-3="PLATE 5" captionStartId-0="28.[151,234,1868,1893]" captionStartId-1="30.[151,234,1897,1922]" captionStartId-2="32.[151,234,1868,1893]" captionStartId-3="34.[151,234,1839,1864]" captionText-0="PLATE 2. Blountia bristolensis Resser, 1938 from the Nolichucky Formation, reservoir on Mumpower Creek, 5.6 km north of Bristol, Virginia (USNM locality 36v). 1–3. Cranidium, holotype of Maryvillia bristolensis Resser, 1938, USMN 94963, dorsal, lateral and anterior views x7.5 (illustrated by Resser 1938, pl. 12, fig. 38). 4–6. Cranidium, USMN 94963, dorsal, lateral and anterior views x12 (previously unillustrated specimen from the type lot of Maryvillia bristolensis Resser, 1938). 7–9. Cranidium, Maryvillia bristolensis Resser, 1938, USMN 94963, dorsal, anterior and lateral views x8.5 (previously unillustrated specimen from the type lot of Maryvillia bristolensis Resser, 1938)." captionText-1="PLATE 3. Blountia bristolensis Resser, 1938 from the Nolichucky Formation, Tennessee and Virginia. 1, 2. Pygidium, holotype, USNM 94942 (illustrated by Resser, 1938, pl. 12, fig. 24), dorsal and posterior views, x16 (reservoir on Mumpower Creek, 5.6 km north of Bristol, Virginia, USNM locality 36v). 3–7. Nearly complete exoskeleton, USNM 144600 (illustrated by Rasetti 1965, pl. 10, figs 1, 2), lateral, posterior, cranidium oriented in dorsal view, pygidium oriented in dorsal view, anterior view, x8 (collection cnp/17, Hurricane Hollow, Norris Reservoir, Union County, Tennessee; Rasetti 1965, p. 20)." captionText-2="PLATE 4. Blountia bristolensis Resser, 1938 from the Pilgrim Formation, Nixon Gulch section, the Horseshoe Hills, Montana, horizon 2.5. 1–3. Cranidium, paratype of Blountia nixonensis Lochman, in Lochman & Duncan, 1944, USNM 511426, dorsal, lateral, and anterior views, x13 (previously unillustrated). 4–6. Cranidium, paratype, of Blountia nixonensis Lochman, in Lochman & Duncan, 1944, USNM 511426, dorsal lateral, and anterior views, x13 (previously unillustrated). 7–9. Cranidium, paratype, of Blountia nixonensis Lochman, in Lochman & Duncan, 1944, USNM 511426, anterior, dorsal, lateral views, x13 (previously unillustrated)." captionText-3="PLATE 5. Blountia bristolensis Resser, 1938 from the Pilgrim Formation, Nixon Gulch section, Horseshoe Hills, Montana, horizon 2.5. 1–3. Pygidium, paratype of Blountia nixonensis Lochman, in Lochman & Duncan, 1944, USNM 511434, dorsal, lateral, and posterior views, x11.5 (previously unillustrated). 4–6. Pygidium, paratype of Blountia nixonensis Lochman, in Lochman & Duncan, 1944, USNM 511432, dorsal, lateral and posterior views, x11.5 (previously unillustrated). 7, 8. Pygidium, paratype of Blountia nixonensis Lochman, in Lochman & Duncan, 1944, USNM 511433, dorsal and posterior views, x11.5 (previously unillustrated). 9–11. Pygidium, paratype of Blountia nixonensis Lochman, in Lochman & Duncan, 1944, USNM 511427, lateral, posterior and dorsal views, x9 (previously unillustrated)." pageId="10" pageNumber="11">Plates 2–5</figureCitation>
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</paragraph>
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<paragraph id="8BE1DC21FFA5FFC26BD2E100FD499756" blockId="10.[151,1180,1335,1483]" box="[151,711,1335,1360]" pageId="10" pageNumber="11">
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1938
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<taxonomicName id="4C5EA7A2FFA5FFC26B95E100FE609749" authority="Resser" authorityName="Resser" authorityYear="1938" box="[208,494,1336,1360]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
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<emphasis id="B92A0033FFA5FFC26B95E100FE2D9756" box="[208,419,1336,1360]" italics="true" pageId="10" pageNumber="11">Blountia bristolensis</emphasis>
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Resser
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</taxonomicName>
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, p. 65, pl. 12, fig. 24.
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</paragraph>
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<paragraph id="8BE1DC21FFA5FFC26BD2E16FFD559769" blockId="10.[151,1180,1335,1483]" box="[151,731,1366,1391]" pageId="10" pageNumber="11">
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1938
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<taxonomicName id="4C5EA7A2FFA5FFC26B95E16FFD8C9768" authority="Resser" authorityName="Resser" authorityYear="1938" box="[208,514,1367,1391]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
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<emphasis id="B92A0033FFA5FFC26B95E16FFE389769" box="[208,438,1367,1391]" italics="true" pageId="10" pageNumber="11">Maryvillia bristolensis</emphasis>
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Resser
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</taxonomicName>
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, p. 87, pl. 12, fig. 38.
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</paragraph>
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<paragraph id="8BE1DC21FFA5FFC26BD2E14DFC6E978B" blockId="10.[151,1180,1335,1483]" box="[151,992,1397,1422]" pageId="10" pageNumber="11">
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1944
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<taxonomicName id="4C5EA7A2FFA5FFC26B95E14DFD79978B" authority="Lochman, in Lochman & Duncan" authorityName="Lochman, in Lochman & Duncan" authorityYear="1944" box="[208,759,1397,1422]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nixonensis">
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<emphasis id="B92A0033FFA5FFC26B95E14DFE17978B" box="[208,409,1397,1421]" italics="true" pageId="10" pageNumber="11">Blountia nixonensis</emphasis>
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Lochman, in Lochman & Duncan
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</taxonomicName>
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, p. 43, pl. 4, figs 7–12.
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</paragraph>
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<paragraph id="8BE1DC21FFA5FFC26BD2E1ABFC5997AA" blockId="10.[151,1180,1335,1483]" box="[151,983,1427,1452]" pageId="10" pageNumber="11">
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1965
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<taxonomicName id="4C5EA7A2FFA5FFC26B95E1ACFE7E97AD" authority="Resser" authorityName="Resser" authorityYear="1938" box="[208,496,1428,1452]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
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<emphasis id="B92A0033FFA5FFC26B95E1ACFE2D97AA" box="[208,419,1428,1452]" italics="true" pageId="10" pageNumber="11">Blountia bristolensis</emphasis>
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Resser
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</taxonomicName>
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; Rasetti, p. 58, pl. 10, figs 1–2, pl. 11 figs 9–12.
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</paragraph>
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<paragraph id="8BE1DC21FFA5FFC26BD2E18BFB1297CC" blockId="10.[151,1180,1335,1483]" box="[151,1180,1458,1483]" pageId="10" pageNumber="11">
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<emphasis id="B92A0033FFA5FFC26BD2E18BFF3397CC" box="[151,189,1459,1482]" italics="true" pageId="10" pageNumber="11">non</emphasis>
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1965
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<taxonomicName id="4C5EA7A2FFA5FFC26BB8E18BFD9297CC" authority="Resser" authorityName="Resser" authorityYear="1938" box="[253,540,1459,1483]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
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<emphasis id="B92A0033FFA5FFC26BB8E18BFE4197CD" box="[253,463,1459,1483]" italics="true" pageId="10" pageNumber="11">Blountia bristolensis</emphasis>
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Resser
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</taxonomicName>
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; Palmer, p. 29, pl. 1, figs 1, 2, 4 [=
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<taxonomicName id="4C5EA7A2FFA5FFC268C7E18BFBC097CD" authority="Westrop & Eoff, 2020" authorityName="Westrop & Eoff" authorityYear="2020" box="[898,1102,1459,1483]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nevadensis" status="sp. nov.">
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<emphasis id="B92A0033FFA5FFC268C7E18BFBC097CD" box="[898,1102,1459,1483]" italics="true" pageId="10" pageNumber="11">Blountia nevadensis</emphasis>
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</taxonomicName>
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<emphasis id="B92A0033FFA5FFC26F10E18BFB0097CD" bold="true" box="[1109,1166,1459,1483]" pageId="10" pageNumber="11">
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<taxonomicNameLabel id="A219BD48FFA5FFC26F10E18BFB0097CD" box="[1109,1166,1459,1483]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="C3448FAAFFA5FFC26BD2E1C0FDBE9479" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="8BE1DC21FFA5FFC26BD2E1C0FDBE9479" blockId="10.[151,1437,1528,2023]" pageId="10" pageNumber="11">
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<emphasis id="B92A0033FFA5FFC26BD2E1C0FE9C9415" bold="true" box="[151,274,1528,1555]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
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Frontal area 25% (23–26) of cranidial length, with sharp change in slope between preglabellar field and anterior border, and well-defined anterior border furrow. Seven thoracic segments. Pygidium semielliptical in outline with inflated pleural field, well-defined border furrow, and down-sloping border of even width occupying 10% (8–12) of pygidial length (sag).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3448FAAFFA5FFC26B82E2B0FCE895E1" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="8BE1DC21FFA5FFC26B82E2B0FBE3957D" blockId="10.[151,1437,1528,2023]" pageId="10" pageNumber="11">
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<emphasis id="B92A0033FFA5FFC26B82E2B0FEB894A5" bold="true" box="[199,310,1672,1699]" pageId="10" pageNumber="11">Material.</emphasis>
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The
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<typeStatus id="54E56283FFA5FFC26A28E2B1FE4194A5" box="[365,463,1673,1699]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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is a pygidium (
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<collectionCode id="ED4F44E4FFA5FFC2693DE2B1FD4594A5" box="[632,715,1673,1699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
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94942; Pl. 5, figs 1, 2) from the Nolichucky Formation, reservoir on Mumpower Creek, 5.6 km north of Bristol,
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<collectingRegion id="499A12C3FFA5FFC269F2E295FC9C94C1" box="[695,786,1709,1735]" country="United States of America" name="Virginia" pageId="10" pageNumber="11">Virginia</collectingRegion>
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(
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<collectionCode id="ED4F44E4FFA5FFC26866E295FCF894C1" box="[803,886,1709,1735]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
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locality 36v). Other material includes the
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<typeStatus id="54E56283FFA5FFC26E24E295FF4994ED" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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||
cranidium of
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<taxonomicName id="4C5EA7A2FFA5FFC26A2CE2E9FD3294ED" authority="Resser" authorityName="Resser" authorityYear="1938" box="[361,700,1745,1771]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
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<emphasis id="B92A0033FFA5FFC26A2CE2E9FDE794ED" box="[361,617,1745,1771]" italics="true" pageId="10" pageNumber="11">Maryvillia bristolensis</emphasis>
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Resser
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</taxonomicName>
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(
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<collectionCode id="ED4F44E4FFA5FFC26989E2E9FC9194ED" box="[716,799,1745,1771]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
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94963, Pl. 2, figs 1–3) and two previously unillustrated cranidia from the type lot (both catalogued as
|
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<collectionCode id="ED4F44E4FFA5FFC269EBE2CDFD719509" box="[686,767,1781,1807]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
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94963, Pl. 2, figs 4–9). Previously unillustrated
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<typeStatus id="54E56283FFA5FFC26E74E2CDFA129509" box="[1329,1436,1781,1807]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
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of
|
||
<taxonomicName id="4C5EA7A2FFA5FFC26BF3E322FD5D9535" authority="Lochman, in Lochman & Duncan" authorityName="Lochman, in Lochman & Duncan" authorityYear="1944" box="[182,723,1817,1843]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nixonensis">
|
||
<emphasis id="B92A0033FFA5FFC26BF3E322FF499535" box="[182,199,1818,1843]" italics="true" pageId="10" pageNumber="11">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFA5FFC26B93E322FEC29535" box="[214,332,1818,1843]" italics="true" pageId="10" pageNumber="11">nixonensis</emphasis>
|
||
Lochman, in Lochman & Duncan
|
||
</taxonomicName>
|
||
, include three cranidia (
|
||
<collectionCode id="ED4F44E4FFA5FFC268A4E321FBBA9535" box="[993,1076,1817,1843]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
|
||
511426, Pl. 4, figs 1–3;
|
||
<collectionCode id="ED4F44E4FFA5FFC26E0EE321FA129535" box="[1355,1436,1817,1843]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
|
||
511429, Pl. 4, figs 4–6;
|
||
<collectionCode id="ED4F44E4FFA5FFC26AE0E305FE789551" box="[421,502,1853,1879]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
|
||
511430, Pl. 4, figs 7–9) and four pygidia (
|
||
<collectionCode id="ED4F44E4FFA5FFC26893E305FBA79551" box="[982,1065,1853,1879]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
|
||
511427, Pl. 5, figs 9–11;
|
||
<collectionCode id="ED4F44E4FFA5FFC26E0EE305FA129551" box="[1355,1436,1853,1879]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
|
||
511432, Pl. 5, figs 4–6;
|
||
<collectionCode id="ED4F44E4FFA5FFC26AE6E359FE7A957D" box="[419,500,1889,1915]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
|
||
511433, Pl. 5, figs 7, 8;
|
||
<collectionCode id="ED4F44E4FFA5FFC26842E359FCD6957D" box="[775,856,1889,1915]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
|
||
511434, Pl. 5, figs 1–3).
|
||
</paragraph>
|
||
<paragraph id="8BE1DC21FFA5FFC26B82E3BCFCE895E1" blockId="10.[151,1437,1528,2023]" pageId="10" pageNumber="11">
|
||
<emphasis id="B92A0033FFA5FFC26B82E3BCFED49599" bold="true" box="[199,346,1924,1951]" pageId="10" pageNumber="11">Occurrence.</emphasis>
|
||
<taxonomicName id="4C5EA7A2FFA5FFC26A25E3BDFCA49599" authority="Zone, Nolichucky Formation" authorityName="Zone, Nolichucky Formation" box="[352,810,1925,1951]" class="Trilobita" family="Aphelaspididae" genus="Aphelaspis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Asaphida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92A0033FFA5FFC26A25E3BDFE559599" box="[352,475,1925,1951]" italics="true" pageId="10" pageNumber="11">Aphelaspis</emphasis>
|
||
Zone, Nolichucky Formation
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation id="EFCFA1D0FFA5FFC26870E3BDFC689599" author="Rasetti, F." box="[821,998,1924,1951]" pageId="10" pageNumber="11" pagination="1 - 127" refId="ref23892" refString="Rasetti, F. (1965) Upper Cambrian trilobite faunas of Northeastern Tennessee. Smithsonian Miscellaneous Collections, 148, 1 - 127." type="journal article" year="1965">Rasetti’s (1965)</bibRefCitation>
|
||
collections cnp/17, cnp/14, and cnq/
|
||
<quantity id="4CA671C4FFA5FFC26EC6E3BDFF2395C5" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="10" pageNumber="11" unit="in" value="14.0">14 in</quantity>
|
||
<collectingRegion id="499A12C3FFA5FFC26BF6E391FEA795C5" box="[179,297,1961,1987]" country="United States of America" name="Tennessee" pageId="10" pageNumber="11">Tennessee</collectingRegion>
|
||
;
|
||
<collectionCode id="ED4F44E4FFA5FFC26A71E391FE0B95C5" box="[308,389,1961,1987]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
|
||
locality 36v near Bristol,
|
||
<collectingRegion id="499A12C3FFA5FFC269E3E391FC8F95C5" box="[678,769,1961,1987]" country="United States of America" name="Virginia" pageId="10" pageNumber="11">Virginia</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFCFA1D0FFA5FFC2684AE391FC1095C5" author="Resser, C. E." box="[783,926,1961,1987]" pageId="10" pageNumber="11" pagination="1 - 140" refId="ref23917" refString="Resser, C. E. (1938) The Cambrian (Restricted) of the southern Appalachians. Geological Society of America Special Papers, 15, 1 - 140." type="journal article" year="1938">Resser 1938</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C5EA7A2FFA5FFC268F4E391FE2E95E1" authority="Zone, Pilgrim Formation, Nixon Gulch, Horseshoe Hills" authorityName="Zone, Pilgrim Formation, Nixon Gulch, Horseshoe Hills" class="Trilobita" family="Aphelaspididae" genus="Aphelaspis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Asaphida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92A0033FFA5FFC268F4E391FBA295C5" box="[945,1068,1961,1987]" italics="true" pageId="10" pageNumber="11">Aphelaspis</emphasis>
|
||
Zone, Pilgrim Formation, Nixon Gulch, Horseshoe Hills
|
||
</taxonomicName>
|
||
,
|
||
<collectingRegion id="499A12C3FFA5FFC26AEEE3F5FD8195E1" box="[427,527,1997,2023]" country="United States of America" name="Montana" pageId="10" pageNumber="11">Montana</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFCFA1D0FFA5FFC26958E3F5FCD795E1" author="Lochman, C. & Duncan, D." box="[541,857,1997,2023]" pageId="10" pageNumber="11" pagination="1 - 172" refId="ref23573" refString="Lochman, C. & Duncan, D. (1944) Early Upper Cambrian faunas of Central Montana. Geological Society of America Special Papers, 54, 1 - 172. https: // doi. org / 10.1130 / SPE 54 - p 1" type="journal article" year="1944">Lochman and Duncan 1944</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3448FAAFFA4FFC36B82E4A0FC4891A1" pageId="11" pageNumber="12" type="discussion">
|
||
<paragraph id="8BE1DC21FFA4FFC36B82E4A0FDC4938C" blockId="11.[151,1437,152,935]" pageId="11" pageNumber="12">
|
||
<emphasis id="B92A0033FFA4FFC36B82E4A0FEC292B5" bold="true" box="[199,332,152,179]" pageId="11" pageNumber="12">Discussion.</emphasis>
|
||
<bibRefCitation id="EFCFA1D0FFA4FFC36A11E4A0FE6292B5" author="Rasetti, F." box="[340,492,152,179]" pageId="11" pageNumber="12" pagination="1 - 127" refId="ref23892" refString="Rasetti, F. (1965) Upper Cambrian trilobite faunas of Northeastern Tennessee. Smithsonian Miscellaneous Collections, 148, 1 - 127." type="journal article" year="1965">Rasetti (1965</bibRefCitation>
|
||
, p. 58) noted that the pygidium and the cranidium that
|
||
<bibRefCitation id="EFCFA1D0FFA4FFC36F1FE4A1FB7892B5" author="Resser, C. E." box="[1114,1270,153,179]" pageId="11" pageNumber="12" pagination="1 - 140" refId="ref23917" refString="Resser, C. E. (1938) The Cambrian (Restricted) of the southern Appalachians. Geological Society of America Special Papers, 15, 1 - 140." type="journal article" year="1938">Resser (1938)</bibRefCitation>
|
||
named
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36E15E4A0FEB092D0" authorityName="Resser" authorityYear="1938" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC36E15E4A0FEB092D0" italics="true" pageId="11" pageNumber="12">Blountia bristolensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36A3DE484FDF992D0" authorityName="Resser" authorityYear="1938" box="[376,631,188,214]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC36A3DE484FDF992D0" box="[376,631,188,214]" italics="true" pageId="11" pageNumber="12">Maryvillia bristolensis</emphasis>
|
||
</taxonomicName>
|
||
, respectively, are from the same collection, and he considered them to represent a single species under the former name. The cranidium of
|
||
<taxonomicName id="4C5EA7A2FFA4FFC368D1E4D8FAAF92FD" authority="Resser (1938)" authorityName="Resser" authorityYear="1938" box="[916,1313,224,251]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC368D1E4D8FBF392FC" box="[916,1149,224,250]" italics="true" pageId="11" pageNumber="12">Blountia bristolensis</emphasis>
|
||
<bibRefCitation id="EFCFA1D0FFA4FFC36FC0E4D9FAAF92FD" author="Resser, C. E." box="[1157,1313,225,251]" pageId="11" pageNumber="12" pagination="1 - 140" refId="ref23917" refString="Resser, C. E. (1938) The Cambrian (Restricted) of the southern Appalachians. Geological Society of America Special Papers, 15, 1 - 140." type="journal article" year="1938">Resser (1938)</bibRefCitation>
|
||
</taxonomicName>
|
||
has a relatively short frontal area that occupies 25% (23–26) of cranidial length, and with a preglabellar field that varies from slightly longer to slightly shorter than the anterior border. There is an abrupt change in slope at border furrow. An exoskeleton assigned to the species by
|
||
<bibRefCitation id="EFCFA1D0FFA4FFC36901E574FD6C9361" author="Rasetti, F." box="[580,738,332,359]" pageId="11" pageNumber="12" pagination="1 - 127" refId="ref23892" refString="Rasetti, F. (1965) Upper Cambrian trilobite faunas of Northeastern Tennessee. Smithsonian Miscellaneous Collections, 148, 1 - 127." type="journal article" year="1965">Rasetti (1965)</bibRefCitation>
|
||
(Pl. 3, figs 3–7) has similar frontal area proportions to Resser’s cranidia and appears to be conspecific.
|
||
</paragraph>
|
||
<paragraph id="8BE1DC21FFA4FFC36B82E5ACFCE390AD" blockId="11.[151,1437,152,935]" pageId="11" pageNumber="12">
|
||
Both
|
||
<bibRefCitation id="EFCFA1D0FFA4FFC36A41E5ACFE2A93A9" author="Palmer, A. R." box="[260,420,404,431]" pageId="11" pageNumber="12" pagination="1 - 105" refId="ref23721" refString="Palmer, A. R. (1965) Trilobites of the Late Cambrian Pterocephaliid Biomere in the Great Basin, United States. U. S. Geological Survey Professional Paper, 463, 1 - 105. https: // doi. org / 10.3133 / pp 493" type="journal article" year="1965">Palmer (1965)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFCFA1D0FFA4FFC36A9CE5ACFDF693A9" author="Rasetti, F." box="[473,632,404,431]" pageId="11" pageNumber="12" pagination="1 - 127" refId="ref23892" refString="Rasetti, F. (1965) Upper Cambrian trilobite faunas of Northeastern Tennessee. Smithsonian Miscellaneous Collections, 148, 1 - 127." type="journal article" year="1965">Rasetti (1965)</bibRefCitation>
|
||
considered
|
||
<taxonomicName id="4C5EA7A2FFA4FFC369BBE5ACFA1293A9" authority="Lochman, in Lochman & Duncan, 1944" authorityName="Lochman, in Lochman & Duncan" authorityYear="1944" box="[766,1436,404,431]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nixonensis">
|
||
<emphasis id="B92A0033FFA4FFC369BBE5ACFC5493A8" box="[766,986,404,430]" italics="true" pageId="11" pageNumber="12">Blountia nixonensis</emphasis>
|
||
Lochman, in
|
||
<bibRefCitation id="EFCFA1D0FFA4FFC36F31E5ACFA1293A9" author="Lochman, C. & Duncan, D." box="[1140,1436,404,431]" pageId="11" pageNumber="12" pagination="1 - 172" refId="ref23573" refString="Lochman, C. & Duncan, D. (1944) Early Upper Cambrian faunas of Central Montana. Geological Society of America Special Papers, 54, 1 - 172. https: // doi. org / 10.1130 / SPE 54 - p 1" type="journal article" year="1944">Lochman & Duncan, 1944</bibRefCitation>
|
||
</taxonomicName>
|
||
to be conspecific with
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36ADAE580FD0493D4" authorityName="Resser" authorityYear="1938" box="[415,650,440,466]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC36ADAE580FD0493D4" box="[415,650,440,466]" italics="true" pageId="11" pageNumber="12">Blountia bristolensis</emphasis>
|
||
</taxonomicName>
|
||
. Specimens figured by
|
||
<bibRefCitation id="EFCFA1D0FFA4FFC368D8E580FB5B93D5" author="Lochman, C. & Duncan, D." box="[925,1237,440,467]" pageId="11" pageNumber="12" pagination="1 - 172" refId="ref23573" refString="Lochman, C. & Duncan, D. (1944) Early Upper Cambrian faunas of Central Montana. Geological Society of America Special Papers, 54, 1 - 172. https: // doi. org / 10.1130 / SPE 54 - p 1" type="journal article" year="1944">Lochman & Duncan (1944</bibRefCitation>
|
||
, pl. 4, figs 7–12) could not be located at the USNM, but their previously unillustrated
|
||
<typeStatus id="54E56283FFA4FFC368EAE5E5FB9493F1" box="[943,1050,477,503]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
|
||
(Pls 4, 5) are closely comparable to
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36BF0E639FEDB901C" authorityName="Resser" authorityYear="1938" box="[181,341,512,539]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC36BF0E639FF48901C" box="[181,198,513,538]" italics="true" pageId="11" pageNumber="12">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFA4FFC36B91E638FEDB901C" box="[212,341,512,538]" italics="true" pageId="11" pageNumber="12">bristolensis</emphasis>
|
||
</taxonomicName>
|
||
, and we agree that they belong to the same species. However, sclerites from the Mendha Formation in
|
||
<collectingRegion id="499A12C3FFA4FFC36BA2E61CFEB39038" box="[231,317,548,574]" country="United States of America" name="Nevada" pageId="11" pageNumber="12">Nevada</collectingRegion>
|
||
that
|
||
<bibRefCitation id="EFCFA1D0FFA4FFC36A33E61CFD9E9039" author="Palmer, A. R." box="[374,528,548,575]" pageId="11" pageNumber="12" pagination="1 - 105" refId="ref23721" refString="Palmer, A. R. (1965) Trilobites of the Late Cambrian Pterocephaliid Biomere in the Great Basin, United States. U. S. Geological Survey Professional Paper, 463, 1 - 105. https: // doi. org / 10.3133 / pp 493" type="journal article" year="1965">Palmer (1965</bibRefCitation>
|
||
, pl. 1, figs 2–3) identified as
|
||
<taxonomicName id="4C5EA7A2FFA4FFC3681EE61CFBCA9038" authorityName="Resser" authorityYear="1938" box="[859,1092,548,574]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC3681EE61CFBCA9038" box="[859,1092,548,574]" italics="true" pageId="11" pageNumber="12">Blountia bristolensis</emphasis>
|
||
</taxonomicName>
|
||
are assigned to a new species,
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36BD2E670FEF49064" authorityName="Westrop & Eoff" authorityYear="2020" box="[151,378,584,610]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nevadensis">
|
||
<emphasis id="B92A0033FFA4FFC36BD2E670FEF49064" box="[151,378,584,610]" italics="true" pageId="11" pageNumber="12">Blountia nevadensis</emphasis>
|
||
</taxonomicName>
|
||
(Pl. 16). Cranidia of the latter (Pl. 16, figs 1–3, 8) differ clearly in having a longer frontal area that features a nasute border that narrows abaxially. The pygidium of
|
||
<taxonomicName id="4C5EA7A2FFA4FFC368EAE655FBC59080" authorityName="Westrop & Eoff" authorityYear="2020" box="[943,1099,620,647]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nevadensis">
|
||
<emphasis id="B92A0033FFA4FFC368EAE655FC4E9080" box="[943,960,621,646]" italics="true" pageId="11" pageNumber="12">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFA4FFC36895E654FBC59080" box="[976,1099,620,646]" italics="true" pageId="11" pageNumber="12">nevadensis</emphasis>
|
||
</taxonomicName>
|
||
is relatively narrower (length equal to 69% of maximum width, versus 55% in
|
||
<taxonomicName id="4C5EA7A2FFA4FFC369F9E6A9FCD290AC" authorityName="Resser" authorityYear="1938" box="[700,860,656,683]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC369F9E6A9FD4390AC" box="[700,717,657,682]" italics="true" pageId="11" pageNumber="12">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFA4FFC3699EE6A8FCD290AC" box="[731,860,656,682]" italics="true" pageId="11" pageNumber="12">bristolensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE1DC21FFA4FFC36B82E68CFC4891A1" blockId="11.[151,1437,152,935]" pageId="11" pageNumber="12">
|
||
Cranidia of
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36A0BE68DFDCB90C8" authority="Westrop & Eoff, 2020" authorityName="Westrop & Eoff" authorityYear="2020" box="[334,581,692,718]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="newfoundlandensis" status="sp. nov.">
|
||
<emphasis id="B92A0033FFA4FFC36A0BE68DFDCB90C8" box="[334,581,692,718]" italics="true" pageId="11" pageNumber="12">B. newfoundlandensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92A0033FFA4FFC36909E68DFD0590C9" bold="true" box="[588,651,693,719]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="A219BD48FFA4FFC36909E68DFD0590C9" box="[588,651,693,719]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(Pl. 13, figs 5–10, Pl. 20, Pl. 21) from the Cow Head Group, western Newfoundland, resembles
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36A82E6E1FDE790F4" authorityName="Resser" authorityYear="1938" box="[455,617,728,755]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC36A82E6E1FE5690F4" box="[455,472,729,754]" italics="true" pageId="11" pageNumber="12">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFA4FFC36AADE6E0FDE790F4" box="[488,617,728,754]" italics="true" pageId="11" pageNumber="12">bristolensis</emphasis>
|
||
</taxonomicName>
|
||
in possessing a relatively short frontal area. The former species differs by having a narrower glabella (glabellar width is 69% of length versus 79% in
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36F55E6C5FB3F9110" authorityName="Resser" authorityYear="1938" box="[1040,1201,764,791]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC36F55E6C5FBAF9110" box="[1040,1057,765,790]" italics="true" pageId="11" pageNumber="12">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFA4FFC36F75E6C4FB3F9110" box="[1072,1201,764,790]" italics="true" pageId="11" pageNumber="12">bristolensis</emphasis>
|
||
</taxonomicName>
|
||
) and palpebral lobes situated slightly farther forward. Pygidial differences are decisive in separating these species. The pygidium of
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36EC1E719FEE09158" authorityName="Westrop & Eoff" authorityYear="2020" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="newfoundlandensis">
|
||
<emphasis id="B92A0033FFA4FFC36EC1E719FA1B913C" box="[1412,1429,801,826]" italics="true" pageId="11" pageNumber="12">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFA4FFC36BD2E77CFEE09158" box="[151,366,836,862]" italics="true" pageId="11" pageNumber="12">newfoundlandensis</emphasis>
|
||
</taxonomicName>
|
||
(Pl. 21) displays a border that expands noticeably towards the axis to occupy18% of pygidial length (sag.) In contrast, the border of
|
||
<taxonomicName id="4C5EA7A2FFA4FFC36902E751FD699184" authorityName="Resser" authorityYear="1938" box="[583,743,872,899]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bristolensis">
|
||
<emphasis id="B92A0033FFA4FFC36902E751FDD69184" box="[583,600,873,898]" italics="true" pageId="11" pageNumber="12">B</emphasis>
|
||
.
|
||
<emphasis id="B92A0033FFA4FFC36923E750FD699184" box="[614,743,872,898]" italics="true" pageId="11" pageNumber="12">bristolensis</emphasis>
|
||
</taxonomicName>
|
||
maintains a nearly even width along the pygidial margin, and is relatively shorter, accounting for 10% (8–12) of pygidial length (sag.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |