425 lines
50 KiB
XML
425 lines
50 KiB
XML
<document id="736DF26F30D187AEF7D1EF92DAA15EB4" ID-CLB-Dataset="45532" ID-DOI="10.5281/zenodo.279945" ID-GBIF-Dataset="29ff95c8-b1d7-4cfa-966c-ca6bd1627974" ID-ISSN="1175-5326" ID-Zenodo-Dep="279945" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460434813551" checkinUser="plazi" docAuthor="Pilato, Giovanni, Kiosya, Yevgen, Lisi, Oscar & Sabella, Giorgio" docDate="2012" docId="03F03A099B5FB461E5F9607BFC842172" docLanguage="en" docName="zt03179p060.pdf" docOrigin="Zootaxa 3179" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Paramacrobiotus klymenki Pilato, Kiosya, Lisi & Sabella, 2012, sp. nov." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="FFC942719B55B46EE56E6649FFE8234B" masterDocTitle="New records of Eutardigrada from Belarus with the description of three new species" masterLastPageNumber="60" masterPageNumber="39" pageNumber="49" updateTime="1698331938802" updateUser="ExternalLinkService">
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<mods:mods id="D27C64DE1F414113B8867CC023E84A97" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CC95652BE3A3418307B1A69077CABBB7">
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<mods:title id="728934F2ABC595B1FB9BD5AE848905A0">New records of Eutardigrada from Belarus with the description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="DEB0032280099E16A22233ABCA23D600" type="personal">
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<mods:roleTerm id="FE9F98772802AECE23C2B1F0BEBB6DEC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3818AC11C4FFD391440478A82782693A">Pilato, Giovanni</mods:namePart>
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</mods:name>
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<mods:name id="4733DFCE021F7DE29AD760BABDE8D7BB" type="personal">
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<mods:roleTerm id="E58EC26555534E0F86F0C2C56D1C695D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1D32A5FB287F9CE9A9247DE2D09818EB">Kiosya, Yevgen</mods:namePart>
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</mods:name>
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<mods:name id="7F570F7284BF6D31FB96E4C3DBF74573" type="personal">
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<mods:role id="068B8CBCE05A4D9EDFAE3528ECB132A5">
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<mods:roleTerm id="CFC303C0D0731ABC1310B47751881463">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A79AEFE8A87C583347FF27EC5A7EAC2E">Lisi, Oscar</mods:namePart>
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</mods:name>
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<mods:name id="D8723A9600174318DE3F914E2EC27C9F" type="personal">
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<mods:roleTerm id="4321E0E05C9810EB3E8C7F899B286B01">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="02D3D23E1E2838BB55D77B5680C1DEE2">Sabella, Giorgio</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="D12C439B9DBE953B56E7A6DAF0AC7C5D">text</mods:typeOfResource>
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<mods:title id="7A3E69A182597B03E1E8D6E4990EF1C6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F8D3F0107CA35E77F2D4814C6493143A">
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<mods:date id="78036BAEB8398CBBA987D51ECB4417BC">2012</mods:date>
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<mods:detail id="3C8F279F09F6432D841CBE2556A8DDE7" type="volume">
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<mods:number id="082EF108A3317A21FC1E8CC0B97F7400">3179</mods:number>
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<mods:start id="226D211DC308791DC6D41902D9DEB19F">39</mods:start>
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<mods:classification id="7A077CCEF6C71E28C0D5FCAACBE6F59D">journal article</mods:classification>
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<mods:identifier id="2C97A8460708727FC450EF37868CB425" type="CLB-Dataset">45532</mods:identifier>
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<mods:identifier id="661E237885E3EDA3DF3EE6E5C7105A41" type="DOI">10.5281/zenodo.279945</mods:identifier>
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<mods:identifier id="A954A6C10AD92385C98CAF2E0E132313" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8ECE90E1F44A26C02F1CA18737A89037" type="Zenodo-Dep">279945</mods:identifier>
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<treatment id="03F03A099B5FB461E5F9607BFC842172" ID-DOI="http://doi.org/10.5281/zenodo.6180891" ID-GBIF-Taxon="119431464" ID-Zenodo-Dep="6180891" LSID="urn:lsid:plazi:treatment:03F03A099B5FB461E5F9607BFC842172" httpUri="http://treatment.plazi.org/id/03F03A099B5FB461E5F9607BFC842172" lastPageId="15" lastPageNumber="54" pageId="10" pageNumber="49">
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<subSubSection id="C343D8949B5FB464E5F9607BFF032526" pageId="10" pageNumber="49" type="nomenclature">
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<paragraph id="8BE68B1F9B5FB464E5F9607BFDAA2500" blockId="10.[151,578,1585,1645]" box="[151,578,1585,1612]" pageId="10" pageNumber="49">
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||
<heading id="D0AE3C739B5FB464E5F9607BFDAA2500" bold="true" box="[151,578,1585,1612]" fontSize="11" level="1" pageId="10" pageNumber="49" reason="1">
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||
<emphasis id="B92D570D9B5FB464E5F9607BFDAA2500" bold="true" box="[151,578,1585,1612]" pageId="10" pageNumber="49">
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||
<taxonomicName id="4C59F09C9B5FB464E5F9607BFE352507" ID-CoL="a620e26c-221e-4aac-a5d3-162cea670366" box="[151,477,1586,1612]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="49" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
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<emphasis id="B92D570D9B5FB464E5F9607BFE352507" bold="true" box="[151,477,1586,1612]" italics="true" pageId="10" pageNumber="49">Paramacrobiotus klymenki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21EEA769B5FB464E48A6078FDAA2500" box="[484,578,1585,1611]" pageId="10" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE68B1F9B5FB464E5F9601CFF032526" blockId="10.[151,578,1585,1645]" box="[151,235,1621,1645]" pageId="10" pageNumber="49">
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||
(
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<figureCitation id="1362979A9B5FB464E5F1601CFF0B2526" box="[159,227,1621,1645]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="10" pageNumber="49">Fig. 6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C343D8949B5FB465E5F960D4FB5323BC" lastPageId="11" lastPageNumber="50" pageId="10" pageNumber="49" type="materials_examined">
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<paragraph id="8BE68B1F9B5FB464E5F960D4FAFE2592" blockId="10.[151,1437,1692,2006]" pageId="10" pageNumber="49">
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<emphasis id="B92D570D9B5FB464E5F960D4FEA225FE" bold="true" box="[151,330,1692,1717]" pageId="10" pageNumber="49">
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<typeStatus id="54E235BD9B5FB464E5F960D4FF3A25FE" box="[151,210,1693,1717]" pageId="10" pageNumber="49">Type</typeStatus>
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material.
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</emphasis>
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Vitebsk Oblast, Sennensky district, spruce forest, vicinity of Margoitsy village (
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<geoCoordinate id="EE6DEDD89B5FB464E07E60D4FA7F25FE" box="[1296,1431,1693,1717]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="54.671947">54°40'19"N</geoCoordinate>
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,
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<geoCoordinate id="EE6DEDD89B5FB464E5F96089FEF32593" box="[151,283,1728,1752]" direction="east" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="29.624166">29°37'27"E</geoCoordinate>
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); moss sample collected (
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<date id="FFE7ADDF9B5FB464E7546089FD582593" box="[570,688,1728,1752]" pageId="10" pageNumber="49" value="2002-06">June 2002</date>
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) by G.O. Mazepa:
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<typeStatus id="54E235BD9B5FB464E6ED6089FC002593" box="[899,1000,1728,1752]" pageId="10" pageNumber="49" type="holotype">holotype</typeStatus>
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, 71
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<typeStatus id="54E235BD9B5FB464E1786088FB692592" box="[1046,1153,1729,1753]" pageId="10" pageNumber="49" type="paratype">paratypes</typeStatus>
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and
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<specimenCount id="9D5F40969B5FB464E1D66089FAFA2592" box="[1208,1298,1728,1753]" pageId="10" pageNumber="49" type="egg">45 eggs</specimenCount>
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.
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</paragraph>
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<paragraph id="8BE68B1F9B5FB464E5A960ACFEEA2423" blockId="10.[151,1437,1692,2006]" pageId="10" pageNumber="49">
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<emphasis id="B92D570D9B5FB464E5A960ACFE6025B6" bold="true" box="[199,392,1764,1789]" pageId="10" pageNumber="49">
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<typeStatus id="54E235BD9B5FB464E5A960ACFEE825B6" box="[199,256,1765,1789]" pageId="10" pageNumber="49">Type</typeStatus>
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repository.
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</emphasis>
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<typeStatus id="54E235BD9B5FB464E4FD60ACFE1325B6" box="[403,507,1765,1789]" pageId="10" pageNumber="49" type="holotype">Holotype</typeStatus>
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(slide No. 5425), 23
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<typeStatus id="54E235BD9B5FB464E79D60AFFCB725B5" box="[755,863,1766,1790]" pageId="10" pageNumber="49" type="paratype">paratypes</typeStatus>
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and
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<specimenCount id="9D5F40969B5FB464E6F660ACFC1825B5" box="[920,1008,1765,1790]" pageId="10" pageNumber="49" type="egg">14 eggs</specimenCount>
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(slide No. 5425) are deposited in the collection of Binda & Pilato (Museum of the Department of Animal Biology “Marcello La Greca”, University of Catania,
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<collectingCountry id="F34ECB8F9B5FB464E5956164FEDB240E" box="[251,307,1837,1861]" name="Italy" pageId="10" pageNumber="49">Italy</collectingCountry>
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). Other specimens and eggs are deposited in the collection of Kiosya (Kharkiv National University,
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<collectingCountry id="F34ECB8F9B5FB464E5F96119FF1F2423" box="[151,247,1872,1896]" name="Ukraine" pageId="10" pageNumber="49">Ukraine</collectingCountry>
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).
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</paragraph>
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<paragraph id="8BE68B1F9B5FB465E5A9613DFB5323BC" blockId="10.[151,1437,1692,2006]" lastBlockId="11.[151,1436,151,501]" lastPageId="11" lastPageNumber="50" pageId="10" pageNumber="49">
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<emphasis id="B92D570D9B5FB464E5A9613DFE4924C6" bold="true" box="[199,417,1908,1933]" pageId="10" pageNumber="49">Specific diagnosis.</emphasis>
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Colourless, cuticle smooth without pores; small dots present on all legs. Eye spots absent. Pharyngeal bulb with three macroplacoids; microplacoid absent; buccal tube not very wide (
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<emphasis id="B92D570D9B5FB464E1D961D3FB2524FA" box="[1207,1229,1946,1969]" italics="true" pageId="10" pageNumber="49">pt</emphasis>
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14.7–16.6); stylet supports inserted on the buccal tube at 79–80%. Claws, of
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<taxonomicName id="4C59F09C9B5FB464E62F61F4FC43249E" box="[833,939,1981,2005]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="49" phylum="Tardigrada" rank="species" species="hufelandi">
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<emphasis id="B92D570D9B5FB464E62F61F4FC43249E" box="[833,939,1981,2005]" italics="true" pageId="10" pageNumber="49">hufelandi</emphasis>
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</taxonomicName>
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-
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<typeStatus id="54E235BD9B5FB464E6DC61F7FC0E249D" box="[946,998,1982,2006]" pageId="10" pageNumber="49">type</typeStatus>
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, slender, with accessory points on the main branches; lunules present. The eggs, freely laid, were spherical, with 10–11 conical processes on the circumference. The processes’ surface had a well visible reticular design consisting of elongate mesh with rectilinear margins. Egg shell areolated; the central area of each areola slightly thickened and not subdivided.
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</paragraph>
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</subSubSection>
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<subSubSection id="C343D8949B5EB463E5A9674AFD0425A3" lastPageId="13" lastPageNumber="52" pageId="11" pageNumber="50" type="description">
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<paragraph id="8BE68B1F9B5EB465E5A9674AFAE622BF" blockId="11.[151,1436,151,501]" pageId="11" pageNumber="50">
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<emphasis id="B92D570D9B5EB465E5A9674AFDFC2257" bold="true" box="[199,532,259,284]" pageId="11" pageNumber="50">
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Description of the
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<typeStatus id="54E235BD9B5EB465E4CA674DFDE72257" box="[420,527,260,284]" pageId="11" pageNumber="50" type="holotype">holotype</typeStatus>
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:
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</emphasis>
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Body length 340 µm; colourless, cuticle smooth without pores; small dots present on the legs. Eye spots absent. Bucco-pharyngeal apparatus of
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<taxonomicName id="4C59F09C9B5EB465E6196761FBED220B" box="[887,1029,296,320]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="50" phylum="Tardigrada" rank="genus">
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<emphasis id="B92D570D9B5EB465E6196761FBED220B" box="[887,1029,296,320]" italics="true" pageId="11" pageNumber="50">Macrobiotus</emphasis>
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</taxonomicName>
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-
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<typeStatus id="54E235BD9B5EB465E1636761FBD7220B" box="[1037,1087,296,320]" pageId="11" pageNumber="50">type</typeStatus>
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(
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<figureCitation id="1362979A9B5EB465E120676EFB632274" box="[1102,1163,295,319]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="11" pageNumber="50">Fig.6</figureCitation>
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A); mouth terminal with ten peribuccal lamellae; buccal armature with an anterior ring of small teeth, a posterior ring of triangular teeth and a system of three dorsal and three ventral transverse ridges. Buccal tube 42.1 µm long and 6.6 µm wide externally (
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<emphasis id="B92D570D9B5EB465E5CE67DCFF5E22E7" box="[160,182,405,428]" italics="true" pageId="11" pageNumber="50">pt</emphasis>
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||
= 15.7). Stylet supports inserted on the buccal tube at 79.8 % of its length (
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<emphasis id="B92D570D9B5EB465E69567DCFBF922E7" box="[1019,1041,405,428]" italics="true" pageId="11" pageNumber="50">pt</emphasis>
|
||
= 79.8). Pharyngeal bulb with apophyses and three macroplacoids; microplacoid absent (
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<figureCitation id="1362979A9B5EB465E66A67FEFCA02284" box="[772,840,439,463]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="11" pageNumber="50">Fig. 6</figureCitation>
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A). Length of first macroplacoid 7.4 µm (
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||
<emphasis id="B92D570D9B5EB465E04A67F0FAD1229B" box="[1316,1337,441,464]" italics="true" pageId="11" pageNumber="50">pt</emphasis>
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||
= 17.6), second 5.2 µm (
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<emphasis id="B92D570D9B5EB465E4226794FE8922BF" box="[332,353,477,500]" italics="true" pageId="11" pageNumber="50">pt</emphasis>
|
||
= 12.4), third 7.3 µm (
|
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<emphasis id="B92D570D9B5EB465E70A6794FD9122BF" box="[612,633,477,500]" italics="true" pageId="11" pageNumber="50">pt</emphasis>
|
||
= 17.3); entire macroplacoid row 21.2 µm long (
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||
<emphasis id="B92D570D9B5EB465E1CE6794FB5E22BF" box="[1184,1206,477,500]" italics="true" pageId="11" pageNumber="50">pt</emphasis>
|
||
= 50.4)
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</paragraph>
|
||
<caption id="DF26DB979B5EB465E5F96175FE5724E5" httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="11" pageNumber="50" targetBox="[191,1386,576,1793]" targetPageId="11">
|
||
<paragraph id="8BE68B1F9B5EB465E5F96175FE5724E5" blockId="11.[151,1436,1852,1966]" pageId="11" pageNumber="50">
|
||
<emphasis id="B92D570D9B5EB465E5F96175FEFD2419" bold="true" box="[151,277,1852,1875]" pageId="11" pageNumber="50">FIGURE 6.</emphasis>
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<taxonomicName id="4C59F09C9B5EB465E4706174FDD92418" box="[286,561,1853,1875]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="50" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5EB465E4706174FDD92418" box="[286,561,1853,1875]" italics="true" pageId="11" pageNumber="50">Paramacrobiotus klymenki</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5EB465E7546174FD642419" box="[570,652,1853,1874]" pageId="11" pageNumber="50" rank="species">
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||
<emphasis id="B92D570D9B5EB465E7546174FD6F2419" bold="true" box="[570,647,1853,1874]" pageId="11" pageNumber="50">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF26DB979B58B463E5F9628DFE402653" httpUri="https://zenodo.org/record/279952/files/figure.png" pageId="13" pageNumber="52" targetBox="[179,1403,219,1169]" targetPageId="13">
|
||
<paragraph id="8BE68B1F9B58B463E5F9628DFE402653" blockId="13.[151,1436,1220,1304]" pageId="13" pageNumber="52">
|
||
<emphasis id="B92D570D9B58B463E5F9628DFEFB2791" bold="true" box="[151,275,1220,1243]" pageId="13" pageNumber="52">FIGURE 7.</emphasis>
|
||
A, B,
|
||
<taxonomicName id="4C59F09C9B58B463E437628CFD872790" box="[345,623,1221,1243]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="52" phylum="Tardigrada" rank="species" species="areolatus">
|
||
<emphasis id="B92D570D9B58B463E437628CFD872790" box="[345,623,1221,1243]" italics="true" pageId="13" pageNumber="52">Paramacrobiotus areolatus</emphasis>
|
||
</taxonomicName>
|
||
(slide No. 3066). A, Bucco-pharyngeal apparatus. B, Claws of the third pair of legs. C, D,
|
||
<taxonomicName id="4C59F09C9B58B463E46762AAFDEF27B2" box="[265,519,1251,1273]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="52" phylum="Tardigrada" rank="species" species="walteri">
|
||
<emphasis id="B92D570D9B58B463E46762AAFDEF27B2" box="[265,519,1251,1273]" italics="true" pageId="13" pageNumber="52">Paramacrobiotus walteri</emphasis>
|
||
</taxonomicName>
|
||
(paratype, slide No 4356). C, Bucco-pharyngeal apparatus. D, Claws of the second pair of legs. (Scale bars = 10 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE68B1F9B58B463E5A9630DFD02255B" blockId="13.[151,1437,1348,2021]" pageId="13" pageNumber="52">
|
||
Claws, of
|
||
<taxonomicName id="4C59F09C9B58B463E457630CFE4B2616" box="[313,419,1349,1373]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="52" phylum="Tardigrada" rank="species" species="hufelandi">
|
||
<emphasis id="B92D570D9B58B463E457630CFE4B2616" box="[313,419,1349,1373]" italics="true" pageId="13" pageNumber="52">hufelandi</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="54E235BD9B58B463E4C4630CFE372616" box="[426,479,1349,1373]" pageId="13" pageNumber="52">type</typeStatus>
|
||
, slender, with accessory points on the main branches (
|
||
<figureCitation id="1362979A9B58B463E153630DFB972617" box="[1085,1151,1348,1372]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="13" pageNumber="52">Fig. 6</figureCitation>
|
||
B–D). External and internal claws of the first pair of legs 10.4 µm (
|
||
<emphasis id="B92D570D9B58B463E71D6323FD6126CA" box="[627,649,1386,1409]" italics="true" pageId="13" pageNumber="52">pt</emphasis>
|
||
= 24.7) and 10.2 µm (
|
||
<emphasis id="B92D570D9B58B463E6ED6323FC7126CA" box="[899,921,1386,1409]" italics="true" pageId="13" pageNumber="52">pt</emphasis>
|
||
= 24.2) long respectively; those of the second and third pair of legs 11.7 µm (
|
||
<emphasis id="B92D570D9B58B463E49D63C7FDE026EE" box="[499,520,1422,1445]" italics="true" pageId="13" pageNumber="52">pt</emphasis>
|
||
= 27.8) and 10.9 µm (
|
||
<emphasis id="B92D570D9B58B463E66A63C7FCF226EE" box="[772,794,1422,1445]" italics="true" pageId="13" pageNumber="52">pt</emphasis>
|
||
= 25.9) long respectively; posterior and anterior claws of the hind legs 13.0 µm (
|
||
<emphasis id="B92D570D9B58B463E4F263FBFE5A2682" box="[412,434,1458,1481]" italics="true" pageId="13" pageNumber="52">pt</emphasis>
|
||
= 30.9) and 11.8 µm (
|
||
<emphasis id="B92D570D9B58B463E7C163FBFD2D2682" box="[687,709,1458,1481]" italics="true" pageId="13" pageNumber="52">pt</emphasis>
|
||
= 28.0) long respectively. Lunules present, small and smooth on the first three pairs of legs, larger and with indented margin on the hind legs. Two cuticular bars present below the claws on the first three pairs of legs (
|
||
<figureCitation id="1362979A9B58B463E75B63B1FD90255B" box="[565,632,1528,1552]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="13" pageNumber="52">Fig. 6</figureCitation>
|
||
B, arrow).
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B58B463E5A96055FD0425A3" blockId="13.[151,1437,1348,2021]" pageId="13" pageNumber="52">
|
||
<emphasis id="B92D570D9B58B463E5A96055FE36257E" bold="true" box="[199,478,1564,1589]" pageId="13" pageNumber="52">Description of the eggs.</emphasis>
|
||
The eggs, freely laid, were spherical, with 10–11 conical processes on the circumference and 22–26 on the hemisphere. The diameter was 69–71 µm excluding the processes, 101–109 µm including. The processes were conical, elongate (sometimes with slightly forked apices), 14.5–18.5 high with basal diameter of 16.4–18.2 µm. The processes surface had a well visible reticular design consisting of elongate mesh with rectilinear margins (
|
||
<figureCitation id="1362979A9B58B463E42660E5FE64258F" box="[328,396,1708,1732]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="13" pageNumber="52">Fig. 6</figureCitation>
|
||
E). A short apical portion of each process was smooth (
|
||
<figureCitation id="1362979A9B58B463E69360E5FBA8258F" box="[1021,1088,1708,1732]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="13" pageNumber="52">Fig. 6</figureCitation>
|
||
E, arrows). Egg shell areolated and the central area of each areola slightly thickened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343D8949B58B463E5A960BCFCB6241F" pageId="13" pageNumber="52" type="discussion">
|
||
<paragraph id="8BE68B1F9B58B463E5A960BCFCB6241F" blockId="13.[151,1437,1348,2021]" pageId="13" pageNumber="52">
|
||
<emphasis id="B92D570D9B58B463E5A960BCFED32446" bold="true" box="[199,315,1781,1805]" pageId="13" pageNumber="52">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="54E235BD9B58B463E41B60BCFE082446" box="[373,480,1781,1805]" pageId="13" pageNumber="52" type="paratype">paratypes</typeStatus>
|
||
were similar to the
|
||
<typeStatus id="54E235BD9B58B463E7AF60BDFCCB2447" box="[705,803,1780,1804]" pageId="13" pageNumber="52" type="holotype">holotype</typeStatus>
|
||
in both qualitative and metric characters (we referred to the
|
||
<emphasis id="B92D570D9B58B463E5AD6153FF30247A" box="[195,216,1818,1841]" italics="true" pageId="13" pageNumber="52">pt</emphasis>
|
||
index values when the specimens had different body length). The measurements of some specimens, the smallest and the largest ones included, are presented in Table 3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C343D8949B58B461E5A96128FC842172" lastPageId="15" lastPageNumber="54" pageId="13" pageNumber="52" type="etymology">
|
||
<paragraph id="8BE68B1F9B58B463E5A96128FD7024D7" blockId="13.[151,1437,1348,2021]" pageId="13" pageNumber="52">
|
||
<emphasis id="B92D570D9B58B463E5A96128FEA62432" bold="true" box="[199,334,1889,1913]" pageId="13" pageNumber="52">Etymology.</emphasis>
|
||
The specific name is in honour to Prof. Vyacheslav Klymenko (Kharkiv National University,
|
||
<collectingCountry id="F34ECB8F9B58B463E5F961CDFF1E24D7" box="[151,246,1924,1948]" name="Ukraine" pageId="13" pageNumber="52">Ukraine</collectingCountry>
|
||
), the scientific adviser of Ye. Kiosya.
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B58B460E5A961E1FF1B23BC" blockId="13.[151,1437,1348,2021]" lastBlockId="14.[151,1436,151,391]" lastPageId="14" lastPageNumber="53" pageId="13" pageNumber="52">
|
||
<emphasis id="B92D570D9B58B463E5A961E1FE27248A" bold="true" box="[199,463,1960,1985]" pageId="13" pageNumber="52">Differential diagnosis.</emphasis>
|
||
Prior to the new species described above, six species of the genus
|
||
<taxonomicName id="4C59F09C9B58B463E1B761E0FA74248A" box="[1241,1436,1961,1985]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="52" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B92D570D9B58B463E1B761E0FA74248A" box="[1241,1436,1961,1985]" italics="true" pageId="13" pageNumber="52">Paramacrobiotus</emphasis>
|
||
</taxonomicName>
|
||
were described with a smooth cuticle, three macroplacoids, lacking the microplacoid and producing eggs with areolated shell and conical processes:
|
||
<taxonomicName id="4C59F09C9B5BB460E75E66D0FD5023FB" box="[560,696,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="areolatus">
|
||
<emphasis id="B92D570D9B5BB460E75E66D0FD5023FB" box="[560,696,152,176]" italics="true" pageId="14" pageNumber="53">P. areolatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B92D570D9B5BB460E7A766D0FC2E23FB" box="[713,966,152,176]" italics="true" pageId="14" pageNumber="53">
|
||
<taxonomicName id="4C59F09C9B5BB460E7A766D0FCA223FB" box="[713,842,153,176]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="crenatus">P. crenatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C59F09C9B5BB460E63766D0FC2E23FB" box="[857,966,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="walteri">P. walteri</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B92D570D9B5BB460E6B666D0FB1923FB" box="[984,1265,152,176]" italics="true" pageId="14" pageNumber="53">
|
||
<taxonomicName id="4C59F09C9B5BB460E6B666D0FB9D23FB" box="[984,1141,153,176]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="centesimus">P. centesimus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C59F09C9B5BB460E1ED66D0FB1923FB" box="[1155,1265,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="huziori">P. huziori</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B92D570D9B5BB460E04066D0FA7923FB" box="[1326,1425,152,176]" italics="true" pageId="14" pageNumber="53">P. de r k a i</emphasis>
|
||
. Comparing adults of, where possible, comparable size the new species can be from the other species (Table 3) as follows:
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5BB460E5A9674DFD5922CC" blockId="14.[151,1436,151,391]" pageId="14" pageNumber="53">
|
||
<taxonomicName id="4C59F09C9B5BB460E5A9674DFE1C2257" box="[199,500,260,284]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5BB460E5A9674DFE1C2257" box="[199,500,260,284]" italics="true" pageId="14" pageNumber="53">Paramacrobiotus klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5BB460E494674DFDBB2257" box="[506,595,260,284]" pageId="14" pageNumber="53" rank="species">
|
||
<emphasis id="B92D570D9B5BB460E494674DFDA42257" bold="true" box="[506,588,260,284]" pageId="14" pageNumber="53">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
was very similar to
|
||
<emphasis id="B92D570D9B5BB460E65B674CFC5D2257" box="[821,949,260,284]" italics="true" pageId="14" pageNumber="53">P. a re o l a t u s</emphasis>
|
||
but differed in lacking eye spots; in having the buccal tube narrower and slightly longer with respect to the body length (Table 3;
|
||
<figureCitation id="1362979A9B5BB460E10B676EFB592274" box="[1125,1201,295,319]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="14" pageNumber="53">Figs. 6</figureCitation>
|
||
A and 7A); and with slightly shorter claws (Table 3) (however, the
|
||
<emphasis id="B92D570D9B5BB460E7CF6704FD5E222F" box="[673,694,333,356]" italics="true" pageId="14" pageNumber="53">pt</emphasis>
|
||
values may appear similar as the buccal tube of
|
||
<emphasis id="B92D570D9B5BB460E18D6704FA8C222F" box="[1251,1380,332,356]" italics="true" pageId="14" pageNumber="53">P. a re o l a t u s</emphasis>
|
||
was slightly shorter with respect to the body length).
|
||
</paragraph>
|
||
<caption id="DF26DB979B5BB460E5F9600FFA9E25D2" httpUri="https://zenodo.org/record/279953/files/figure.png" pageId="14" pageNumber="53" targetBox="[186,1403,450,1557]" targetPageId="14">
|
||
<paragraph id="8BE68B1F9B5BB460E5F9600FFA9E25D2" blockId="14.[151,1435,1606,1690]" pageId="14" pageNumber="53">
|
||
<emphasis id="B92D570D9B5BB460E5F9600FFEFA2510" bold="true" box="[151,274,1606,1629]" pageId="14" pageNumber="53">FIGURE 8.</emphasis>
|
||
A, B,
|
||
<taxonomicName id="4C59F09C9B5BB460E43B600FFD9F2517" box="[341,631,1606,1628]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="centesimus">
|
||
<emphasis id="B92D570D9B5BB460E43B600FFD9F2517" box="[341,631,1606,1628]" italics="true" pageId="14" pageNumber="53">Paramacrobiotus centesimus</emphasis>
|
||
</taxonomicName>
|
||
(slide No. 4740). A, Bucco-pharyngeal apparatus: B, Detail of an egg. The very dense reticular design of the processes surface is visible. C, D,
|
||
<taxonomicName id="4C59F09C9B5BB460E67F602CFBE52530" box="[785,1037,1637,1659]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="huziori">
|
||
<emphasis id="B92D570D9B5BB460E67F602CFBE52530" box="[785,1037,1637,1659]" italics="true" pageId="14" pageNumber="53">Paramacrobiotus huziori</emphasis>
|
||
</taxonomicName>
|
||
(slide No. 5408). Details of an egg: the dense reticular design of the processes (arrows ‘a’) and the subdivided areolae (arrows ‘b’) are visible. (Scale bars = 10 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE68B1F9B5BB460E5A96089FC10240E" blockId="14.[151,1437,1728,2005]" pageId="14" pageNumber="53">
|
||
The new species differed from
|
||
<taxonomicName id="4C59F09C9B5BB460E75A608BFD5A2592" box="[564,690,1730,1753]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="crenatus">
|
||
<emphasis id="B92D570D9B5BB460E75A608BFD5A2592" box="[564,690,1730,1753]" italics="true" pageId="14" pageNumber="53">P. crenatus</emphasis>
|
||
</taxonomicName>
|
||
in having a narrower buccal tube; lunules on the hind legs less developed; smaller eggs (diameter 69–71 µm excluding the processes in
|
||
<taxonomicName id="4C59F09C9B5BB460E6AA60AFFBAB25B6" box="[964,1091,1765,1789]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5BB460E6AA60AFFBAB25B6" box="[964,1091,1765,1789]" italics="true" pageId="14" pageNumber="53">P. klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5BB460E12460ACFB4A25B6" box="[1098,1186,1765,1789]" pageId="14" pageNumber="53" rank="species">
|
||
<emphasis id="B92D570D9B5BB460E12460ACFB7425B6" bold="true" box="[1098,1180,1765,1789]" pageId="14" pageNumber="53">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, 92–128 µm in
|
||
<emphasis id="B92D570D9B5BB460E03A60AFFF3C246A" italics="true" pageId="14" pageNumber="53">P. c re - natus</emphasis>
|
||
); smaller egg processes (14.5–18.5 µm high in
|
||
<taxonomicName id="4C59F09C9B5BB460E66E6143FC6A246A" box="[768,898,1801,1825]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5BB460E66E6143FC6A246A" box="[768,898,1801,1825]" italics="true" pageId="14" pageNumber="53">P. klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5BB460E6E26140FC0F246B" box="[908,999,1801,1825]" pageId="14" pageNumber="53" rank="species">
|
||
<emphasis id="B92D570D9B5BB460E6E26140FC09246A" bold="true" box="[908,993,1801,1825]" pageId="14" pageNumber="53">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, 40–44 µm in
|
||
<taxonomicName id="4C59F09C9B5BB460E1F66143FAFD246A" box="[1176,1301,1801,1825]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="crenatus">
|
||
<emphasis id="B92D570D9B5BB460E1F66143FB4E246A" box="[1176,1190,1802,1825]" italics="true" pageId="14" pageNumber="53">P</emphasis>
|
||
.
|
||
<emphasis id="B92D570D9B5BB460E1D86143FAFD246A" box="[1206,1301,1802,1825]" italics="true" pageId="14" pageNumber="53">crenatus</emphasis>
|
||
</taxonomicName>
|
||
, with basal diameter 16.4–18.2 µm in
|
||
<taxonomicName id="4C59F09C9B5BB460E4AE6167FDA8240E" box="[448,576,1837,1861]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5BB460E4AE6167FDA8240E" box="[448,576,1837,1861]" italics="true" pageId="14" pageNumber="53">P. klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5BB460E7286164FD48240E" box="[582,672,1837,1861]" pageId="14" pageNumber="53" rank="species">
|
||
<emphasis id="B92D570D9B5BB460E7286164FD71240E" bold="true" box="[582,665,1837,1861]" pageId="14" pageNumber="53">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and 24–28 µm in
|
||
<taxonomicName id="4C59F09C9B5BB460E6026167FC0F240E" box="[876,999,1838,1861]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="crenatus">
|
||
<emphasis id="B92D570D9B5BB460E6026167FC0F240E" box="[876,999,1838,1861]" italics="true" pageId="14" pageNumber="53">P. crenatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5BB460E5A96118FF11249E" blockId="14.[151,1437,1728,2005]" pageId="14" pageNumber="53">
|
||
<taxonomicName id="4C59F09C9B5BB460E5A96118FE1D2422" box="[199,501,1873,1897]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5BB460E5A96118FE1D2422" box="[199,501,1873,1897]" italics="true" pageId="14" pageNumber="53">Paramacrobiotus klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5BB460E4926118FDBE2423" box="[508,598,1873,1897]" pageId="14" pageNumber="53" rank="species">
|
||
<emphasis id="B92D570D9B5BB460E4926118FDA72422" bold="true" box="[508,591,1873,1897]" pageId="14" pageNumber="53">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differed from
|
||
<emphasis id="B92D570D9B5BB460E792611BFC8B2422" box="[764,867,1873,1897]" italics="true" pageId="14" pageNumber="53">P. w al te r i</emphasis>
|
||
in lacking eye spots; in having the stylet supports inserted on the buccal tube in a more caudal position (Table 3), shorter claws (Table 3), also different in shape (in
|
||
<taxonomicName id="4C59F09C9B5BB460E0E9613FFF0F24FA" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="53" phylum="Tardigrada" rank="species" species="walteri">
|
||
<emphasis id="B92D570D9B5BB460E0E9613FFF0F24FA" italics="true" pageId="14" pageNumber="53">P. walteri</emphasis>
|
||
</taxonomicName>
|
||
the difference in length between main and secondary branch was markedly higher, as seen in
|
||
<figureCitation id="1362979A9B5BB460E07761D1FA8E24FB" box="[1305,1382,1944,1968]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="14" pageNumber="53">Figs. 6</figureCitation>
|
||
B–D and 7D).
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5AB461E5A966D1FD922274" blockId="15.[151,1436,151,569]" pageId="15" pageNumber="54">
|
||
<taxonomicName id="4C59F09C9B5AB461E5A966D1FE1F23FB" box="[199,503,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="54" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5AB461E5A966D1FE1F23FB" box="[199,503,152,176]" italics="true" pageId="15" pageNumber="54">Paramacrobiotus klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5AB461E76F66D1FDB523E4" box="[513,605,152,176]" pageId="15" pageNumber="54" rank="species">
|
||
<emphasis id="B92D570D9B5AB461E76F66D1FDBE23FB" bold="true" box="[513,598,152,176]" pageId="15" pageNumber="54">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differed from
|
||
<taxonomicName id="4C59F09C9B5AB461E66466D0FC4B23FB" box="[778,931,153,176]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="54" phylum="Tardigrada" rank="species" species="centesimus">
|
||
<emphasis id="B92D570D9B5AB461E66466D0FC4B23FB" box="[778,931,153,176]" italics="true" pageId="15" pageNumber="54">P. centesimus</emphasis>
|
||
</taxonomicName>
|
||
in having a narrower buccal tube (Table 3;
|
||
<figureCitation id="1362979A9B5AB461E5F966F2FF0B2398" box="[151,227,187,211]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="15" pageNumber="54">Figs. 6</figureCitation>
|
||
A and 8A), lunules on the hind legs indented; longer egg processes (14.5–18.5 µm in
|
||
<taxonomicName id="4C59F09C9B5AB461E1DD66F4FADB239F" box="[1203,1331,188,212]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="54" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5AB461E1DD66F4FADB239F" box="[1203,1331,188,212]" italics="true" pageId="15" pageNumber="54">P. klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5AB461E05566F5FA7C2398" box="[1339,1428,188,212]" pageId="15" pageNumber="54" rank="species">
|
||
<emphasis id="B92D570D9B5AB461E05566F5FA66239F" bold="true" box="[1339,1422,188,212]" pageId="15" pageNumber="54">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, 7–11 µm in
|
||
<taxonomicName id="4C59F09C9B5AB461E44D66A9FE5323BC" box="[291,443,224,247]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="54" phylum="Tardigrada" rank="species" species="centesimus">
|
||
<emphasis id="B92D570D9B5AB461E44D66A9FE5323BC" box="[291,443,224,247]" italics="true" pageId="15" pageNumber="54">P. centesimus</emphasis>
|
||
</taxonomicName>
|
||
), slightly different reticular design of the processes (the design was less dense, and the mesh larger and more elongate in
|
||
<taxonomicName id="4C59F09C9B5AB461E77F674DFD782250" box="[529,656,259,283]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="54" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5AB461E77F674DFD782250" box="[529,656,259,283]" italics="true" pageId="15" pageNumber="54">P. klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5AB461E7FB674AFD052250" box="[661,749,259,283]" pageId="15" pageNumber="54" rank="species">
|
||
<emphasis id="B92D570D9B5AB461E7FB674AFD0F2250" bold="true" box="[661,743,259,283]" pageId="15" pageNumber="54">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, while the design was more dense and the mesh almost isodiametric in
|
||
<taxonomicName id="4C59F09C9B5AB461E4606761FE4D2274" box="[270,421,296,319]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="54" phylum="Tardigrada" rank="species" species="centesimus">
|
||
<emphasis id="B92D570D9B5AB461E4606761FE4D2274" box="[270,421,296,319]" italics="true" pageId="15" pageNumber="54">P. centesimus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1362979A9B5AB461E4DA676EFDE92274" box="[436,513,295,319]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="15" pageNumber="54">Figs. 6</figureCitation>
|
||
E and 8B).
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5AB461E5A96702FD8E2285" blockId="15.[151,1436,151,569]" pageId="15" pageNumber="54">
|
||
<taxonomicName id="4C59F09C9B5AB461E5A96702FE1E2228" box="[199,502,331,355]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="54" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5AB461E5A96702FE1E2228" box="[199,502,331,355]" italics="true" pageId="15" pageNumber="54">Paramacrobiotus klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5AB461E4936702FDBF2229" box="[509,599,331,355]" pageId="15" pageNumber="54" rank="species">
|
||
<emphasis id="B92D570D9B5AB461E4936702FDB82228" bold="true" box="[509,592,331,355]" pageId="15" pageNumber="54">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differed from
|
||
<emphasis id="B92D570D9B5AB461E7936705FC8D2228" box="[765,869,331,355]" italics="true" pageId="15" pageNumber="54">P. h u z i o r i</emphasis>
|
||
in lacking eye spots, in having a narrower buccal tube, lunules on the hind legs indented, egg processes with less dense reticular design and more clearly elongate mesh with more rectilinear margins (
|
||
<figureCitation id="1362979A9B5AB461E75667DBFD6D22E1" box="[568,645,402,426]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="15" pageNumber="54">Figs. 6</figureCitation>
|
||
E and 8C, D, arrow a); and egg shell areolae not subdivided (
|
||
<figureCitation id="1362979A9B5AB461E05367DBFA6222E1" box="[1341,1418,402,426]" captionStart="FIGURE 6" captionStartId="11.[151,250,1852,1875]" captionTargetBox="[191,1386,576,1793]" captionTargetId="figure@11.[160,1427,553,1831]" captionTargetPageId="11" captionText="FIGURE 6. Paramacrobiotus klymenki sp. nov. A, Bucco-pharyngeal apparatus of the holotype (slide No. 5425). B, C, D, Claws of the second, third and fourth pairs of legs respectively of the holotype (slide No. 5425). The arrow in B indicates the cuticular bar below the lunules. E, Detail of an egg (slide No. 5425). The arrows indicate the unsculptured apices of egg processes. (Scale bars = 10 µm)." httpUri="https://zenodo.org/record/279951/files/figure.png" pageId="15" pageNumber="54">Figs. 6</figureCitation>
|
||
E and 8C, D, arrow b, and 8C,D, arrows b).
|
||
</paragraph>
|
||
<paragraph id="8BE68B1F9B5AB461E5A96790FC842172" blockId="15.[151,1436,151,569]" pageId="15" pageNumber="54">
|
||
Lack of eye spots, indented lunules on hind legs, claws with less developed accessory points, egg shell areolae not subdivided, egg processes with basal diameter longer than the process height and reticular design less dense, distinguishes
|
||
<taxonomicName id="4C59F09C9B5AB461E45E6468FDB62172" box="[304,606,545,569]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="54" phylum="Tardigrada" rank="species" species="klymenki" status="sp. nov.">
|
||
<emphasis id="B92D570D9B5AB461E45E6468FDB62172" box="[304,606,545,569]" italics="true" pageId="15" pageNumber="54">Paramacrobiotus klymenki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21EEA769B5AB461E70A6468FD562172" box="[612,702,545,569]" pageId="15" pageNumber="54" rank="species">
|
||
<emphasis id="B92D570D9B5AB461E70A6468FD5F2172" bold="true" box="[612,695,545,569]" pageId="15" pageNumber="54">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from
|
||
<taxonomicName id="4C59F09C9B5AB461E66C646BFC8D2172" box="[770,869,545,569]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="54" phylum="Tardigrada" rank="species" species="derkai">
|
||
<emphasis id="B92D570D9B5AB461E66C646BFC8D2172" box="[770,869,545,569]" italics="true" pageId="15" pageNumber="54">P. derkai</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |