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<document id="242BB3B0C9FDA2204B66307761037F78" ID-DOI="10.5281/zenodo.188725" ID-GBIF-Dataset="97a0d3d1-80c3-450b-9404-255b5a20c7f5" ID-ISSN="1175-5326" ID-Zenodo-Dep="188725" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461096479258" checkinUser="plazi" docAuthor="Shear, William A., Taylor, Steven J., Wynne, Judson &amp; Krejca, Jean K." docDate="2009" docId="03DD87D0AB1ECC24FF2AF9D4FEF3FC34" docLanguage="en" docName="zt02151p065.pdf" docOrigin="Zootaxa 2151" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pratherodesmus despaini Shear, new species" docType="treatment" docUuid="390D8938-B824-467D-88FC-6AB62969AB82" docUuidSource="ZooBank" docVersion="10" lastPageNumber="58" masterDocId="FFE4FFA8AB17CC2FFFBDFFF5FFAEFFA4" masterDocTitle="Cave millipeds of the United States. VIII. New genera and species of polydesmidan millipeds from caves in the southwestern United States (Diplopoda, Polydesmida, Macrosternodesmidae)" masterLastPageNumber="65" masterPageNumber="47" pageNumber="56" updateTime="1698589473289" updateUser="plazi">
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<mods:title id="C3F8BB9F47859410640918D67CA3A068">Cave millipeds of the United States. VIII. New genera and species of polydesmidan millipeds from caves in the southwestern United States (Diplopoda, Polydesmida, Macrosternodesmidae)</mods:title>
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<mods:namePart id="B63B4814F0C0BD2B59719F0E1E73B4B9">Shear, William A.</mods:namePart>
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<mods:namePart id="49A8F355F6C85C034FEA81E6D56A14A0">Taylor, Steven J.</mods:namePart>
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<mods:namePart id="FC7A2379894B352AFB0E54016E787275">Wynne, Judson</mods:namePart>
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<mods:namePart id="350372E39B2A043F8091F17D0BF4F613">Krejca, Jean K.</mods:namePart>
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<treatment id="03DD87D0AB1ECC24FF2AF9D4FEF3FC34" ID-DOI="http://doi.org/10.5281/zenodo.6213340" ID-GBIF-Taxon="119620484" ID-Zenodo-Dep="6213340" LSID="urn:lsid:zoobank.org:act:390D8938-B824-467D-88FC-6AB62969AB82" httpUri="http://treatment.plazi.org/id/03DD87D0AB1ECC24FF2AF9D4FEF3FC34" lastPageId="11" lastPageNumber="58" pageId="9" pageNumber="56">
<subSubSection id="C36E654DAB1ECC26FF2AF9D4FD86F9F9" pageId="9" pageNumber="56" type="nomenclature">
<paragraph id="8BCB36C6AB1ECC26FF2AF9D4FD1AF99E" blockId="9.[151,692,1568,1629]" box="[151,692,1568,1595]" pageId="9" pageNumber="56">
<heading id="D08381AAAB1ECC26FF2AF9D4FD1AF99E" bold="true" box="[151,692,1568,1595]" fontSize="11" level="1" pageId="9" pageNumber="56" reason="1">
<emphasis id="B900EAD4AB1ECC26FF2AF9D4FD1AF99E" bold="true" box="[151,692,1568,1595]" pageId="9" pageNumber="56">
<taxonomicName id="4C744D45AB1ECC26FF2AF9D4FDB9F99E" authority="Shear" authorityName="Shear" box="[151,535,1568,1595]" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="despaini" status="sp. nov.">
<emphasis id="B900EAD4AB1ECC26FF2AF9D4FE66F99F" bold="true" box="[151,456,1569,1595]" italics="true" pageId="9" pageNumber="56">Pratherodesmus despaini</emphasis>
Shear
</taxonomicName>
,
<taxonomicNameLabel id="A23357AFAB1ECC26FD98F9D5FD1AF99E" box="[549,692,1568,1594]" pageId="9" pageNumber="56" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCB36C6AB1ECC26FF2AF9B6FD86F9F9" blockId="9.[151,692,1568,1629]" box="[151,552,1603,1629]" pageId="9" pageNumber="56">
<figureCitation id="134F2A43AB1ECC26FF2AF9B6FF49F9F9" box="[151,231,1603,1629]" captionStart="FIGURES 1, 2" captionStartId="4.[151,269,906,930]" captionTargetBox="[154,1436,197,890]" captionTargetId="figure@4.[154,1439,197,891]" captionTargetPageId="4" captionText="FIGURES 1, 2. Heads, collums and first two segments of males, dorsal view. Fig. 1. Pratherodesmus despaini. Fig. 2. P. voylesi." httpUri="https://zenodo.org/record/188726/files/figure.png" pageId="9" pageNumber="56">Figs. 1</figureCitation>
,
<figureCitation id="134F2A43AB1ECC26FF48F9B6FEABF9F9" box="[245,261,1603,1629]" captionStart="FIGURES 3 6" captionStartId="5.[151,270,1440,1464]" captionTargetBox="[151,1434,310,1416]" captionTargetId="figure@5.[151,1436,310,1417]" captionTargetPageId="5" captionText="FIGURES 3 6. Segments 10, dorsal view. Fig. 3. Nevadesmus ophimontis. Fig. 2. Pratherodesmus ecclesia. Fig. 3. P. despaini. Fig. 6. P. voylesi." httpUri="https://zenodo.org/record/188727/files/figure.png" pageId="9" pageNumber="56">5</figureCitation>
,
<figureCitation id="134F2A43AB1ECC26FEAFF9B6FE8CF9F9" box="[274,290,1603,1629]" captionStart="FIGURES 7 10" captionStartId="6.[151,270,1492,1516]" captionTargetBox="[153,1436,349,1459]" captionTargetId="figure@6.[153,1438,349,1461]" captionTargetPageId="6" captionText="FIGURES 7 10. Figs. 7 9. Pygidia, dorsal view. Fig. 7. Nevadesmus ophimontis. Fig. 8. Pratherodesmus despaini. Fig. 9. P. voylesi. Fig. 10. Metazonital setae anterior to ozopore of segment 10, Nevadesmus ophimontis." httpUri="https://zenodo.org/record/188728/files/figure.png" pageId="9" pageNumber="56">8</figureCitation>
, 2024, 32, 35, 36, 39
</paragraph>
</subSubSection>
<subSubSection id="C36E654DAB1ECC25FF2AF965FD37FD90" lastPageId="10" lastPageNumber="57" pageId="9" pageNumber="56" type="description">
<paragraph id="8BCB36C6AB1ECC26FF2AF965FEBCF8BB" blockId="9.[151,1437,1680,2016]" pageId="9" pageNumber="56">
<emphasis id="B900EAD4AB1ECC26FF2AF965FF43F90E" bold="true" box="[151,237,1680,1706]" pageId="9" pageNumber="56">
<typeStatus id="54CF8864AB1ECC26FF2AF965FF4CF90E" box="[151,226,1680,1706]" pageId="9" pageNumber="56">Types</typeStatus>
:
</emphasis>
Male
<typeStatus id="54CF8864AB1ECC26FEFDF964FE1FF90F" box="[320,433,1681,1707]" pageId="9" pageNumber="56" type="holotype">holotype</typeStatus>
, two male and one female
<typeStatus id="54CF8864AB1ECC26FCB8F964FCDFF90F" box="[773,881,1681,1707]" pageId="9" pageNumber="56" type="paratype">paratype</typeStatus>
from Kaweah Cave, Tulare Co., California, collected
<date id="FFCA1006AB1ECC26FEB4F94DFE1EF976" box="[265,432,1720,1746]" pageId="9" pageNumber="56" value="2004-04-28">28 April 2004</date>
by J. Krejca, P. Sprouse, S. Fryer, D. Ubick, P. Paquin and W. Savary; one male and two female
<typeStatus id="54CF8864AB1ECC26FE9CF92BFE3FF95C" box="[289,401,1758,1784]" pageId="9" pageNumber="56" type="paratype">paratypes</typeStatus>
from the same locality, collected
<date id="FFCA1006AB1ECC26FCA1F92BFC79F95C" box="[796,983,1758,1784]" pageId="9" pageNumber="56" value="2006-08-10">10 August 2006</date>
by J. Krejca and J. Despain, deposited in FMNH.
</paragraph>
<paragraph id="8BCB36C6AB1ECC25FF7BF8DEFA35FF5B" blockId="9.[151,1437,1680,2016]" lastBlockId="10.[151,1437,152,564]" lastPageId="10" lastPageNumber="57" pageId="9" pageNumber="56">
<emphasis id="B900EAD4AB1ECC26FF7BF8DEFECFF8E1" bold="true" box="[198,353,1835,1861]" pageId="9" pageNumber="56">Description:</emphasis>
<emphasis id="B900EAD4AB1ECC26FED5F8D9FE0AF8E1" box="[360,420,1836,1861]" italics="true" pageId="9" pageNumber="56">Male</emphasis>
. Length 9.0 mm, width 0.6 mm (
<figureCitation id="134F2A43AB1ECC26FC94F8DEFC2FF8E1" box="[809,897,1835,1861]" captionStart="FIGURE 39" captionStartId="17.[151,255,987,1011]" captionTargetBox="[151,1435,390,962]" captionTargetId="figure@17.[151,1436,390,963]" captionTargetPageId="17" captionText="FIGURE 39. Living example of Pratherodesmus despaini, photographed in Kaweah Cave (photo by J. Krejca)." httpUri="https://zenodo.org/record/188736/files/figure.png" pageId="9" pageNumber="56">Fig. 39</figureCitation>
). Head about 50% wider than collum (
<figureCitation id="134F2A43AB1ECC26FAF8F8DEFA24F8E1" box="[1349,1418,1835,1861]" captionStart="FIGURES 1, 2" captionStartId="4.[151,269,906,930]" captionTargetBox="[154,1436,197,890]" captionTargetId="figure@4.[154,1439,197,891]" captionTargetPageId="4" captionText="FIGURES 1, 2. Heads, collums and first two segments of males, dorsal view. Fig. 1. Pratherodesmus despaini. Fig. 2. P. voylesi." httpUri="https://zenodo.org/record/188726/files/figure.png" pageId="9" pageNumber="56">Fig. 1</figureCitation>
). Antennae long, extending back to posterior border of fifth segment. Collum with anterior margin arcuate, with 12 marginal setae, posterior margin straight to slightly sinuate, with eight setae, middle row with six setae; posteriolateral angles slightly produced (
<figureCitation id="134F2A43AB1ECC26FD3EF855FD64F81E" box="[643,714,1952,1978]" captionStart="FIGURES 1, 2" captionStartId="4.[151,269,906,930]" captionTargetBox="[154,1436,197,890]" captionTargetId="figure@4.[154,1439,197,891]" captionTargetPageId="4" captionText="FIGURES 1, 2. Heads, collums and first two segments of males, dorsal view. Fig. 1. Pratherodesmus despaini. Fig. 2. P. voylesi." httpUri="https://zenodo.org/record/188726/files/figure.png" pageId="9" pageNumber="56">Fig. 1</figureCitation>
). Typical midbody segment (
<figureCitation id="134F2A43AB1ECC26FB93F855FBD6F81E" box="[1070,1144,1952,1978]" captionStart="FIGURES 3 6" captionStartId="5.[151,270,1440,1464]" captionTargetBox="[151,1434,310,1416]" captionTargetId="figure@5.[151,1436,310,1417]" captionTargetPageId="5" captionText="FIGURES 3 6. Segments 10, dorsal view. Fig. 3. Nevadesmus ophimontis. Fig. 2. Pratherodesmus ecclesia. Fig. 3. P. despaini. Fig. 6. P. voylesi." httpUri="https://zenodo.org/record/188727/files/figure.png" pageId="9" pageNumber="56">Fig. 5</figureCitation>
, segment 10) with three lateral marginal teeth, each subtended by a seta; anterior row of six setae very strongly procurved, lateralmost seta of row widely separated, not appearing to be part of row; middle row of six setae slightly recurved; posterior marginal row with six setae. Posteriolateral metazonital corners strongly produced beyond ozopores, ozopores subtended by usual three setae. Pygidium as described for
<taxonomicName id="4C744D45AB1DCC25FC08FF10FB8DFF5A" box="[949,1059,229,254]" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="voylesi">
<emphasis id="B900EAD4AB1DCC25FC08FF10FB8DFF5A" box="[949,1059,229,254]" italics="true" pageId="10" pageNumber="57">P. voylesi</emphasis>
</taxonomicName>
, but with 12 long setae (
<figureCitation id="134F2A43AB1DCC25FAF9FF10FA25FF5B" box="[1348,1419,229,255]" captionStart="FIGURES 7 10" captionStartId="6.[151,270,1492,1516]" captionTargetBox="[153,1436,349,1459]" captionTargetId="figure@6.[153,1438,349,1461]" captionTargetPageId="6" captionText="FIGURES 7 10. Figs. 7 9. Pygidia, dorsal view. Fig. 7. Nevadesmus ophimontis. Fig. 8. Pratherodesmus despaini. Fig. 9. P. voylesi. Fig. 10. Metazonital setae anterior to ozopore of segment 10, Nevadesmus ophimontis." httpUri="https://zenodo.org/record/188728/files/figure.png" pageId="10" pageNumber="57">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BCB36C6AB1DCC25FF7BFEF9FDA5FE43" blockId="10.[151,1437,152,564]" pageId="10" pageNumber="57">
Pregonopodal legs markedly encrassate (
<figureCitation id="134F2A43AB1DCC25FD09FEF9FD52FE82" box="[692,764,268,294]" captionStart="FIGURES 1, 2" captionStartId="4.[151,269,906,930]" captionTargetBox="[154,1436,197,890]" captionTargetId="figure@4.[154,1439,197,891]" captionTargetPageId="4" captionText="FIGURES 1, 2. Heads, collums and first two segments of males, dorsal view. Fig. 1. Pratherodesmus despaini. Fig. 2. P. voylesi." httpUri="https://zenodo.org/record/188726/files/figure.png" pageId="10" pageNumber="57">Fig. 1</figureCitation>
). Gonopods (Figs. 2024, 32, 35) with gonocoxae and gonostome as described for
<taxonomicName id="4C744D45AB1DCC25FE57FEC7FDF5FEEF" box="[490,603,306,331]" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="voylesi">
<emphasis id="B900EAD4AB1DCC25FE57FEC7FDF5FEEF" box="[490,603,306,331]" italics="true" pageId="10" pageNumber="57">P. voylesi</emphasis>
</taxonomicName>
. Prefemora strongly transverse, articulating process narrow, pointed; prefemoral stem not marked. Exomere present, small, slightly curved, acute. Endomerite large, trullate, apically with two small lobes set with prominent, regular, small warts. Solenomere longer than in other species, with fewer cuticular scales; subtending process short, acute. tibiotarsus not flattened, as acute process similar in length to solenomere.
</paragraph>
<paragraph id="8BCB36C6AB1DCC25FF7BFE01FD37FD90" blockId="10.[151,1437,152,564]" pageId="10" pageNumber="57">
<emphasis id="B900EAD4AB1DCC25FF7BFE01FEB0FDA9" box="[198,286,500,525]" italics="true" pageId="10" pageNumber="57">Female</emphasis>
: Length, 9.0 mm, width 0.70 mm. Nonsexual characters similar to male. Cyphopods not observed, not extruded in available female specimens.
</paragraph>
</subSubSection>
<caption id="DF0B664EAB1DCC25FF2AF88EFCB5F811" httpUri="https://zenodo.org/record/188731/files/figure.png" pageId="10" pageNumber="57" targetBox="[151,1434,626,1890]" targetPageId="10">
<paragraph id="8BCB36C6AB1DCC25FF2AF88EFCB5F811" blockId="10.[151,1437,1915,1973]" pageId="10" pageNumber="57">
<emphasis id="B900EAD4AB1DCC25FF2AF88EFECBF837" bold="true" box="[151,357,1915,1939]" pageId="10" pageNumber="57">FIGURES 2124.</emphasis>
Gonopods of
<taxonomicName id="4C744D45AB1DCC25FDB8F88EFC8EF836" box="[517,800,1915,1938]" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="despaini">
<emphasis id="B900EAD4AB1DCC25FDB8F88EFC8EF836" box="[517,800,1915,1938]" italics="true" pageId="10" pageNumber="57">Pratherodesmus despaini</emphasis>
</taxonomicName>
. Fig. 21. Gonopods in gonostome, ventral view. Fig. 22. Lateral view. Fig. 23. Anterior view. Fig. 24. Posterior view.
</paragraph>
</caption>
<subSubSection id="C36E654DAB1CCC24FF7BFF62FB81FD26" pageId="11" pageNumber="58" type="distribution">
<paragraph id="8BCB36C6AB1CCC24FF7BFF62FB81FD26" blockId="11.[151,1438,151,912]" pageId="11" pageNumber="58">
<emphasis id="B900EAD4AB1CCC24FF7BFF62FDA4FF15" bold="true" box="[198,522,151,177]" pageId="11" pageNumber="58">Distribution and habitat:</emphasis>
Known only from the
<typeStatus id="54CF8864AB1CCC24FC92FF6DFCCFFF16" box="[815,865,152,178]" pageId="11" pageNumber="58">type</typeStatus>
locality. Kaweah Cave is located close to the western boundary of Sequoia Kings Canyon National Park, and is unique among the caves in the park because it occurs at low elevation and has greater biodiversity than is found among the higher altitude caves (for locations of caves in the park, see the map in
<bibRefCitation id="EFE54B37AB1CCC24FD02FEF9FC63FE82" author="Shear" box="[703,973,268,294]" pageId="11" pageNumber="65" refString="Shear, W. A., &amp; Shelley, R. M. (2008) Cave millipeds of the United States. VI. Sequoiadesmus krejcae, n. gen., n. sp., from Sequoia and Kings Canyon National Parks, California, USA (Diplopoda, Polyesmida, Trichopolydesmidea, Macrosternodesmidae). Zootaxa, 1693, 41 - 48." type="journal article" year="2008">Shear &amp; Shelley 2008</bibRefCitation>
). Four troglobitic species, including
<taxonomicName id="4C744D45AB1CCC24FA3BFEF9FF55FEEF" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="despaini">
<emphasis id="B900EAD4AB1CCC24FA3BFEF9FF55FEEF" italics="true" pageId="11" pageNumber="58">P. despaini</emphasis>
</taxonomicName>
, and two other endemic troglophiles are to be found there. The other troglobites are an unidentified trichoniscid isopod, a cambalid millipede, and a pseudoscorpion, probably a species of
<taxonomicName id="4C744D45AB1CCC24FB12FEACFAF0FED6" box="[1199,1374,345,370]" class="Arachnida" family="Chernetidae" genus="Tuberochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAD4AB1CCC24FB12FEACFAF0FED6" box="[1199,1374,345,370]" italics="true" pageId="11" pageNumber="58">Tuberochernes</emphasis>
</taxonomicName>
. The endemic troglophiles include a harvestmen in the genus
<taxonomicName id="4C744D45AB1CCC24FC9BFE75FC25FE3D" box="[806,907,384,409]" class="Arachnida" family="Neopilionidae" genus="Calicina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAD4AB1CCC24FC9BFE75FC25FE3D" box="[806,907,384,409]" italics="true" pageId="11" pageNumber="58">Calicina</emphasis>
</taxonomicName>
and a spider in the genus
<taxonomicName id="4C744D45AB1CCC24FB05FE75FAA0FE3D" box="[1208,1294,384,409]" class="Arachnida" family="Telemidae" genus="Usofila" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAD4AB1CCC24FB05FE75FAA0FE3D" box="[1208,1294,384,409]" italics="true" pageId="11" pageNumber="58">Usofila</emphasis>
</taxonomicName>
. These may all represent new unnamed species. The caves accessible portion is a sinuous crawl less than
<quantity id="4C8C9B23AB1CCC24FAB7FE53FAF9FE64" box="[1290,1367,422,448]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="58" unit="m" value="100.0">100 m</quantity>
long, ending in a large room where the animals were collected. One male was collected from a root, and the other specimens were associated with unidentified guano (probably rodent or bat) on the floor. The temperature was not measured on the day of collections, but during an earlier visit to the same room of the cave the temperature was recorded at 11 degrees Celsius. The surrounding vegetation is California lower montane blue oak-foothill pine woodland and savanna (SWReGAP land cover
<typeStatus id="54CF8864AB1CCC24FD40FD9DFC9DFD26" box="[765,819,616,642]" pageId="11" pageNumber="58">type</typeStatus>
; Lowery
<emphasis id="B900EAD4AB1CCC24FC1CFD9DFC72FD25" box="[929,988,616,641]" italics="true" pageId="11" pageNumber="58">et al.</emphasis>
2006).
</paragraph>
</subSubSection>
<subSubSection id="C36E654DAB1CCC24FF7BFD7BFEF3FC34" pageId="11" pageNumber="58" type="etymology">
<paragraph id="8BCB36C6AB1CCC24FF7BFD7BFEC6FD52" blockId="11.[151,1438,151,912]" pageId="11" pageNumber="58">
<emphasis id="B900EAD4AB1CCC24FF7BFD7BFEF9FD0C" bold="true" box="[198,343,654,680]" pageId="11" pageNumber="58">Etymology:</emphasis>
Named for Joel Despain, Cave Management Specialist at Sequoia and Kings Canyon National Parks, who organized and performed much of the sampling that resulted in the discovery of many new cave species.
</paragraph>
<paragraph id="8BCB36C6AB1CCC24FF7BFCF7FEF3FC34" blockId="11.[151,1438,151,912]" pageId="11" pageNumber="58">
<emphasis id="B900EAD4AB1CCC24FF7BFCF7FEB8FCB8" bold="true" box="[198,278,770,796]" pageId="11" pageNumber="58">Notes:</emphasis>
This species is more distant from the closely related
<taxonomicName id="4C744D45AB1CCC24FC3FFCF7FC52FCBF" box="[898,1020,770,795]" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="ecclesia">
<emphasis id="B900EAD4AB1CCC24FC3FFCF7FC52FCBF" box="[898,1020,770,795]" italics="true" pageId="11" pageNumber="58">P. ecclesia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D45AB1CCC24FB89FCF7FB0CFCBF" box="[1076,1186,770,795]" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="voylesi">
<emphasis id="B900EAD4AB1CCC24FB89FCF7FB0CFCBF" box="[1076,1186,770,795]" italics="true" pageId="11" pageNumber="58">P. voylesi</emphasis>
</taxonomicName>
. While the gonopods are built along the same basic plan, the presence of a small exomere in
<taxonomicName id="4C744D45AB1CCC24FC4FFCDCFBDBFCE6" box="[1010,1141,809,834]" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="despaini">
<emphasis id="B900EAD4AB1CCC24FC4FFCDCFBDBFCE6" box="[1010,1141,809,834]" italics="true" pageId="11" pageNumber="58">P. despaini</emphasis>
</taxonomicName>
suggests, in comparison with
<taxonomicName id="4C744D45AB1CCC24FF66FCA5FED0FCCD" box="[219,382,848,873]" class="Diplopoda" family="Macrosternodesmidae" genus="Nevadesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAD4AB1CCC24FF66FCA5FED0FCCD" box="[219,382,848,873]" italics="true" pageId="11" pageNumber="58">Nevadesmus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D45AB1CCC24FE7CFCA5FD95FCCD" box="[449,571,848,873]" class="Diplopoda" genus="Tidesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAD4AB1CCC24FE7CFCA5FD95FCCD" box="[449,571,848,873]" italics="true" pageId="11" pageNumber="58">Tidesmus</emphasis>
</taxonomicName>
, that it would occupy a more basal phylogenetic position within
<taxonomicName id="4C744D45AB1CCC24FF2AFC82FEF8FC34" box="[151,342,887,912]" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAD4AB1CCC24FF2AFC82FEF8FC34" box="[151,342,887,912]" italics="true" pageId="11" pageNumber="58">Pratherodesmus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>