350 lines
45 KiB
XML
350 lines
45 KiB
XML
<document id="9783130CCE8B51EE8AF6D12493687D01" ID-CLB-Dataset="8140" ID-DOI="10.2478/aemnp-2019-0021" ID-GBIF-Dataset="726682f3-9eac-4961-ac4d-d8b8679e8211" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488950" ID-ZooBank="ABBA2B4F-8B60-41E2-B80B-45861F974B23" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219567173" checkinUser="felipe" docAuthor="Ribera, Ignacio, Hernando, Carles & Cieslak, Alexandra" docDate="2019" docId="03DD87A0FFD6FFD7FC2EFB22DAFCFD8B" docLanguage="en" docName="ActaEntMusNatPra.59.1.253-272.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Hydraena (Hydraena) naja Ribera & Hernando & Cieslak 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="259" masterDocId="FFE4FFD8FFD3FFD1FF86FF93D922FFFE" masterDocTitle="Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae" masterLastPageNumber="272" masterPageNumber="253" pageNumber="258" updateTime="1698926098198" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:mods id="E572FBD1491FBCD4595FC1E9C0559D1D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="54C7C865067046A390B8D927D9C33A0E">
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<mods:title id="88B6DBA2850447D8CCBF1D42732AF1C9">Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae</mods:title>
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</mods:titleInfo>
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<mods:name id="F3C84F3AE4171C3C5E1AAB00E505B1B1" type="personal">
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<mods:role id="A5FA85166EFD82D1F4B6F87086706BAD">
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<mods:roleTerm id="C9EAEA90B3624D4AA5863B4CC5CC2C8C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="72A1B158C7FF3DA43BEAF0BE45988964">Ribera, Ignacio</mods:namePart>
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<mods:affiliation id="42C737FDC28C70B2E2129EA0CFF19DEC">Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37, E- 08003 Barcelona, Spain;</mods:affiliation>
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</mods:name>
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<mods:name id="05465A5B9D2AFCA46F10089C088C61EE" type="personal">
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<mods:role id="86A2E87DF4807AEE14843DD54204F05F">
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<mods:roleTerm id="DDA410A119320D11B90815F2C0D376AD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D9049F73E8930C1507C8EEB1035BE754">Hernando, Carles</mods:namePart>
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<mods:affiliation id="773597AFC19BFAE020F1BD645BC936DC">P. O. box 118, E- 08911 Badalona, Catalonia, Spain; e-mail: leptotyphlus @ gmail. com</mods:affiliation>
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</mods:name>
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<mods:name id="2494C8C3B48A8398E1DED1FA45BBDEB4" type="personal">
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<mods:role id="FB7FBA19BCCB40C1EF30FB7466F29D99">
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<mods:roleTerm id="6925771D6DF1D2D4063817EC1788F189">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="691A863CFDED4B6C9E647C89A3569362">Cieslak, Alexandra</mods:namePart>
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<mods:affiliation id="00EF9B5DFFCC475D8E8B0C0F339850A8">Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37, E- 08003 Barcelona, Spain;</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="C467649673AB29343D5644D974FAFC25">text</mods:typeOfResource>
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<mods:titleInfo id="26FFB8C596BB7ED566DE69395B679ACB">
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<mods:title id="C0F6E467C4B02FF41B6F3864C9CE03FE">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="C2D1FE2AF5409A9534B0D9D2E04D3348">
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<mods:date id="79B2287F2CD6E9BE808B3646880B46E2">2019</mods:date>
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<mods:detail id="2DF5912C97FFBE9BD22F7FB3A9319434" type="pubDate">
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<mods:number id="E38C8FE5E957BF47E989092AB758C0D1">2019-06-24</mods:number>
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<mods:detail id="8C3A96275E65FB9094212E52993170E2" type="volume">
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<mods:number id="1585776EB34B9BA103B7FF8296E429E5">59</mods:number>
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</mods:detail>
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<mods:number id="DFBDEE82363CCF03F71D5D575632C042">1</mods:number>
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<mods:start id="BF223A83A825DB728882F7E2FFFBEAD9">253</mods:start>
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<mods:classification id="50483B5495C2012EF9DC20810A935430">journal article</mods:classification>
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<mods:identifier id="9CF0BFE4A1E29366BAA7E2C30CE99A45" type="CLB-Dataset">8140</mods:identifier>
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<mods:identifier id="76844202A02D03E82E3680151AEB862C" type="DOI">10.2478/aemnp-2019-0021</mods:identifier>
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<mods:identifier id="673462CD7C05CE4B1B91D07E8B888F42" type="GBIF-Dataset">726682f3-9eac-4961-ac4d-d8b8679e8211</mods:identifier>
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</mods:mods>
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<treatment id="03DD87A0FFD6FFD7FC2EFB22DAFCFD8B" ID-DOI="http://doi.org/10.5281/zenodo.4549006" ID-GBIF-Taxon="177423638" ID-Zenodo-Dep="4549006" LSID="urn:lsid:plazi:treatment:03DD87A0FFD6FFD7FC2EFB22DAFCFD8B" httpUri="http://treatment.plazi.org/id/03DD87A0FFD6FFD7FC2EFB22DAFCFD8B" lastPageId="6" lastPageNumber="259" pageId="5" pageNumber="258">
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<subSubSection id="C36E653DFFD6FFD4FC2EFB22DC12FB36" box="[936,1328,1201,1224]" pageId="5" pageNumber="258" type="nomenclature">
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<paragraph id="8BCB36B6FFD6FFD4FC2EFB22DC12FB36" blockId="5.[936,1328,1201,1224]" box="[936,1328,1201,1224]" pageId="5" pageNumber="258">
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<heading id="D08381DAFFD6FFD4FC2EFB22DC12FB36" box="[936,1328,1201,1224]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="258" reason="3">
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<emphasis id="B900EAA4FFD6FFD4FC2EFB22DC12FB36" bold="true" box="[936,1328,1201,1224]" pageId="5" pageNumber="258">
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<taxonomicName id="4C744D35FFD6FFD4FC2EFB22DDF6FB36" ID-CoL="8Y4N3" authority="Ribera & Hernando & Cieslak, 2019" authorityName="Ribera & Hernando & Cieslak" authorityYear="2019" box="[936,1236,1201,1224]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="naja" status="sp. nov." subGenus="Hydraena">
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<emphasis id="B900EAA4FFD6FFD4FC2EFB22DD34FB36" bold="true" box="[936,1046,1201,1224]" italics="true" pageId="5" pageNumber="258">Hydraena</emphasis>
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||
(
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<emphasis id="B900EAA4FFD6FFD4FBA3FB22DDB1FB36" bold="true" box="[1061,1171,1201,1224]" italics="true" pageId="5" pageNumber="258">Hydraena</emphasis>
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)
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<emphasis id="B900EAA4FFD6FFD4FB25FB22DDF6FB36" bold="true" box="[1187,1236,1201,1224]" italics="true" pageId="5" pageNumber="258">naja</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23357DFFFD6FFD4FB5CFB22DC12FB36" box="[1242,1328,1201,1224]" pageId="5" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E653DFFD6FFD4FBB3FB5CDD81FB1C" box="[1077,1187,1231,1250]" pageId="5" pageNumber="258" type="description">
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<paragraph id="8BCB36B6FFD6FFD4FBB3FB5CDD81FB1C" blockId="5.[1077,1187,1231,1250]" box="[1077,1187,1231,1250]" pageId="5" pageNumber="258">
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(
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<figureCitation id="134F2A33FFD6FFD4FBBDFB5CDD5FFB1C" box="[1083,1149,1231,1250]" captionStart="Figs 14–17" captionStartId="8.[151,188,2019,2038]" captionTargetBox="[179,1369,223,1980]" captionTargetId="figure-13@8.[151,1436,189,1999]" captionTargetPageId="8" captionText="Figs 14–17. Habitus of new species: 14 – Hydraena naja sp. nov.; 15 – Ochthebius alhajarensis sp. nov.; 16 – Ochthebius bernard sp. nov.; 17 – Agraphydrus elongatus sp. nov. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4488958" httpUri="https://zenodo.org/record/4488958/files/figure.png" pageId="5" pageNumber="258">Figs 14</figureCitation>
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,
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<figureCitation id="134F2A33FFD6FFD4FB00FB5CDDBEFB1C" box="[1158,1180,1231,1250]" captionStart="Figs 18–21" captionStartId="9.[151,188,2012,2031]" captionTargetBox="[257,1306,228,1943]" captionTargetId="figure-12@9.[224,1364,189,1988]" captionTargetPageId="9" captionText="Figs 18–21. Aedeagus of new species: 18 – Hydraena naja sp. nov.; 19 – Ochthebius alhajarensis sp. nov.; 20 – Ochthebius bernard sp. nov.; 21 – Agraphydrus elongatus sp. nov. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4488962" httpUri="https://zenodo.org/record/4488962/files/figure.png" pageId="5" pageNumber="258">18</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36E653DFFD6FFD4FCBAFB6ADA9CFAD8" pageId="5" pageNumber="258" type="materials_examined">
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||
<paragraph id="8BCB36B6FFD6FFD4FCBAFB6ADA9CFAD8" blockId="5.[828,1437,1273,1638]" pageId="5" pageNumber="258">
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||
<emphasis id="B900EAA4FFD6FFD4FCBAFB6ADA97FAF2" bold="true" box="[828,949,1273,1292]" pageId="5" pageNumber="258">
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||
<typeStatus id="54CF8814FFD6FFD4FCBAFB6ADA4BFAF2" box="[828,873,1273,1292]" pageId="5" pageNumber="258">Type</typeStatus>
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locality.
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</emphasis>
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Source of wadi Bani Awf in Jebel Al-Hajar,
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<collectingCountry id="F3637626FFD6FFD4FAB3FB6ADC4BFAF2" box="[1333,1385,1273,1292]" name="Oman" pageId="5" pageNumber="258">Oman</collectingCountry>
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(Loc. 4;
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<figureCitation id="134F2A33FFD6FFD4FCD4FA80DAA5FAD8" box="[850,903,1299,1318]" captionStart="Fig" captionStartId="1.[1088,1120,2029,2048]" captionTargetBox="[152,1071,1387,2079]" captionTargetId="figure-691@1.[152,1074,1387,2079]" captionTargetPageId="1" captionText="Fig. 1. Sampled localities (see Table 1 for details)." figureDoi="http://doi.org/10.5281/zenodo.4488952" httpUri="https://zenodo.org/record/4488952/files/figure.png" pageId="5" pageNumber="258">Figs 1</figureCitation>
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,
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<figureCitation id="134F2A33FFD6FFD4FC17FA80DA91FAD8" box="[913,947,1299,1318]" captionStart="Figs 2–7" captionStartId="2.[151,188,2009,2028]" captionTargetBox="[151,1436,546,1992]" captionTargetId="figure-13@2.[151,1436,546,1992]" captionTargetPageId="2" captionText="Figs 2–7. Some of the sampled localities in north Oman. 2 – Loc. No. 1: Ghul spring with pools; 3 – No. 2: residual pools in wadi in Bahla city; 4 – Loc. No. 3: residual pools in wadi; 5 – Loc. No. 4: source of wadi Bani Awf; 6 – Loc. No. 4: source of wadi Bani Awf; 7 – Loc. No. 5: wadi Bani Awf. See Table 1 for details on the localities, and Fig. 1 for their geographical location." figureDoi="http://doi.org/10.5281/zenodo.4488954" httpUri="https://zenodo.org/record/4488954/files/figure.png" pageId="5" pageNumber="258">5, 6</figureCitation>
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||
).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36E653DFFD6FFD4FCBAFABDDC09F998" pageId="5" pageNumber="258" type="materials_examined">
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||
<paragraph id="8BCB36B6FFD6FFD4FCBAFABDDC09F998" blockId="5.[828,1437,1273,1638]" pageId="5" pageNumber="258">
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<emphasis id="B900EAA4FFD6FFD4FCBAFABDDA9DFABF" bold="true" box="[828,959,1326,1345]" pageId="5" pageNumber="258">Type material.</emphasis>
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||
<materialsCitation id="3B1C3CEBFFD6FFD4FC44FABDDDFEFA6F" ID-GBIF-Occurrence="3038225306" collectingDate="2010-04-06" collectionCode="NHMW" collectorName="J. Al-Akhdar & Ribera, Cieslak & Hernando" country="Oman" elevation="1300" latitude="23.176722" location="wadi Bani Awf" longLatPrecision="1" longitude="57.409473" pageId="5" pageNumber="258" specimenCount="2" typeStatus="holotype">
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||
<typeStatus id="54CF8814FFD6FFD4FC44FABDDD34FAC1" box="[962,1046,1326,1345]" pageId="5" pageNumber="258" type="holotype">HOLOTYPE</typeStatus>
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: ♁ (
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||
<collectionCode id="ED65AE73FFD6FFD4FBBFFABDDDA1FABF" box="[1081,1155,1326,1345]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="258" type="Museum">NHMW</collectionCode>
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), “4
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||
<collectingCountry id="F3637626FFD6FFD4FB2FFABDDDFFFABF" box="[1193,1245,1326,1345]" name="Oman" pageId="5" pageNumber="258">Oman</collectingCountry>
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<date id="FFCA1076FFD6FFD4FB67FABDDC0EFABF" box="[1249,1324,1326,1345]" pageId="5" pageNumber="258" value="2010-04-06">
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<collectingDate id="EF8EE99EFFD6FFD4FB67FABDDC0EFABF" box="[1249,1324,1326,1345]" pageId="5" pageNumber="258" value="2010-04-06">6.4.2010</collectingDate>
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||
</date>
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||
<collectorName id="26815360FFD6FFD4FAA9FABDDCBEFABF" box="[1327,1436,1326,1345]" pageId="5" pageNumber="258">J.Al-Akhdar</collectorName>
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||
// source of
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<location id="8EAB606DFFD6FFD4FC27FADADD01FAA2" LSID="urn:lsid:plazi:treatment:03DD87A0FFD6FFD7FC2EFB22DAFCFD8B:8EAB606DFFD6FFD4FC27FADADD01FAA2" box="[929,1059,1353,1372]" country="Oman" latitude="23.176722" longLatPrecision="1" longitude="57.409473" name="wadi Bani Awf" pageId="5" pageNumber="258">wadi Bani Awf</location>
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||
, on rock //
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||
<geoCoordinate id="EE405071FFD6FFD4FB05FADADDCCFAA2" box="[1155,1262,1353,1372]" degrees="23" direction="north" minutes="10" orientation="latitude" pageId="5" pageNumber="258" precision="1" seconds="36.2" value="23.176722">N23 10 36.2</geoCoordinate>
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<geoCoordinate id="EE405071FFD6FFD4FB75FADADC79FAA2" box="[1267,1371,1353,1372]" degrees="57" direction="east" minutes="24" orientation="longitude" pageId="5" pageNumber="258" precision="1" seconds="34.1" value="57.409473">E57 24 34.1</geoCoordinate>
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<quantity id="4C8C9B53FFD6FFD4FAE7FADADCBEFAA2" box="[1377,1436,1353,1372]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="258" unit="m" value="1300.0">
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<elevation id="0059D185FFD6FFD4FAE7FADADCBEFAA2" box="[1377,1436,1353,1372]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="258" unit="m" value="1300.0">1300m</elevation>
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||
</quantity>
|
||
//
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||
<collectorName id="26815360FFD6FFD4FCCAFAF0DD1EFA88" box="[844,1084,1379,1398]" pageId="5" pageNumber="258">Ribera, Cieslak & Hernando</collectorName>
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||
leg.”, aedeagus dissected and mounted in DMHF on a transparent card, with holotype label
|
||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B1C3CEBFFD6FFD4FB65FAEDDDE9F9CF" ID-GBIF-Occurrence="3038225303" pageId="5" pageNumber="258" typeStatus="paratype">
|
||
<typeStatus id="54CF8814FFD6FFD4FB65FAEDDC19FA71" box="[1251,1339,1406,1425]" pageId="5" pageNumber="258" type="paratype">PARATYPES</typeStatus>
|
||
(
|
||
<specimenCount id="9D72FD3FFFD6FFD4FAC2FAEDDCB5FA6F" box="[1348,1431,1406,1425]" pageId="5" pageNumber="258" type="generic">210 spec.</specimenCount>
|
||
) (
|
||
<collectionCode id="ED65AE73FFD6FFD4FCC4FA0ADA5DFA52" box="[834,895,1433,1452]" pageId="5" pageNumber="258">CCHB</collectionCode>
|
||
,
|
||
<collectionCode id="ED65AE73FFD6FFD4FC0EFA0ADA99FA52" box="[904,955,1433,1452]" pageId="5" pageNumber="258">IBEB</collectionCode>
|
||
,
|
||
<collectionCode id="ED65AE73FFD6FFD4FC45FA0ADD27FA52" box="[963,1029,1433,1452]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="258" type="Museum">MNCN</collectionCode>
|
||
,
|
||
<collectionCode id="ED65AE73FFD6FFD4FB88FA0ADD76FA52" box="[1038,1108,1433,1452]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="258" type="Museum">NHMW</collectionCode>
|
||
,
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||
<collectionCode id="ED65AE73FFD6FFD4FBDBFA0ADDBCFA52" box="[1117,1182,1433,1452]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="258">NMPC</collectionCode>
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||
): 15 ♁♁
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<specimenCount id="9D72FD3FFFD6FFD4FB6DFA0ADC07FA52" box="[1259,1317,1429,1452]" pageId="5" pageNumber="258" type="female">13 ♀♀</specimenCount>
|
||
, same data as holotype, with
|
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<typeStatus id="54CF8814FFD6FFD4FC31FA20DADDFA38" box="[951,1023,1459,1478]" pageId="5" pageNumber="258" type="paratype">paratype</typeStatus>
|
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labels;
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<specimenCount id="9D72FD3FFFD6FFD4FBB8FA20DDADFA38" box="[1086,1167,1459,1478]" pageId="5" pageNumber="258" type="generic">181 spec.</specimenCount>
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||
, “3
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<collectingCountry id="F3637626FFD6FFD4FB2AFA20DDC2FA38" box="[1196,1248,1459,1478]" name="Oman" pageId="5" pageNumber="258">Oman</collectingCountry>
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||
<date id="FFCA1076FFD6FFD4FB62FA20DC0FFA38" box="[1252,1325,1459,1478]" pageId="5" pageNumber="258" value="2010-04-06">6.4.2010</date>
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J.Al-Akhdar // rd. Tanuf-Hat,residual pools in wadi //
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<geoCoordinate id="EE405071FFD6FFD4FB09FA5DDDD7FA1F" box="[1167,1269,1486,1505]" degrees="23" direction="north" minutes="05" orientation="latitude" pageId="5" pageNumber="258" precision="1" seconds="36.2" value="23.09339">N23 05 36.2</geoCoordinate>
|
||
<geoCoordinate id="EE405071FFD6FFD4FB7FFA5DDC7FFA1F" box="[1273,1373,1486,1505]" degrees="57" direction="east" minutes="25" orientation="longitude" pageId="5" pageNumber="258" precision="1" seconds="56.6" value="57.43239">E57 25 56.6</geoCoordinate>
|
||
<quantity id="4C8C9B53FFD6FFD4FAE7FA5DDCBEFA1F" box="[1377,1436,1486,1505]" metricMagnitude="3" metricUnit="m" metricValue="1.307" pageId="5" pageNumber="258" unit="m" value="1307.0">1307m</quantity>
|
||
// Ribera, Cieslak & Hernando leg.” (
|
||
<specimenCount id="9D72FD3FFFD6FFD4FB07FA7ADD9FFA02" box="[1153,1213,1513,1532]" pageId="5" pageNumber="258" type="generic">1 spec.</specimenCount>
|
||
used for DNA extraction, voucher number IBE-RA97, sequences published in
|
||
<bibRefCitation id="EFE54B47FFD6FFD4FB71F990DCBEF9E8" author="TRIZZINO" box="[1271,1436,1539,1558]" etAl="et al." firstAuthor="TRIZZINO" pageId="5" pageNumber="258" pagination="192 - 208" refId="ref18053" refString="TRIZZINO M., JACH M. A., AUDISIO P., ALONSO R. & RIBERA I. 2013: A molecular phylogeny of the cosmopolitan hiperdiverse genus Hydraena Kugelann (Coleoptera, Hydraenidae). Systematic Entomology 38: 192 - 208." type="journal article" year="2013">TRIZZINO et al. 2013</bibRefCitation>
|
||
as “
|
||
<taxonomicName id="4C744D35FFD6FFD4FCDCF98DDA8CF9CF" box="[858,942,1566,1585]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="genus">Hydraena</taxonomicName>
|
||
sp.
|
||
<collectingCountry id="F3637626FFD6FFD4FC49F98DDD26F9CF" box="[975,1028,1566,1585]" name="Oman" pageId="5" pageNumber="258">OMA</collectingCountry>
|
||
”), with
|
||
<typeStatus id="54CF8814FFD6FFD4FBC3F98DDDACF9CF" box="[1093,1166,1566,1585]" pageId="5" pageNumber="258" type="paratype">paratype</typeStatus>
|
||
labels;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1C3CEBFFD6FFD4FB48F98DDC09F998" ID-GBIF-Occurrence="3038225305" pageId="5" pageNumber="258" typeStatus="paratype">
|
||
1 ♁, “12
|
||
<collectingCountry id="F3637626FFD6FFD4FA9FF98DDC6FF9CF" box="[1305,1357,1566,1585]" name="Oman" pageId="5" pageNumber="258">Oman</collectingCountry>
|
||
<date id="FFCA1076FFD6FFD4FAD7F98DDCBEF9CF" box="[1361,1436,1566,1585]" pageId="5" pageNumber="258" value="2010-04-09">9.4.2010</date>
|
||
<quantity id="4C8C9B53FFD6FFD4FCBAF9AADA44F9B2" box="[828,870,1593,1612]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="258" unit="km" value="1.0">1 km</quantity>
|
||
W Qalhat // residual pools in wadi //
|
||
<geoCoordinate id="EE405071FFD6FFD4FB19F9AADC25F9B2" box="[1183,1287,1593,1612]" degrees="22" direction="north" minutes="41" orientation="latitude" pageId="5" pageNumber="258" precision="1" seconds="25.4" value="22.690388">N22 41 25.4</geoCoordinate>
|
||
<geoCoordinate id="EE405071FFD6FFD4FA8AF9AADC53F9B2" box="[1292,1393,1593,1612]" degrees="59" direction="east" minutes="22" orientation="longitude" pageId="5" pageNumber="258" precision="1" seconds="03.0" value="59.367496">E59 22 03.0</geoCoordinate>
|
||
<quantity id="4C8C9B53FFD6FFD4FAF0F9AADCBEF9B2" box="[1398,1436,1593,1612]" metricMagnitude="1" metricUnit="m" metricValue="8.8" pageId="5" pageNumber="258" unit="m" value="88.0">88m</quantity>
|
||
// Ribera, Cieslak & Hernando leg.”, with
|
||
<typeStatus id="54CF8814FFD6FFD4FB22F9C0DDCFF998" box="[1188,1261,1619,1638]" pageId="5" pageNumber="258" type="paratype">paratype</typeStatus>
|
||
labels.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD6FFD7FCBAF9ECD897FCCB" lastPageId="6" lastPageNumber="259" pageId="5" pageNumber="258" type="description">
|
||
<paragraph id="8BCB36B6FFD6FFD4FCBAF9ECDDF3F948" blockId="5.[828,1437,1663,2070]" pageId="5" pageNumber="258">
|
||
<emphasis id="B900EAA4FFD6FFD4FCBAF9ECDAE5F968" bold="true" box="[828,967,1663,1686]" pageId="5" pageNumber="258">Description.</emphasis>
|
||
Habitus of male as in
|
||
<figureCitation id="134F2A33FFD6FFD4FB3CF9ECDC2BF968" box="[1210,1289,1663,1686]" captionStart="Figs 14–17" captionStartId="8.[151,188,2019,2038]" captionTargetBox="[179,1369,223,1980]" captionTargetId="figure-13@8.[151,1436,189,1999]" captionTargetPageId="8" captionText="Figs 14–17. Habitus of new species: 14 – Hydraena naja sp. nov.; 15 – Ochthebius alhajarensis sp. nov.; 16 – Ochthebius bernard sp. nov.; 17 – Agraphydrus elongatus sp. nov. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4488958" httpUri="https://zenodo.org/record/4488958/files/figure.png" pageId="5" pageNumber="258">Fig. 14</figureCitation>
|
||
; body length:
|
||
<quantity id="4C8C9B53FFD6FFD4FCBAF90CDAFBF948" box="[828,985,1695,1718]" metricMagnitude="-3" metricUnit="m" metricValue="2.525" metricValueMax="2.75" metricValueMin="2.3" pageId="5" pageNumber="258" unit="mm" value="2.525" valueMax="2.75" valueMin="2.3">2.30–2.75 mm</quantity>
|
||
, width:
|
||
<quantity id="4C8C9B53FFD6FFD4FBA9F90CDDEFF948" box="[1071,1229,1695,1718]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="5" pageNumber="258" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.90–1.10 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCB36B6FFD6FFD4FCDCF92CDD94F848" blockId="5.[828,1437,1663,2070]" pageId="5" pageNumber="258">Elytra, legs, palpi and antennae reddish brown; pronotum darker except for anterior margin, head almost black. Apex of maxillary palpi dark brown. Anterior margin of labrum deeply excised, with coarse punctures. Clypeus entirely densely punctate, matt. Central area of frons less densely punctate, with fine sparse pubescence between punctures; lateral parts of frons (ocular groove) deeply impressed, densely micropunctate.</paragraph>
|
||
<paragraph id="8BCB36B6FFD6FFD7FCDCF82CD806FF0B" blockId="5.[828,1437,1663,2070]" lastBlockId="6.[151,760,190,2069]" lastPageId="6" lastPageNumber="259" pageId="5" pageNumber="258">Pronotum distinctly cordiform, anterior margin concave. Surface very densely punctate, interstices densely micropunctate. Median longitudinal impression and oblique posterior admedian grooves shallow. Sublateral groove deep.</paragraph>
|
||
<paragraph id="8BCB36B6FFD5FFD7FF33FF6DD84CFDEB" blockId="6.[151,760,190,2069]" pageId="6" pageNumber="259">Elytra elongate, subparallel-sided, with about 15 rows of punctures, with nine to ten rows between suture and shoulder; rows usually very regular, except for an admedian area in anterior third and apical area; punctures small, very densely arranged with small recumbent whitish setae; intervals narrow, very slightly convex, glabrous. Explanate margin of elytra well developed, not reaching elytral apex, weakly serrate; elytral apices more or less separately rounded.</paragraph>
|
||
<paragraph id="8BCB36B6FFD5FFD7FF33FD8DD829FD8B" blockId="6.[151,760,190,2069]" pageId="6" pageNumber="259">Male meso- and metatibiae with apical expansions on ventral side, larger on metatibiae. Female with unmodified tibiae.</paragraph>
|
||
<paragraph id="8BCB36B6FFD5FFD7FF33FDEDD857FCEB" blockId="6.[151,760,190,2069]" pageId="6" pageNumber="259">Ventral surface covered with a very dense short pubescence, forming a plastron except for the last abdominal ventrites and two longitudinal glabrous areas on metaventrite; abdominal ventrites with a dense fringe of setae on posterior margin.</paragraph>
|
||
<paragraph id="8BCB36B6FFD5FFD7FF33FC8DD897FCCB" blockId="6.[151,760,190,2069]" box="[181,437,798,821]" pageId="6" pageNumber="259">
|
||
Aedeagus as in
|
||
<figureCitation id="134F2A33FFD5FFD7FEE7FC8DD890FCCB" box="[353,434,798,821]" captionStart="Figs 18–21" captionStartId="9.[151,188,2012,2031]" captionTargetBox="[257,1306,228,1943]" captionTargetId="figure-12@9.[224,1364,189,1988]" captionTargetPageId="9" captionText="Figs 18–21. Aedeagus of new species: 18 – Hydraena naja sp. nov.; 19 – Ochthebius alhajarensis sp. nov.; 20 – Ochthebius bernard sp. nov.; 21 – Agraphydrus elongatus sp. nov. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4488962" httpUri="https://zenodo.org/record/4488962/files/figure.png" pageId="6" pageNumber="259">Fig. 18</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD5FFD7FF11FCADD88BFAAB" pageId="6" pageNumber="259" type="diagnosis">
|
||
<paragraph id="8BCB36B6FFD5FFD7FF11FCADD88BFAAB" blockId="6.[151,760,190,2069]" pageId="6" pageNumber="259">
|
||
<emphasis id="B900EAA4FFD5FFD7FF11FCADD8B9FCAB" bold="true" box="[151,411,830,853]" pageId="6" pageNumber="259">Differential diagnosis.</emphasis>
|
||
The species seems to be most closely related to
|
||
<taxonomicName id="4C744D35FFD5FFD7FEF6FCCDDB97FC8B" authority="Ferro, 1984" authorityName="Ferro" authorityYear="1984" box="[368,693,862,885]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="verstraeteni">
|
||
<emphasis id="B900EAA4FFD5FFD7FEF6FCCDDB02FC8B" box="[368,544,862,885]" italics="true" pageId="6" pageNumber="259">H. verstraeteni</emphasis>
|
||
<bibRefCitation id="EFE54B47FFD5FFD7FDAAFCCDDB97FC8B" author="Ferro" box="[556,693,862,885]" firstAuthor="Ferro" pageId="6" pageNumber="259" pagination="61 - 71" refId="ref15964" refString="FERRO G. 1984: Hydraenidae nuovi ed interessanti del Museo di Storia Naturale di Praga. III contributo (Coleoptera Hydraenidae). Bulletin et Annales de la SocieteRoyale Belge d'Entomologie 120: 61 - 71." type="journal article" year="1984">Ferro, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
from south
|
||
<collectingCountry id="F3637626FFD5FFD7FF5FFCEDD827FC6B" box="[217,261,894,917]" name="Iran" pageId="6" pageNumber="259">Iran</collectingCountry>
|
||
(
|
||
<collectingRegion id="49B0F854FFD5FFD7FE94FCEDD8B4FC6B" box="[274,406,894,917]" country="Iran" name="Hormozgan" pageId="6" pageNumber="259">Hormozgan</collectingRegion>
|
||
) (
|
||
<bibRefCitation id="EFE54B47FFD5FFD7FE2AFCEDDB0CFC6B" author="FERRO" box="[428,558,894,917]" firstAuthor="FERRO" pageId="6" pageNumber="259" pagination="61 - 71" refId="ref15964" refString="FERRO G. 1984: Hydraenidae nuovi ed interessanti del Museo di Storia Naturale di Praga. III contributo (Coleoptera Hydraenidae). Bulletin et Annales de la SocieteRoyale Belge d'Entomologie 120: 61 - 71." type="journal article" year="1984">FERRO 1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE54B47FFD5FFD7FDBFFCEDDB89FC6B" author="JACH" box="[569,683,894,917]" firstAuthor="JACH" pageId="6" pageNumber="259" pagination="77 - 125" refId="ref16713" refString="JACH M. A. 1992: New and little known Palearctic species of the genus Hydraena (s. l.) Kugelann (Coleoptera: Hydraenidae). Koleopterologische Rundschau 62: 77 - 125." type="journal article" year="1992">JÄCH 1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE54B47FFD5FFD7FD30FCEDD80FFC4B" author="SKALE & JACH" firstAuthor="SKALE" pageId="6" pageNumber="259" pagination="185 - 216" refId="ref17942" refString="SKALE A. & JACH M. A. 2011: Updating the Hydraena fauna of Iran, with descriptions of eight new species (Insecta: Coleoptera: Hydraenidae). Vernate 30: 185 - 216." type="journal article" year="2011">SKALE & JÄCH 2011</bibRefCitation>
|
||
), based on the external morphology and the male genitalia. Based on the studied material both species cannot be separated on its external morphology, and only differ in the shape of the median lobe and associated appendages of the male genitalia (see fig.
|
||
<quantity id="4C8C9B53FFD5FFD7FD5BFB8DD98EFBAB" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" pageId="6" pageNumber="259" unit="in" value="41.0">41 in</quantity>
|
||
<bibRefCitation id="EFE54B47FFD5FFD7FF33FBADD809FBAB" author="JACH" box="[181,299,1086,1109]" firstAuthor="JACH" pageId="6" pageNumber="259" pagination="77 - 125" refId="ref16713" refString="JACH M. A. 1992: New and little known Palearctic species of the genus Hydraena (s. l.) Kugelann (Coleoptera: Hydraenidae). Koleopterologische Rundschau 62: 77 - 125." type="journal article" year="1992">JÄCH 1992</bibRefCitation>
|
||
). Both species differ 5.3% in their COI-5 gene, based on a specimen of
|
||
<taxonomicName id="4C744D35FFD5FFD7FE5FFBCDDB59FB8B" authorityName="Ferro" authorityYear="1984" box="[473,635,1118,1141]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="verstraeteni">
|
||
<emphasis id="B900EAA4FFD5FFD7FE5FFBCDD8CEFB8B" box="[473,492,1118,1141]" italics="true" pageId="6" pageNumber="259">H</emphasis>
|
||
.
|
||
<emphasis id="B900EAA4FFD5FFD7FE7FFBCDDB59FB8B" box="[505,635,1118,1141]" italics="true" pageId="6" pageNumber="259">verstraeteni</emphasis>
|
||
</taxonomicName>
|
||
from south
|
||
<collectingCountry id="F3637626FFD5FFD7FF11FBEDD9E1FB6B" box="[151,195,1150,1173]" name="Iran" pageId="6" pageNumber="259">Iran</collectingCountry>
|
||
(voucher IBE-AN461, prov.
|
||
<collectingRegion id="49B0F854FFD5FFD7FD85FBEDDB5BFB6B" box="[515,633,1150,1173]" country="Iran" name="Khuzestan" pageId="6" pageNumber="259">Khuzestan</collectingRegion>
|
||
, Behbahan, Garmabeh river,
|
||
<date id="FFCA1076FFD5FFD7FECAFB0DD88AFB4B" box="[332,424,1182,1205]" pageId="6" pageNumber="259" value="2011-05-05">5.v.2011</date>
|
||
E. Irani leg.). Phylogenetically both species are included in a clade with
|
||
<taxonomicName id="4C744D35FFD5FFD7FDCDFB2DD82CFB0B" authority="Janssens, 1981" authorityName="Janssens" authorityYear="1981" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="persica">
|
||
<emphasis id="B900EAA4FFD5FFD7FDCDFB2DDB98FB2B" box="[587,698,1214,1237]" italics="true" pageId="6" pageNumber="259">H. persica</emphasis>
|
||
Janssens, 1981
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D35FFD5FFD7FE9DFB4DDB57FB0B" authority="Jach & Skale, 2009" authorityName="Jach & Skale" authorityYear="2009" box="[283,629,1246,1269]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="dochula">
|
||
<emphasis id="B900EAA4FFD5FFD7FE9DFB4DD8B4FB0B" box="[283,406,1246,1269]" italics="true" pageId="6" pageNumber="259">H. dochula</emphasis>
|
||
Jäch & Skale, 2009
|
||
</taxonomicName>
|
||
and related species, in turn related with the species of the
|
||
<taxonomicName id="4C744D35FFD5FFD7FDF8FB6DDBD3FAEB" authorityName="Reitter" authorityYear="1885" box="[638,753,1278,1301]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="grandis">
|
||
<emphasis id="B900EAA4FFD5FFD7FDF8FB6DDBD3FAEB" box="[638,753,1278,1301]" italics="true" pageId="6" pageNumber="259">H. grandis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D35FFD5FFD7FF11FA8DD820FACB" authorityName="Curtis" authorityYear="1830" box="[151,258,1310,1333]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="rufipes">
|
||
<emphasis id="B900EAA4FFD5FFD7FF11FA8DD820FACB" box="[151,258,1310,1333]" italics="true" pageId="6" pageNumber="259">H. rufipes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D35FFD5FFD7FEBEFA8DD89DFACB" box="[312,447,1310,1333]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="pulchella">
|
||
<emphasis id="B900EAA4FFD5FFD7FEBEFA8DD89DFACB" box="[312,447,1310,1333]" italics="true" pageId="6" pageNumber="259">H. pulchella</emphasis>
|
||
</taxonomicName>
|
||
groups (
|
||
<bibRefCitation id="EFE54B47FFD5FFD7FDA6FA8DDBDAFACB" author="TRIZZINO" box="[544,760,1310,1333]" etAl="et al." firstAuthor="TRIZZINO" pageId="6" pageNumber="259" pagination="192 - 208" refId="ref18053" refString="TRIZZINO M., JACH M. A., AUDISIO P., ALONSO R. & RIBERA I. 2013: A molecular phylogeny of the cosmopolitan hiperdiverse genus Hydraena Kugelann (Coleoptera, Hydraenidae). Systematic Entomology 38: 192 - 208." type="journal article" year="2013">TRIZZINO et al. 2013</bibRefCitation>
|
||
and unpublished results).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD5FFD7FF11FACDDC0DFDEB" pageId="6" pageNumber="259" type="etymology">
|
||
<paragraph id="8BCB36B6FFD5FFD7FF11FACDD8B2FA0B" blockId="6.[151,760,190,2069]" pageId="6" pageNumber="259">
|
||
<emphasis id="B900EAA4FFD5FFD7FF11FACDD838FA8B" bold="true" box="[151,282,1374,1397]" pageId="6" pageNumber="259">Etymology.</emphasis>
|
||
Named after the elapid snake genus
|
||
<emphasis id="B900EAA4FFD5FFD7FD45FACDDBDAFA8B" box="[707,760,1374,1397]" italics="true" pageId="6" pageNumber="259">Naja</emphasis>
|
||
(cobras), in reference to the shape of the enlarged setae of the aedeagus (
|
||
<figureCitation id="134F2A33FFD5FFD7FEABFA0DD859FA4B" box="[301,379,1438,1461]" captionStart="Figs 18–21" captionStartId="9.[151,188,2012,2031]" captionTargetBox="[257,1306,228,1943]" captionTargetId="figure-12@9.[224,1364,189,1988]" captionTargetPageId="9" captionText="Figs 18–21. Aedeagus of new species: 18 – Hydraena naja sp. nov.; 19 – Ochthebius alhajarensis sp. nov.; 20 – Ochthebius bernard sp. nov.; 21 – Agraphydrus elongatus sp. nov. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4488962" httpUri="https://zenodo.org/record/4488962/files/figure.png" pageId="6" pageNumber="259">Fig. 18</figureCitation>
|
||
), resembling a cobra in its characteristic threatening position. Noun in singular nominative, standing in apposition.
|
||
</paragraph>
|
||
<paragraph id="8BCB36B6FFD5FFD7FF11FA6DDC0DFDEB" blockId="6.[151,760,190,2069]" lastBlockId="6.[828,1436,190,629]" pageId="6" pageNumber="259">
|
||
<emphasis id="B900EAA4FFD5FFD7FF11FA6DD8AAF9EB" bold="true" box="[151,392,1534,1557]" pageId="6" pageNumber="259">Notes on the habitat.</emphasis>
|
||
The species was most commonly found in two localities, residual pools in a wadi (locality No. 3) and the source of wadi Bani Awf (locality No. 4). The residual pool (
|
||
<figureCitation id="134F2A33FFD5FFD7FEEBF9CDD893F98B" box="[365,433,1630,1653]" captionStart="Figs 2–7" captionStartId="2.[151,188,2009,2028]" captionTargetBox="[151,1436,546,1992]" captionTargetId="figure-13@2.[151,1436,546,1992]" captionTargetPageId="2" captionText="Figs 2–7. Some of the sampled localities in north Oman. 2 – Loc. No. 1: Ghul spring with pools; 3 – No. 2: residual pools in wadi in Bahla city; 4 – Loc. No. 3: residual pools in wadi; 5 – Loc. No. 4: source of wadi Bani Awf; 6 – Loc. No. 4: source of wadi Bani Awf; 7 – Loc. No. 5: wadi Bani Awf. See Table 1 for details on the localities, and Fig. 1 for their geographical location." figureDoi="http://doi.org/10.5281/zenodo.4488954" httpUri="https://zenodo.org/record/4488954/files/figure.png" pageId="6" pageNumber="259">Fig. 4</figureCitation>
|
||
) was the result of disruption by road works of a (by then) dry wadi, producing a pool with a diameter of ca.
|
||
<quantity id="4C8C9B53FFD5FFD7FE00F90DD8EBF94B" box="[390,457,1694,1717]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="6" pageNumber="259" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">3–4 m</quantity>
|
||
with muddy and sandy substratum and very turbid water. The species was here very abundant, together with
|
||
<taxonomicName id="4C744D35FFD5FFD7FE1AF94DDB51F90B" authority="Ribera & Hernando & Cieslak, 2019" authorityName="Ribera & Hernando & Cieslak" authorityYear="2019" box="[412,627,1758,1781]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="bernard" status="sp. nov.">
|
||
<emphasis id="B900EAA4FFD5FFD7FE1AF94DDB51F90B" box="[412,627,1758,1781]" italics="true" pageId="6" pageNumber="259">Ochthebius bernard</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357DFFFD5FFD7FDFFF94DDBEEF90B" box="[633,716,1758,1781]" pageId="6" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C744D35FFD5FFD7FF11F96DDB1CF8EB" authority="Wollaston, 1864" authorityName="Wollaston" authorityYear="1864" box="[151,574,1790,1813]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="quadricollis">
|
||
<emphasis id="B900EAA4FFD5FFD7FF11F96DD8A9F8EB" box="[151,395,1790,1813]" italics="true" pageId="6" pageNumber="259">Hydraena quadricollis</emphasis>
|
||
<bibRefCitation id="EFE54B47FFD5FFD7FE09F96DDB1CF8EB" author="Wollaston" box="[399,574,1790,1813]" firstAuthor="Wollaston" pageId="6" pageNumber="259" refId="ref18306" refString="WOLLASTON T. V. 1864: Catalogue of the coleopterous insects of the Canaries in the collection of the British Museum. London, British Museum, xiii + 648 pp." type="book" year="1864">Wollaston, 1864</bibRefCitation>
|
||
</taxonomicName>
|
||
(see below), plus other species in lower numbers. The source of wadi Bani Awf (
|
||
<figureCitation id="134F2A33FFD5FFD7FF54F8ADD813F8AB" box="[210,305,1854,1877]" captionStart="Figs 2–7" captionStartId="2.[151,188,2009,2028]" captionTargetBox="[151,1436,546,1992]" captionTargetId="figure-13@2.[151,1436,546,1992]" captionTargetPageId="2" captionText="Figs 2–7. Some of the sampled localities in north Oman. 2 – Loc. No. 1: Ghul spring with pools; 3 – No. 2: residual pools in wadi in Bahla city; 4 – Loc. No. 3: residual pools in wadi; 5 – Loc. No. 4: source of wadi Bani Awf; 6 – Loc. No. 4: source of wadi Bani Awf; 7 – Loc. No. 5: wadi Bani Awf. See Table 1 for details on the localities, and Fig. 1 for their geographical location." figureDoi="http://doi.org/10.5281/zenodo.4488954" httpUri="https://zenodo.org/record/4488954/files/figure.png" pageId="6" pageNumber="259">Figs 5, 6</figureCitation>
|
||
) is a spring on a rocky surface, apparently at least partially artificially excavated to form a small tunnel through which the water is diverted to an artificial open channel that goes down the valley. When the locality was visited (6
|
||
<superScript id="7C019BFEFFD5FFD7FEABF82FD81BF834" attach="left" box="[301,313,1980,1994]" fontSize="6" pageId="6" pageNumber="259">th</superScript>
|
||
April) all the water was diverted through the channel, but occasionally the water should overflow it and run through a natural rocky bed, that at the time of our visit was completely dry. The distance between the spring and the channel was at most ca.
|
||
<quantity id="4C8C9B53FFD5FFD7FA95FF4DDC6EFF0B" box="[1299,1356,222,245]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="259" unit="m" value="10.0">10 m</quantity>
|
||
, with a rocky substratum with a thin layer of sand and gravel, with some green filamentous algae. Despite the reduced dimensions of the habitat the diversity was remarkably high, with six species of
|
||
<taxonomicName id="4C744D35FFD5FFD7FBBDFECDDDE7FE8B" authorityName="Mulsant" authorityYear="1844" box="[1083,1221,350,373]" class="Insecta" family="Hydraenidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="family">Hydraenidae</taxonomicName>
|
||
, all of them endemic to
|
||
<collectingCountry id="F3637626FFD5FFD7FCD0FEEDDABAFE6B" box="[854,920,382,405]" name="Oman" pageId="6" pageNumber="259">Oman</collectingCountry>
|
||
and the
|
||
<collectingCountry id="F3637626FFD5FFD7FC76FEEDDD05FE6B" box="[1008,1063,382,405]" name="United Arab Emirates" pageId="6" pageNumber="259">UAE</collectingCountry>
|
||
and three of them newly described here (
|
||
<taxonomicName id="4C744D35FFD5FFD7FCFEFE0DDD3AFE4B" authority="Ribera & Hernando & Cieslak, 2019" authorityName="Ribera & Hernando & Cieslak" authorityYear="2019" box="[888,1048,414,437]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="naja" status="sp. nov.">
|
||
<emphasis id="B900EAA4FFD5FFD7FCFEFE0DDD3AFE4B" box="[888,1048,414,437]" italics="true" pageId="6" pageNumber="259">Hydraena naja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357DFFFD5FFD7FB9BFE0DDD52FE4B" box="[1053,1136,414,437]" pageId="6" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C744D35FFD5FFD7FBFAFE0DDC27FE4B" authorityName="Jach & Delgado" authorityYear="2010" box="[1148,1285,414,437]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="gattolliati">
|
||
<emphasis id="B900EAA4FFD5FFD7FBFAFE0DDC27FE4B" box="[1148,1285,414,437]" italics="true" pageId="6" pageNumber="259">H. gattolliati</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D35FFD5FFD7FA97FE0DDCB7FE4B" authorityName="Jach & Diaz" authorityYear="2000" box="[1297,1429,414,437]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="putearius">
|
||
<emphasis id="B900EAA4FFD5FFD7FA97FE0DDCB7FE4B" box="[1297,1429,414,437]" italics="true" pageId="6" pageNumber="259">H. putearius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D35FFD5FFD7FCBAFE2DDD30FE2B" authority="Ribera & Hernando & Cieslak, 2019" authorityName="Ribera & Hernando & Cieslak" authorityYear="2019" box="[828,1042,446,469]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="bernard" status="sp. nov.">
|
||
<emphasis id="B900EAA4FFD5FFD7FCBAFE2DDD30FE2B" box="[828,1042,446,469]" italics="true" pageId="6" pageNumber="259">Ochthebius bernard</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357DFFFD5FFD7FB9EFE2DDD4EFE2B" box="[1048,1132,446,469]" pageId="6" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C744D35FFD5FFD7FBFEFE2DDC3FFE2B" authority="Ribera & Hernando & Cieslak, 2019" authorityName="Ribera & Hernando & Cieslak" authorityYear="2019" box="[1144,1309,446,469]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="alhajarensis" status="sp. nov.">
|
||
<emphasis id="B900EAA4FFD5FFD7FBFEFE2DDC3FFE2B" box="[1144,1309,446,469]" italics="true" pageId="6" pageNumber="259">O. alhajarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357DFFFD5FFD7FAA4FE2DDC54FE2B" box="[1314,1398,446,469]" pageId="6" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C744D35FFD5FFD7FA04FE2DDAB9FE0B" authorityName="Jach & Delgado" authorityYear="2010" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="wurayah">
|
||
<emphasis id="B900EAA4FFD5FFD7FA04FE2DDAB9FE0B" italics="true" pageId="6" pageNumber="259">O. wurayah</emphasis>
|
||
</taxonomicName>
|
||
), in addition to
|
||
<taxonomicName id="4C744D35FFD5FFD7FBC0FE4DDC24FE0B" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[1094,1286,478,501]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="gestroi">
|
||
<emphasis id="B900EAA4FFD5FFD7FBC0FE4DDC24FE0B" box="[1094,1286,478,501]" italics="true" pageId="6" pageNumber="259">Copelatus gestroi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C744D35FFD5FFD7FA95FE4DDAE2FDEB" baseAuthorityName="Regimbart" baseAuthorityYear="1897" class="Insecta" family="Dytiscidae" genus="Nebrioporus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="mascatensis">
|
||
<emphasis id="B900EAA4FFD5FFD7FA95FE4DDAE2FDEB" italics="true" pageId="6" pageNumber="259">Nebrioporus mascatensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D35FFD5FFD7FC73FE6DDC0AFDEB" authorityName="Hajek & Wewalka" authorityYear="2009" box="[1013,1320,510,533]" class="Insecta" family="Dytiscidae" genus="Hydroglyphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="sinuspersicus">
|
||
<emphasis id="B900EAA4FFD5FFD7FC73FE6DDC0AFDEB" box="[1013,1320,510,533]" italics="true" pageId="6" pageNumber="259">Hydroglyphus sinuspersicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD5FFD7FCBAFD8DDAFCFD8B" pageId="6" pageNumber="259" type="distribution">
|
||
<paragraph id="8BCB36B6FFD5FFD7FCBAFD8DDAFCFD8B" blockId="6.[828,1436,190,629]" pageId="6" pageNumber="259">
|
||
<emphasis id="B900EAA4FFD5FFD7FCBAFD8DDAF3FDCB" bold="true" box="[828,977,542,565]" pageId="6" pageNumber="259">Distribution.</emphasis>
|
||
Found in the Al Hajar mountains, with an isolated specimen in a wadi in the coast of the Gulf of
|
||
<collectingCountry id="F3637626FFD5FFD7FCBAFDCDDA5CFD8B" box="[828,894,606,629]" name="Oman" pageId="6" pageNumber="259">Oman</collectingCountry>
|
||
(
|
||
<figureCitation id="134F2A33FFD5FFD7FC0BFDCDDAEFFD8B" box="[909,973,606,629]" captionStart="Fig" captionStartId="1.[1088,1120,2029,2048]" captionTargetBox="[152,1071,1387,2079]" captionTargetId="figure-691@1.[152,1074,1387,2079]" captionTargetPageId="1" captionText="Fig. 1. Sampled localities (see Table 1 for details)." figureDoi="http://doi.org/10.5281/zenodo.4488952" httpUri="https://zenodo.org/record/4488952/files/figure.png" pageId="6" pageNumber="259">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |