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<document id="23F26A18483ED34250C85A0D5B977E1B" ID-CLB-Dataset="46164" ID-DOI="10.11646/zootaxa.3780.2.4" ID-GBIF-Dataset="3391e352-6b3a-4b69-9be8-ee1e301ed70b" ID-ISSN="1175-5326" ID-Zenodo-Dep="227958" ID-ZooBank="AD3F93D5-BC85-4DA5-9957-6EA5673A8B42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460400159163" checkinUser="plazi" docAuthor="Huemer, Peter &amp; Timossi, Giovanni" docDate="2014" docId="03BC4C4BFFB6EE3162D7F8D31A0E21F8" docLanguage="en" docName="zt03780p296.pdf" docOrigin="Zootaxa 3780 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sattleria haemusi Huemer, sp. nov." docType="treatment" docVersion="9" lastPageNumber="293" masterDocId="FF853433FFBFEE3A6240FFCB1844260C" masterDocTitle="Sattleria revisited: unexpected cryptic diversity on the Balkan Peninsula and in the south-eastern Alps (Lepidoptera: Gelechiidae)" masterLastPageNumber="296" masterPageNumber="282" pageNumber="291" updateTime="1698299150435" updateUser="ExternalLinkService">
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<mods:title id="080610F82B75989A7E70E09770F47548">Sattleria revisited: unexpected cryptic diversity on the Balkan Peninsula and in the south-eastern Alps (Lepidoptera: Gelechiidae)</mods:title>
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<mods:namePart id="58EBB1094391F183B83B8DFF81A6F365">Huemer, Peter</mods:namePart>
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<mods:namePart id="554E0F255EC329E938B73494E9FB1A7F">Timossi, Giovanni</mods:namePart>
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<mods:date id="143DE47135C8101A6869B34E3864ADC6">2014</mods:date>
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<treatment id="03BC4C4BFFB6EE3162D7F8D31A0E21F8" ID-DOI="http://doi.org/10.5281/zenodo.6139563" ID-GBIF-Taxon="119419407" ID-Zenodo-Dep="6139563" LSID="urn:lsid:plazi:treatment:03BC4C4BFFB6EE3162D7F8D31A0E21F8" httpUri="http://treatment.plazi.org/id/03BC4C4BFFB6EE3162D7F8D31A0E21F8" lastPageId="11" lastPageNumber="293" pageId="9" pageNumber="291">
<subSubSection id="C30FAED6FFB6EE3362D7F8D31A06213E" box="[151,578,1816,1842]" pageId="9" pageNumber="291" type="nomenclature">
<paragraph id="8BAAFD5DFFB6EE3362D7F8D31A06213E" blockId="9.[151,578,1816,1842]" box="[151,578,1816,1842]" pageId="9" pageNumber="291">
<heading id="D0E24A31FFB6EE3362D7F8D31A06213E" bold="true" box="[151,578,1816,1842]" fontSize="11" level="1" pageId="9" pageNumber="291" reason="1">
<emphasis id="B961214FFFB6EE3362D7F8D31A06213E" bold="true" box="[151,578,1816,1842]" pageId="9" pageNumber="291">
<taxonomicName id="4C1586DEFFB6EE3362D7F8D3199D213E" ID-CoL="93KF6" authority="Huemer" authorityName="Huemer" box="[151,473,1816,1842]" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="291" phylum="Arthropoda" rank="species" species="haemusi" status="sp. nov.">
<emphasis id="B961214FFFB6EE3362D7F8D3192F213E" bold="true" box="[151,363,1816,1842]" italics="true" pageId="9" pageNumber="291">Sattleria haemusi</emphasis>
Huemer
</taxonomicName>
,
<taxonomicNameLabel id="A2529C34FFB6EE3363A5F8D31A06213E" box="[485,578,1816,1842]" pageId="9" pageNumber="291" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FAED6FFB6EE3362D7F8951B9D21EF" pageId="9" pageNumber="291" type="materials_examined">
<paragraph id="8BAAFD5DFFB6EE3362D7F8951DD22197" blockId="9.[151,1436,1886,2019]" pageId="9" pageNumber="291">
<emphasis id="B961214FFFB6EE3362D7F8951909217B" bold="true" box="[151,333,1886,1911]" pageId="9" pageNumber="291">
<typeStatus id="54AE43FFFFB6EE3362D7F8951896217B" box="[151,210,1886,1911]" pageId="9" pageNumber="291">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54AE43FFFFB6EE33631CF8951980217B" box="[348,452,1886,1911]" pageId="9" pageNumber="291" type="holotype">Holotype</typeStatus>
♂, SW
<collectingCountry id="F302BDCDFFB6EE336000F8951AE7217B" box="[576,675,1886,1911]" name="Bulgaria" pageId="9" pageNumber="291">Bulgaria</collectingCountry>
, Rila Mts. Granchar Circus
<quantity id="4CED50B8FFB6EE336645F8941C26217B" box="[1029,1122,1887,1911]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="291" unit="m" value="2200.0">2200 m</quantity>
<geoCoordinate id="EE219B9AFFB6EE336631F8951D44217B" box="[1137,1280,1886,1911]" direction="north" orientation="latitude" pageId="9" pageNumber="291" precision="15" value="42.12111">N42°07´16´´</geoCoordinate>
<geoCoordinate id="EE219B9AFFB6EE336750F8951DD8217B" box="[1296,1436,1886,1911]" direction="east" orientation="longitude" pageId="9" pageNumber="291" precision="15" value="23.591112">E23°35´28´´</geoCoordinate>
<date id="FFABDB9DFFB6EE3362D7F84819532197" box="[151,279,1923,1947]" pageId="9" pageNumber="291" value="2009-08-27">27.08.2009</date>
, at light leg. B. Zlatkov &amp; Y. Mutafchiev GU 13/1356 ♂ P. Huemer
<collectionCode id="ED046598FFB6EE33666DF8481C132197" box="[1069,1111,1923,1947]" pageId="9" pageNumber="291">BC</collectionCode>
<collectionCode id="ED046598FFB6EE33661DF8481CEC2196" box="[1117,1192,1923,1946]" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="9" pageNumber="291">TLMF</collectionCode>
Lep 08944 (
<collectionCode id="ED046598FFB6EE33677EF8481DCE2197" box="[1342,1418,1923,1947]" pageId="9" pageNumber="291">FBBZ</collectionCode>
).
</paragraph>
<paragraph id="8BAAFD5DFFB6EE336287F86D1B9D21EF" blockId="9.[151,1436,1886,2019]" pageId="9" pageNumber="291">
<emphasis id="B961214FFFB6EE336287F86D199421B3" bold="true" box="[199,464,1958,1983]" pageId="9" pageNumber="291">
<typeStatus id="54AE43FFFFB6EE336287F86D197221B3" box="[199,310,1958,1983]" pageId="9" pageNumber="291" type="paratype">Paratype</typeStatus>
.
<collectingCountry id="F302BDCDFFB6EE336302F86D198E21B3" box="[322,458,1958,1983]" name="Macedonia" pageId="9" pageNumber="291">Macedonia</collectingCountry>
:
</emphasis>
<specimenCount id="9D1336D4FFB6EE336397F86C1A4721B3" box="[471,515,1959,1983]" pageId="9" pageNumber="291" type="generic">1 ♂</specimenCount>
, Tetovo, Povoa Sapka, Felskar, W Tetovo,
<quantity id="4CED50B8FFB6EE3361AAF86C1C0621B3" box="[1002,1090,1959,1983]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="9" pageNumber="291" unit="m" value="2130.0">2130 m</quantity>
,
<date id="FFABDB9DFFB6EE33660EF86C1CF721B3" box="[1102,1203,1959,1983]" pageId="9" pageNumber="291" value="2012-08-07">7.8.2012</date>
, leg. C. Wieser, gen. slide GU 13/1360 ♂ P. Huemer
<collectionCode id="ED046598FFB6EE3363BCF8001A7D21EF" LSID="urn:lsid:biocol.org:col:15536" box="[508,569,1995,2019]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="9" pageNumber="291">DNA</collectionCode>
barcode id KML Lep 0 0 484 (LMK).
</paragraph>
</subSubSection>
<subSubSection id="C30FAED6FFB5EE306287FF5C1A412519" pageId="10" pageNumber="292" type="description">
<paragraph id="8BAAFD5DFFB5EE306287FF5C1D3E2784" blockId="10.[151,1436,151,789]" pageId="10" pageNumber="292">
<emphasis id="B961214FFFB5EE306287FF5C191D26BC" bold="true" box="[199,345,151,176]" pageId="10" pageNumber="292">Description.</emphasis>
Adult (
<figureCitation id="132EE1D8FFB5EE3063EDFF5C19B626BC" box="[429,498,151,176]" captionStart="FIGURES 1 4" captionStartId="3.[151,263,1902,1925]" captionTargetBox="[212,1381,1044,1867]" captionTargetId="figure@3.[187,1400,1014,1879]" captionTargetPageId="3" captionText="FIGURES 1 4. Adults of Sattleria. 1, S. sophiae, sp. nov., holotype; 2, S. dolomitica, sp. nov., holotype; 3, S. dinarica, sp. nov., holotype; 4, S. haemusi, sp. nov., holotype." httpUri="https://zenodo.org/record/227959/files/figure.png" pageId="10" pageNumber="292">Fig. 4</figureCitation>
). Male. Head brown with cream-coloured face; labial palpus cream, mottled brown; antenna dark brown, scapus and flagellum covered with cream scales on lower surface; thorax and tegula brown. Wingspan 16.017.0 mm; forewing light brown with extended dark brown and whitish mottling, whitish costal and tornal spots ill defined; costa from base to terminal area dark brown; fold with two oblique black-brown spots, separated by light brown, a dot-shaped black spot at 1/2 and an ill-defined angulated spot at 3/5; termen with some black scales, fringes concolorous with ground colour, fringe line present; hindwing light grey brown with concolorous fringes. Forelegs and middle legs brown with white scales, hindlegs white with long white bristles.
</paragraph>
<paragraph id="8BAAFD5DFFB5EE306287FE5F1D1F27A0" blockId="10.[151,1436,151,789]" box="[199,1371,404,429]" pageId="10" pageNumber="292">The colour and wing markings may differ slightly in fresh specimens as the type-material is rather worn.</paragraph>
<paragraph id="8BAAFD5DFFB5EE306287FE7C19D427DC" blockId="10.[151,1436,151,789]" box="[199,400,439,464]" pageId="10" pageNumber="292">Female unknown.</paragraph>
<paragraph id="8BAAFD5DFFB5EE306287FE171AEE24FC" blockId="10.[151,1436,151,789]" pageId="10" pageNumber="292">
Male genitalia (
<figureCitation id="132EE1D8FFB5EE30633FFE17199D27F8" box="[383,473,476,501]" captionStart="FIGURES 9 12" captionStartId="7.[151,263,1899,1922]" captionTargetBox="[265,1330,718,1868]" captionTargetId="figure@7.[250,1337,700,1878]" captionTargetPageId="7" captionText="FIGURES 9 12. Male genitalia of Sattleria. 9, S. dinarica, sp. nov., paratype, slide GEL 1129; 10, S. triglavica Povolný, paratype, slide Mus. Vind. 11202; 11, S. haemusi, sp. nov., holotype, slide GU 13 / 1356 P. Huemer; 12, S. dzieduszyckii (Nowicki), slide L. M. P. 97." httpUri="https://zenodo.org/record/227961/files/figure.png" pageId="10" pageNumber="292">Figs 11</figureCitation>
,
<figureCitation id="132EE1D8FFB5EE3063AAFE171A4D27F8" box="[490,521,476,500]" captionStart="FIGURES 17 20" captionStartId="9.[151,263,1662,1685]" captionTargetBox="[237,1353,186,1621]" captionTargetId="figure@9.[220,1366,171,1641]" captionTargetPageId="9" captionText="FIGURES 17 20. Male genitalia of Sattleria, details of vincular processes (seconardy process marked with arrow) and valva. 17, S. dinarica, sp. nov., paratype, slide GEL 1129; 18, S. triglavica Povolný, paratype, slide Mus. Vind. 11202; 19, S. haemusi, sp. nov., holotype, slide GU 13 / 1356 P. Huemer; 20, S. dzieduszyckii (Nowicki), slide L. M. P. 97." httpUri="https://zenodo.org/record/227963/files/figure.png" pageId="10" pageNumber="292">19</figureCitation>
). Uncus with rounded apex, culcitula moderately large, gnathos a large hook; tegumen anteriorly widened, broadly and deeply emarginated anterior margin; pedunculi long, slender; valva long, slender, extending almost to apex of uncus, distally slightly inflated with pointed tip and long apical setae; sacculus slender, hardly inflated, with acute apex; primary process of vinculum long, moderately broad, needle shaped, about level with sacculus; secondary process of vinculum fused with primary process, extending from base to about 2/5 of primary process, broadly arched, sub-oval with outer margins slightly serrate; saccus long, moderately slender; phallus moderately long and slender, straight, without medial projection, coecum weakly inflated with two minute basal sclerites, apex with small hooklet.
</paragraph>
<paragraph id="8BAAFD5DFFB5EE306287FD371A412519" blockId="10.[151,1436,151,789]" box="[199,517,764,789]" pageId="10" pageNumber="292">Female genitalia. Unknown.</paragraph>
</subSubSection>
<caption id="DF6AADD5FFB5EE3062D7F8951ACB21E3" httpUri="https://zenodo.org/record/227964/files/figure.png" pageId="10" pageNumber="292" targetBox="[172,1431,852,1858]" targetPageId="10">
<paragraph id="8BAAFD5DFFB5EE3062D7F8951ACB21E3" blockId="10.[151,1436,1886,2031]" pageId="10" pageNumber="292">
<emphasis id="B961214FFFB5EE3062D7F89519602178" bold="true" box="[151,292,1886,1908]" pageId="10" pageNumber="292">FIGURE 21.</emphasis>
Neighbour joining tree of
<taxonomicName id="4C1586DEFFB5EE306007F8951ADA2178" box="[583,670,1886,1908]" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="B961214FFFB5EE306007F8951ADA2178" box="[583,670,1886,1908]" italics="true" pageId="10" pageNumber="292">Sattleria</emphasis>
</taxonomicName>
from central and south-eastern Europe (Kimura 2 parameter, built with MEGA 5; cf. Tamura
<emphasis id="B961214FFFB5EE30633CF8B519F5219F" box="[380,433,1917,1939]" italics="true" pageId="10" pageNumber="292">et al.</emphasis>
2011) based on sequences of the mtDNA COI gene (barcode fragment 5, 658 bp), except for
<taxonomicName id="4C1586DEFFB5EE3067C9F8B5195221BE" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="292" phylum="Arthropoda" rank="species" species="basistrigella">
<emphasis id="B961214FFFB5EE3067C9F8B5195221BE" italics="true" pageId="10" pageNumber="292">S. basistrigella</emphasis>
</taxonomicName>
(599 bp, 1 specimen),
<taxonomicName id="4C1586DEFFB5EE3063BCF8571ADE21BE" box="[508,666,1948,1970]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="292" phylum="Arthropoda" rank="species" species="dzieduszyckii">
<emphasis id="B961214FFFB5EE3063BCF8571ADE21BE" box="[508,666,1948,1970]" italics="true" pageId="10" pageNumber="292">S. dzieduszyckii</emphasis>
</taxonomicName>
(623 bp, 1 specimen),
<taxonomicName id="4C1586DEFFB5EE3061C3F8571BBA21BE" box="[899,1022,1948,1970]" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="292" phylum="Arthropoda" rank="species" species="triglavica">
<emphasis id="B961214FFFB5EE3061C3F8571BBA21BE" box="[899,1022,1948,1970]" italics="true" pageId="10" pageNumber="292">S. triglavica</emphasis>
</taxonomicName>
(540 bp, 1 specimen). Width of triangles represent sample size, depth the molecular variation within the cluster. Source: DNA Barcode data from BOLD (Barcode of Life Database, cf. Ratnasingham &amp; Hebert 2007).
</paragraph>
</caption>
<subSubSection id="C30FAED6FFB4EE316287FF5C1AB027EB" pageId="11" pageNumber="293" type="diagnosis">
<paragraph id="8BAAFD5DFFB4EE316287FF5C1D4A2751" blockId="11.[151,1436,151,662]" pageId="11" pageNumber="293">
<emphasis id="B961214FFFB4EE316287FF5C190526BC" bold="true" box="[199,321,151,176]" pageId="11" pageNumber="293">Diagnosis.</emphasis>
<taxonomicName id="4C1586DEFFB4EE31630AFF531A5426BC" box="[330,528,152,176]" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="293" phylum="Arthropoda" rank="species" species="haemusi">
<emphasis id="B961214FFFB4EE31630AFF531A5426BC" box="[330,528,152,176]" italics="true" pageId="11" pageNumber="293">Sattleria haemusi</emphasis>
</taxonomicName>
is externally very similar to other strictly allopatric species of the genus with a divided basal streak of the forewing, i.e.,
<taxonomicName id="4C1586DEFFB4EE316032FF701B5A26DF" box="[626,798,187,211]" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="293" phylum="Arthropoda" rank="species" species="melaleucella">
<emphasis id="B961214FFFB4EE316032FF701B5A26DF" box="[626,798,187,211]" italics="true" pageId="11" pageNumber="293">S. melaleucella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1586DEFFB4EE316118FF701C4D26DF" box="[856,1033,187,211]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="293" phylum="Arthropoda" rank="species" species="dzieduszyckii">
<emphasis id="B961214FFFB4EE316118FF701C4D26DF" box="[856,1033,187,211]" italics="true" pageId="11" pageNumber="293">S. dzieduszyckii</emphasis>
</taxonomicName>
, but is smaller than the former and with more distinct black-brown markings on the forewings compared to the latter. Dissection of genitalia is necessary for safe identification. The male genitalia closely match those of
<taxonomicName id="4C1586DEFFB4EE3161B7FECB1C362714" box="[1015,1138,256,280]" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="293" phylum="Arthropoda" rank="species" species="dinarica">
<emphasis id="B961214FFFB4EE3161B7FECB1C362714" box="[1015,1138,256,280]" italics="true" pageId="11" pageNumber="293">S. dinarica</emphasis>
</taxonomicName>
(
<figureCitation id="132EE1D8FFB4EE3166C4FF341C8A2714" box="[1156,1230,255,280]" captionStart="FIGURES 9 12" captionStartId="7.[151,263,1899,1922]" captionTargetBox="[265,1330,718,1868]" captionTargetId="figure@7.[250,1337,700,1878]" captionTargetPageId="7" captionText="FIGURES 9 12. Male genitalia of Sattleria. 9, S. dinarica, sp. nov., paratype, slide GEL 1129; 10, S. triglavica Povolný, paratype, slide Mus. Vind. 11202; 11, S. haemusi, sp. nov., holotype, slide GU 13 / 1356 P. Huemer; 12, S. dzieduszyckii (Nowicki), slide L. M. P. 97." httpUri="https://zenodo.org/record/227961/files/figure.png" pageId="11" pageNumber="293">Figs 9</figureCitation>
,
<figureCitation id="132EE1D8FFB4EE31669CFECB1CBE2714" box="[1244,1274,256,280]" captionStart="FIGURES 17 20" captionStartId="9.[151,263,1662,1685]" captionTargetBox="[237,1353,186,1621]" captionTargetId="figure@9.[220,1366,171,1641]" captionTargetPageId="9" captionText="FIGURES 17 20. Male genitalia of Sattleria, details of vincular processes (seconardy process marked with arrow) and valva. 17, S. dinarica, sp. nov., paratype, slide GEL 1129; 18, S. triglavica Povolný, paratype, slide Mus. Vind. 11202; 19, S. haemusi, sp. nov., holotype, slide GU 13 / 1356 P. Huemer; 20, S. dzieduszyckii (Nowicki), slide L. M. P. 97." httpUri="https://zenodo.org/record/227963/files/figure.png" pageId="11" pageNumber="293">17</figureCitation>
),
<taxonomicName id="4C1586DEFFB4EE316751FECB1DD82714" box="[1297,1436,256,280]" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="293" phylum="Arthropoda" rank="species" species="triglavica">
<emphasis id="B961214FFFB4EE316751FECB1DD82714" box="[1297,1436,256,280]" italics="true" pageId="11" pageNumber="293">S. triglavica</emphasis>
</taxonomicName>
(
<figureCitation id="132EE1D8FFB4EE3162DFFEE918B32737" box="[159,247,290,315]" captionStart="FIGURES 9 12" captionStartId="7.[151,263,1899,1922]" captionTargetBox="[265,1330,718,1868]" captionTargetId="figure@7.[250,1337,700,1878]" captionTargetPageId="7" captionText="FIGURES 9 12. Male genitalia of Sattleria. 9, S. dinarica, sp. nov., paratype, slide GEL 1129; 10, S. triglavica Povolný, paratype, slide Mus. Vind. 11202; 11, S. haemusi, sp. nov., holotype, slide GU 13 / 1356 P. Huemer; 12, S. dzieduszyckii (Nowicki), slide L. M. P. 97." httpUri="https://zenodo.org/record/227961/files/figure.png" pageId="11" pageNumber="293">Figs 10</figureCitation>
,
<figureCitation id="132EE1D8FFB4EE316345FEE819652737" box="[261,289,291,315]" captionStart="FIGURES 17 20" captionStartId="9.[151,263,1662,1685]" captionTargetBox="[237,1353,186,1621]" captionTargetId="figure@9.[220,1366,171,1641]" captionTargetPageId="9" captionText="FIGURES 17 20. Male genitalia of Sattleria, details of vincular processes (seconardy process marked with arrow) and valva. 17, S. dinarica, sp. nov., paratype, slide GEL 1129; 18, S. triglavica Povolný, paratype, slide Mus. Vind. 11202; 19, S. haemusi, sp. nov., holotype, slide GU 13 / 1356 P. Huemer; 20, S. dzieduszyckii (Nowicki), slide L. M. P. 97." httpUri="https://zenodo.org/record/227963/files/figure.png" pageId="11" pageNumber="293">18</figureCitation>
) and
<taxonomicName id="4C1586DEFFB4EE316324FEE81A522737" box="[356,534,291,315]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="293" phylum="Arthropoda" rank="species" species="dzieduszyckii">
<emphasis id="B961214FFFB4EE316324FEE81A522737" box="[356,534,291,315]" italics="true" pageId="11" pageNumber="293">S. dzieduszyckii</emphasis>
</taxonomicName>
(
<figureCitation id="132EE1D8FFB4EE316067FEE91A3B2737" box="[551,639,290,315]" captionStart="FIGURES 9 12" captionStartId="7.[151,263,1899,1922]" captionTargetBox="[265,1330,718,1868]" captionTargetId="figure@7.[250,1337,700,1878]" captionTargetPageId="7" captionText="FIGURES 9 12. Male genitalia of Sattleria. 9, S. dinarica, sp. nov., paratype, slide GEL 1129; 10, S. triglavica Povolný, paratype, slide Mus. Vind. 11202; 11, S. haemusi, sp. nov., holotype, slide GU 13 / 1356 P. Huemer; 12, S. dzieduszyckii (Nowicki), slide L. M. P. 97." httpUri="https://zenodo.org/record/227961/files/figure.png" pageId="11" pageNumber="293">Figs 12</figureCitation>
,
<figureCitation id="132EE1D8FFB4EE3160CDFEE81AE82737" box="[653,684,291,315]" captionStart="FIGURES 17 20" captionStartId="9.[151,263,1662,1685]" captionTargetBox="[237,1353,186,1621]" captionTargetId="figure@9.[220,1366,171,1641]" captionTargetPageId="9" captionText="FIGURES 17 20. Male genitalia of Sattleria, details of vincular processes (seconardy process marked with arrow) and valva. 17, S. dinarica, sp. nov., paratype, slide GEL 1129; 18, S. triglavica Povolný, paratype, slide Mus. Vind. 11202; 19, S. haemusi, sp. nov., holotype, slide GU 13 / 1356 P. Huemer; 20, S. dzieduszyckii (Nowicki), slide L. M. P. 97." httpUri="https://zenodo.org/record/227963/files/figure.png" pageId="11" pageNumber="293">20</figureCitation>
), but differ by the evenly curved sub-oval shape of the secondary process of the vinculum, from the former two also by the broader primary process with stronger setae.
</paragraph>
<paragraph id="8BAAFD5DFFB4EE316287FEAC1B8C27AF" blockId="11.[151,1436,151,662]" pageId="11" pageNumber="293">
<emphasis id="B961214FFFB4EE316287FEAC1939278C" bold="true" box="[199,381,359,384]" pageId="11" pageNumber="293">Molecular data</emphasis>
(
<figureCitation id="132EE1D8FFB4EE3163CCFEAC199B278C" box="[396,479,359,384]" captionStart="FIGURE 21" captionStartId="10.[151,249,1886,1908]" captionTargetBox="[172,1431,852,1858]" captionTargetId="figure@10.[151,1436,809,1865]" captionTargetPageId="10" captionText="FIGURE 21. Neighbour joining tree of Sattleria from central and south-eastern Europe (Kimura 2 parameter, built with MEGA 5; cf. Tamura et al. 2011) based on sequences of the mtDNA COI gene (barcode fragment 5 , 658 bp), except for S. basistrigella (599 bp, 1 specimen), S. dzieduszyckii (623 bp, 1 specimen), S. triglavica (540 bp, 1 specimen). Width of triangles represent sample size, depth the molecular variation within the cluster. Source: DNA Barcode data from BOLD (Barcode of Life Database, cf. Ratnasingham &amp; Hebert 2007)." httpUri="https://zenodo.org/record/227964/files/figure.png" pageId="11" pageNumber="293">Fig. 21</figureCitation>
). The intraspecific divergence of the barcode region is low with 0.15% (p-dist) (n=2). The distance to the nearest neighbour
<taxonomicName id="4C1586DEFFB4EE316001FE401AFD27AF" box="[577,697,395,419]" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="293" phylum="Arthropoda" rank="species" species="cottiella">
<emphasis id="B961214FFFB4EE316001FE401AFD27AF" box="[577,697,395,419]" italics="true" pageId="11" pageNumber="293">S. cottiella</emphasis>
</taxonomicName>
is 3.69% (p-dist) (n=5).
</paragraph>
<paragraph id="8BAAFD5DFFB4EE316287FE661AB027EB" blockId="11.[151,1436,151,662]" pageId="11" pageNumber="293">
<emphasis id="B961214FFFB4EE316287FE66190E27CA" bold="true" box="[199,330,429,454]" pageId="11" pageNumber="293">Bionomics.</emphasis>
Host-plants and early stages are unknown. The few adults known to date have been collected in August at light at elevations of about
<quantity id="4CED50B8FFB4EE31607CFE1B1AB427EB" box="[572,752,464,488]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="11" pageNumber="293" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">
<date id="FFABDB9DFFB4EE31607CFE1B1A8B27E4" box="[572,719,464,488]" pageId="11" pageNumber="293" value="2100" valueMax="2200">2100 to 2200</date>
m
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FAED6FFB4EE316287FE39195B245C" pageId="11" pageNumber="293" type="distribution">
<paragraph id="8BAAFD5DFFB4EE316287FE39195B245C" blockId="11.[151,1436,151,662]" pageId="11" pageNumber="293">
<emphasis id="B961214FFFB4EE316287FE3919242407" bold="true" box="[199,352,498,523]" pageId="11" pageNumber="293">Distribution.</emphasis>
Known with certainty only from the Rila Mts. (
<collectingCountry id="F302BDCDFFB4EE316139FE391B992407" box="[889,989,498,523]" name="Bulgaria" pageId="11" pageNumber="293">Bulgaria</collectingCountry>
) and the Šar Planina Mts. (
<collectingCountry id="F302BDCDFFB4EE31674DFE391DD42407" box="[1293,1424,498,523]" name="Macedonia" pageId="11" pageNumber="293">Macedonia</collectingCountry>
), but from biogeographical considerations, unverified records from
<collectingCountry id="F302BDCDFFB4EE3161C9FDDE1BA62422" box="[905,994,533,558]" name="Albania" pageId="11" pageNumber="293">Albania</collectingCountry>
(
<bibRefCitation id="EF8480ACFFB4EE3161B3FDDE1CD02422" author="Povolny" box="[1011,1172,533,558]" pageId="11" pageNumber="293" refString="Povolny, D. (2001) A review of the taxa hierarchy in the euro-alpine genus Sattleria Povolny, 1965 (Lepidoptera, Gelechiidae, Gnorimoschemini). Acta Universitatis Agriculturae Brno, 49, 39 - 58." type="journal article" year="2001">Povolný 2001</bibRefCitation>
) probably also refer to this species.
</paragraph>
</subSubSection>
<subSubSection id="C30FAED6FFB4EE316287FD9118AB249A" pageId="11" pageNumber="293" type="etymology">
<paragraph id="8BAAFD5DFFB4EE316287FD9118AB249A" blockId="11.[151,1436,151,662]" pageId="11" pageNumber="293">
<emphasis id="B961214FFFB4EE316287FD91190A247F" bold="true" box="[199,334,602,627]" pageId="11" pageNumber="293">Etymology.</emphasis>
The name is derived from the latin noun
<emphasis id="B961214FFFB4EE316174FD901BCD247F" box="[820,905,603,627]" italics="true" pageId="11" pageNumber="293">haemus</emphasis>
= Balkans, referring to the distribution of the species.
</paragraph>
</subSubSection>
<subSubSection id="C30FAED6FFB4EE3162D7F8741A0E21F8" pageId="11" pageNumber="293" type="materials_examined">
<paragraph id="8BAAFD5DFFB4EE3162D7F8741A0E21F8" blockId="11.[151,1436,1983,2036]" pageId="11" pageNumber="293">
<emphasis id="B961214FFFB4EE3162D7F87418A821D8" bold="true" box="[151,236,1983,2004]" pageId="11" pageNumber="293">MAP 1.</emphasis>
The distribution of taxa of
<taxonomicName id="4C1586DEFFB4EE31604FF8741A2221D9" box="[527,614,1983,2005]" class="Insecta" family="Gelechiidae" genus="Sattleria" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B961214FFFB4EE31604FF8741A2221D9" box="[527,614,1983,2005]" italics="true" pageId="11" pageNumber="293">Sattleria</emphasis>
</taxonomicName>
in central and south-eastern Europe (exclusively based on examined material); altitudinal zones above
<quantity id="4CED50B8FFB4EE3163C6F815199721F8" box="[390,467,2014,2036]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="293" unit="m" value="1600.0">1600 m</quantity>
s.l. in blue.
</paragraph>
</subSubSection>
</treatment>
</document>