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<document id="B860440C1C7BB80C1962544E2BBBB6D0" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629656228228" checkinUser="felipe" docAuthor="Fancello, Luca, Hernando, Carles &amp; Leo, Piero" docDate="2009" docId="03A387F2FFCBFFCFD2E7FE63FAD85FCB" docLanguage="en" docName="zt02318p338.pdf" docOrigin="Zootaxa 2318" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptotyphlus nardii Fancello &amp; Hernando &amp; Leo 2009, sp. nov." docType="treatment" docVersion="2" lastPageNumber="325" masterDocId="FF9AFF8AFFCCFFC7D270FFC5FFC75F79" masterDocTitle="The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *" masterLastPageNumber="338" masterPageNumber="317" pageNumber="324" updateTime="1699090143914" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="3EFF86A7CFFCEE67A240B1379AE759DB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9CC12535ACCB3ED2A138423847AAC21A">
<mods:title id="9C2583FE4704A95CFC7C1EE3EB4EF10E">The endogean beetle fauna of the Marganai-Oridda-Valle del Leni area (SW Sardinia), with description of seven new species of Staphylinidae Leptotyphlinae (Coleoptera) *</mods:title>
</mods:titleInfo>
<mods:name id="F8D6CEE25AC8D12651C862D8A7FAF127" type="personal">
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<mods:namePart id="FCDD2608A6221FCC01A541D26B2CEB6E">Fancello, Luca</mods:namePart>
<mods:affiliation id="745CF454FD8D20BA39862EEA9AA43DAF">Via Bainsizza 12, I- 09123 Cagliari, Italy. E-mail: L. fancello @ hotmail. it Museu de Ciènces Naturals de la Ciutadella (Zoologia), Passeig Picasso s / n, E- 08003 Barcelona, Spain. E-mail: c _ hernando @ telefonica. net Via Tola 21, I- 09128 Cagliari, Italy. E-mail: piero. leo @ tiscali. it</mods:affiliation>
</mods:name>
<mods:name id="2E03EA35EC8C24D67A0443018A50CE61" type="personal">
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<mods:namePart id="B3265E2918809A894FF4977BB1D068C6">Hernando, Carles</mods:namePart>
</mods:name>
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<mods:namePart id="B8926A4FE64634FADDEE7BA85B478AFC">Leo, Piero</mods:namePart>
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<mods:title id="284A8A9DC898C3E26248974F92EB2F13">Zootaxa</mods:title>
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<mods:part id="59AE0F26EC1C44960A649A3627799D53">
<mods:date id="37A8B79508D2C7B15EF6887AB65B0777">2009</mods:date>
<mods:detail id="40C90E37D79B0632A19E42A3F7135885" type="pubDate">
<mods:number id="A08D325A349A76E006C386ABFAAF236F">2009-12-22</mods:number>
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<mods:detail id="BAE0006122038417B847AF0C7A4789B7" type="volume">
<mods:number id="ABBA005E5E36AEDC77EB8A8BF0F802C2">2318</mods:number>
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<mods:classification id="E5C9109BB45DE446E26F78969941DE98">journal article</mods:classification>
<mods:identifier id="EA7999C438DC83D02946672BDB861E59" type="ISSN">1175-5326</mods:identifier>
</mods:mods>
<treatment id="03A387F2FFCBFFCFD2E7FE63FAD85FCB" LSID="urn:lsid:plazi:treatment:03A387F2FFCBFFCFD2E7FE63FAD85FCB" httpUri="http://treatment.plazi.org/id/03A387F2FFCBFFCFD2E7FE63FAD85FCB" lastPageId="8" lastPageNumber="325" pageId="7" pageNumber="324">
<subSubSection id="C310656FFFCBFFC0D2E7FE63FE2F5EB9" box="[151,488,422,448]" pageId="7" pageNumber="324" type="nomenclature">
<paragraph id="8BB536E4FFCBFFC0D2E7FE63FE2F5EB9" blockId="7.[151,488,422,483]" box="[151,488,422,448]" pageId="7" pageNumber="324">
<heading id="D0FD8188FFCBFFC0D2E7FE63FE2F5EB9" bold="true" box="[151,488,422,448]" fontSize="11" level="1" pageId="7" pageNumber="324" reason="1">
<emphasis id="B97EEAF6FFCBFFC0D2E7FE63FE2F5EB9" bold="true" box="[151,488,422,448]" pageId="7" pageNumber="324">
<taxonomicName id="4C0A4D67FFCBFFC0D2E7FE63FE445EB9" ID-CoL="3TZ9G" authority="Fancello &amp; Hernando &amp; Leo, 2009" authorityName="Fancello &amp; Hernando &amp; Leo" authorityYear="2009" box="[151,387,422,448]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="nardii" status="sp. nov.">
<emphasis id="B97EEAF6FFCBFFC0D2E7FE63FE445EB9" bold="true" box="[151,387,422,448]" italics="true" pageId="7" pageNumber="324">Leptotyphlus nardii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D578DFFCBFFC0D3FBFE63FE2F5EB9" box="[395,488,422,448]" pageId="7" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFCBFFC0D2E7FE0CFE9E5E9A" box="[151,345,457,483]" pageId="7" pageNumber="324" type="description">
<paragraph id="8BB536E4FFCBFFC0D2E7FE0CFE9E5E9A" blockId="7.[151,488,422,483]" box="[151,345,457,483]" pageId="7" pageNumber="324">
(
<figureCitation id="13312A61FFCBFFC0D2EFFE0CFF3D5E9A" box="[159,250,457,483]" captionStart="FIGURES 15–17" captionStartId="8.[151,271,995,1019]" captionTargetBox="[157,1424,241,965]" captionTargetId="figure-374@8.[151,1436,235,969]" captionTargetPageId="8" captionText="FIGURES 15–17. Male copulatory organ of Leptotyphlus spp. in lateral view, scale bars: 0.05 mm. 15. Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro). 16. Leptotyphlus nardii sp. nov. (paratype, Nebida). 17. Leptotyphlus minator sp. nov. (paratype, Nebida)." pageId="7" pageNumber="324">Figs 16</figureCitation>
,
<figureCitation id="13312A61FFCBFFC0D376FE0CFE965E9A" box="[262,337,457,483]" captionStart="FIGURES 18–26" captionStartId="9.[151,270,1788,1812]" captionTargetBox="[151,1432,278,1760]" captionTargetId="figure-132@9.[151,1436,274,1763]" captionTargetPageId="9" captionText="FIGURES 18–26. Leptotyphlus spp., scale bars 0.05 mm. 18–20. Leptotyphlus nardii sp. nov. (paratype, Nebida). 18. Labrum in dorsal view. 19. Right mandible in dorsal view. 20. Female genital armature in ventral view. 21–23. Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro). 21. Labrum in dorsal view. 22. Right mandible in dorsal view. 23. Female genital armature of Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro) in ventral view. 24–26. Leptotyphlus minator sp. nov. (paratype, Gutturu di Monte Nieddu). 24. Female genital armature in ventral view. 25. Labrum in dorsal view. 26. Right mandible in dorsal view." pageId="7" pageNumber="324">18–20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFCBFFC0D2E7FDD3FA415DDD" pageId="7" pageNumber="324" type="diagnosis">
<paragraph id="8BB536E4FFCBFFC0D2E7FDD3FA415DDD" blockId="7.[151,1437,534,2030]" pageId="7" pageNumber="324">
<emphasis id="B97EEAF6FFCBFFC0D2E7FDD3FED95D49" bold="true" box="[151,286,534,560]" pageId="7" pageNumber="324">Diagnosis.</emphasis>
A
<taxonomicName id="4C0A4D67FFCBFFC0D337FDD2FD535D49" authority="Fauvel, 1874" authorityName="Fauvel" authorityYear="1874" box="[327,660,534,560]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAF6FFCBFFC0D337FDD2FE2F5D49" box="[327,488,535,560]" italics="true" pageId="7" pageNumber="324">Leptotyphlus</emphasis>
<bibRefCitation id="EF9B4B15FFCBFFC0D381FDD3FD535D49" author="Fauvel, A." box="[497,660,534,560]" pageId="7" pageNumber="324" pagination="167 - 340" refId="ref12217" refString="Fauvel, A. (1874) Faune Gallo-Rhenane ou description des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhenanes et la Valais avec tableaux synoptyque et planches gravees. Bulletin de la Societe Linneenne de Normandie (2), 8, 167 - 340." type="journal article" year="1874">Fauvel, 1874</bibRefCitation>
</taxonomicName>
of medium size, with rather poor punctuation and superficial microreticulation. Differs from its congeners by the characters of the copulatory organ, but is undoubtedly akin to
<taxonomicName id="4C0A4D67FFCBFFC0D283FDA1FE495D04" authorityName="Fauvel" authorityYear="1874" box="[243,398,612,637]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B97EEAF6FFCBFFC0D283FDA1FE495D04" box="[243,398,612,637]" italics="true" pageId="7" pageNumber="324">Leptotyphlus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D578DFFCBFFC0D3E7FDA1FE105D07" box="[407,471,612,638]" pageId="7" pageNumber="324" sensu="stricto">s. str.</taxonomicNameLabel>
species of the
<taxonomicName id="4C0A4D67FFCBFFC0D0FEFDA1FC7A5D07" authority="Coiffait, 1955" authorityName="Coiffait" authorityYear="1955" box="[654,957,612,638]" class="Insecta" family="Brachyceridae" genus="Raymondiellus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="sardous">
<emphasis id="B97EEAF6FFCBFFC0D0FEFDA1FD595D04" box="[654,670,612,637]" italics="true" pageId="7" pageNumber="324">L</emphasis>
.
<emphasis id="B97EEAF6FFCBFFC0D0DEFDA1FCCB5D04" box="[686,780,612,637]" italics="true" pageId="7" pageNumber="324">sardous</emphasis>
Coiffait, 1955
</taxonomicName>
a-group (
<emphasis id="B97EEAF6FFCBFFC0D65EFDA1FBB75D04" box="[1070,1136,612,637]" italics="true" pageId="7" pageNumber="324">sensu</emphasis>
<bibRefCitation id="EF9B4B15FFCBFFC0D60AFDA1FB3F5D07" author="Pace R. &amp; Fauna d'Italia, XXXIV" box="[1146,1272,612,638]" pageId="7" pageNumber="324" refId="ref13459" refString="Pace R., (1996). Fauna d'Italia, XXXIV. Coleoptera Staphylinidae Leptotyphlinae. Calderini, Bologna, 328 pp." type="book" year="1996">Pace 1996</bibRefCitation>
) and to some peninsular taxa of the
<taxonomicName id="4C0A4D67FFCBFFC0D3EBFD4FFD735DDD" authority="Pace, 1973" authorityName="Pace" authorityYear="1973" box="[411,692,650,676]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="lessinicus">
<emphasis id="B97EEAF6FFCBFFC0D3EBFD4FFE6C5DDA" box="[411,427,650,675]" italics="true" pageId="7" pageNumber="324">L</emphasis>
.
<emphasis id="B97EEAF6FFCBFFC0D3CAFD4FFDEB5DDA" box="[442,556,650,675]" italics="true" pageId="7" pageNumber="324">lessinicus</emphasis>
<bibRefCitation id="EF9B4B15FFCBFFC0D043FD4FFD735DDD" author="Pace, R." box="[563,692,650,676]" pageId="7" pageNumber="324" pagination="313 - 324" refId="ref13267" refString="Pace, R. (1973) Tre nuove specie di Leptotyphlinae Scheepeltz e descrizione del maschio di Metrotyphlus castrii Coiffait (Coleoptera Staphylinidae). Memorie del Museo Civico di Storia Naturale di Verona, 20 [1972], 313 - 324." type="journal article" year="1973">Pace, 1973</bibRefCitation>
</taxonomicName>
-group, belonging to subgenus
<taxonomicName id="4C0A4D67FFCBFFC0D650FD4FFA445DDD" authority="Coiffait, 1955 b" authorityName="Coiffait" authorityYear="1955" box="[1056,1411,650,676]" class="Insecta" family="Staphylinidae" genus="Stigmotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAF6FFCBFFC0D650FD4FFB025DDA" box="[1056,1221,650,675]" italics="true" pageId="7" pageNumber="324">Stigmotyphlus</emphasis>
<bibRefCitation id="EF9B4B15FFCBFFC0D6BCFD4FFA445DDD" author="Coiffait, H." box="[1228,1411,650,676]" pageId="7" pageNumber="324" pagination="262 - 269" refId="ref11730" refString="Coiffait, H. (1955 b) Contribution a la connaisance des Osoriini anophtalmes (Col. Staphylinidae). Revue Francaise d'Entomologie, 22, 262 - 269." type="journal article" year="1955">Coiffait, 1955b</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFCBFFC0D2B6FD75FA575CCA" pageId="7" pageNumber="324" type="materials_examined">
<paragraph id="8BB536E4FFCBFFC0D2B6FD75FC3D5DB2" blockId="7.[151,1437,534,2030]" box="[198,1018,688,715]" pageId="7" pageNumber="324">
<materialsCitation id="3B623CB9FFCBFFC0D2B6FD75FC305DB2" box="[198,1015,688,715]" county="Carbonia-Iglesias prov." location="Nebida" municipality="Iglesias" pageId="7" pageNumber="324" specimenCount="1" stateProvince="Sardegna" typeStatus="holotype">
<emphasis id="B97EEAF6FFCBFFC0D2B6FD75FEAB5DB3" bold="true" box="[198,364,688,714]" pageId="7" pageNumber="324">
<typeStatus id="54B18846FFCBFFC0D2B6FD75FEC35DB3" box="[198,260,688,714]" pageId="7" pageNumber="324">Type</typeStatus>
locality.
</emphasis>
SW
<collectingRegion id="49CEF806FFCBFFC0D3D6FD74FDCC5DB2" box="[422,523,689,715]" country="Italy" name="Sardegna" pageId="7" pageNumber="324">Sardinia</collectingRegion>
,
<collectingCounty id="62D44E68FFCBFFC0D066FD74FCED5DB2" box="[534,810,689,715]" pageId="7" pageNumber="324">Carbonia-Iglesias prov.</collectingCounty>
,
<collectingMunicipality id="6BD1AC9EFFCBFFC0D146FD74FC555DB2" box="[822,914,689,715]" pageId="7" pageNumber="324">Iglesias</collectingMunicipality>
,
<location id="8ED5603FFFCBFFC0D1EEFD74FC305DB2" LSID="urn:lsid:plazi:treatment:03A387F2FFCBFFCFD2E7FE63FAD85FCB:8ED5603FFFCBFFC0D1EEFD74FC305DB2" box="[926,1015,689,715]" county="Carbonia-Iglesias prov." municipality="Iglesias" name="Nebida" pageId="7" pageNumber="324" stateProvince="Sardegna">Nebida</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB536E4FFCBFFC0D2B6FD12FA575CCA" blockId="7.[151,1437,534,2030]" pageId="7" pageNumber="324">
<emphasis id="B97EEAF6FFCBFFC0D2B6FD12FE9D5D88" bold="true" box="[198,346,727,753]" pageId="7" pageNumber="324">Type series.</emphasis>
<materialsCitation id="3B623CB9FFCBFFC0D314FD1DFE2D5C61" collectingDate="1987-01-25" collectionCode="MSNG" collectorName="L. Fancello &amp; P. Leo" county="Iglesias" elevation="120" location="Nebida" municipality="Carbonia-Iglesias" pageId="7" pageNumber="324" specimenCount="1" specimenCount-male="1" stateProvince="Sardegna" typeStatus="holotype">
<typeStatus id="54B18846FFCBFFC0D314FD1DFE165D8B" box="[356,465,728,754]" pageId="7" pageNumber="324" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0CFD6DFFCBFFC0D3ACFD1CFE295D89" box="[476,494,729,752]" pageId="7" pageNumber="324" type="male">♂</specimenCount>
: “SW
<collectingRegion id="49CEF806FFCBFFC0D032FD1DFD615D8B" box="[578,678,728,754]" country="Italy" name="Sardegna" pageId="7" pageNumber="324">Sardinia</collectingRegion>
,
<collectingCounty id="62D44E68FFCBFFC0D0C5FD1DFCC95D8B" box="[693,782,728,754]" pageId="7" pageNumber="324">Iglesias</collectingCounty>
(
<collectingMunicipality id="6BD1AC9EFFCBFFC0D16FFD1DFC335D8B" box="[799,1012,728,754]" pageId="7" pageNumber="324">Carbonia-Iglesias</collectingMunicipality>
),
<location id="8ED5603FFFCBFFC0D67AFD1DFBA65D8B" LSID="urn:lsid:plazi:treatment:03A387F2FFCBFFCFD2E7FE63FAD85FCB:8ED5603FFFCBFFC0D67AFD1DFBA65D8B" box="[1034,1121,728,754]" county="Iglesias" municipality="Carbonia-Iglesias" name="Nebida" pageId="7" pageNumber="324" stateProvince="Sardegna">Nebida</location>
,
<quantity id="4CF29B01FFCBFFC0D61FFD1DFB795D8B" box="[1135,1214,728,754]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="324" unit="m" value="120.0">
<elevation id="0027D1D7FFCBFFC0D61FFD1DFB795D8B" box="[1135,1214,728,754]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="324" unit="m" value="120.0">120 m</elevation>
</quantity>
,
<date id="FFB41024FFCBFFC0D6BDFD1DFAF85D8B" box="[1229,1343,728,754]" pageId="7" pageNumber="324" value="1987-01-25">
<collectingDate id="EFF0E9CCFFCBFFC0D6BDFD1DFAF85D8B" box="[1229,1343,728,754]" pageId="7" pageNumber="324" value="1987-01-25">25.I.1987</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFCBFFC0D7F2FD1DFF3B5C61" pageId="7" pageNumber="324">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFCBFFC0D351FD3BFEAB5C61" box="[289,364,766,792]" pageId="7" pageNumber="324">P. Leo</collectorName>
” (
<collectionCode id="ED1BAE21FFCBFFC0D3F6FD3BFE255C61" box="[390,482,766,792]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="7" pageNumber="324">MSNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B623CB9FFCBFFC0D385FD3BFBC35CF5" collectingDate="1986-11-11" collectingDateMax="1990-04-13" collectingDateMin="1986-11-11" collectionCode="CLF-PL, CCH, CNBFVR" collectorName="L. Fancello &amp; P. Leo" county="Gutturu di Monte Nieddu" elevation="130" location="Carbonia" municipality="Carbonia-Iglesias prov." pageId="7" pageNumber="324" specimenCount="32" specimenCount-female="25" specimenCount-male="7" stateProvince="Sardegna" typeStatus="paratype">
<typeStatus id="54B18846FFCBFFC0D385FD3BFDAB5C61" box="[501,620,766,792]" pageId="7" pageNumber="324" type="paratype">Paratypes</typeStatus>
:
<location id="8ED5603FFFCBFFC0D006FD3BFD265C61" LSID="urn:lsid:plazi:treatment:03A387F2FFCBFFCFD2E7FE63FAD85FCB:8ED5603FFFCBFFC0D006FD3BFD265C61" box="[630,737,766,792]" county="Gutturu di Monte Nieddu" municipality="Carbonia-Iglesias prov." name="Carbonia" pageId="7" pageNumber="324" stateProvince="Sardegna">Carbonia</location>
(
<location id="8ED5603FFFCBFFC0D081FD3BFBCF5C61" LSID="urn:lsid:plazi:treatment:03A387F2FFCBFFCFD2E7FE63FAD85FCB:8ED5603FFFCBFFC0D081FD3BFBCF5C61" box="[753,1032,766,792]" county="Gutturu di Monte Nieddu" municipality="Carbonia-Iglesias prov." name="Carbonia-Iglesias prov." pageId="7" pageNumber="324" stateProvince="Sardegna">Carbonia-Iglesias prov.</location>
),
<location id="8ED5603FFFCBFFC0D66CFD3BFBBD5C61" LSID="urn:lsid:plazi:treatment:03A387F2FFCBFFCFD2E7FE63FAD85FCB:8ED5603FFFCBFFC0D66CFD3BFBBD5C61" box="[1052,1146,766,792]" county="Gutturu di Monte Nieddu" municipality="Carbonia-Iglesias prov." name="Barbusi" pageId="7" pageNumber="324" stateProvince="Sardegna">Barbusi</location>
,
<quantity id="4CF29B01FFCBFFC0D6F5FD3BFB155C61" box="[1157,1234,766,792]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="324" unit="m" value="130.0">
<elevation id="0027D1D7FFCBFFC0D6F5FD3BFB155C61" box="[1157,1234,766,792]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="324" unit="m" value="130.0">130 m</elevation>
</quantity>
,
<date id="FFB41024FFCBFFC0D6AEFD3BFAA25C61" box="[1246,1381,766,792]" pageId="7" pageNumber="324" value="1986-11-11">
<collectingDate id="EFF0E9CCFFCBFFC0D6AEFD3BFAA25C61" box="[1246,1381,766,792]" pageId="7" pageNumber="324" value="1986-11-11">11.XI.1986</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFCBFFC0D2E7FCE0FED85C46" box="[151,287,805,831]" pageId="7" pageNumber="324">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFCBFFC0D337FCE0FE525C46" box="[327,405,805,831]" pageId="7" pageNumber="324">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFCBFFC0D3D2FCE0FE255C47" box="[418,482,805,831]" pageId="7" pageNumber="324" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFCBFFC0D06DFCE0FD8E5C47" box="[541,585,805,831]" pageId="7" pageNumber="324" type="female">2 ♀</specimenCount>
(
<collectionCode id="ED1BAE21FFCBFFC0D01DFCE0FD105C46" box="[621,727,805,831]" pageId="7" pageNumber="324">CLF-PL</collectionCode>
). Domusnovas (
<collectingMunicipality id="6BD1AC9EFFCBFFC0D1E7FCE0FB735C46" box="[919,1204,805,831]" pageId="7" pageNumber="324">Carbonia-Iglesias prov.</collectingMunicipality>
),
<collectingCounty id="62D44E68FFCBFFC0D6BAFCE0FF375C1F" pageId="7" pageNumber="324">Gutturu di Monte Nieddu</collectingCounty>
,
<quantity id="4CF29B01FFCBFFC0D28BFC89FE8F5C1F" box="[251,328,844,870]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="324" unit="m" value="290.0">
<elevation id="0027D1D7FFCBFFC0D28BFC89FE8F5C1F" box="[251,328,844,870]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="324" unit="m" value="290.0">290 m</elevation>
</quantity>
,
<date id="FFB41024FFCBFFC0D323FC89FE015C1F" box="[339,454,844,870]" pageId="7" pageNumber="324" value="1990-01-12">
<collectingDate id="EFF0E9CCFFCBFFC0D323FC89FE015C1F" box="[339,454,844,870]" pageId="7" pageNumber="324" value="1990-01-12">12.I.1990</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFCBFFC0D073FC89FD4F5C1F" box="[515,648,844,870]" pageId="7" pageNumber="324">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFCBFFC0D0DDFC89FD3E5C1F" box="[685,761,844,870]" pageId="7" pageNumber="324">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFCBFFC0D175FC89FCE95C1D" box="[773,814,844,870]" pageId="7" pageNumber="324" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFCBFFC0D116FC89FC575C1D" box="[870,912,844,870]" pageId="7" pageNumber="324" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED1BAE21FFCBFFC0D1EFFC89FBCF5C1F" box="[927,1032,844,870]" pageId="7" pageNumber="324">CLF-PL</collectionCode>
);
<date id="FFB41024FFCBFFC0D66BFC89FB675C1F" box="[1051,1184,844,870]" pageId="7" pageNumber="324" value="1990-04-13">
<collectingDate id="EFF0E9CCFFCBFFC0D66BFC89FB675C1F" box="[1051,1184,844,870]" pageId="7" pageNumber="324" value="1990-04-13">13.IV.1990</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFCBFFC0D6AEFC89FAA45C1F" box="[1246,1379,844,870]" pageId="7" pageNumber="324">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFCBFFC0D7F7FC89FF005CF5" pageId="7" pageNumber="324">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFCBFFC0D2A3FCB7FED65CF2" box="[211,273,882,908]" pageId="7" pageNumber="324" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFCBFFC0D33AFCB7FE455CF2" box="[330,386,882,908]" pageId="7" pageNumber="324" type="female">11 ♀</specimenCount>
(
<specimenCount id="9D0CFD6DFFCBFFC0D3D9FCB7FE165CF2" box="[425,465,882,908]" pageId="7" pageNumber="324" type="female">1 ♀</specimenCount>
<collectionCode id="ED1BAE21FFCBFFC0D3A8FCB7FDD05CF5" box="[472,535,882,908]" pageId="7" pageNumber="324">CCH</collectionCode>
,
<specimenCount id="9D0CFD6DFFCBFFC0D053FCB7FDA65CF2" box="[547,609,882,908]" pageId="7" pageNumber="324" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFCBFFC0D0EAFCB7FD035CF2" box="[666,708,882,908]" pageId="7" pageNumber="324" type="female">9 ♀</specimenCount>
<collectionCode id="ED1BAE21FFCBFFC0D0AFFCB7FC835CF5" box="[735,836,882,908]" pageId="7" pageNumber="324">CLF-PL</collectionCode>
,
<specimenCount id="9D0CFD6DFFCBFFC0D120FCB7FCBD5CF2" box="[848,890,882,908]" pageId="7" pageNumber="324" type="female">1 ♀</specimenCount>
<collectionCode id="ED1BAE21FFCBFFC0D1F1FCB7FC3B5CF5" box="[897,1020,882,908]" pageId="7" pageNumber="324">CNBFVR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B623CB9FFCBFFC0D67FFCB7FC6F5CC8" collectingDate="1987-01-25" collectorName="L. Fancello &amp; P. Leo" county="Nebida" elevation="120" location="Iglesias" municipality="Carbonia-Iglesias prov." pageId="7" pageNumber="324" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Sardegna" typeStatus="paratype">
<location id="8ED5603FFFCBFFC0D67FFCB7FBAF5CF5" LSID="urn:lsid:plazi:treatment:03A387F2FFCBFFCFD2E7FE63FAD85FCB:8ED5603FFFCBFFC0D67FFCB7FBAF5CF5" box="[1039,1128,882,908]" county="Nebida" municipality="Carbonia-Iglesias prov." name="Iglesias" pageId="7" pageNumber="324" stateProvince="Sardegna">Iglesias</location>
(
<collectingMunicipality id="6BD1AC9EFFCBFFC0D608FCB7FA485CF5" box="[1144,1423,882,908]" pageId="7" pageNumber="324">Carbonia-Iglesias prov.</collectingMunicipality>
),
<collectingCounty id="62D44E68FFCBFFC0D2E7FC5CFF285CCA" box="[151,239,921,947]" pageId="7" pageNumber="324">Nebida</collectingCounty>
,
<quantity id="4CF29B01FFCBFFC0D28AFC5CFE815CCA" box="[250,326,921,947]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="324" unit="m" value="120.0">
<elevation id="0027D1D7FFCBFFC0D28AFC5CFE815CCA" box="[250,326,921,947]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="324" unit="m" value="120.0">120 m</elevation>
</quantity>
,
<date id="FFB41024FFCBFFC0D323FC5CFE025CCA" box="[339,453,921,947]" pageId="7" pageNumber="324" value="1987-01-25">
<collectingDate id="EFF0E9CCFFCBFFC0D323FC5CFE025CCA" box="[339,453,921,947]" pageId="7" pageNumber="324" value="1987-01-25">25.I.1987</collectingDate>
</date>
, leg.
<collectorName id="26FF5332FFCBFFC0D074FC5CFD4D5CCA" box="[516,650,921,947]" pageId="7" pageNumber="324">L. Fancello</collectorName>
&amp;
<collectorName id="26FF5332FFCBFFC0D0DFFC5CFD3C5CCA" box="[687,763,921,947]" pageId="7" pageNumber="324">P. Leo</collectorName>
,
<specimenCount id="9D0CFD6DFFCBFFC0D177FC5CFC825CC8" box="[775,837,921,947]" pageId="7" pageNumber="324" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFCBFFC0D10EFC5CFC6F5CC8" box="[894,936,921,947]" pageId="7" pageNumber="324" type="female">3 ♀</specimenCount>
</materialsCitation>
<materialsCitation id="3B623CB9FFCBFFC0D1B3FC5CFA4B5CCA" box="[963,1420,921,947]" collectionCode="CLF-PL, MSNG" pageId="7" pageNumber="324" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Sardegna" typeStatus="paratype">
(
<specimenCount id="9D0CFD6DFFCBFFC0D1BCFC5CFBCC5CC8" box="[972,1035,921,947]" pageId="7" pageNumber="324" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="9D0CFD6DFFCBFFC0D634FC5CFBA95CC8" box="[1092,1134,921,947]" pageId="7" pageNumber="324" type="female">2 ♀</specimenCount>
<collectionCode id="ED1BAE21FFCBFFC0D6F9FC5CFB295CCA" box="[1161,1262,921,947]" pageId="7" pageNumber="324">CLF-PL</collectionCode>
,
<specimenCount id="9D0CFD6DFFCBFFC0D689FC5CFAE45CC8" box="[1273,1315,921,947]" pageId="7" pageNumber="324" type="female">1 ♀</specimenCount>
<collectionCode id="ED1BAE21FFCBFFC0D75AFC5CFA435CCA" box="[1322,1412,921,947]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="7" pageNumber="324">MSNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFCBFFC0D2B6FC7AFDA15897" pageId="7" pageNumber="324" type="description">
<paragraph id="8BB536E4FFCBFFC0D2B6FC7AFE6C5A4F" blockId="7.[151,1437,534,2030]" pageId="7" pageNumber="324">
<emphasis id="B97EEAF6FFCBFFC0D2B6FC7AFE995CA0" bold="true" box="[198,350,959,985]" pageId="7" pageNumber="324">Description.</emphasis>
Total length 1.10–1.20 mm. Body of pale testaceous colour, clearly shiny, with superficial microsculpture on all integuments. Head slightly wider than pronotum, with moderately rounded sides and only slightly outlined frontal carinas; punctuation poor; labrum (
<figureCitation id="13312A61FFCBFFC0D1F9FBC8FC1A5B5E" box="[905,989,1037,1063]" captionStart="FIGURES 18–26" captionStartId="9.[151,270,1788,1812]" captionTargetBox="[151,1432,278,1760]" captionTargetId="figure-132@9.[151,1436,274,1763]" captionTargetPageId="9" captionText="FIGURES 18–26. Leptotyphlus spp., scale bars 0.05 mm. 18–20. Leptotyphlus nardii sp. nov. (paratype, Nebida). 18. Labrum in dorsal view. 19. Right mandible in dorsal view. 20. Female genital armature in ventral view. 21–23. Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro). 21. Labrum in dorsal view. 22. Right mandible in dorsal view. 23. Female genital armature of Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro) in ventral view. 24–26. Leptotyphlus minator sp. nov. (paratype, Gutturu di Monte Nieddu). 24. Female genital armature in ventral view. 25. Labrum in dorsal view. 26. Right mandible in dorsal view." pageId="7" pageNumber="324">Fig. 18</figureCitation>
) with three weakly developed median teeth; mandibles (
<figureCitation id="13312A61FFCBFFC0D31EFBF1FE025B37" box="[366,453,1076,1102]" captionStart="FIGURES 18–26" captionStartId="9.[151,270,1788,1812]" captionTargetBox="[151,1432,278,1760]" captionTargetId="figure-132@9.[151,1436,274,1763]" captionTargetPageId="9" captionText="FIGURES 18–26. Leptotyphlus spp., scale bars 0.05 mm. 18–20. Leptotyphlus nardii sp. nov. (paratype, Nebida). 18. Labrum in dorsal view. 19. Right mandible in dorsal view. 20. Female genital armature in ventral view. 21–23. Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro). 21. Labrum in dorsal view. 22. Right mandible in dorsal view. 23. Female genital armature of Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro) in ventral view. 24–26. Leptotyphlus minator sp. nov. (paratype, Gutturu di Monte Nieddu). 24. Female genital armature in ventral view. 25. Labrum in dorsal view. 26. Right mandible in dorsal view." pageId="7" pageNumber="324">Fig. 19</figureCitation>
) bifid with robust retinaculum and premolar tooth; antennae relatively short, just longer than head. Pronotum slightly longer than wide, with two longitudinal superficial grooves, slightly converging anteriorly; punctuation minute and sparser than on head. Elytral punctuation imperceptible. Convexity of abdominal tergites normal. Aedeagus as in
<figureCitation id="13312A61FFCBFFC0D143FB6DFC4F5BBB" box="[819,904,1192,1218]" captionStart="FIGURES 15–17" captionStartId="8.[151,271,995,1019]" captionTargetBox="[157,1424,241,965]" captionTargetId="figure-374@8.[151,1436,235,969]" captionTargetPageId="8" captionText="FIGURES 15–17. Male copulatory organ of Leptotyphlus spp. in lateral view, scale bars: 0.05 mm. 15. Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro). 16. Leptotyphlus nardii sp. nov. (paratype, Nebida). 17. Leptotyphlus minator sp. nov. (paratype, Nebida)." pageId="7" pageNumber="324">Fig. 16</figureCitation>
, lying on the left side of the abdomen, with a robust and arched dorsal copulatory piece; sternal lamina distally divided into two branches; tip of ventral branch in lateral view with a distinct downturned lobiform appendix. Female genital armature asymmetrical, structured as in
<figureCitation id="13312A61FFCBFFC0D321FAD9FE6F5A4F" box="[337,424,1308,1334]" captionStart="FIGURES 18–26" captionStartId="9.[151,270,1788,1812]" captionTargetBox="[151,1432,278,1760]" captionTargetId="figure-132@9.[151,1436,274,1763]" captionTargetPageId="9" captionText="FIGURES 18–26. Leptotyphlus spp., scale bars 0.05 mm. 18–20. Leptotyphlus nardii sp. nov. (paratype, Nebida). 18. Labrum in dorsal view. 19. Right mandible in dorsal view. 20. Female genital armature in ventral view. 21–23. Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro). 21. Labrum in dorsal view. 22. Right mandible in dorsal view. 23. Female genital armature of Leptotyphlus villacidrinus sp. nov. (paratype, Villacidro) in ventral view. 24–26. Leptotyphlus minator sp. nov. (paratype, Gutturu di Monte Nieddu). 24. Female genital armature in ventral view. 25. Labrum in dorsal view. 26. Right mandible in dorsal view." pageId="7" pageNumber="324">Fig. 20</figureCitation>
.
</paragraph>
<paragraph id="8BB536E4FFCBFFC0D2B6FA87FC295855" blockId="7.[151,1437,534,2030]" pageId="7" pageNumber="324">
<emphasis id="B97EEAF6FFCBFFC0D2B6FA87FEB45A25" bold="true" box="[198,371,1346,1372]" pageId="7" pageNumber="324">Observations.</emphasis>
As hinted in the diagnosis, the phyletic relationships of the new species seem complex. This forced us to describe the new species and the following two without subgeneric assignment. Besides, the subdivision of
<taxonomicName id="4C0A4D67FFCBFFC0D33FFA55FE2A5AD0" authorityName="Fauvel" authorityYear="1874" box="[335,493,1424,1449]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAF6FFCBFFC0D33FFA55FE2A5AD0" box="[335,493,1424,1449]" italics="true" pageId="7" pageNumber="324">Leptotyphlus</emphasis>
</taxonomicName>
is still debated:
<bibRefCitation id="EF9B4B15FFCBFFC0D0CBFA55FCB05AD3" author="Herman, L. H." box="[699,887,1424,1450]" pageId="7" pageNumber="324" pagination="1807 - 2440" refId="ref12405" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. 4. Staphylininae group (part 1), Euaesthetinae, Leptotyphlinae, Megalopsidinae, Oxyporinae, Pseudopsinae, Solieriinae, Steninae. Bullettin of the American Museum of Natural History, 265, 1807 - 2440." type="journal article" year="2001">Herman (2001)</bibRefCitation>
considers all described subgenera simply as synonyms of
<taxonomicName id="4C0A4D67FFCBFFC0D348FA72FE165AA9" authorityName="Fauvel" authorityYear="1874" box="[312,465,1463,1488]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAF6FFCBFFC0D348FA72FE165AA9" box="[312,465,1463,1488]" italics="true" pageId="7" pageNumber="324">Leptotyphlus</emphasis>
</taxonomicName>
, whereas
<bibRefCitation id="EF9B4B15FFCBFFC0D039FA73FCCB5AA9" author="Smetana, A." box="[585,780,1462,1488]" pageId="7" pageNumber="324" refId="ref13998" refString="Smetana, A. (2004 a) Fauna Europaea: Staphylinidae Leptotyphlinae. In: Alonso-Zarazaga, M. A. (Ed.), Fauna Europaea: Coleoptera 1, Beetles. Fauna Europaea version 1.1, available at http: // www. faunaeur. org [accessed September 2009 as version 1.3 of April 19 th 2007.]" type="url" year="2004">
Smetana (2004
<bibRefCitation id="EF9B4B15FFCBFFC0D08FFA73FCCB5AA9" author="Smetana, A." box="[767,780,1462,1488]" pageId="7" pageNumber="324" refId="ref14062" refString="Smetana, A. (2004 b) Staphylinidae Leptotyphlinae. In: Lobl, I. &amp; Smetana, A. (Eds), Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 568 - 578." type="book" year="2004">a</bibRefCitation>
</bibRefCitation>
, 2004b) continues to follow the classic arrangement of the genus proposed by
<bibRefCitation id="EF9B4B15FFCBFFC0D3DEFA18FDA45A8E" author="Coiffait, H." box="[430,611,1501,1527]" pageId="7" pageNumber="324" pagination="1 - 651" refId="ref11852" refString="Coiffait, H. (1972) Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites, Sousfamilles: Xantholininae et Leptotyphlinae. Nouvelle Revue d'Entomologie, Suppl. 2 (2), 1 - 651." type="journal article" year="1972">Coiffait (1972)</bibRefCitation>
and
<bibRefCitation id="EF9B4B15FFCBFFC0D0EFFA18FCF45A8E" author="Pace R. &amp; Fauna d'Italia, XXXIV" box="[671,819,1501,1527]" pageId="7" pageNumber="324" refId="ref13459" refString="Pace R., (1996). Fauna d'Italia, XXXIV. Coleoptera Staphylinidae Leptotyphlinae. Calderini, Bologna, 328 pp." type="book" year="1996">Pace (1996)</bibRefCitation>
. In our opinion the characters proposed by
<bibRefCitation id="EF9B4B15FFCBFFC0D74FFA18FF215967" author="Coiffait, H." pageId="7" pageNumber="324" pagination="1 - 651" refId="ref11852" refString="Coiffait, H. (1972) Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites, Sousfamilles: Xantholininae et Leptotyphlinae. Nouvelle Revue d'Entomologie, Suppl. 2 (2), 1 - 651." type="journal article" year="1972">Coiffait (1972)</bibRefCitation>
and
<bibRefCitation id="EF9B4B15FFCBFFC0D352F9C1FE755967" author="Pace R. &amp; Fauna d'Italia, XXXIV" box="[290,434,1540,1566]" pageId="7" pageNumber="324" refId="ref13459" refString="Pace R., (1996). Fauna d'Italia, XXXIV. Coleoptera Staphylinidae Leptotyphlinae. Calderini, Bologna, 328 pp." type="book" year="1996">Pace (1996)</bibRefCitation>
sometimes cause the placement of clearly allied species within separate subgenera, like in the emblematic case of the new species described above.
<taxonomicName id="4C0A4D67FFCBFFC0D1F8F9EEFBAB593D" authority="Fancello &amp; Hernando &amp; Leo, 2009" authorityName="Fancello &amp; Hernando &amp; Leo" authorityYear="2009" box="[904,1132,1579,1604]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="nardii" status="sp. nov.">
<emphasis id="B97EEAF6FFCBFFC0D1F8F9EEFBAB593D" box="[904,1132,1579,1604]" italics="true" pageId="7" pageNumber="324">Leptotyphlus nardii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D578DFFCBFFC0D604F9EFFB09593D" box="[1140,1230,1578,1604]" pageId="7" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
should indeed be ascribed to subgenus
<taxonomicName id="4C0A4D67FFCBFFC0D3E3F994FDFF5913" authorityName="Coiffait" authorityYear="1955" box="[403,568,1617,1642]" class="Insecta" family="Staphylinidae" genus="Stigmotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAF6FFCBFFC0D3E3F994FDFF5913" box="[403,568,1617,1642]" italics="true" pageId="7" pageNumber="324">Stigmotyphlus</emphasis>
</taxonomicName>
for the asymmetrical female genital armature and for the aedeagus with a large coil-shaped dorsal copulatory piece; however, from further examination it appears that
<taxonomicName id="4C0A4D67FFCBFFC0D6A5F9BDFAFD59E8" authority="Fancello &amp; Hernando &amp; Leo, 2009" authorityName="Fancello &amp; Hernando &amp; Leo" authorityYear="2009" box="[1237,1338,1656,1682]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="nardii" status="sp. nov.">
<emphasis id="B97EEAF6FFCBFFC0D6A5F9BDFB2259E8" box="[1237,1253,1656,1681]" italics="true" pageId="7" pageNumber="324">L</emphasis>
.
<emphasis id="B97EEAF6FFCBFFC0D684F9BDFAFD59E8" box="[1268,1338,1656,1681]" italics="true" pageId="7" pageNumber="324">nardii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D578DFFCBFFC0D732F9BDFA5B59EB" box="[1346,1436,1656,1682]" pageId="7" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
is clearly related to some peninsular species of
<taxonomicName id="4C0A4D67FFCBFFC0D0BBF95AFCB659C1" authorityName="Coiffait" authorityYear="1955" box="[715,881,1695,1720]" class="Insecta" family="Staphylinidae" genus="Stigmotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEAF6FFCBFFC0D0BBF95AFCB659C1" box="[715,881,1695,1720]" italics="true" pageId="7" pageNumber="324">Stigmotyphlus</emphasis>
</taxonomicName>
of the
<taxonomicName id="4C0A4D67FFCBFFC0D1B6F95AFB9E59C1" authorityName="Pace" authorityYear="1973" box="[966,1113,1694,1720]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="lessinicus">
<emphasis id="B97EEAF6FFCBFFC0D1B6F95AFC1159C1" box="[966,982,1695,1720]" italics="true" pageId="7" pageNumber="324">L</emphasis>
.
<emphasis id="B97EEAF6FFCBFFC0D196F95AFB9E59C1" box="[998,1113,1695,1720]" italics="true" pageId="7" pageNumber="324">lessinicus</emphasis>
</taxonomicName>
-group (
<taxonomicName id="4C0A4D67FFCBFFC0D6CAF95AFF1659A6" authority="Pace, 1977" authorityName="Pace" authorityYear="1977" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="molisensis">
<emphasis id="B97EEAF6FFCBFFC0D6CAF95AFB0D59C1" box="[1210,1226,1695,1720]" italics="true" pageId="7" pageNumber="324">L</emphasis>
.
<emphasis id="B97EEAF6FFCBFFC0D6AAF95AFA9259C1" box="[1242,1365,1695,1720]" italics="true" pageId="7" pageNumber="324">molisensis</emphasis>
<bibRefCitation id="EF9B4B15FFCBFFC0D72DF95BFF1659A6" author="Pace, R." pageId="7" pageNumber="324" pagination="155 - 201" refId="ref13312" refString="Pace, R. (1977) Dodici nuove specie di Leptotyphlinae dell'Italia peninsulare (Coleoptera Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 4, 155 - 201." type="journal article" year="1977">Pace, 1977</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0A4D67FFCBFFC0D37BF900FDEA59A6" authority="Pace, 1977" authorityName="Pace" authorityYear="1977" box="[267,557,1733,1759]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="campanus">
<emphasis id="B97EEAF6FFCBFFC0D37BF900FEDC59A7" box="[267,283,1733,1758]" italics="true" pageId="7" pageNumber="324">L</emphasis>
.
<emphasis id="B97EEAF6FFCBFFC0D35BF900FE6559A7" box="[299,418,1733,1758]" italics="true" pageId="7" pageNumber="324">campanus</emphasis>
<bibRefCitation id="EF9B4B15FFCBFFC0D3DAF900FDEA59A6" author="Pace, R." box="[426,557,1733,1759]" pageId="7" pageNumber="324" pagination="155 - 201" refId="ref13312" refString="Pace, R. (1977) Dodici nuove specie di Leptotyphlinae dell'Italia peninsulare (Coleoptera Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 4, 155 - 201." type="journal article" year="1977">Pace, 1977</bibRefCitation>
</taxonomicName>
, extremely similar with regard to all structures of the copulatory organ:
<emphasis id="B97EEAF6FFCBFFC0D70FF900FA5A59A7" box="[1407,1437,1733,1758]" italics="true" pageId="7" pageNumber="324">cf.</emphasis>
<bibRefCitation id="EF9B4B15FFCBFFC0D2E7F929FED3587F" author="Pace, R." box="[151,276,1772,1798]" pageId="7" pageNumber="324" pagination="155 - 201" refId="ref13312" refString="Pace, R. (1977) Dodici nuove specie di Leptotyphlinae dell'Italia peninsulare (Coleoptera Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 4, 155 - 201." type="journal article" year="1977">Pace 1977</bibRefCitation>
, figs 190–191), and to Sardo-Corsican species of
<taxonomicName id="4C0A4D67FFCBFFC0D111F929FC3F587C" authorityName="Fauvel" authorityYear="1874" box="[865,1016,1772,1797]" class="Insecta" family="Staphylinidae" genus="Leptotyphlus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B97EEAF6FFCBFFC0D111F929FC3F587C" box="[865,1016,1772,1797]" italics="true" pageId="7" pageNumber="324">Leptotyphlus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D578DFFCBFFC0D671F929FBF8587F" box="[1025,1087,1772,1798]" pageId="7" pageNumber="324" sensu="stricto">s. str.</taxonomicNameLabel>
of the
<taxonomicName id="4C0A4D67FFCBFFC0D6E4F929FAC8587C" authorityName="Coiffait" authorityYear="1955" box="[1172,1295,1772,1798]" class="Insecta" family="Brachyceridae" genus="Raymondiellus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="sardous">
<emphasis id="B97EEAF6FFCBFFC0D6E4F929FB63587C" box="[1172,1188,1772,1797]" italics="true" pageId="7" pageNumber="324">L</emphasis>
.
<emphasis id="B97EEAF6FFCBFFC0D6C3F929FAC8587C" box="[1203,1295,1772,1797]" italics="true" pageId="7" pageNumber="324">sardous</emphasis>
</taxonomicName>
-group, also with a coil-shaped dorsal copulatory piece (
<emphasis id="B97EEAF6FFCBFFC0D0E7F8D6FD6B5855" box="[663,684,1811,1836]" italics="true" pageId="7" pageNumber="324">cf</emphasis>
.
<bibRefCitation id="EF9B4B15FFCBFFC0D0CBF8D7FCF05855" author="Pace R. &amp; Fauna d'Italia, XXXIV" box="[699,823,1810,1836]" pageId="7" pageNumber="324" refId="ref13459" refString="Pace R., (1996). Fauna d'Italia, XXXIV. Coleoptera Staphylinidae Leptotyphlinae. Calderini, Bologna, 328 pp." type="book" year="1996">Pace 1996</bibRefCitation>
, figs 163–165).
</paragraph>
<paragraph id="8BB536E4FFCBFFC0D2B6F8FDFDA15897" blockId="7.[151,1437,534,2030]" pageId="7" pageNumber="324">
<emphasis id="B97EEAF6FFCBFFC0D2B6F8FDFE50582B" bold="true" box="[198,407,1848,1874]" pageId="7" pageNumber="324">Ecological notes.</emphasis>
The species was collected by washing soil in both Mediterranean maquis with
<taxonomicName id="4C0A4D67FFCBFFC0D74CF8FCFF395800" class="Magnoliopsida" family="Anacardiaceae" genus="Pistacia" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="324" phylum="Tracheophyta" rank="species" species="lentiscus">
<emphasis id="B97EEAF6FFCBFFC0D74CF8FCFF395800" italics="true" pageId="7" pageNumber="324">Pistacia lentiscus</emphasis>
</taxonomicName>
dominance and
<taxonomicName id="4C0A4D67FFCBFFC0D3B3F8A5FD9F5800" box="[451,600,1888,1913]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="324" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis id="B97EEAF6FFCBFFC0D3B3F8A5FD9F5800" box="[451,600,1888,1913]" italics="true" pageId="7" pageNumber="324">Quercus ilex</emphasis>
</taxonomicName>
woodland. Collecting sites are situated on Palaeozoic limestone soils. The following endogean beetles were collected together with the new species:
<taxonomicName id="4C0A4D67FFCBFFC0D65FF842FA5058D9" box="[1071,1431,1926,1952]" class="Insecta" family="Staphylinidae" genus="Batysciola" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="damryi">
<emphasis id="B97EEAF6FFCBFFC0D65FF842FAC258D9" box="[1071,1285,1927,1952]" italics="true" pageId="7" pageNumber="324">Batysciola damryi</emphasis>
(Leiodidae)
</taxonomicName>
,
<taxonomicName id="4C0A4D67FFCBFFC0D2E7F868FDF858BE" authorityName="Belon" authorityYear="1882" box="[151,575,1965,1991]" class="Insecta" family="Zopheridae" genus="Langelandia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="B97EEAF6FFCBFFC0D2E7F868FE5658BF" box="[151,401,1965,1990]" italics="true" pageId="7" pageNumber="324">Langelandia reitteri</emphasis>
(Zopheridae)
</taxonomicName>
,
<taxonomicName id="4C0A4D67FFCBFFC0D03EF868FC0158BE" box="[590,966,1965,1991]" class="Insecta" family="Curculionidae" genus="Torneuma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97EEAF6FFCBFFC0D03EF868FD0E58BF" box="[590,713,1965,1990]" italics="true" pageId="7" pageNumber="324">Torneuma</emphasis>
sp. (Curculionidae)
</taxonomicName>
,
<taxonomicName id="4C0A4D67FFCBFFC0D1A5F868FB1458BF" authorityName="Perris" authorityYear="1869" box="[981,1235,1965,1990]" class="Insecta" family="Brachyceridae" genus="Alaocyba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="species" species="carinulata">
<emphasis id="B97EEAF6FFCBFFC0D1A5F868FB1458BF" box="[981,1235,1965,1990]" italics="true" pageId="7" pageNumber="324">Alaocyba carinulata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0A4D67FFCBFFC0D695F868FDA55897" authority="sardous" authorityName="sardous (Perris" authorityYear="1869" class="Insecta" family="Brachyceridae" genus="Raymondiellus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="324" phylum="Arthropoda" rank="subSpecies" species="sardous" subSpecies="sardous">
<emphasis id="B97EEAF6FFCBFFC0D695F868FE925894" italics="true" pageId="7" pageNumber="324">Raymondiellus sardous sardous</emphasis>
(Raymondionymidae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C310656FFFC4FFCFD2B6FF52FAD85FCB" box="[198,1311,151,178]" pageId="8" pageNumber="325" type="etymology">
<paragraph id="8BB536E4FFC4FFCFD2B6FF52FAD85FCB" blockId="8.[198,1311,151,178]" box="[198,1311,151,178]" pageId="8" pageNumber="325">
<emphasis id="B97EEAF6FFC4FFCFD2B6FF52FE945FC8" bold="true" box="[198,339,151,177]" pageId="8" pageNumber="325">Etymology.</emphasis>
It is our pleasure to dedicate this species to our friend Gianluca Nardi (CNBFVR).
</paragraph>
</subSubSection>
</treatment>
</document>
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