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<document id="03C21947EC36D8629CC6A1AAD003DD2D" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629685057509" checkinUser="felipe" docAuthor="Scarbrough, A. G." docDate="2010" docId="038C87F8FFBCFFC8FF7B6E39E727FCDA" docLanguage="en" docName="zt02540p047.pdf" docOrigin="Zootaxa 2540" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ommatius prolixus Scarbrough 2010, n. sp." docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="FFB5FF80FF9CFFEAFFEC6A75E64FFF88" masterDocTitle="An overview of the Afrotropical Ommatiinae (Diptera: Asilidae) with a key to genera" masterLastPageNumber="47" masterPageNumber="1" pageNumber="33" updateTime="1699095850413" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0646276DE9062B24298619BA7C25031B">An overview of the Afrotropical Ommatiinae (Diptera: Asilidae) with a key to genera</mods:title>
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<treatment id="038C87F8FFBCFFC8FF7B6E39E727FCDA" ID-DOI="http://doi.org/10.5281/zenodo.5310864" ID-Zenodo-Dep="5310864" LSID="urn:lsid:plazi:treatment:038C87F8FFBCFFC8FF7B6E39E727FCDA" httpUri="http://treatment.plazi.org/id/038C87F8FFBCFFC8FF7B6E39E727FCDA" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
<subSubSection id="C33F6565FFBCFFCAFF7B6E39E79FFBED" box="[151,464,1099,1126]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8B9A36EEFFBCFFCAFF7B6E39E79FFBED" blockId="32.[151,464,1099,1160]" box="[151,464,1099,1126]" pageId="32" pageNumber="33">
<heading id="D0D28182FFBCFFCAFF7B6E39E79FFBED" bold="true" box="[151,464,1099,1126]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="B951EAFCFFBCFFCAFF7B6E39E79FFBED" bold="true" box="[151,464,1099,1126]" pageId="32" pageNumber="33">
<emphasis id="B951EAFCFFBCFFCAFF7B6E39E7C9FBEE" bold="true" box="[151,390,1100,1126]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="4C254D6DFFBCFFCAFF7B6E39E7CFFBEE" ID-CoL="8D3F6" authority="Scarbrough, 2010" authorityName="Scarbrough" authorityYear="2010" box="[151,384,1100,1126]" class="Insecta" family="Asilidae" genus="Ommatius" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="prolixus" status="sp. nov.">Ommatius prolixus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2625787FFBCFFCAFE616E3EE79FFBED" box="[397,464,1099,1125]" pageId="32" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FFBCFFC8FF7B6E1BE7EAFDAF" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
<paragraph id="8B9A36EEFFBCFFCAFF7B6E1BE76EFB00" blockId="32.[151,464,1099,1160]" box="[151,289,1134,1160]" pageId="32" pageNumber="33">
<figureCitation id="131E2A6BFFBCFFCAFF7B6E1BE76EFB00" box="[151,289,1134,1160]" captionStart="FIGURES 8388" captionStartId="33.[151,271,1971,1995]" captionTargetBox="[233,1373,489,1938]" captionTargetId="figure-152@33.[215,1380,451,1964]" captionTargetPageId="33" captionText="FIGURES 8388. Ommatius prolixus n. sp., male, wing and terminalia. 83. Right wing. 8486. Lateral, dorsal, and ventral views; 87. Aedeagus, lateral view; 88. Gonostylus." pageId="32" pageNumber="33">Figs. 8388</figureCitation>
</paragraph>
<paragraph id="8B9A36EEFFBCFFCAFF7B6ECDE260FA3B" blockId="32.[151,1436,1208,2025]" pageId="32" pageNumber="33">
<emphasis id="B951EAFCFFBCFFCAFF7B6ECDE767FB5A" bold="true" box="[151,296,1208,1234]" pageId="32" pageNumber="33">Description</emphasis>
, male. Body
<quantity id="4CDD9B0BFFBCFFCAFE2F6ECCE462FB5B" box="[451,557,1209,1235]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="32" pageNumber="33" unit="mm" value="11.3">11.3 mm</quantity>
; wing
<quantity id="4CDD9B0BFFBCFFCAFD976ECCE498FB5B" box="[635,727,1209,1235]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="32" pageNumber="33" unit="mm" value="8.3">8.3 mm</quantity>
.
<emphasis id="B951EAFCFFBCFFCAFD0E6ECCE56EFB5A" box="[738,801,1209,1234]" italics="true" pageId="32" pageNumber="33">Head</emphasis>
: Face gray tomentose with scattered vestiture; mystax pale yellow, sparse setae, 45 pencil-like bristles plus 9 longer, thinner, brown bristles; FHWR 1.0:7.5. Palpus, proboscis, and frons yellowish or whitish setose. Antenna brownish, scape entirely and base of pedicel brownish-yellow; brown setose; flagellum slightly shorter than scape and pedicel together; FWLR 1.0:1.9. Frons yellowish tomentose. Ocellar tubercle brown setose, 2 setae slightly shorter than scape and pedicel combined. Occiput yellowish tomentose with yellowish and white setae; postocular bristles mostly yellow, 1 3 dorsal bristles brown, apex of longest about 1/3 distance beyond eye margin.
</paragraph>
<paragraph id="8B9A36EEFFBCFFCBFF296FCAE5AFFE3F" blockId="32.[151,1436,1208,2025]" lastBlockId="33.[151,1436,152,439]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="B951EAFCFFBCFFCAFF296FCAE753FA50" box="[197,284,1471,1496]" italics="true" pageId="32" pageNumber="33">Thorax</emphasis>
: Pronotum brownish-gray or gray tomentose with whitish or yellowish vestiture. Mesonotum brown, postalar callus brownish-yellow, tomentum dense, yellowish or grayish; brown as follows: wide brown median stripe and 2 spots laterally, streak divided longitudinally and spots separated by yellowish tomentum narrowly; sparse brown setae medially, whitish setae laterally; 3 dorsocentral and 58 lateral bristles black. Scutellum gray tomentose with scattered white setae and 2 long black marginal bristles. Pleuron yellowish-gray or gray tomentose, vestiture pale yellow or white; anepimeral bristles thin, seta-like; ground color of metacoxa and metepisternum yellow. Halter yellow.
<emphasis id="B951EAFCFFBCFFCAFC826CEAE5E4F930" box="[878,939,1695,1720]" italics="true" pageId="32" pageNumber="33">Wing</emphasis>
(
<figureCitation id="131E2A6BFFBCFFCAFC576CEBE25AF930" box="[955,1045,1694,1720]" captionStart="FIGURES 8388" captionStartId="33.[151,271,1971,1995]" captionTargetBox="[233,1373,489,1938]" captionTargetId="figure-152@33.[215,1380,451,1964]" captionTargetPageId="33" captionText="FIGURES 8388. Ommatius prolixus n. sp., male, wing and terminalia. 83. Right wing. 8486. Lateral, dorsal, and ventral views; 87. Aedeagus, lateral view; 88. Gonostylus." pageId="32" pageNumber="33">Fig. 83</figureCitation>
): Hyaline; apical 1/3 and narrow posterior margin with dense microtrichia, sparse or absent otherwise. Cell r
<subScript id="17A134ABFFBCFFCAFBD86CA1E273F96A" attach="left" box="[1076,1084,1748,1762]" fontSize="6" pageId="32" pageNumber="33">4</subScript>
narrowly triangular, slightly sigmoid apically, base well beyond apex of cell d. Crossvein r-m beyond middle of cell d, about as long as CuA
<subScript id="17A134ABFFBCFFCAFF236D54E698F8A7" attach="left" box="[207,215,1825,1839]" fontSize="6" pageId="32" pageNumber="33">1</subScript>
+M
<subScript id="17A134ABFFBCFFCAFEEF6D54E744F8A7" attach="right" box="[259,267,1825,1839]" fontSize="6" pageId="32" pageNumber="33">3</subScript>
. Cell m
<subScript id="17A134ABFFBCFFCAFE816D54E73AF8A7" attach="left" box="[365,373,1825,1839]" fontSize="6" pageId="32" pageNumber="33">1</subScript>
wide, narrow base, vein M
<subScript id="17A134ABFFBCFFCAFD2D6D54E486F8A7" attach="left" box="[705,713,1825,1839]" fontSize="6" pageId="32" pageNumber="33">1</subScript>
strongly arched anteriorly; m
<subScript id="17A134ABFFBCFFCAFBDE6D54E275F8A7" attach="both" box="[1074,1082,1825,1839]" fontSize="6" pageId="32" pageNumber="33">1</subScript>
WR 1.0:2.6:2.5.
<emphasis id="B951EAFCFFBCFFCAFAE86D64E308F8A2" box="[1284,1351,1809,1834]" italics="true" pageId="32" pageNumber="33">Legs:</emphasis>
Coxae yellowish tomentose with pale yellow or whitish vestiture; coxae yellow, meso- and metathoracic coxae brownish-yellow anteriorly. Trochanters yellow, apical margin blackish; metathoracic trochanter with whitish bristles. Femora slender; anterior bristles mostly brown. Pro- and mesothoracic femora yellow, slightly brownish-yellow anteriorly; prothoracic femur posteriorly and ventrally with abundant, long, wavy, yellowish setae, including a ventral row, sparse brown setae posterodorsally; basal 1/3 of prothoracic femur with 23 bristles posteriorly. Mesothoracic femur with thin bristles ventrally, those posteroventrally slightly thicker; sparse long, thin, wavy setae posteriorly; preapical posterodorsal bristle brown. Metathoracic femur mostly brown anteriorly and dorsally, apical 1/2 ventrally and posteriorly brown, yellow or brownish-yellow elsewhere; MFWLR 1.0:6.9. Pro- and mesothoracic tibiae yellow with mostly yellow vestiture; prothoracic tibia with a fringe of abundant, yellowish setae and long, thin, yellow bristles laterally, setae not unusually dense. Metathoracic tibia with apical 1/4 brown, and only brown bristles. Tarsi with basal tarsomeres yellow, narrow apices brown; apical 4 tarsomeres brown with narrow yellow bases, bristles brown; basal tarsomere of prothoracic tarsus with 2 yellow bristles and thin fringe of yellow setae.
</paragraph>
<caption id="DF5A6666FFBDFFCBFF7B6DC6E549F862" pageId="33" pageNumber="34" startId="33.[151,271,1971,1995]" targetBox="[233,1373,489,1938]" targetPageId="33">
<paragraph id="8B9A36EEFFBDFFCBFF7B6DC6E549F862" blockId="33.[151,1436,1971,2026]" pageId="33" pageNumber="34">
<emphasis id="B951EAFCFFBDFFCBFF7B6DC6E72DF843" bold="true" box="[151,354,1971,1995]" pageId="33" pageNumber="34">FIGURES 8388.</emphasis>
<taxonomicName id="4C254D6DFFBDFFCBFE866DC6E474F842" authority="Scarbrough, 2010" authorityName="Scarbrough" authorityYear="2010" box="[362,571,1971,1994]" class="Insecta" family="Asilidae" genus="Ommatius" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="prolixus" status="sp. nov.">
<emphasis id="B951EAFCFFBDFFCBFE866DC6E474F842" box="[362,571,1971,1994]" italics="true" pageId="33" pageNumber="34">Ommatius prolixus</emphasis>
</taxonomicName>
<emphasis id="B951EAFCFFBDFFCBFDAF6DC6E4C7F843" bold="true" box="[579,648,1971,1995]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="A2625787FFBDFFCBFDAF6DC6E4CDF843" box="[579,642,1971,1995]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
male, wing and terminalia. 83. Right wing. 8486. Lateral, dorsal, and ventral views; 87. Aedeagus, lateral view; 88. Gonostylus.
</paragraph>
</caption>
<paragraph id="8B9A36EEFFBEFFC8FF296AEDE581FD8A" blockId="34.[151,1436,152,850]" pageId="34" pageNumber="35">
<emphasis id="B951EAFCFFBEFFC8FF296AEDE77BFF39" box="[197,308,152,177]" italics="true" pageId="34" pageNumber="35">Abdomen</emphasis>
: Dark brown with narrow apical margins of segments yellow; mostly yellowish-gray tomentose, white setose; tomentum becoming slightly brownish-yellow and thin on apical 34 segments, apical 12 tergites sunshiny; apical 45 tergites dorsally and sternites 78 with numerous brown setae, brown setae sparse on preceding segments; apical margin of tergites 16 with a row of yellow setae,
<quantity id="4CDD9B0BFFBEFFC8FB4A6B7DE2BEFEAA" box="[1190,1265,264,290]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="34" pageNumber="35" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">23 in</quantity>
apical corners longest and thickest.
<emphasis id="B951EAFCFFBEFFC8FE786B58E454FECE" box="[404,539,301,326]" italics="true" pageId="34" pageNumber="35">Terminalia</emphasis>
(
<figureCitation id="131E2A6BFFBEFFC8FDC06B58E48CFECF" box="[556,707,301,327]" captionStart="FIGURES 8388" captionStartId="33.[151,271,1971,1995]" captionTargetBox="[233,1373,489,1938]" captionTargetId="figure-152@33.[215,1380,451,1964]" captionTargetPageId="33" captionText="FIGURES 8388. Ommatius prolixus n. sp., male, wing and terminalia. 83. Right wing. 8486. Lateral, dorsal, and ventral views; 87. Aedeagus, lateral view; 88. Gonostylus." pageId="34" pageNumber="35">Figs. 8488</figureCitation>
): Dark brown, wider than abdominal segment 8, as long as combined length of segments 78. Epandrium wide in lateral view curved or slightly angled posteriorly, apices contrastingly narrow and short, somewhat truncate in dorsal view. Gonostylus subrectangular, wide basally, narrow and hooked apically; concave laterally. Gonocoxite fused with hypandrium basally, inner margin elongate with 12 yellow setae apically. Hypandrium with apical margin broadly rounded, slightly angled preapically, with narrow U-shaped band of dense, brown setae.
</paragraph>
<paragraph id="8B9A36EEFFBEFFC8FF296878E7EAFDAF" blockId="34.[151,1436,152,850]" box="[197,421,525,551]" pageId="34" pageNumber="35">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C33F6565FFBEFFC8FF296847E4F7FDFA" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8B9A36EEFFBEFFC8FF296847E4F7FDFA" blockId="34.[151,1436,152,850]" pageId="34" pageNumber="35">
<emphasis id="B951EAFCFFBEFFC8FF296847E74CFDC4" bold="true" box="[197,259,562,588]" pageId="34" pageNumber="35">Type</emphasis>
.
<materialsCitation id="3B4D3CB3FFBEFFC8FEFF6847E4FBFDFA" collectingDate="1980-11-24" collectingDateMax="1980-11-27" collectingDateMin="1980-11-24" collectionCode="NMSA" collectorName="Stuckenberg &amp; Londt &amp; Ad" country="Malawi" elevation="1500" location="Plateau" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Zomba" typeStatus="holotype">
<emphasis id="B951EAFCFFBEFFC8FEFF6847E7DCFDC4" bold="true" box="[275,403,562,588]" pageId="34" pageNumber="35">
<collectingCountry id="F332767EFFBEFFC8FEFF6847E7DCFDC4" box="[275,403,562,588]" name="Malawi" pageId="34" pageNumber="35">MALAWI</collectingCountry>
</emphasis>
:
<collectingRegion id="49E1F80CFFBEFFC8FE4F6847E7B8FDC4" box="[419,503,562,588]" country="Malawi" name="Zomba" pageId="34" pageNumber="35">Zomba</collectingRegion>
<location id="8EFA6035FFBEFFC8FE136847E419FDC4" LSID="urn:lsid:plazi:treatment:038C87F8FFBCFFC8FF7B6E39E727FCDA:8EFA6035FFBEFFC8FE136847E419FDC4" box="[511,598,562,588]" country="Malawi" name="Plateau" pageId="34" pageNumber="35" stateProvince="Zomba">Plateau</location>
[1523ʹS 3520ʹE _] /
<date id="FF9B102EFFBEFFC8FCB76847E248FDC4" box="[859,1031,562,588]" pageId="34" pageNumber="35" value="1980-11-24" valueMax="1980-11-27" valueMin="1980-11-24">
<collectingDate id="EFDFE9C6FFBEFFC8FCB76847E248FDC4" box="[859,1031,562,588]" pageId="34" pageNumber="35" value="1980-11-24" valueMax="1980-11-27" valueMin="1980-11-24">2427.xi.1980</collectingDate>
</date>
<quantity id="4CDD9B0BFFBEFFC8FBE36847E222FDC4" box="[1039,1133,562,588]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="m" value="1500.0">
<elevation id="0008D1DDFFBEFFC8FBE36847E222FDC4" box="[1039,1133,562,588]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="m" value="1500.0">1500 m</elevation>
</quantity>
, /
<collectorName id="26D05338FFBEFFC8FB666847E352FDC4" box="[1162,1309,562,588]" pageId="34" pageNumber="35">Stuckenberg</collectorName>
&amp;
<collectorName id="26D05338FFBEFFC8FAA96847E3C3FDC4" box="[1349,1420,562,588]" pageId="34" pageNumber="35">Londt</collectorName>
/ 1535
<collectorName id="26D05338FFBEFFC8FF3C682DE6BAFDFA" box="[208,245,600,626]" pageId="34" pageNumber="35">Ad</collectorName>
montane forest (
<emphasis id="B951EAFCFFBEFFC8FE506822E468FDF9" bold="true" box="[444,551,599,625]" pageId="34" pageNumber="35">
<typeStatus id="549E884CFFBEFFC8FE506822E468FDF9" box="[444,551,599,625]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
</emphasis>
<specimenCount id="9D23FD67FFBEFFC8FDC36822E40CFDE3" box="[559,579,599,619]" pageId="34" pageNumber="35" type="male"></specimenCount>
,
<collectionCode id="ED34AE2BFFBEFFC8FDBE682DE4E3FDFA" box="[594,684,600,626]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="34" pageNumber="35" type="Museum">NMSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FFBEFFC8FF296808E3C7FD1F" box="[197,1416,637,663]" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8B9A36EEFFBEFFC8FF296808E3C7FD1F" blockId="34.[151,1436,152,850]" box="[197,1416,637,663]" pageId="34" pageNumber="35">
<emphasis id="B951EAFCFFBEFFC8FF296808E705FD1F" bold="true" box="[197,330,637,663]" pageId="34" pageNumber="35">Etymology</emphasis>
. Latin
<taxonomicName id="4C254D6DFFBEFFC8FE726808E7B2FD1E" authorityName="Scarbrough" authorityYear="2010" box="[414,509,637,662]" class="Insecta" family="Asilidae" genus="Ommatius" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="B951EAFCFFBEFFC8FE726808E7B2FD1E" box="[414,509,637,662]" italics="true" pageId="34" pageNumber="35">prolixus</emphasis>
</taxonomicName>
, for broad or wide, referring to the diameter of the epandrium in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FFBEFFC8FF2968D7E298FD34" box="[197,1239,674,700]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8B9A36EEFFBEFFC8FF2968D7E298FD34" blockId="34.[151,1436,152,850]" box="[197,1239,674,700]" pageId="34" pageNumber="35">
<emphasis id="B951EAFCFFBEFFC8FF2968D7E711FD34" bold="true" box="[197,350,674,700]" pageId="34" pageNumber="35">Distribution</emphasis>
.Collected in November in a montane forest on the
<collectingRegion id="49E1F80CFFBEFFC8FC5A68D7E246FD34" box="[950,1033,674,700]" country="Malawi" name="Zomba" pageId="34" pageNumber="35">Zomba</collectingRegion>
Plateau,
<collectingCountry id="F332767EFFBEFFC8FB9A68D7E29CFD34" box="[1142,1235,674,700]" name="Malawi" pageId="34" pageNumber="35">Malawi</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F6565FFBEFFC8FF2968B2E727FCDA" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8B9A36EEFFBEFFC8FF2968B2E727FCDA" blockId="34.[151,1436,152,850]" pageId="34" pageNumber="35">
<emphasis id="B951EAFCFFBEFFC8FF2968B2E778FD69" bold="true" box="[197,311,711,737]" pageId="34" pageNumber="35">Remarks</emphasis>
. The sparse vestiture of the lower face, fringe of setae on the prothoracic leg, and characters of the terminalia (
<figureCitation id="131E2A6BFFBEFFC8FEBB6898E7A2FC8F" box="[343,493,749,775]" captionStart="FIGURES 8388" captionStartId="33.[151,271,1971,1995]" captionTargetBox="[233,1373,489,1938]" captionTargetId="figure-152@33.[215,1380,451,1964]" captionTargetPageId="33" captionText="FIGURES 8388. Ommatius prolixus n. sp., male, wing and terminalia. 83. Right wing. 8486. Lateral, dorsal, and ventral views; 87. Aedeagus, lateral view; 88. Gonostylus." pageId="34" pageNumber="35">Figs. 8488</figureCitation>
) distinguishes the male of
<taxonomicName id="4C254D6DFFBEFFC8FCAF6898E581FC8E" authorityName="Scarbrough" authorityYear="2010" box="[835,974,749,774]" class="Insecta" family="Asilidae" genus="Ommatius" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="prolixus">
<emphasis id="B951EAFCFFBEFFC8FCAF6898E581FC8E" box="[835,974,749,774]" italics="true" pageId="34" pageNumber="35">O. prolixus</emphasis>
</taxonomicName>
from congeners. The male is further distinguished by the wide epandrium in lateral view and blunt apex, broadly rounded hypandrium, and shape of the gonostylus.
</paragraph>
</subSubSection>
</treatment>
</document>