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<document id="CC9FE54781023E2659D96B6524826618" ID-CLB-Dataset="8176" ID-DOI="10.5852/ejt.2021.734.1233" ID-GBIF-Dataset="fd8f5cb9-13e9-4c90-ac15-502fa7ee8d38" ID-ISSN="2118-9773" ID-Zenodo-Dep="4542107" ID-ZooBank="663A0C2D-E3CF-4B0C-8BA1-566B174829A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613423098920" checkinUser="plazi" docAuthor="Montoya-Molina, Santiago &amp; Vaz-De, Fernando Z." docDate="2021" docId="03850669FFBEFFE1FE0D225338ABCCBE" docLanguage="en" docName="ejt-734_1-64.pdf" docOrigin="European Journal of Taxonomy 734" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius (Luederwaldtinia) triquetrus" docType="treatment" docVersion="9" lastPageNumber="48" masterDocId="FFBC7E11FF90FFCEFFAD26453B59CB56" masterDocTitle="Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="47" updateTime="1698929631379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BC14856ABCB26A13768BBE788380FAC9">Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="13309B76D205FDA50530940A1F524EEA">Montoya-Molina, Santiago</mods:namePart>
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<mods:affiliation id="8E68094D2E30795CA3CC7273952EC239">Federal University of Mato Grosso, Institute of Biosciences, Programa de Pós Graduaç „ o em Ecología e Conservaç „ o da Biodiversidade, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil. Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 129, Praha-Suchdol, 16500, Czech Republic. Federal University of Mato Grosso, Institute of Biosciences, Department of Biology and Zoology, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil.</mods:affiliation>
<mods:nameIdentifier id="308FB68488712F97A2CC213AC38BEC73" type="email">horomo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="23C6DCEF2CD8EA9136F4450BC610DF77">Vaz-De, Fernando Z.</mods:namePart>
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<mods:title id="60089E0964B4A28094444F792A03CD30">European Journal of Taxonomy</mods:title>
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<mods:date id="E4A0647587BBD5EF974A06CDB4F3BAD8">2021</mods:date>
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<treatment id="03850669FFBEFFE1FE0D225338ABCCBE" ID-DOI="http://doi.org/10.5281/zenodo.4545863" ID-GBIF-Taxon="177420802" ID-Zenodo-Dep="4545863" LSID="urn:lsid:plazi:treatment:03850669FFBEFFE1FE0D225338ABCCBE" httpUri="http://treatment.plazi.org/id/03850669FFBEFFE1FE0D225338ABCCBE" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection id="C336E4F4FFBEFFE0FE0D2253382ECF00" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8B93B77FFFBEFFE0FE0D2253382ECF00" blockId="46.[416,1171,1046,1110]" pageId="46" pageNumber="47">
<taxonomicName id="4C2CCCFCFFBEFFE0FE0D22533FCACF67" ID-CoL="35ND7" authority="(Luederwaldt, 1923)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1923" box="[416,1171,1046,1073]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="triquetrus" subGenus="Luederwaldtinia">
<emphasis id="B9586B6DFFBEFFE0FE0D22533962CF67" bold="true" box="[416,571,1046,1073]" italics="true" pageId="46" pageNumber="47">Dichotomius</emphasis>
(
<emphasis id="B9586B6DFFBEFFE0FDE12253384ACF67" bold="true" box="[588,787,1046,1073]" italics="true" pageId="46" pageNumber="47">Luederwaldtinia</emphasis>
)
<emphasis id="B9586B6DFFBEFFE0FC89225238C2CF67" bold="true" box="[804,923,1047,1073]" italics="true" pageId="46" pageNumber="47">triquetrus</emphasis>
(
<bibRefCitation id="EFBDCA8EFFBEFFE0FC0722523FD2CF67" author="Luederwaldt H." box="[938,1163,1047,1073]" pageId="46" pageNumber="47" pagination="1 - 13" refId="ref30108" refString="Luederwaldt H. 1923. Neue Pinotus - Arten. Revista do Museu Paulista 14: 1 - 13." type="journal article" year="1923">Luederwaldt, 1923</bibRefCitation>
)
</taxonomicName>
<figureCitation id="1317ABFAFFBEFFE0FD1122793848CF00" box="[700,785,1084,1110]" captionStart="Fig" captionStartId="48.[189,232,1928,1954]" captionTargetBox="[193,1370,275,1892]" captionTargetId="figure-63@48.[189,1374,265,1894]" captionTargetPageId="48" captionText="Fig. 16. Dichotomius triquetrus (Luederwaldt, 1923). AB. Dorsal habitus. A. ♂. B. ♀. D. Lectotype, ♀ (MZSP). C. Lateral view of the aedeagus. D. Dorsal view of the head. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Elytra." figureDoi="http://doi.org/10.5281/zenodo.4542139" httpUri="https://zenodo.org/record/4542139/files/figure.png" pageId="46" pageNumber="47">Fig. 16</figureCitation>
, Map 14
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBEFFE0FF1022C5388CCE54" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="8B93B77FFFBEFFE0FF1022C538FACFCC" blockId="46.[189,931,1151,1178]" box="[189,931,1151,1178]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB3C9051FFBEFFE0FF1022C538FACFCC" box="[189,931,1151,1178]" pageId="46" pageNumber="47">
<treatmentCitation id="0A8D916EFFBEFFE0FF1022C538FACFCC" author="Luederwaldt" box="[189,931,1151,1178]" page="11" pageId="46" pageNumber="47" year="1923">
<taxonomicName id="4C2CCCFCFFBEFFE0FF1022C539C2CFCF" ID-CoL="4HZWV" authority="Luederwaldt, 1923: 11" authorityName="Luederwaldt" authorityPageNumber="11" authorityYear="1923" box="[189,667,1151,1177]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="triquetrus">
<emphasis id="B9586B6DFFBEFFE0FF1022C53ACBCFCF" box="[189,402,1152,1177]" italics="true" pageId="46" pageNumber="47">Pinotus triquetrus</emphasis>
<bibRefCitation id="EFBDCA8EFFBEFFE0FE34223A39C2CFCF" author="Luederwaldt H." box="[409,667,1151,1177]" pageId="46" pageNumber="47" pagination="1 - 13" refId="ref30108" refString="Luederwaldt H. 1923. Neue Pinotus - Arten. Revista do Museu Paulista 14: 1 - 13." type="journal article" year="1923">Luederwaldt, 1923:11</bibRefCitation>
</taxonomicName>
(original description).
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF1022863991CF8A" blockId="46.[189,981,1218,1282]" box="[189,712,1218,1245]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB3C9051FFBEFFE0FF1022863991CF8A" box="[189,712,1218,1245]" pageId="46" pageNumber="47">
<taxonomicName id="4C2CCCFCFFBEFFE0FF1022863ACBCF8A" box="[189,402,1219,1244]" class="Insecta" family="Scarabaeidae" genus="Pintous" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="triquetrus">
<emphasis id="B9586B6DFFBEFFE0FF1022863ACBCF8A" box="[189,402,1219,1244]" italics="true" pageId="46" pageNumber="47">Pintous triquetrus</emphasis>
</taxonomicName>
<treatmentCitation id="0A8D916EFFBEFFE0FE022287399CCF8A" author="Blackwelder R. E." box="[431,709,1218,1244]" page="208" pageId="46" pageNumber="47" year="1944">
<bibRefCitation id="EFBDCA8EFFBEFFE0FE022287399CCF8A" author="Blackwelder R. E." box="[431,709,1218,1244]" pageId="46" pageNumber="47" pagination="189 - 341" refId="ref29526" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 208</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF1022A2388CCE54" blockId="46.[189,981,1218,1282]" box="[189,981,1255,1282]" pageId="46" pageNumber="47">
<taxonomicName id="4C2CCCFCFFBEFFE0FF1022A23A97CE57" ID-CoL="35ND7" baseAuthorityName="Luederwaldt" baseAuthorityYear="1923" box="[189,462,1255,1281]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="triquetrus">
<emphasis id="B9586B6DFFBEFFE0FF1022A23A97CE57" box="[189,462,1255,1281]" italics="true" pageId="46" pageNumber="47">Dichotomius triquetrus</emphasis>
</taxonomicName>
<bibRefCitation id="EFBDCA8EFFBEFFE0FE4622A239BFCE57" author="Schoolmeesters P." box="[491,742,1255,1281]" pageId="46" pageNumber="47" pagination="2016" refId="ref30865" refString="Schoolmeesters P. 2018. Scarabs: World Scarabaeidae Database. In: Roskov Y., Abucay L., Orrell T., Nicolson D., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R. E., Decock W. &amp; De Wever A. (eds) Species 2000 &amp; ITIS Catalogue of Life, 2016 Annual Checklist. Available from www. catalogueoflife. org / annual-checklist / 2016 [accessed 29 Jan. 2021]." type="book chapter" year="2018">Schoolmeesters 2018</bibRefCitation>
(
<emphasis id="B9586B6DFFBEFFE0FD5B22A2389DCE57" box="[758,964,1255,1281]" italics="true" pageId="46" pageNumber="47">Catalogue of Life</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBEFFE0FF10236F392DCD78" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8B93B77FFFBEFFE0FF10236F3A6FCE12" blockId="46.[189,310,1322,1348]" box="[189,310,1322,1348]" pageId="46" pageNumber="47">
<heading id="D0DB0013FFBEFFE0FF10236F3A6FCE12" bold="true" box="[189,310,1322,1348]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B9586B6DFFBEFFE0FF10236F3A6FCE12" bold="true" box="[189,310,1322,1348]" pageId="46" pageNumber="47">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF10231E392DCD78" blockId="46.[188,1399,1371,1582]" pageId="46" pageNumber="47">
<taxonomicName id="4C2CCCFCFFBEFFE0FF10231E3A8ACE23" baseAuthorityName="Luederwaldt" baseAuthorityYear="1923" box="[189,467,1371,1397]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="triquetrus">
<emphasis id="B9586B6DFFBEFFE0FF10231E3A8ACE23" box="[189,467,1371,1397]" italics="true" pageId="46" pageNumber="47">Dichotomius triquetrus</emphasis>
</taxonomicName>
is separated from other species in the group by the following combination of characters: dorsal and ventral surface reddish to dark brown. Anterior clypeal margin with deep emargination separating two obtuse teeth. Head interocular space displaying flat punctures, resembling rough; distinct ocellate punctuation near eyes. Pronotum medial disc with fine punctures, anterolateral edges with well-impressed punctures, ocellate punctuation along margins. Elytra striae strongly marked, interstriae microsculpture shagreened
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBEFFE1FF10201238E3CA30" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8B93B77FFFBEFFE0FF1020123AF3CD27" blockId="46.[189,426,1623,1649]" box="[189,426,1623,1649]" pageId="46" pageNumber="47">
<heading id="D0DB0013FFBEFFE0FF1020123AF3CD27" bold="true" box="[189,426,1623,1649]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B9586B6DFFBEFFE0FF1020123AF3CD27" bold="true" box="[189,426,1623,1649]" pageId="46" pageNumber="47">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF4720CD3961CDF5" blockId="46.[189,1399,1672,1772]" box="[234,568,1672,1699]" pageId="46" pageNumber="47">
<emphasis id="B9586B6DFFBEFFE0FF4720CD3A3FCDF4" bold="true" box="[234,358,1672,1698]" pageId="46" pageNumber="47">
<typeStatus id="549709DDFFBEFFE0FF4720CD3A3FCDF4" box="[234,358,1672,1698]" pageId="46" pageNumber="47" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF1020E83F31CDBA" blockId="46.[189,1399,1672,1772]" pageId="46" pageNumber="47">
<materialsCitation id="3B44BD22FFBEFFE0FF1020E83F31CDBA" ID-GBIF-Occurrence="3037418471" collectionCode="MZSP" country="Trinidad and Tobago" location="TRINIDAD AND TOBAGO" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<collectingCountry id="F33BF7EFFFBEFFE0FF1020E83955CD91" box="[189,524,1709,1735]" name="Trinidad and Tobago" pageId="46" pageNumber="47">TRINIDAD AND TOBAGO</collectingCountry>
<specimenCount id="9D2A7CF6FFBEFFE0FD8D20E8396ECD91" box="[544,567,1709,1735]" pageId="46" pageNumber="47" type="female"></specimenCount>
; “Caparo Trinidad Heyne, Berlin-Wilm.” [white, p]/ “
<taxonomicName id="4C2CCCFCFFBEFFE0FB0920EB3E2ECD91" authority="Montoya-Molina &amp; Vaz-De, 2021" authorityName="Montoya-Molina &amp; Vaz-De" authorityYear="2021" box="[1188,1399,1710,1735]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="triquetrus" status="sp. nov.">
<emphasis id="B9586B6DFFBEFFE0FB0920EB3E2ECD91" box="[1188,1399,1710,1735]" italics="true" pageId="46" pageNumber="47">Pinotus triquetrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BD616FFBEFFE0FF1020973BA5CDBA" box="[189,252,1746,1772]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
L̹d.det.23” [white, hw]/ “18188” [white, p] /[blue rounded label];
<collectionCode id="ED3D2FBAFFBEFFE0FBBA20973F3DCDBA" box="[1047,1124,1746,1772]" collectionName="Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="46" pageNumber="47" type="Museum">MZSP</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF4721503AB8CC79" blockId="46.[189,1399,1813,2025]" box="[234,481,1813,1839]" pageId="46" pageNumber="47">
<emphasis id="B9586B6DFFBEFFE0FF4721503AB8CC79" bold="true" box="[234,481,1813,1839]" pageId="46" pageNumber="47">Additional material</emphasis>
</paragraph>
<paragraph id="8B93B77FFFBEFFE0FF10217F3A9ECCBE" blockId="46.[189,1399,1813,2025]" pageId="46" pageNumber="47">
<materialsCitation id="3B44BD22FFBEFFE0FF10217F38F4CCC8" ID-GBIF-Occurrence="3037418559" collectingDate="2000-06-03" collectingMethod="flight interception trap" collectionCode="CEMT" collectorName="S. Alistair &amp; Ramsdale" country="Trinidad and Tobago" county="St. George Co." elevation="244" latitude="10.692778" location="Simla Research Station" longLatPrecision="21" longitude="-61.289444" municipality="Range" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="1" stateProvince="Arima">
<collectingCountry id="F33BF7EFFFBEFFE0FF10217F3948CC02" box="[189,529,1850,1876]" name="Trinidad and Tobago" pageId="46" pageNumber="47">TRINIDAD AND TOBAGO</collectingCountry>
<specimenCount id="9D2A7CF6FFBEFFE0FD84217F390CCC02" box="[553,597,1848,1876]" pageId="46" pageNumber="47">1 ♂</specimenCount>
,
<specimenCount id="9D2A7CF6FFBEFFE0FDC9217F39C8CC03" box="[612,657,1850,1877]" pageId="46" pageNumber="47" type="female">1 ♀</specimenCount>
;
<collectingCounty id="62F2CFF3FFBEFFE0FD0D217F3809CC02" box="[672,848,1850,1876]" pageId="46" pageNumber="47">St. George Co.</collectingCounty>
,
<collectingMunicipality id="6BF72D05FFBEFFE0FCF1217E38F3CC03" box="[860,938,1851,1877]" pageId="46" pageNumber="47">Range</collectingMunicipality>
,
<location id="8EF3E1A4FFBEFFE0FC1B217F3F93CC02" LSID="urn:lsid:plazi:treatment:03850669FFBEFFE1FE0D225338ABCCBE:8EF3E1A4FFBEFFE0FC1B217F3F93CC02" box="[950,1226,1850,1876]" country="Trinidad and Tobago" county="St. George Co." latitude="10.692778" longLatPrecision="21" longitude="-61.289444" municipality="Range" name="Simla Research Station" pageId="46" pageNumber="47" stateProvince="Arima">Simla Research Station</location>
,
<location id="8EF3E1A4FFBEFFE0FB79217F3E2BCC02" LSID="urn:lsid:plazi:treatment:03850669FFBEFFE1FE0D225338ABCCBE:8EF3E1A4FFBEFFE0FB79217F3E2BCC02" box="[1236,1394,1850,1876]" country="Trinidad and Tobago" county="St. George Co." latitude="10.692778" longLatPrecision="21" longitude="-61.289444" municipality="Range" name="Arima Valley" pageId="46" pageNumber="47" stateProvince="Arima">Arima Valley</location>
,
<location id="8EF3E1A4FFBEFFE0FF1021253A6BCC2C" LSID="urn:lsid:plazi:treatment:03850669FFBEFFE1FE0D225338ABCCBE:8EF3E1A4FFBEFFE0FF1021253A6BCC2C" box="[189,306,1888,1914]" country="Trinidad and Tobago" county="St. George Co." latitude="10.692778" longLatPrecision="21" longitude="-61.289444" municipality="Range" name="N. Range" pageId="46" pageNumber="47" stateProvince="Arima">N. Range</location>
;
<elevation id="0001504CFFBEFFE0FE92211A3A90CC2F" box="[319,457,1887,1913]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="46" pageNumber="47" unit="ft" value="800.0">
<quantity id="4CD41A9AFFBEFFE0FE92211A3ADECC2F" box="[319,391,1887,1913]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="46" pageNumber="47" unit="ft" value="800.0">800 ft</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE18D1B8FFBEFFE0FE75211B3937CC2C" box="[472,622,1886,1914]" degrees="10" direction="north" minutes="41" orientation="latitude" pageId="46" pageNumber="47" precision="15" seconds="34" value="10.692778">10º4134″ N</geoCoordinate>
,
<geoCoordinate id="EE18D1B8FFBEFFE0FDD1211B384FCC2C" box="[636,790,1886,1914]" degrees="61" direction="west" minutes="17" orientation="longitude" pageId="46" pageNumber="47" precision="15" seconds="22" value="-61.289444">61º1722″ W</geoCoordinate>
; FIT [
<collectingMethod id="526DCF68FFBEFFE0FCCD211A3F2DCC2C" box="[864,1140,1887,1914]" pageId="46" pageNumber="47">flight interception trap</collectingMethod>
]; premontane tropical rainforest;
<date id="FF9291BFFFBEFFE0FEEC21C13A8DCCC8" box="[321,468,1924,1951]" pageId="46" pageNumber="47" value="2000-06-03">
<collectingDate id="EFD66857FFBEFFE0FEEC21C13A8DCCC8" box="[321,468,1924,1951]" pageId="46" pageNumber="47" value="2000-06-03">3 Jun. 2000</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBEFFE0FE4921C13907CCC8" box="[484,606,1924,1950]" pageId="46" pageNumber="47">S. Alistair</collectorName>
and
<collectorName id="26D9D2A9FFBEFFE0FD3321C1384BCCC8" box="[670,786,1924,1950]" pageId="46" pageNumber="47">Ramsdale</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBEFFE0FCF121C138F4CCC8" box="[860,941,1924,1950]" pageId="46" pageNumber="47">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBEFFE0FC6321C13A4CCC92" ID-GBIF-Occurrence="3037418340" collectionCode="BMNH" collectorName="F. Birch" country="Trinidad and Tobago" location="Trinidad" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D2A7CF6FFBEFFE0FC6321C138A6CCC9" box="[974,1023,1924,1951]" pageId="46" pageNumber="47" type="female">1 ♀</specimenCount>
;
<location id="8EF3E1A4FFBEFFE0FBBC21C13F20CCC8" LSID="urn:lsid:plazi:treatment:03850669FFBEFFE1FE0D225338ABCCBE:8EF3E1A4FFBEFFE0FBBC21C13F20CCC8" box="[1041,1145,1924,1950]" country="Trinidad and Tobago" name="Trinidad" pageId="46" pageNumber="47">Trinidad</location>
; 1904;
<collectorName id="26D9D2A9FFBEFFE0FB7B21C03E61CCC8" box="[1238,1336,1924,1951]" pageId="46" pageNumber="47">F. Birch</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBEFFE0FF1021EF3A4CCC92" box="[189,277,1962,1988]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="46" pageNumber="47">BMNH</collectionCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBEFFE0FE9D21EC38DBCC92" ID-GBIF-Occurrence="3037418593" box="[304,898,1959,1988]" collectionCode="BMNH" collectorName="Fry" country="Trinidad and Tobago" location="Antilles Trinidad" pageId="46" pageNumber="47" specimenCode="BMNH 1905-100" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D2A7CF6FFBEFFE0FE9D21EC3A07CC95" box="[304,350,1959,1987]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
;
<location id="8EF3E1A4FFBEFFE0FEC121EC3961CC95" LSID="urn:lsid:plazi:treatment:03850669FFBEFFE1FE0D225338ABCCBE:8EF3E1A4FFBEFFE0FEC121EC3961CC95" box="[364,568,1961,1987]" country="Trinidad and Tobago" name="Antilles Trinidad" pageId="46" pageNumber="47">Antilles Trinidad</location>
;
<collectorName id="26D9D2A9FFBEFFE0FDE821EF3937CC92" box="[581,622,1962,1988]" pageId="46" pageNumber="47">Fry</collectorName>
leg.;
<specimenCode id="DB8A1F04FFBEFFE0FD1C21EF38DBCC92" box="[689,898,1961,1988]" collectionCode="BMNH" pageId="46" pageNumber="47">BMNH 1905-100</specimenCode>
</materialsCitation>
<materialsCitation id="3B44BD22FFBEFFE0FC3021EC3A9BCCBE" ID-GBIF-Occurrence="3037418338" collectingDate="1912-12" collectingDateMax="1913-05" collectingDateMin="1912-12" collectionCode="MZSP" collectorName="R. Thaiter" country="Trinidad and Tobago" location="Porto Spain" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D2A7CF6FFBEFFE0FC3021EC3892CC95" box="[925,971,1959,1987]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
;
<location id="8EF3E1A4FFBEFFE0FC7721EF3F31CC95" LSID="urn:lsid:plazi:treatment:03850669FFBEFFE1FE0D225338ABCCBE:8EF3E1A4FFBEFFE0FC7721EF3F31CC95" box="[986,1128,1961,1988]" country="Trinidad and Tobago" name="Porto Spain" pageId="46" pageNumber="47">Porto Spain</location>
,
<date id="FF9291BFFFBEFFE0FBD821EF3FBCCC95" box="[1141,1253,1961,1988]" pageId="46" pageNumber="47" value="1912-12">
<collectingDate id="EFD66857FFBEFFE0FBD821EF3FBCCC95" box="[1141,1253,1961,1988]" pageId="46" pageNumber="47" value="1912-12">Dec 1912</collectingDate>
</date>
<date id="FF9291BFFFBEFFE0FB5821EC3E2BCC95" box="[1269,1394,1961,1987]" pageId="46" pageNumber="47" value="1913-05">
<collectingDate id="EFD66857FFBEFFE0FB5821EC3E2BCC95" box="[1269,1394,1961,1987]" pageId="46" pageNumber="47" value="1913-05">May 1913</collectingDate>
</date>
;
<collectorName id="26D9D2A9FFBEFFE0FF10218A3A6DCCBE" box="[189,308,1998,2025]" pageId="46" pageNumber="47">R. Thaiter</collectorName>
leg.;
<collectionCode id="ED3D2FBAFFBEFFE0FED8218B3A9BCCBE" box="[373,450,1998,2024]" collectionName="Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="46" pageNumber="47" type="Museum">MZSP</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF10274C3947CA75" blockId="47.[189,542,263,291]" box="[189,542,263,291]" pageId="47" pageNumber="48">
<materialsCitation id="3B44BD22FFBFFFE1FF10274C3943CA75" ID-GBIF-Occurrence="3037418510" box="[189,538,263,291]" collectionCode="BMNH" country="Venezuela" location="VENEZUELA" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F33BF7EFFFBFFFE1FF10274C3A32CA75" box="[189,363,265,291]" name="Venezuela" pageId="47" pageNumber="48">VENEZUELA</collectingCountry>
<specimenCount id="9D2A7CF6FFBFFFE1FE2E274C3AE9CA75" box="[387,432,263,291]" pageId="47" pageNumber="48" type="male">1 ♂</specimenCount>
;
<collectionCode id="ED3D2FBAFFBFFFE1FE13274C3943CA75" box="[446,538,265,291]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="47" pageNumber="48">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF10270938E3CA30" blockId="47.[189,954,332,359]" box="[189,954,332,359]" pageId="47" pageNumber="48">
COUNTRY UNKNOWN •
<materialsCitation id="3B44BD22FFBFFFE1FDAE270938EFCA30" ID-GBIF-Occurrence="3037418564" box="[515,950,332,359]" collectionCode="BMNH" collectorName="Nevinson" pageId="47" pageNumber="48" specimenCode="BMNH 1918-14" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D2A7CF6FFBFFFE1FDAE27093969CA31" box="[515,560,332,359]" pageId="47" pageNumber="48" type="female">1 ♀</specimenCount>
;
<collectorName id="26D9D2A9FFBFFFE1FD92270939F6CA30" box="[575,687,332,358]" pageId="47" pageNumber="48">Nevinson</collectorName>
leg.;
<specimenCode id="DB8A1F04FFBFFFE1FD5C270838EFCA30" box="[753,950,332,359]" collectionCode="BMNH" pageId="47" pageNumber="48">BMNH 1918-14</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBFFFE1FF1027CA381CCDB7" pageId="47" pageNumber="48" type="description">
<paragraph id="8B93B77FFFBFFFE1FF1027CA3A32CAFF" blockId="47.[189,363,399,425]" box="[189,363,399,425]" pageId="47" pageNumber="48">
<heading id="D0DB0013FFBFFFE1FF1027CA3A32CAFF" bold="true" box="[189,363,399,425]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B9586B6DFFBFFFE1FF1027CA3A32CAFF" bold="true" box="[189,363,399,425]" pageId="47" pageNumber="48">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF4727853A1CCA8C" blockId="47.[189,1205,448,511]" box="[234,325,448,474]" pageId="47" pageNumber="48">
<emphasis id="B9586B6DFFBFFFE1FF4727853A1CCA8C" bold="true" box="[234,325,448,474]" pageId="47" pageNumber="48">Female</emphasis>
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF1027A03FECCAA8" blockId="47.[189,1205,448,511]" box="[189,1205,484,511]" pageId="47" pageNumber="48">
MEASUREMENTS. Length:
<quantity id="4CD41A9AFFBFFFE1FE7327A03901CAA9" box="[478,600,485,511]" metricMagnitude="-2" metricUnit="m" metricValue="1.789" pageId="47" pageNumber="48" unit="mm" value="17.89">17.89 mm</quantity>
. Dorsal and ventral surfaces reddish to dark brown.
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF10246D389FC9E7" blockId="47.[189,1400,552,689]" pageId="47" pageNumber="48">
HEAD. Anterior edge surface horizontally rugose, interocular space displaying flat punctures, resembling rough; distinct ocellate punctuation near eyes. Clypeus with central deep emargination separating two acute clypeal teeth (
<figureCitation id="1317ABFAFFBFFFE1FE142434394BC9DA" box="[441,530,625,652]" captionStart="Fig" captionStartId="48.[189,232,1928,1954]" captionTargetBox="[193,1370,275,1892]" captionTargetId="figure-63@48.[189,1374,265,1894]" captionTargetPageId="48" captionText="Fig. 16. Dichotomius triquetrus (Luederwaldt, 1923). AB. Dorsal habitus. A. ♂. B. ♀. D. Lectotype, ♀ (MZSP). C. Lateral view of the aedeagus. D. Dorsal view of the head. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Elytra." figureDoi="http://doi.org/10.5281/zenodo.4542139" httpUri="https://zenodo.org/record/4542139/files/figure.png" pageId="47" pageNumber="48">Fig. 16</figureCitation>
BC). Cephalic carina bituberculate, wider than high, tubercles obtuse apically (
<figureCitation id="1317ABFAFFBFFFE1FE8724D33ACFC9E7" box="[298,406,662,689]" captionStart="Fig" captionStartId="48.[189,232,1928,1954]" captionTargetBox="[193,1370,275,1892]" captionTargetId="figure-63@48.[189,1374,265,1894]" captionTargetPageId="48" captionText="Fig. 16. Dichotomius triquetrus (Luederwaldt, 1923). AB. Dorsal habitus. A. ♂. B. ♀. D. Lectotype, ♀ (MZSP). C. Lateral view of the aedeagus. D. Dorsal view of the head. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Elytra." figureDoi="http://doi.org/10.5281/zenodo.4542139" httpUri="https://zenodo.org/record/4542139/files/figure.png" pageId="47" pageNumber="48">Fig. 16C</figureCitation>
). Ventral clypeal process truncate (see
<figureCitation id="1317ABFAFFBFFFE1FCF024D338E0C9E7" box="[861,953,662,689]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. AD. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. EF. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). GH. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). IJ. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="47" pageNumber="48">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF10249F3998C835" blockId="47.[189,1398,730,867]" pageId="47" pageNumber="48">
PRONOTUM. Gibbous, wider than long. Pronotal anterior disc shiny and smooth with fine, dense and wellimpressed punctures near antero-lateral sides. Uninterrupted ocellate punctures along pronotal margins (
<figureCitation id="1317ABFAFFBFFFE1FF6B25663A03C868" box="[198,346,803,830]" captionStart="Fig" captionStartId="48.[189,232,1928,1954]" captionTargetBox="[193,1370,275,1892]" captionTargetId="figure-63@48.[189,1374,265,1894]" captionTargetPageId="48" captionText="Fig. 16. Dichotomius triquetrus (Luederwaldt, 1923). AB. Dorsal habitus. A. ♂. B. ♀. D. Lectotype, ♀ (MZSP). C. Lateral view of the aedeagus. D. Dorsal view of the head. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Elytra." figureDoi="http://doi.org/10.5281/zenodo.4542139" httpUri="https://zenodo.org/record/4542139/files/figure.png" pageId="47" pageNumber="48">Fig. 16B, D</figureCitation>
). Hypomeral disc punctate; external margin bearing single row of organized long setae, approximately of same length (see
<figureCitation id="1317ABFAFFBFFFE1FDF6250D39EAC835" box="[603,691,840,867]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. AD. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. EF. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). GH. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). IJ. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="47" pageNumber="48">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF1025C93910CFFE" blockId="47.[189,1400,908,1192]" pageId="47" pageNumber="48">
ABDOMEN. Prosternum surface with fine microsculptured, glabrous. Mesosternum and metepisternum surface microsculpture shagreened with ocellate setigerous punctures. Metasternum setigerous punctures restricted mainly to anterior and lateral edges of mestasternal lobe, puncture diameter variable. Punctuation of antero-median edge connecting with lateral ones through margin of mesocoxa (see
<figureCitation id="1317ABFAFFBFFFE1FF5B225A3A0CCF6C" box="[246,341,1055,1082]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. AD. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. EF. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). GH. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). IJ. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="47" pageNumber="48">Fig. 2G</figureCitation>
). Lateral edges of metasternum with ocellate, setigerous punctures separated by width of a puncture, pilosity sparse, setae short, not reaching metepisternum suture. Longitudinal sulcus clearly visible. Ocellate punctures along anterior margins of sternites. Pygidium ocellate punctation on basal edge and through lateral margins.
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF1022973E1ACE47" blockId="47.[189,1398,1233,1297]" pageId="47" pageNumber="48">
ELYTRA. Interstriae microsculpture shagreened (see
<figureCitation id="1317ABFAFFBFFFE1FCB522943832CFBA" box="[792,875,1233,1260]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. AD. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. EF. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). GH. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). IJ. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="47" pageNumber="48">Fig. 2J</figureCitation>
), fine punctures randomly distributed. Striae strongly marked, ocellate punctures separated by diameter of a puncture on elytral disc (
<figureCitation id="1317ABFAFFBFFFE1FB6522B33E6FCE47" box="[1224,1334,1270,1297]" captionStart="Fig" captionStartId="48.[189,232,1928,1954]" captionTargetBox="[193,1370,275,1892]" captionTargetId="figure-63@48.[189,1374,265,1894]" captionTargetPageId="48" captionText="Fig. 16. Dichotomius triquetrus (Luederwaldt, 1923). AB. Dorsal habitus. A. ♂. B. ♀. D. Lectotype, ♀ (MZSP). C. Lateral view of the aedeagus. D. Dorsal view of the head. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Elytra." figureDoi="http://doi.org/10.5281/zenodo.4542139" httpUri="https://zenodo.org/record/4542139/files/figure.png" pageId="47" pageNumber="48">Fig. 16D</figureCitation>
).
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF10237F3893CE2F" blockId="47.[187,1399,1337,1401]" pageId="47" pageNumber="48">LEGS. Metacoxa posterior edge bearing single row of organized ocellate punctures, separated by diameter of a puncture. Posterior margin of meta-trochanter with long setae.</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF1023E43AB7CEED" blockId="47.[189,494,1441,1467]" box="[189,494,1441,1467]" pageId="47" pageNumber="48">
<heading id="D0DB0013FFBFFFE1FF1023E43AB7CEED" bold="true" box="[189,494,1441,1467]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B9586B6DFFBFFFE1FF1023E43AB7CEED" bold="true" box="[189,494,1441,1467]" pageId="47" pageNumber="48">Morphological variation</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF47239738EBCEBA" blockId="47.[189,1398,1490,1590]" box="[234,946,1490,1517]" pageId="47" pageNumber="48">
<emphasis id="B9586B6DFFBFFFE1FF4723973A6CCEBA" bold="true" box="[234,309,1490,1516]" pageId="47" pageNumber="48">Males</emphasis>
(
<figureCitation id="1317ABFAFFBFFFE1FEE823973AE9CEBB" box="[325,432,1490,1517]" captionStart="Fig" captionStartId="48.[189,232,1928,1954]" captionTargetBox="[193,1370,275,1892]" captionTargetId="figure-63@48.[189,1374,265,1894]" captionTargetPageId="48" captionText="Fig. 16. Dichotomius triquetrus (Luederwaldt, 1923). AB. Dorsal habitus. A. ♂. B. ♀. D. Lectotype, ♀ (MZSP). C. Lateral view of the aedeagus. D. Dorsal view of the head. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Elytra." figureDoi="http://doi.org/10.5281/zenodo.4542139" httpUri="https://zenodo.org/record/4542139/files/figure.png" pageId="47" pageNumber="48">Fig. 16A</figureCitation>
) are distinguished from females as follows:
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF1023B23E0CCD60" blockId="47.[189,1398,1490,1590]" pageId="47" pageNumber="48">HEAD. Cephalic carina higher than wide, conical in shape. Median tubercle twice as high as lateral tubercles, rounded, weakly bifurcate apically and curved posteriorly. Apex of lateral tubercles acute.</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF1020253A73CDC8" blockId="47.[189,1399,1631,1694]" pageId="47" pageNumber="48">PRONOTUM. Surface covered by fine punctures, denser at antero-lateral areas; anterior disc with weak declivity.</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF10208D381CCDB7" blockId="47.[189,837,1735,1762]" box="[189,837,1735,1762]" pageId="47" pageNumber="48">
PARAMERES AND LAMELLA COPULATRIX. As in
<figureCitation id="1317ABFAFFBFFFE1FD182082385FCDB7" box="[693,774,1735,1761]" captionStart="Fig" captionStartId="48.[189,232,1928,1954]" captionTargetBox="[193,1370,275,1892]" captionTargetId="figure-63@48.[189,1374,265,1894]" captionTargetPageId="48" captionText="Fig. 16. Dichotomius triquetrus (Luederwaldt, 1923). AB. Dorsal habitus. A. ♂. B. ♀. D. Lectotype, ♀ (MZSP). C. Lateral view of the aedeagus. D. Dorsal view of the head. E. Ventral view of the aedeagus. FG. Lamella copulatrix. H. Elytra." figureDoi="http://doi.org/10.5281/zenodo.4542139" httpUri="https://zenodo.org/record/4542139/files/figure.png" pageId="47" pageNumber="48">Fig. 16</figureCitation>
EG.
</paragraph>
</subSubSection>
<subSubSection id="C336E4F4FFBFFFE1FF10214F38ABCCBE" pageId="47" pageNumber="48" type="discussion">
<paragraph id="8B93B77FFFBFFFE1FF10214F3A76CC72" blockId="47.[189,303,1802,1828]" box="[189,303,1802,1828]" pageId="47" pageNumber="48">
<heading id="D0DB0013FFBFFFE1FF10214F3A76CC72" bold="true" box="[189,303,1802,1828]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B9586B6DFFBFFFE1FF10214F3A76CC72" bold="true" box="[189,303,1802,1828]" pageId="47" pageNumber="48">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B93B77FFFBFFFE1FF10217F38ABCCBE" blockId="47.[186,1399,1850,2025]" pageId="47" pageNumber="48">
This is an endemic species from
<collectingCountry id="F33BF7EFFFBFFFE1FD84217F39C8CC02" box="[553,657,1850,1876]" name="Trinidad and Tobago" pageId="47" pageNumber="48">Trinidad</collectingCountry>
.The redescription was based on the well-developed female of the MZSP and the underdeveloped female at the CEMT. The male redescription was based on the specimen in the MZSP wrongly determined by Pereira as “
<taxonomicName id="4C2CCCFCFFBFFFE1FCAC21C038F8CCC8" authorityName="Harold" authorityYear="1880" box="[769,929,1924,1950]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="belus">
<emphasis id="B9586B6DFFBFFFE1FCAC21C038F8CCC8" box="[769,929,1924,1950]" italics="true" pageId="47" pageNumber="48">Pinotus belus</emphasis>
</taxonomicName>
” and an underdeveloped male from the CEMT. Male genitalia were extracted from the underdeveloped male at CEMT.
<bibRefCitation id="EFBDCA8EFFBFFFE1FBF421EC3E1BCC95" author="Luederwaldt H." box="[1113,1346,1961,1987]" pageId="47" pageNumber="48" pagination="1 - 13" refId="ref30108" refString="Luederwaldt H. 1923. Neue Pinotus - Arten. Revista do Museu Paulista 14: 1 - 13." type="journal article" year="1923">Luederwaldt (1923)</bibRefCitation>
also mentioned a male specimen in the
<typeStatus id="549709DDFFBFFFE1FDFE218A39DFCCBF" box="[595,646,1999,2025]" pageId="47" pageNumber="48">type</typeStatus>
series that could not be traced.
</paragraph>
</subSubSection>
</treatment>
</document>