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<document id="38C5526508861A7895B1674DCFE8BEBB" ID-CLB-Dataset="22642" ID-DOI="10.11646/zootaxa.4514.3.3" ID-GBIF-Dataset="54115d80-05ed-4ee7-b035-1ac714f9c31d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3770231" ID-ZooBank="8F3F3680-5950-4FAB-B8DC-455FAC27535A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588001511746" checkinUser="plazi" docAuthor="Abdel-Dayem, Mahmoud S., Rasool, Iftekhar, Elgharbawy, Ali A., Nagel, Peter &amp; Aldhafer, Hathal M." docDate="2018" docId="03811B61FF9BFF9DA5BCFD8CFBB1FEB5" docLanguage="en" docName="zootaxa.4514.3.3.pdf" docOrigin="Zootaxa 4514 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paussus (Hylotorus) cephalotes Raffray 1886" docType="treatment" docVersion="7" lastPageNumber="360" masterDocId="FFB86319FF89FF8EA52BFB78FFF4FFD0" masterDocTitle="Faunistic inventory and zoogeographical analysis of the ground beetles (Coleoptera, Carabidae) of Garf Raydah Nature Reserve, Southwestern of Saudi Arabia, and description of a new species of Paussinae" masterLastPageNumber="371" masterPageNumber="341" pageNumber="359" updateTime="1698837116288" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0923EA641970BA4490DB725741BC91F5">Faunistic inventory and zoogeographical analysis of the ground beetles (Coleoptera, Carabidae) of Garf Raydah Nature Reserve, Southwestern of Saudi Arabia, and description of a new species of Paussinae</mods:title>
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<mods:namePart id="68E0623C66355E15EC91772E9839A586">Abdel-Dayem, Mahmoud S.</mods:namePart>
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<mods:namePart id="09C4020E251FFC152DF233D61C92D1FC">Rasool, Iftekhar</mods:namePart>
<mods:affiliation id="B95A901CC7B556ED5B5EC15FE4763B9E">King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia.</mods:affiliation>
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<mods:namePart id="62CCC11DE61C660E8604E73A237E0621">Elgharbawy, Ali A.</mods:namePart>
<mods:affiliation id="CE4FE1C95AF98DE47C2B74F0AFC1523C">Zoology Department, Faculty of Science, Al Azhar University, Nasr City, Cairo, Egypt.</mods:affiliation>
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<mods:namePart id="9A157E8BCAFB3120408EA371ECE8DC9C">Nagel, Peter</mods:namePart>
<mods:affiliation id="645BCB5C01378DBAAE95564B7D9DA3A1">Biogeography Research Group, Department of Environmental Sciences, Faculty of Science, University of Basel, St. Johanns-Vorstadt 10, 4056 Basel, Switzerland</mods:affiliation>
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<mods:namePart id="DE73F6D6A6108084C199F28C8A882E4D">Aldhafer, Hathal M.</mods:namePart>
<mods:affiliation id="D4B91D21E0C1AD3BAA3E8CE9C912EB7A">King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agriculture Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia.</mods:affiliation>
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<heading id="D0DF1D1BFF9BFF9CA5BCFD8CFD31F8DE" bold="true" box="[151,709,1780,1806]" fontSize="11" level="1" pageId="18" pageNumber="359" reason="1">
<taxonomicName id="4C28D1F4FF9BFF9CA5BCFD8CFD31F8DE" ID-CoL="8MCMK" authority="Raffray, 1886" authorityName="Raffray" authorityYear="1886" box="[151,709,1780,1806]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="359" phylum="Arthropoda" rank="species" species="cephalotes" subGenus="Hylotorus">
<emphasis id="B95C7665FF9BFF9CA5BCFD8CFD31F8DE" bold="true" box="[151,709,1780,1806]" pageId="18" pageNumber="359">
<emphasis id="B95C7665FF9BFF9CA5BCFD8CFF0FF8DE" bold="true" box="[151,251,1780,1806]" italics="true" pageId="18" pageNumber="359">Paussus</emphasis>
(
<emphasis id="B95C7665FF9BFF9CA420FD8CFE77F8DE" bold="true" box="[267,387,1780,1806]" italics="true" pageId="18" pageNumber="359">Hylotorus</emphasis>
)
<emphasis id="B95C7665FF9BFF9CA4BEFD8CFDE0F8DE" bold="true" box="[405,532,1780,1806]" italics="true" pageId="18" pageNumber="359">cephalotes</emphasis>
<bibRefCitation id="EFB9D786FF9BFF9CA730FD8CFD31F8DE" author="Raffray, A." box="[539,709,1780,1806]" pageId="18" pageNumber="359" pagination="307 - 359" refId="ref23048" refString="Raffray, A. (1886) Materiaux pour servir a 1 ' etude des coleopteres de la famille des Paussides. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 2 (8), 307 - 359, pls. 15 - 19." type="journal article" year="1886">Raffray, 1886</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<emphasis id="B95C7665FF9BFF9CA5BCFC42FE74F883" bold="true" box="[151,384,1850,1875]" pageId="18" pageNumber="359">Material examined.</emphasis>
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<collectionCode id="ED3932B2FF9BFF9CA4ACFC43FE2DF883" box="[391,473,1851,1875]" pageId="18" pageNumber="359">GRNR</collectionCode>
:
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<elevation id="00054D44FF9BFF9CA4CFFC43FDC9F882" box="[484,573,1851,1875]" metricMagnitude="3" metricUnit="m" metricValue="1.7719999999999998" pageId="18" pageNumber="359" unit="m" value="1772.0">1772 m</elevation>
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:
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<collectingDate id="EFD2755FFF9BFF9CA76CFC43FD31F883" box="[583,709,1851,1875]" pageId="18" pageNumber="359" value="2017-04-04">04.IV.2017</collectingDate>
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, HP,
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.
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<subSubSection id="C332F9FCFF9BFF9DA5ECFC26FBB1FEB5" lastPageId="19" lastPageNumber="360" pageId="18" pageNumber="359" type="distribution">
<paragraph id="8B97AA77FF9BFF9CA5ECFC26FB5FF8A7" blockId="18.[151,1436,1850,2019]" box="[199,1195,1886,1911]" pageId="18" pageNumber="359">
<emphasis id="B95C7665FF9BFF9CA5ECFC26FE94F8A7" bold="true" box="[199,352,1886,1911]" pageId="18" pageNumber="359">Distribution.</emphasis>
END_AR species, occurring in southwestern Arabian Peninsula (SA, YE).
</paragraph>
<paragraph id="8B97AA77FF9BFF9CA5ECFCFAFA68F832" blockId="18.[151,1436,1850,2019]" pageId="18" pageNumber="359">
<emphasis id="B95C7665FF9BFF9CA5ECFCFAFE01F84B" bold="true" box="[199,501,1922,1947]" pageId="18" pageNumber="359">Life forms and Remarks.</emphasis>
Myrmecophilous (M) species.
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<emphasis id="B95C7665FF9BFF9CA67CFCFCFBC6F84B" box="[855,1074,1922,1947]" italics="true" pageId="18" pageNumber="359">Paussus cephalotes</emphasis>
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was described from a specimen collected by Dr. Millingen from Hejaz Mountains of western
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without specific locality (
<bibRefCitation id="EFB9D786FF9BFF9CA1FCFCDEFA97F86F" author="Nagel, P." box="[1239,1379,1958,1983]" pageId="18" pageNumber="359" pagination="99 - 107" refId="ref22579" refString="Nagel, P. (1982) Insects of Saudi Arabia. Coleoptera: Fam. Carabidae, subfam. Paussinae. Fauna of Saudi Arabia, 4, 99 - 107." type="journal article" year="1982">Nagel 1982</bibRefCitation>
). No specimen matching the original description (
<bibRefCitation id="EFB9D786FF9BFF9CA78FFCB2FCB4F833" author="Raffray, A." box="[676,832,1994,2019]" pageId="18" pageNumber="359" pagination="307 - 359" refId="ref23048" refString="Raffray, A. (1886) Materiaux pour servir a 1 ' etude des coleopteres de la famille des Paussides. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 2 (8), 307 - 359, pls. 15 - 19." type="journal article" year="1886">Raffray 1886</bibRefCitation>
) has subsequently been collected until Van Harten
</paragraph>
<paragraph id="8B97AA77FF9AFF9DA5BCFBEFFB7AFEEF" blockId="19.[151,1436,151,357]" pageId="19" pageNumber="360">
collected two specimens from
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(Radaa and
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) in 1998 (
<bibRefCitation id="EFB9D786FF9AFF9DA68FFBE0FB3EFF7F" author="Luna de Carvalho, E." box="[932,1226,151,176]" pageId="19" pageNumber="360" pagination="35 - 38" refId="ref22078" refString="Luna de Carvalho, E. (2000 a) Notas sobre o genero Cephalotipaussus (Luna de Carvalho, 1987) e redescoberta no Yemen de C. cephalotes (Raffray, 1885) (Coleoptera: Carabidae: Paussinae). Elytron, 13, (1999), 35 - 38." type="journal article" year="2000">Luna de Carvalho 2000 a</bibRefCitation>
, b). During recent entomological survey in
<collectingRegion id="49EC6495FF9AFF9DA485FBC4FDA1FF05" box="[430,597,188,213]" country="Saudi Arabia" name="Al Bahah" pageId="19" pageNumber="360">Baha Province</collectingRegion>
, southwestern
<collectingCountry id="F33FEAE7FF9AFF9DA7D6FBC4FC60FF05" box="[765,916,188,213]" name="Saudi Arabia" pageId="19" pageNumber="360">Saudi Arabia</collectingCountry>
,
<bibRefCitation id="EFB9D786FF9AFF9DA68BFBC4FB4FFF04" author="El-Hawagry, M. S. &amp; Khalil, M. W. &amp; Sharaf, M. R. &amp; Fadl, H. H. &amp; Aldawood, A. S." box="[928,1211,188,213]" pageId="19" pageNumber="360" pagination="1 - 88" refId="ref20525" refString="El-Hawagry, M. S., Khalil, M. W., Sharaf, M. R., Fadl, H. H. &amp; Aldawood, A. S. (2013) A preliminary study on the insect fauna of Al-Baha Province, Saudi Arabia, with descriptions of two new species. ZooKeys, 274, 1 - 88. https: // doi. org / 10.3897 / zookeys. 274.4529" type="journal article" year="2013">
El-Hawagry
<emphasis id="B95C7665FF9AFF9DA11BFBC5FB9DFF05" box="[1072,1129,188,213]" italics="true" pageId="19" pageNumber="360">et al.</emphasis>
(2013)
</bibRefCitation>
reported specimens of
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<emphasis id="B95C7665FF9AFF9DA59EFB99FEBDFF28" box="[181,329,223,248]" italics="true" pageId="19" pageNumber="360">P. cephalotes</emphasis>
</taxonomicName>
from Jebel Shada during April-June at
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1600
<bibRefCitation id="EFB9D786FF9AFF9DA66CFB98FCA9FF27" author="Kataev, B. M." box="[839,861,224,247]" pageId="19" pageNumber="360" pagination="247 - 250" refId="ref21632" refString="Kataev, B. M. (2014) On the taxonomic status of Idiomelas himalayensis Della Beffa (Coleoptera, Carabidae: Harpalini). Entomological Review, 94 (2), 247 - 250. https: // doi. org / 10.1134 / S 0013873814020134" type="journal article" year="2014">m</bibRefCitation>
</quantity>
<materialsCitation id="3B40A02AFF9AFF9DA64EFBA7FB7DFEEF" ID-GBIF-Occurrence="2608963698" county="Baljurashi forest" elevation="1772" location="The" municipality="The" pageId="19" pageNumber="360" specimenCount="1" stateProvince="Al Bahah">
and Moore &amp; Robertson (2014) report this species from
<collectingRegion id="49EC6495FF9AFF9DA5FDFA7CFEB0FECD" box="[214,324,260,285]" country="Saudi Arabia" name="Al Bahah" pageId="19" pageNumber="360">Al Bahah</collectingRegion>
,
<collectingCounty id="62F6D2FBFF9AFF9DA47AFA7CFDE4FECD" box="[337,528,260,285]" pageId="19" pageNumber="360">Baljurashi forest</collectingCounty>
.
<collectingMunicipality id="6BF3300DFF9AFF9DA737FA7CFDBCFECD" box="[540,584,260,285]" pageId="19" pageNumber="360">The</collectingMunicipality>
species is recorded for first time from
<collectingRegion id="49EC6495FF9AFF9DA12CFA7CFB53FECD" box="[1031,1191,260,285]" country="Saudi Arabia" name="'Asir" pageId="19" pageNumber="360">Asir Province</collectingRegion>
.
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single specimen was collected under stones in an
<taxonomicName id="4C28D1F4FF9AFF9DA72CFA50FD1AFE90" box="[519,750,295,320]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="19" pageNumber="360" phylum="Tracheophyta" rank="species" species="ficus-indica">
<emphasis id="B95C7665FF9AFF9DA72CFA50FD1AFE90" box="[519,750,295,320]" italics="true" pageId="19" pageNumber="360">Opuntia ficus-indica</emphasis>
</taxonomicName>
community at an altitude of
<quantity id="4CD00792FF9AFF9DA119FA50FB7DFEEF" box="[1074,1161,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.7719999999999998" pageId="19" pageNumber="360" unit="m" value="1772.0">
<elevation id="00054D44FF9AFF9DA119FA50FB7DFEEF" box="[1074,1161,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.7719999999999998" pageId="19" pageNumber="360" unit="m" value="1772.0">1772 m</elevation>
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.
</paragraph>
<paragraph id="8B97AA77FF9AFF9DA5BCFA34FBB1FEB5" blockId="19.[151,1436,151,357]" box="[151,1093,332,357]" pageId="19" pageNumber="360">
The subgenus assignment
<emphasis id="B95C7665FF9AFF9DA496FA34FDB9FEB5" box="[445,589,332,357]" italics="true" pageId="19" pageNumber="360">
<taxonomicName id="4C28D1F4FF9AFF9DA4EEFA34FDB0FEB5" authorityName="Dalman" authorityYear="1823" box="[453,580,332,357]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="358" phylum="Arthropoda" rank="subGenus" subGenus="Hylotorus">Hylotorus</taxonomicName>
</emphasis>
is adopted from
<bibRefCitation id="EFB9D786FF9AFF9DA621FA34FBB5FEB5" author="Robertson, J. &amp; Moore, W." box="[778,1089,332,357]" pageId="19" pageNumber="360" pagination="134 - 170" refId="ref23316" refString="Robertson, J. A &amp; Moore, W. (2016) Phylogeny of Paussus L. (Carabidae: Paussinae): unravelling morphological convergence associated with myrmecophilous life histories. Systematic Entomology, 42 (1), 134 - 170, figs. S 1 - S 2, tabs. S 1 - S 5 [(January 2017): 28 Sep 2016: version of record online, first published, pp. 170, end of text] https: // doi. org / 10.1111 / syen. 12205" type="journal article" year="2016">Robertson &amp; Moore (2016)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>