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<mods:title id="DEA96F41062C828E186A1D6DC917D9D2">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<treatment id="038B1D1D6478FFA3FCA1EA04D47DFCC7" ID-DOI="http://doi.org/10.5281/zenodo.13127606" ID-Zenodo-Dep="13127606" LSID="urn:lsid:plazi:treatment:038B1D1D6478FFA3FCA1EA04D47DFCC7" httpUri="http://treatment.plazi.org/id/038B1D1D6478FFA3FCA1EA04D47DFCC7" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="C338FF806478FFA0FCA1EA04D517F8C8" box="[813,1318,1812,1836]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8B9DAC0B6478FFA0FCA1EA04D517F8C8" blockId="17.[813,1318,1812,1868]" box="[813,1318,1812,1836]" pageId="17" pageNumber="18">
<heading id="D0D51B676478FFA0FCA1EA04D517F8C8" box="[813,1318,1812,1836]" fontSize="9" level="2" pageId="17" pageNumber="18" reason="8">
<taxonomicName id="4C22D7886478FFA0FCA1EA04D4B1F8C8" authority="Chen et al." authorityName="Chen" box="[813,1152,1812,1836]" family="Gondolellidae" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="laii">
<emphasis id="B95670196478FFA0FCA1EA04D435F8C8" box="[813,1028,1812,1836]" italics="true" pageId="17" pageNumber="18">Gladigondolella laii</emphasis>
Chen et al.
</taxonomicName>
morphotype A
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF806478FFA0FCA1EA24D3FCF8A8" box="[813,973,1844,1868]" pageId="17" pageNumber="18" type="description">
<paragraph id="8B9DAC0B6478FFA0FCA1EA24D3FCF8A8" blockId="17.[813,1318,1812,1868]" box="[813,973,1844,1868]" pageId="17" pageNumber="18">
<figureCitation id="1319B08E6478FFA0FCA1EA24D3F2F8A8" box="[813,963,1844,1868]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.13127504" httpUri="https://zenodo.org/record/13127504/files/figure.png" pageId="17" pageNumber="18">Fig. 13B, C, G</figureCitation>
?
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647BFFA3FF1BEDFBD120FE67" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8B9DAC0B647BFFA3FF1BEDFBD2B6FEE7" blockId="18.[151,775,235,387]" box="[151,647,235,259]" pageId="18" pageNumber="19">
2013
<taxonomicName id="4C22D788647BFFA3FF5EEDFBD1A3FEE7" box="[210,402,235,259]" genus="Neogondolella" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B9567019647BFFA3FF5EEDFBD15DFEE7" box="[210,364,235,259]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
sp.
</taxonomicName>
; Yan et al., fig. 6, nr. X.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FF1BEC1BD120FE67" blockId="18.[151,775,235,387]" pageId="18" pageNumber="19">
2015
<taxonomicName id="4C22D788647BFFA3FF5EEC1BD15DFEC7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[210,364,267,291]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FF5EEC1BD15DFEC7" box="[210,364,267,291]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
</taxonomicName>
? n. sp. A; Chen et al., figs. 9.5
<date id="FF9C8ACB647BFFA3FD31EC1BD0FAFEA7" pageId="18" pageNumber="19" value="2019-08-09">9.8. 2019</date>
<taxonomicName id="4C22D788647BFFA3FF5EEC3BD15DFEA7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[210,364,299,323]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FF5EEC3BD15DFEA7" box="[210,364,299,323]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
</taxonomicName>
? n. sp. A; Chen et al., figs. 6.8,
<date id="FF9C8ACB647BFFA3FD35EC3BD0FAFE87" pageId="18" pageNumber="19" value="2021-09-06">6.9. 2021</date>
<taxonomicName id="4C22D788647BFFA3FF58EC5BD19FFE87" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[212,430,331,355]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii" status="sp. nov.">
<emphasis id="B9567019647BFFA3FF58EC5BD19FFE87" box="[212,430,331,355]" italics="true" pageId="18" pageNumber="19">Gladigondolella laii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647BFFA3FE3AEC5CD23DFE87" box="[438,524,332,355]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
; Chen et al., figs. 4.20, 4.21 (only).
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647BFFA3FF1BECBBD1F2FE27" box="[151,451,427,451]" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8B9DAC0B647BFFA3FF1BECBBD1F2FE27" blockId="18.[151,451,427,451]" box="[151,451,427,451]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF1BECBBD0C9FE27" box="[151,248,427,451]" italics="true" pageId="18" pageNumber="19">Material</emphasis>
. ca.
<specimenCount id="9D246782647BFFA3FEA5ECBBD18EFE27" box="[297,447,427,451]" count="15" pageId="18" pageNumber="19" type="generic">15 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647BFFA3FF1BECFCD274FD87" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8B9DAC0B647BFFA3FF1BECFCD274FD87" blockId="18.[151,776,491,1667]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF1BECFCD139FDE7" box="[151,264,492,515]" italics="true" pageId="18" pageNumber="19">Diagnosis.</emphasis>
Segminiplanate P
<subScript id="17A6AE4E647BFFA3FE60ECE7D1C4FDEC" attach="left" box="[492,501,503,520]" fontSize="7" pageId="18" pageNumber="19">1</subScript>
element. Platform is arched, thickened and the platform margins are oriented orally. Denticles slightly fused at mid-length, larger and more discrete towards the anterior end.
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647BFFA3FF21EF7CD335FC67" pageId="18" pageNumber="19" type="description">
<paragraph id="8B9DAC0B647BFFA3FF21EF7CD335FC67" blockId="18.[151,776,491,1667]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF21EF7CD101FD67" box="[173,304,620,643]" italics="true" pageId="18" pageNumber="19">Description.</emphasis>
Te element is arched in lateral view. In oral view, the margins of the thickened platform are subparallel at mid-length, and taper anteriorly. Te platform is V-shaped, and the margin is directed orally. Te cusp may be conspicuous in well preserved specimens. A posterior process may be present, which bears additional posterior denticles, otherwise the posterior margin is rounded. Te carina denticles are partly fused at mid-length, taller and more discrete towards the anterior end.
</paragraph>
</subSubSection>
<subSubSection id="C338FF80647BFFA3FF21EE9BD47DFCC7" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8B9DAC0B647BFFA3FF21EE9BD4FCFD47" blockId="18.[151,776,491,1667]" lastBlockId="18.[813,1437,235,803]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FF21EE9BD1C0FC47" box="[173,497,907,931]" italics="true" pageId="18" pageNumber="19">
Remarks.
<taxonomicName id="4C22D788647BFFA3FE94EE9BD1C0FC47" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[280,497,907,931]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii">Gladigondolella laii</taxonomicName>
</emphasis>
differs from
<taxonomicName id="4C22D788647BFFA3FD0DEE9BD2D4FC47" baseAuthorityName="Orchard" baseAuthorityYear="2007" box="[641,741,907,931]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FD0DEE9BD2D4FC47" box="[641,741,907,931]" italics="true" pageId="18" pageNumber="19">Borinella</emphasis>
</taxonomicName>
by having a more fused carina and an arched and thick platform whose margin are oriented orally. In comparison with the
<typeStatus id="549912A9647BFFA3FF75EEFCD119FBE0" box="[249,296,1004,1028]" pageId="18" pageNumber="19">type</typeStatus>
species of
<taxonomicName id="4C22D788647BFFA3FE2EEEFBD27EFBE7" authorityName="Muller" authorityYear="1962" box="[418,591,1003,1027]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FE2EEEFBD27EFBE7" box="[418,591,1003,1027]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
, a Middle Triassic species, the posterior process is much less developed and less arcuated in the carinal axis.
<bibRefCitation id="EFB3D1FA647BFFA3FDB0E93BD336FBA7" author="Chen" box="[572,775,1067,1091]" etAl="et al." firstAuthor="Chen" pageId="18" pageNumber="19" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al. (2015)</bibRefCitation>
rejected the assignment of their
<emphasis id="B9567019647BFFA3FE75E95CD226FB87" box="[505,535,1100,1123]" italics="true" pageId="18" pageNumber="19">Ng</emphasis>
. n. sp. A to
<taxonomicName id="4C22D788647BFFA3FD16E95BD0D0FB67" authorityName="Muller" authorityYear="1962" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FD16E95BD0D0FB67" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
on the basis that posterior denticles are never present behind the cusp. In our material (e.g.
<figureCitation id="1319B08E647BFFA3FD2CE99BD2CEFB47" box="[672,767,1163,1187]" captionStart="Fig" captionStartId="18.[169,200,1743,1763]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@19.[228,1362,254,1764]" captionTargetPageId="19" captionText="Fig. 13 Gladigondolellinae, Neogondolellinae and Cornudininae from Qiakong, Laren, Shanggang and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A Borinella buurensis (Dagis), LAR212, PIMUZ 39103. B, C, G Gladigondolella laii (morphotype A); B LAR231C, PIMUZ 39132; C BAN2, PIMUZ 39133;G LAR227C, PIMUZ 39134. D, sp.indet.(P2 element?), SHA319, PIMUZ 39293.E, F, H, I Gladigondolella laii (morphotype B); E LIL510,PIMUZ 39135; F LIL512, PIMUZ 39136; H QIA140, PIMUZ 39137;I LIL510,PIMUZ 39138. JL, O, Spathicuspus spathi (Sweet); J QIA141, PIMUZ 39300; K QIA144, PIMUZ 39301; L BAN1, PIMUZ 39302; O SHA322,PIMUZ 39303. M, P Spathicuspus n. sp. A; M QIA144, PIMUZ 39297; P, QIA143, PIMUZ 39298. N, R Spathicuspus spathi (juvenile) (Sweet); N, LIL515D, PIMUZ 39304; R QIA143,PIMUZ 39305. Q Spathicuspus n. sp.B; LAR226C, PIMUZ 39299" figureDoi="http://doi.org/10.5281/zenodo.13127504" httpUri="https://zenodo.org/record/13127504/files/figure.png" pageId="18" pageNumber="19">Fig. 13C</figureCitation>
) small denticles are present occasionally. With the exception of these additional posterior denticles, our specimens resemble those of
<bibRefCitation id="EFB3D1FA647BFFA3FE20E9FBD246FAE7" author="Chen" box="[428,631,1259,1283]" etAl="et al." firstAuthor="Chen" pageId="18" pageNumber="19" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al. (2015)</bibRefCitation>
very closely, hence they are synonymized here. Implicitly we assume there is variation within the species, a small posterior process being present or not. Tis taxon is thought to be intermediary between
<taxonomicName id="4C22D788647BFFA3FE02E87BD1C3FA67" baseAuthorityName="Orchard" baseAuthorityYear="2007" box="[398,498,1387,1411]" family="Gondolellidae" genus="Borinella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FE02E87BD1C3FA67" box="[398,498,1387,1411]" italics="true" pageId="18" pageNumber="19">Borinella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788647BFFA3FDA4E87BD2E4FA67" authorityName="Muller" authorityYear="1962" box="[552,725,1387,1411]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FDA4E87BD2E4FA67" box="[552,725,1387,1411]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
, but through the shared derived characteristic of the posterior cusp we determine it as the former genus. Chen et al., (2019; figs. 6.8, 6.9) illustrated similar and coeval (associated with
<taxonomicName id="4C22D788647BFFA3FEADE8FCD19DF9E7" baseAuthorityName="Solien" baseAuthorityYear="1979" box="[289,428,1515,1539]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="collinsoni">
<emphasis id="B9567019647BFFA3FEADE8FCD19DF9E7" box="[289,428,1515,1539]" italics="true" pageId="18" pageNumber="19">Ic. collinsoni</emphasis>
</taxonomicName>
) elements from
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, which they assigned to
<taxonomicName id="4C22D788647BFFA3FEDDEB1BD1DAF9C7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[337,491,1547,1571]" genus="Neogondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B9567019647BFFA3FEDDEB1BD1DAF9C7" box="[337,491,1547,1571]" italics="true" pageId="18" pageNumber="19">Neogondolella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62647BFFA3FE7FEB1CD21FF9C7" box="[499,558,1548,1571]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
A. Teir illustrated specimens more closely resemble
<taxonomicName id="4C22D788647BFFA3FD9EEB3BD174F987" authority="(Huckriede)" baseAuthorityName="Huckriede" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="tethydis">
<emphasis id="B9567019647BFFA3FD9EEB3BD087F987" italics="true" pageId="18" pageNumber="19">Gladigondolella tethydis</emphasis>
(Huckriede)
</taxonomicName>
and were later assigned to a new species of
<taxonomicName id="4C22D788647BFFA3FF69EB7BD1A3F967" authorityName="Muller" authorityYear="1962" box="[229,402,1643,1667]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FF69EB7BD1A3F967" box="[229,402,1643,1667]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
:
<taxonomicName id="4C22D788647BFFA3FE2CEB7BD35AFEE7" authority="(Chen et al., 2021)" baseAuthorityName="Chen" baseAuthorityYear="2021" box="[416,875,235,1667]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="laii">
<emphasis id="B9567019647BFFA3FE2CEB7BD248F967" box="[416,633,1643,1667]" italics="true" pageId="18" pageNumber="19">Gladigondolella laii</emphasis>
(
<bibRefCitation id="EFB3D1FA647BFFA3FD04EB7BD350FEE7" author="Chen" box="[648,865,235,1667]" etAl="et al." firstAuthor="Chen" pageId="18" pageNumber="19" refId="ref39445" refString="Chen, Y., Joachimski, M. M., Richoz, S., Krystyn, L., AljinoviC, D., SmirCiC, D., Kolar-Jurkovsek, T., Lai, X., &amp; Zhang, Z. (2021). Smithian and Spathian (Early Triassic) conodonts from Oman and Croatia and their depth habitat revealed. Global and Planetary Change, 196, 103362. https: // doi. org / 10. 1016 / j. gloplacha. 2020.103362" type="journal volume" year="2021">Chen et al., 2021</bibRefCitation>
)
</taxonomicName>
. To date
<taxonomicName id="4C22D788647BFFA3FC43EDFBD44DFEE7" authorityName="Muller" authorityYear="1962" box="[975,1148,235,259]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FC43EDFBD44DFEE7" box="[975,1148,235,259]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
is mostly known from the Middle Triassic (e.g.
<bibRefCitation id="EFB3D1FA647BFFA3FB94EC1BD501FEC7" author="Goudemand" box="[1048,1328,267,291]" etAl="et al." firstAuthor="Goudemand" pageId="18" pageNumber="19" pagination="199 - 207" refId="ref41061" refString="Goudemand, N., Orchard, M. J., Bucher, H., &amp; Jenks, J. (2012 a). The elusive origin of Chiosella timorensis (Conodont Triassic). Geobios, 45 (2), 199 - 207." type="journal article" year="2012" yearSuffix="a">Goudemand et al., 2012a</bibRefCitation>
;
<bibRefCitation id="EFB3D1FA647BFFA3FACCEC1BD382FEA7" author="Orchard" etAl="et al." firstAuthor="Orchard" pageId="18" pageNumber="19" refId="ref45799" refString="Orchard, M. J., Gradinaru, E., &amp; Nicora, A. (2007 a). A summary of the conodont succession around the Olenekian-Anisian boundary at Desli Caira, North Dobrogea, Romania." type="book" year="2007" yearSuffix="a">Orchard et al., 2007a</bibRefCitation>
,
<bibRefCitation id="EFB3D1FA647BFFA3FC4FEC3BD434FEA7" author="Orchard" box="[963,1029,299,323]" firstAuthor="Orchard" pageId="18" pageNumber="19" pagination="347 - 354" refId="ref45840" refString="Orchard, M. J., Lehrmann, D. J., Wei, J., Wang, H., &amp; Taylor, H. J. (2007 b). Conodonts from the Olenekian-Anisian boundary beds, Guandao, Guizhou Province, China. New Mexico Museum of Natural History and Science Bulletin, 41, 347 - 354." type="journal article" year="2007" yearSuffix="b">2007b</bibRefCitation>
). If, as it seems, the here illustrated specimens and/or those from Chen et al., (2019, 2021) do indeed belong to
<taxonomicName id="4C22D788647BFFA3FBAEEC7BD4FEFE67" authorityName="Muller" authorityYear="1962" box="[1058,1231,363,387]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FBAEEC7BD4FEFE67" box="[1058,1231,363,387]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
, then they represent their oldest known representatives of this genus so far. A few unpublished specimens from middle Smithian rocks of
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may also belong to this genus (Leu et al., submitted; L. Dudit, pers. comm.), which further question the origin and age of
<taxonomicName id="4C22D788647BFFA3FBC0EF1BD4C8FDC7" authorityName="Muller" authorityYear="1962" box="[1100,1273,523,547]" genus="Gladigondolella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B9567019647BFFA3FBC0EF1BD4C8FDC7" box="[1100,1273,523,547]" italics="true" pageId="18" pageNumber="19">Gladigondolella</emphasis>
</taxonomicName>
. Its preference for deeper, colder habitats (
<bibRefCitation id="EFB3D1FA647BFFA3FBD5EF3BD517FDA7" author="Trotter" box="[1113,1318,555,579]" etAl="et al." firstAuthor="Trotter" pageId="18" pageNumber="19" pagination="165 - 174" refId="ref47635" refString="Trotter, J. A., Williams, I. S., Nicora, A., Mazza, M., &amp; Rigo, M. (2015). Long-term cycles of Triassic climate change: A new δ 18 O record from conodont apatite. Earth and Planetary Science Letters, 415, 165 - 174. https: // doi. org / 10.1016 / j. epsl. 2015.01. 038" type="journal article" year="2015">Trotter et al., 2015</bibRefCitation>
) may then explain its spotty occurrences within the Early Triassic, preferably during relatively colder intervals and/or in deeper, colder refuge areas like
<collectingCountry id="F335EC9B647BFFA3FB0DEF9BD4F8FD47" box="[1153,1225,651,675]" name="Oman" pageId="18" pageNumber="19">Oman</collectingCountry>
.
</paragraph>
<paragraph id="8B9DAC0B647BFFA3FCCEEFBCD47DFCC7" blockId="18.[813,1437,235,803]" pageId="18" pageNumber="19">
<emphasis id="B9567019647BFFA3FCCEEFBCD38CFD27" box="[834,957,684,707]" italics="true" pageId="18" pageNumber="19">Occurrence</emphasis>
. South
<collectingCountry id="F335EC9B647BFFA3FB99EFBBD468FD27" box="[1045,1113,683,707]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
;
<taxonomicName id="4C22D788647BFFA3FBEAEFBCD4DEFD27" baseAuthorityName="Solien" baseAuthorityYear="1979" box="[1126,1263,683,707]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="collinsoni">
<emphasis id="B9567019647BFFA3FBEAEFBCD4DEFD27" box="[1126,1263,683,707]" italics="true" pageId="18" pageNumber="19">Ic. collinsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788647BFFA3FAA5EFBCD5ADFD27" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[1321,1436,683,707]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B9567019647BFFA3FAA5EFBCD5ADFD27" box="[1321,1436,683,707]" italics="true" pageId="18" pageNumber="19">Tr. homeri</emphasis>
</taxonomicName>
zone, Jiarong (
<bibRefCitation id="EFB3D1FA647BFFA3FC40EFDBD4B8FD07" author="Chen" box="[972,1161,715,739]" etAl="et al." firstAuthor="Chen" pageId="18" pageNumber="19" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., &amp; Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al., 2015</bibRefCitation>
). Bed 54, Luolou formation,
<taxonomicName id="4C22D788647BFFA3FCEAEFFCD3DCFCE7" baseAuthorityName="Solien" baseAuthorityYear="1979" box="[870,1005,747,771]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="collinsoni">
<emphasis id="B9567019647BFFA3FCEAEFFCD3DCFCE7" box="[870,1005,747,771]" italics="true" pageId="18" pageNumber="19">Ic. collinsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C22D788647BFFA3FBAFEFFCD4A4FCE7" baseAuthorityName="Bender" baseAuthorityYear="1970" box="[1059,1173,747,771]" genus="Triassospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="homeri">
<emphasis id="B9567019647BFFA3FBAFEFFCD4A4FCE7" box="[1059,1173,747,771]" italics="true" pageId="18" pageNumber="19">Tr. homeri</emphasis>
</taxonomicName>
zone, Bianyang section,
<collectingRegion id="49E662E9647BFFA3FCA1EE1BD3BDFCC7" box="[813,908,779,803]" country="China" name="Guizhou" pageId="18" pageNumber="19">Guizhou</collectingRegion>
(
<bibRefCitation id="EFB3D1FA647BFFA3FC15EE1BD40FFCC7" author="Yan" box="[921,1086,779,803]" etAl="et al." firstAuthor="Yan" pageId="18" pageNumber="19" pagination="509 - 522" refId="ref48484" refString="Yan, C., Wang, L., Jiang, H., Wignall, P. B., Sun, Y., Chen, Y., &amp; Lai, X. (2013). Uppermost Permian to Lower Triassic conodonts at Bianyang section, Guizhou province, South China. Palaios, 28 (8), 509 - 522. https: // doi. org / 10.2110 / palo. 2012. p 12 - 077 r" type="journal article" year="2013">Yan et al., 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>