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<document ID-DOI="10.5281/zenodo.3382576" ID-GBIF-Dataset="48817a66-49e4-4e6f-b6c6-879db64b9ec9" ID-Zenodo-Dep="3382576" approvalRequired="52" approvalRequired_for_bibRefs="8" approvalRequired_for_illustrations="24" approvalRequired_for_taxonomicNames="20" checkinTime="1567247981485" checkinUser="plazi" docAuthor="Rauhut, Oliver W. M." docDate="2003" docId="77323C29FFD4B422FF109F78FDEFFBF2" docLanguage="en" docName="Rauhut2003BasalTheropod_ocr.pdf" docOrigin="Special papers in palaeontology 69" docStyle="DocumentStyle{}" docTitle="Dilophosaurus wetherilli Welles 1954" docType="treatment" docVersion="14" lastPageNumber="19" masterDocId="8B0B4451FFC7B431FFF59E70FFF0FFFF" masterDocTitle="The interrelationships and evolution of basal theropod dinosaurs" masterLastPageNumber="213" masterPageNumber="1" pageNumber="19" updateTime="1643591701704" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The interrelationships and evolution of basal theropod dinosaurs</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rauhut, Oliver W. M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Special papers in palaeontology</mods:title>
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<mods:part>
<mods:date>2003</mods:date>
<mods:detail type="pubDate">
<mods:number>2003-05-31</mods:number>
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<mods:detail type="volume">
<mods:number>69</mods:number>
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<mods:start>1</mods:start>
<mods:end>213</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.3382576</mods:identifier>
<mods:identifier type="GBIF-Dataset">48817a66-49e4-4e6f-b6c6-879db64b9ec9</mods:identifier>
<mods:identifier type="Zenodo-Dep">3382576</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5123153" ID-GBIF-Taxon="159404376" ID-Zenodo-Dep="5123153" LSID="urn:lsid:plazi:treatment:77323C29FFD4B422FF109F78FDEFFBF2" httpUri="http://treatment.plazi.org/id/77323C29FFD4B422FF109F78FDEFFBF2" lastPageNumber="19" pageId="19" pageNumber="19">
<subSubSection box="[229,916,264,300]" pageId="19" pageNumber="19" type="nomenclature">
<paragraph blockId="19.[227,916,206,358]" box="[229,916,264,300]" pageId="19" pageNumber="19">
<taxonomicName authority="(Welles, 1954)" authorityName="Welles" authorityYear="1954" baseAuthorityName="Welles" baseAuthorityYear="1954" box="[229,916,264,300]" class="Reptilia" family="Troodontidae" genus="Dilophosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="19" pageNumber="19" phylum="Chordata" rank="species" species="wetherilli">
<emphasis box="[229,655,264,300]" italics="true" pageId="19" pageNumber="19">Dilophosaurus wetherilli</emphasis>
(
<bibRefCitation author="Welles" box="[679,905,264,300]" firstAuthor="Welles" pageId="19" pageNumber="19" pagination="591 - 598" refId="ref144302" refString="WELLES, s. P. 1954. New Jurassic dinosaur from the Kayenta Formation of Arizona. Bulletin ofthe Geological Society of America, 65, 591 - 598." type="journal article" year="1954">Welles, 1954</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="19" type="distribution">
<paragraph blockId="19.[227,916,206,358]" box="[228,739,322,358]" pageId="19" pageNumber="19">
<emphasis box="[228,296,322,358]" italics="true" pageId="19" pageNumber="19">
<taxonomicName authorityName="Diakonoff" authorityYear="1982" box="[228,291,322,358]" class="Insecta" family="Tortricidae" genus="Age" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="19" phylum="Arthropoda" rank="genus">Age</taxonomicName>
.
</emphasis>
?Sinemurian-Pliensbachian.
</paragraph>
<paragraph blockId="19.[231,967,411,445]" box="[231,967,411,445]" pageId="19" pageNumber="19">
<emphasis box="[231,415,411,445]" italics="true" pageId="19" pageNumber="19">Occurrence.</emphasis>
Kayenta Formation, Arizona, USA.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="19" type="diagnosis">
<paragraph blockId="19.[229,1996,499,582]" pageId="19" pageNumber="19">
<emphasis box="[229,408,499,535]" italics="true" pageId="19" pageNumber="19">Diagnosis.</emphasis>
Lacrimal with thickened dorsoposterior rim; cervical neural spines with a distinct central cap and an anterior and posterior shoulder ; scapular blade with squared distal expansion.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="19" type="discussion">
<paragraph blockId="19.[229,1999,636,1039]" pageId="19" pageNumber="19">
<emphasis box="[229,830,636,672]" italics="true" pageId="19" pageNumber="19">
Remarks.
<taxonomicName baseAuthorityName="Welles" baseAuthorityYear="1954" box="[403,830,636,672]" class="Reptilia" family="Not" genus="Dilophosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="19" pageNumber="19" phylum="Chordata" rank="species" species="wetherilli">Dilophosaurus wetherilli</taxonomicName>
</emphasis>
(
<figureCitation box="[859,1066,636,672]" captionStart="text-fig. 5" captionText="text-fig. 5. Skull reconstructions of representatives of Jurassic OTUs in left lateral view, a, Dilophosaurus wetherilli, Early Jurassic (Sinemurian-Pliensbachian), Kayenta Formation, Arizona, USA; based on UCMP V 4214 and V 6468. B, Syntarsus rhodesiensis, Early Jurassic (Hettangian-Sinemurian), Forest Sandstone, Zimbabwe; composite reconstruction based on many isolated skull elements from the National Museum of Natural History in Harare (see Appendix), c, Magnosaurus oxoniensis, Middle Jurassic (Callovian), Oxford Clay, England; based on OUM J 13558, unpreserved elements shaded. D, Monolophosaurus jiangi, Middle Jurassic, Wucaiwan Formation, China; redrawn from Zhao and Currie (1993b). E, Allosaurus fragilis, Late Jurassic (Kimmeridgian-Tithonian), Morrison Formation, USA; based on MOR 693. f, basal bird Archaeopteryx sp., Late Jurassic (Tithonian), lithographic limestones of Solnhofen, Germany; based on Wellnhofer (1974), Elzanowski and Wellnhofer (1996), and the Berlin, Eichstätt, and Munich specimens. G, Ceratosaurus sp., Late Jurassic (Kimmeridgian-Tithonian), Morrison Formation, USA; based on USNM 4735 and UMNH VP 5278. H, Ornitholestes hermanni, Late Jurassic (Kimmeridgian-Tithonian), Morrison Formation, USA, based on AMNH 619. Abbreviations as in Text-figure 4, and: If, lacrimal fenestra; mf, maxillary fenestra; nf, nasal foramen; pmf, promaxillary fenestra. Scale bars represent 10 mm (b, f, h), 50 mm (c) and 100 mm (a, d, e, g)." httpUri="https://zenodo.org/record/3382584/files/figure.png" pageId="19" pageNumber="19" targetBox="[379,1958,617,2559]" targetPageId="24">Text-fig. 5a</figureCitation>
) was originally described as a new species of
<taxonomicName class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="19" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="19" pageNumber="19">Megalosaurus</emphasis>
</taxonomicName>
by Welles in 1954, but it was not until 1984 that a detailed description of the holotype was published (Welles 1984). The most striking feature of the skull are the two large, very thin cranial crests, best preserved in a so-far undescribed skull (UCMP V6468). However, since similar crests have been reported for
<taxonomicName authority="(Rowe 1989)" authorityName="Rowe" authorityYear="1989" baseAuthorityName="Rowe" baseAuthorityYear="1989" box="[444,837,819,855]" class="Reptilia" family="Coelophysidae" genus="Syntarsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="19" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis box="[444,606,819,855]" italics="true" pageId="19" pageNumber="19">Syntarsus</emphasis>
(
<bibRefCitation author="Rowe" box="[631,826,819,855]" firstAuthor="Rowe" pageId="19" pageNumber="19" pagination="125 - 136" refId="ref141428" refString="ROWE,. 1989. A new species of the theropod dinosaur Syntarsus from the Early Jurassic Kayenta Formation of Arizona. Journal of Vertebrate Paleontology, 9, 125 - 136." type="journal article" year="1989">Rowe 1989</bibRefCitation>
)
</taxonomicName>
and another species from the Lower Jurassic of China [based on the presence of these crests, this species was originally referred to the genus
<taxonomicName authorityName="Welles" authorityYear="1970" box="[1440,1689,865,901]" class="Reptilia" family="Troodontidae" genus="Dilophosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="19" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis box="[1440,1689,865,901]" italics="true" pageId="19" pageNumber="19">Dilophosaurus</emphasis>
</taxonomicName>
as
<taxonomicName authority="(Hu 1993)" authorityName="Hu" authorityYear="1993" baseAuthorityName="Hu" baseAuthorityYear="1993" class="Reptilia" family="Not" genus="Dilophosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="19" pageNumber="19" phylum="Chordata" rank="species" species="sinensis">
<emphasis box="[1743,1925,865,901]" italics="true" pageId="19" pageNumber="19">D. sinensis</emphasis>
(
<bibRefCitation author="Hu" firstAuthor="Hu" journalOrPublisher="Vertebrata PalAsiatica" pageId="19" pageNumber="19" pagination="65 - 69" part="31" refId="ref134763" refString="HU, s. 1993. A new Theropoda (Dilophosaurus sinensis sp. nov.) from Yunnan, China. Vertebrata PalAsiatica, 31, 65 - 69. [In Chinese]." title="A new Theropoda (Dilophosaurus sinensis sp. nov.) from Yunnan, China" type="journal article" year="1993">Hu 1993</bibRefCitation>
)
</taxonomicName>
, but this referral has recently been doubted (
<bibRefCitation author="Lamanna" box="[1078,1431,910,946]" etAl="et al." firstAuthor="Lamanna" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="19" pageNumber="19" pagination="57 A" part="18" refId="ref136159" refString="LAMANNA, M. c., smith, J. b., you, H., H oltz,. R. J. and dodson, p. 1998. A reassessment of the Chinese theropod dinosaur Dilophosaurus sinensis. Journal of Vertebrate Paleontology, 18 (Supplement to No. 3), 57 A." title="A reassessment of the Chinese theropod dinosaur Dilophosaurus sinensis" type="journal volume" year="1998">
Lamanna
<emphasis box="[1246,1327,910,946]" italics="true" pageId="19" pageNumber="19">et al.</emphasis>
1998
</bibRefCitation>
)], this character cannot be used to define the genus or species, but
<taxonomicName baseAuthorityName="Welles" baseAuthorityYear="1954" box="[782,996,955,991]" class="Reptilia" family="Not" genus="Dilophosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="19" pageNumber="19" phylum="Chordata" rank="species" species="wetherilli">
<emphasis box="[782,996,955,991]" italics="true" pageId="19" pageNumber="19">D. wetherilli</emphasis>
</taxonomicName>
shows enough other apomorphies to establish it as a valid genus and species.
</paragraph>
</subSubSection>
</treatment>
</document>