treatments-xml/data/03/D7/87/03D787E6FFE2FFE6FF6BA2FC3D5CBFBF.xml
2024-06-21 12:22:17 +02:00

218 lines
26 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="81FE1000B8FA8DED5308E16999D691C4" ID-CLB-Dataset="45540" ID-DOI="10.11646/zootaxa.3814.3.2" ID-GBIF-Dataset="708ea530-4c8d-473b-8f29-1a54dea92f28" ID-ISSN="1175-5326" ID-Zenodo-Dep="231107" ID-ZooBank="5391C375-896D-4794-84A6-1846DF9E407D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460434473942" checkinUser="plazi" docAuthor="Kania, Iwona" docDate="2014" docId="03D787E6FFE2FFE6FF6BA2FC3D5CBFBF" docLanguage="en" docName="zt03814p352.pdf" docOrigin="Zootaxa 3814 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helius formosus Krzeminski 1993" docType="treatment" docVersion="9" lastPageNumber="337" masterDocId="FFEEFF9EFFE1FFE2FFFCA26B3C50BD09" masterDocTitle="Subfamily Limoniinae Speiser, 1909 (Diptera, Limoniidae) from Baltic amber (Eocene): the genus Helius Lepeletier &amp; Serville, 1828" masterLastPageNumber="352" masterPageNumber="333" pageNumber="336" updateTime="1698331894480" updateUser="ExternalLinkService">
<mods:mods id="97CA495FCFD24C112CE1FCA7ED03D543" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3415344EB264069F1628F3DD419503A5">
<mods:title id="53F751BF2D9A2D09C3BB7B4B06D59CD7">Subfamily Limoniinae Speiser, 1909 (Diptera, Limoniidae) from Baltic amber (Eocene): the genus Helius Lepeletier &amp; Serville, 1828</mods:title>
</mods:titleInfo>
<mods:name id="3B5EE328AFD731769E42166D5DA123FA" type="personal">
<mods:role id="0D1063D1E16B4F23A5E35C47F38CAEC3">
<mods:roleTerm id="04314E1619EE6451FF2806D44F944415">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13F73E1359A6C8B35D5A264C40425EE0">Kania, Iwona</mods:namePart>
</mods:name>
<mods:typeOfResource id="EB6563A16974836B0BBD75368A21C87A">text</mods:typeOfResource>
<mods:relatedItem id="7C1BDDA7E6DDC6034C6FB6790CF76402" type="host">
<mods:titleInfo id="2CA609244D241C0220EB3AD055347282">
<mods:title id="CFDB06687FA651005557276FF6684DD6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5421B76CF4CFA957D52BDBA81ACB96A3">
<mods:date id="DEBE42456465ABD6E4E0DFE46B2057D2">2014</mods:date>
<mods:detail id="B084B313C707651AD9A01C47D17BB059" type="volume">
<mods:number id="C278B0E9880CBF694D3AB1C0A0B72A56">3814</mods:number>
</mods:detail>
<mods:detail id="824C59A40633443799C7B47559A904B5" type="issue">
<mods:number id="77C64097E2C65EB39DCB32B69E4C5AEC">3</mods:number>
</mods:detail>
<mods:extent id="E5703C832A1B010466B967F6E282448C" unit="page">
<mods:start id="A824295713A2600A6522B769112CBCA8">333</mods:start>
<mods:end id="B25B0294F2E653DA3B060E50AB9A5F51">352</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="814CC362F62BCD4409F89A10E0B2D1B4">journal article</mods:classification>
<mods:identifier id="D33C49AC7A91BB0D64205293B843F78D" type="CLB-Dataset">45540</mods:identifier>
<mods:identifier id="B79411D210F3ABC3A843B5A1B65AEEFC" type="DOI">10.11646/zootaxa.3814.3.2</mods:identifier>
<mods:identifier id="CAA400C4C9D273469E583422499E29FC" type="GBIF-Dataset">708ea530-4c8d-473b-8f29-1a54dea92f28</mods:identifier>
<mods:identifier id="10159A9702161D679F613E7DEF05E2FA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6882F3EDB575814970779185345B9852" type="Zenodo-Dep">231107</mods:identifier>
<mods:identifier id="0FB364B14EB2DB5C3E5861A3AA7730B0" type="ZooBank">5391C375-896D-4794-84A6-1846DF9E407D</mods:identifier>
</mods:mods>
<treatment id="03D787E6FFE2FFE6FF6BA2FC3D5CBFBF" ID-DOI="http://doi.org/10.5281/zenodo.6138156" ID-GBIF-Taxon="119431375" ID-Zenodo-Dep="6138156" LSID="urn:lsid:plazi:treatment:03D787E6FFE2FFE6FF6BA2FC3D5CBFBF" httpUri="http://treatment.plazi.org/id/03D787E6FFE2FFE6FF6BA2FC3D5CBFBF" lastPageId="4" lastPageNumber="337" pageId="3" pageNumber="336">
<subSubSection id="C364657BFFE2FFE1FF6BA2FC3E99BC1E" pageId="3" pageNumber="336" type="nomenclature">
<paragraph id="8BC136F0FFE2FFE1FF6BA2FC3E12BDBB" blockId="3.[151,578,151,211]" box="[151,578,151,178]" pageId="3" pageNumber="336">
<heading id="D089819CFFE2FFE1FF6BA2FC3E12BDBB" bold="true" box="[151,578,151,178]" fontSize="11" level="1" pageId="3" pageNumber="336" reason="1">
<taxonomicName id="4C7E4D73FFE2FFE1FF6BA2FC3E12BDBB" authority="Krzeminski, 1993" authorityName="Krzeminski" authorityYear="1993" box="[151,578,151,178]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B90AEAE2FFE2FFE1FF6BA2FC3E12BDBB" bold="true" box="[151,578,151,178]" pageId="3" pageNumber="336">
<emphasis id="B90AEAE2FFE2FFE1FF6BA2FC3D0EBDB8" bold="true" box="[151,350,151,177]" italics="true" pageId="3" pageNumber="336">Helius formosus</emphasis>
<bibRefCitation id="EFEF4B01FFE2FFE1FE99A2F33E12BDBB" author="Krzeminski" box="[357,578,152,178]" pageId="3" pageNumber="336" refString="Krzeminski, W. (1993) Fossil Tipulomorpha (Diptera, Nematocera) from Baltic amber (Upper Eocene). - Revision of the genus Helius Lepeletier et Serville (Limoniidae). Acta Zoologica Cracoviensia, 35, 597 - 601." type="journal article" year="1993">Krzemiński, 1993</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC136F0FFE2FFE1FF6BA2D13D74BDDA" blockId="3.[151,578,151,211]" box="[151,292,186,211]" pageId="3" pageNumber="336">
(
<figureCitation id="13452A75FFE2FFE1FF63A2D13CB5BDDA" box="[159,229,186,211]" captionStart="FIGURE 2" captionStartId="3.[151,250,1525,1547]" captionTargetBox="[343,1248,589,1504]" captionTargetId="figure@3.[336,1255,581,1510]" captionTargetPageId="3" captionText="FIGURE 2. Helius formosus Krzemiński 1993, No. 225 (female), holotype; A, head; B, relation between the length of palpi (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231109/files/figure.png" pageId="3" pageNumber="336">Figs 2</figureCitation>
AC)
</paragraph>
<paragraph id="8BC136F0FFE2FFE1FF6BA36A3E99BC1E" blockId="3.[151,713,257,279]" box="[151,713,257,279]" pageId="3" pageNumber="336">
v*.1993
<taxonomicName id="4C7E4D73FFE2FFE1FF13A36A3E44BC1E" authority="Krzeminski" authorityName="Krzeminski" box="[239,532,257,279]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B90AEAE2FFE2FFE1FF13A36A3DC5BC1E" box="[239,405,257,279]" italics="true" pageId="3" pageNumber="336">Helius formosus</emphasis>
Krzemiński
</taxonomicName>
, p. 600, figs. 89.
</paragraph>
</subSubSection>
<subSubSection id="C364657BFFE2FFE1FF6BA32F3F50BF19" pageId="3" pageNumber="336" type="materials_examined">
<paragraph id="8BC136F0FFE2FFE1FF6BA32F3EADBC89" blockId="3.[151,1436,324,528]" pageId="3" pageNumber="336">
<emphasis id="B90AEAE2FFE2FFE1FF6BA32F3DD2BC54" bold="true" box="[151,386,324,349]" pageId="3" pageNumber="336">Material examined.</emphasis>
<typeStatus id="54C58852FFE2FFE1FE77A32F3DA8BC54" box="[395,504,324,349]" pageId="3" pageNumber="336" type="holotype">Holotype</typeStatus>
: No. 225 (female) (
<collectionCode id="ED6FAE35FFE2FFE1FD24A32F3F7FBC55" box="[728,815,324,348]" pageId="3" pageNumber="336">ZMHB</collectionCode>
); additional material: No. 212/116 (female) (
<collectionCode id="ED6FAE35FFE2FFE1FAD3A32F39C1BC55" box="[1327,1425,324,348]" pageId="3" pageNumber="336">MEPAS</collectionCode>
); No. MP/3340 (female); No. MP/3341 (female) (
<collectionCode id="ED6FAE35FFE2FFE1FD4DA3033EA1BC89" box="[689,753,360,384]" httpUri="http://grbio.org/cool/hv18-23j2" name="Institute of Systematic Zoology" pageId="3" pageNumber="336">ISZP</collectionCode>
).
</paragraph>
<paragraph id="8BC136F0FFE2FFE1FF3BA3E73F50BF19" blockId="3.[151,1436,324,528]" pageId="3" pageNumber="336">
<emphasis id="B90AEAE2FFE2FFE1FF3BA3E73D82BCAC" bold="true" box="[199,466,396,421]" pageId="3" pageNumber="336">Diagnostic characters.</emphasis>
Rostrum less than twice as long as the head, longer than palpus, only slightly shorter than antenna; antenna only 1.2x the length of the rostrum, palpus 0.750.8x the length of the rostrum; the last palpal segment almost 1.25x the length of all the preceding segments together; vein r-m rather short, cross-vein mcu in approximately proximal 1/3 of d-cell base length.
</paragraph>
</subSubSection>
<caption id="DF016678FFE2FFE1FF6BA79E3EA4BB23" httpUri="https://zenodo.org/record/231109/files/figure.png" pageId="3" pageNumber="336" targetBox="[343,1248,589,1504]" targetPageId="3">
<paragraph id="8BC136F0FFE2FFE1FF6BA79E3EA4BB23" blockId="3.[151,1436,1524,1578]" pageId="3" pageNumber="336">
<emphasis id="B90AEAE2FFE2FFE1FF6BA79E3D43BB03" bold="true" box="[151,275,1525,1547]" pageId="3" pageNumber="336">FIGURE 2.</emphasis>
<taxonomicName id="4C7E4D73FFE2FFE1FEE7A79F3ED2BB02" authority="Krzeminski 1993" authorityName="Krzeminski" authorityYear="1993" box="[283,642,1524,1547]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B90AEAE2FFE2FFE1FEE7A79F3D98BB02" bold="true" box="[283,456,1524,1547]" italics="true" pageId="3" pageNumber="336">Helius formosus</emphasis>
Krzemiński 1993
</taxonomicName>
, No. 225 (female), holotype; A, head; B, relation between the length of palpi (p), antenna (a), rostrum (r) and head (h); C, wing venation.
</paragraph>
</caption>
<subSubSection id="C364657BFFE2FFE6FF3BA43A3FCBBC66" lastPageId="4" lastPageNumber="337" pageId="3" pageNumber="336" type="description">
<paragraph id="8BC136F0FFE2FFE1FF3BA43A3831BB63" blockId="3.[151,1437,1617,2002]" box="[199,1121,1617,1642]" pageId="3" pageNumber="336">
<emphasis id="B90AEAE2FFE2FFE1FF3BA43A3D24BB63" bold="true" box="[199,372,1617,1642]" pageId="3" pageNumber="336">Redescription.</emphasis>
Body: the length of specimens
<quantity id="4C869B15FFE2FFE1FD2BA43A3F2CBB60" box="[727,892,1617,1642]" metricMagnitude="-3" metricUnit="m" metricValue="6.4399999999999995" metricValueMax="7.54" metricValueMin="5.34" pageId="3" pageNumber="336" unit="mm" value="6.4399999999999995" valueMax="7.54" valueMin="5.34">5.347.54 mm</quantity>
, (
<quantity id="4C869B15FFE2FFE1FC73A43A3FB7BB60" box="[911,999,1617,1642]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="336" unit="mm" value="5.7">5.7 mm</quantity>
<typeStatus id="54C58852FFE2FFE1FC11A43A3804BB63" box="[1005,1108,1617,1642]" pageId="3" pageNumber="336" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BC136F0FFE2FFE1FF3BA41E3E65BA6F" blockId="3.[151,1437,1617,2002]" pageId="3" pageNumber="336">
Head (
<figureCitation id="13452A75FFE2FFE1FEE8A41E3D0ABB87" box="[276,346,1653,1678]" captionStart="FIGURE 2" captionStartId="3.[151,250,1525,1547]" captionTargetBox="[343,1248,589,1504]" captionTargetId="figure@3.[336,1255,581,1510]" captionTargetPageId="3" captionText="FIGURE 2. Helius formosus Krzemiński 1993, No. 225 (female), holotype; A, head; B, relation between the length of palpi (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231109/files/figure.png" pageId="3" pageNumber="336">Fig. 2</figureCitation>
A): rostrum
<quantity id="4C869B15FFE2FFE1FE14A41E3EDEBB84" box="[488,654,1653,1678]" metricMagnitude="-3" metricUnit="m" metricValue="1.025" metricValueMax="1.16" metricValueMin="0.89" pageId="3" pageNumber="336" unit="mm" value="1.025" valueMax="1.16" valueMin="0.89">0.891.16 mm</quantity>
long,
<quantity id="4C869B15FFE2FFE1FD25A41D3F20BB84" box="[729,880,1654,1678]" metricMagnitude="-4" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="3" pageNumber="336" unit="mm" value="0.105" valueMax="0.11" valueMin="0.1">0.10.11 mm</quantity>
wide,
<quantity id="4C869B15FFE2FFE1FC3CA41D3877BB84" box="[960,1063,1654,1678]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="3" pageNumber="336" unit="mm" value="1.09">1.09 mm</quantity>
long, 0.09 wide (
<typeStatus id="54C58852FFE2FFE1FB09A41E390DBB87" box="[1269,1373,1653,1678]" pageId="3" pageNumber="336" type="holotype">holotype</typeStatus>
), less than twice as long as the head; antenna 16-segmented,
<quantity id="4C869B15FFE2FFE1FCF1A4F13FE2BBB8" box="[781,946,1690,1714]" metricMagnitude="-3" metricUnit="m" metricValue="1.165" metricValueMax="1.24" metricValueMin="1.09" pageId="3" pageNumber="336" unit="mm" value="1.165" valueMax="1.24" valueMin="1.09">1.091.24 mm</quantity>
long (
<typeStatus id="54C58852FFE2FFE1FC00A4F23834BBBB" box="[1020,1124,1689,1714]" pageId="3" pageNumber="336" type="holotype">holotype</typeStatus>
), short, approximately 1.2x the length of the rostrum (
<figureCitation id="13452A75FFE2FFE1FE39A4D63E5BBBDF" box="[453,523,1725,1750]" captionStart="FIGURE 2" captionStartId="3.[151,250,1525,1547]" captionTargetBox="[343,1248,589,1504]" captionTargetId="figure@3.[336,1255,581,1510]" captionTargetPageId="3" captionText="FIGURE 2. Helius formosus Krzemiński 1993, No. 225 (female), holotype; A, head; B, relation between the length of palpi (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231109/files/figure.png" pageId="3" pageNumber="336">Fig. 2</figureCitation>
B); scape cylindrical, large; pedicel large, its distal part conspicuously widened; first flagellomere barrel-shaped, flagellomeres 25 oval, short, flagellomeres 610 cylindrical, elongated; flagellomeres 28 with two elongated setae; flagellomeres 913 with three additional setae, much longer than the length of flagellomeres which they occur, 23 times longer than the segments bearing them (
<figureCitation id="13452A75FFE2FFE1FB51A54238ACBA4B" box="[1197,1276,1833,1858]" captionStart="FIGURE 11" captionStartId="14.[151,250,1881,1903]" captionTargetBox="[201,1391,193,1853]" captionTargetId="figure@14.[192,1394,193,1860]" captionTargetPageId="14" captionText="FIGURE 11. Morphology of Helius from Baltic amber. A B H. fossilis sp. nov., No. 587 (RS 64) (male), holotype, A. the body, lateral view; B. head, lateral view C. H. hoffeinsorum sp. nov., No. 1462 (male), hypopygium; D E H. formosus Krzemiński, 1993, No. 225 (female), holotype, D. antennae, E. wing. Abbreviation of male hypopygium: gx—gonocoxite, ing—inner gonostylus, oug—outer gonostylus, aed—aedeagus, p—paramere." httpUri="https://zenodo.org/record/231118/files/figure.png" pageId="3" pageNumber="336">Fig. 11</figureCitation>
D); last one at least twice the length of penultimate.
</paragraph>
<paragraph id="8BC136F0FFE2FFE1FF3BA51A387FBADB" blockId="3.[151,1437,1617,2002]" pageId="3" pageNumber="336">
Palpus (
<figureCitation id="13452A75FFE2FFE1FEDFA51A3D38BA83" box="[291,360,1905,1930]" captionStart="FIGURE 2" captionStartId="3.[151,250,1525,1547]" captionTargetBox="[343,1248,589,1504]" captionTargetId="figure@3.[336,1255,581,1510]" captionTargetPageId="3" captionText="FIGURE 2. Helius formosus Krzemiński 1993, No. 225 (female), holotype; A, head; B, relation between the length of palpi (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231109/files/figure.png" pageId="3" pageNumber="336">Fig. 2</figureCitation>
A):
<quantity id="4C869B15FFE2FFE1FE69A51A3E6ABA80" box="[405,570,1905,1930]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" metricValueMax="8.1" metricValueMin="6.1" pageId="3" pageNumber="336" unit="mm" value="0.71" valueMax="0.81" valueMin="0.61">0.610.81 mm</quantity>
long (
<typeStatus id="54C58852FFE2FFE1FD7AA51A3EBEBA83" box="[646,750,1905,1930]" pageId="3" pageNumber="336" type="holotype">holotype</typeStatus>
); the length of last palpal segment
<quantity id="4C869B15FFE2FFE1FB7EA51A3979BA80" box="[1154,1321,1905,1930]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="3" pageNumber="336" unit="mm" value="0.375" valueMax="0.4" valueMin="0.35">0.350.40 mm</quantity>
,
<quantity id="4C869B15FFE2FFE1FACBA51939CDBA80" box="[1335,1437,1906,1930]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="336" unit="mm" value="0.38">0.38 mm</quantity>
(
<typeStatus id="54C58852FFE2FFE1FF63A5FE3D57BAA7" box="[159,263,1941,1966]" pageId="3" pageNumber="336" type="holotype">holotype</typeStatus>
), penultimate
<quantity id="4C869B15FFE2FFE1FE59A5FE3E16BAA4" box="[421,582,1941,1966]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="3" pageNumber="336" unit="mm" value="0.13" valueMax="0.15" valueMin="0.11">0.110.15 mm</quantity>
(
<typeStatus id="54C58852FFE2FFE1FDA9A5FE3EEDBAA7" box="[597,701,1941,1966]" pageId="3" pageNumber="336" type="holotype">holotype</typeStatus>
), first and the second segments
<quantity id="4C869B15FFE2FFE1FBDCA5FD3892BAA4" box="[1056,1218,1942,1966]" metricMagnitude="-5" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="3" pageNumber="336" unit="mm" value="0.095" valueMax="0.1" valueMin="0.09">0.090.10 mm</quantity>
(
<typeStatus id="54C58852FFE2FFE1FB2DA5FE3969BAA7" box="[1233,1337,1941,1966]" pageId="3" pageNumber="336" type="holotype">holotype</typeStatus>
); the last palpal segment almost 1.25x the length of the all the preceding segments together.
</paragraph>
<paragraph id="8BC136F0FFE5FFE6FF3BA2FC3E36BC40" blockId="4.[151,1436,151,699]" pageId="4" pageNumber="337">
Wing (
<figureCitation id="13452A75FFE5FFE6FEEEA2FC3D0ABDB9" box="[274,346,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1525,1547]" captionTargetBox="[343,1248,589,1504]" captionTargetId="figure@3.[336,1255,581,1510]" captionTargetPageId="3" captionText="FIGURE 2. Helius formosus Krzemiński 1993, No. 225 (female), holotype; A, head; B, relation between the length of palpi (p), antenna (a), rostrum (r) and head (h); C, wing venation." httpUri="https://zenodo.org/record/231109/files/figure.png" pageId="4" pageNumber="337">Figs 2</figureCitation>
C, 11E): wing
<quantity id="4C869B15FFE5FFE6FE00A2FC3ECEBDA6" box="[508,670,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.43" metricValueMax="7.52" metricValueMin="5.34" pageId="4" pageNumber="337" unit="mm" value="6.43" valueMax="7.52" valueMin="5.34">5.347.52 mm</quantity>
length and
<quantity id="4C869B15FFE5FFE6FCDCA2FC3F93BDA6" box="[800,963,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" metricValueMax="1.97" metricValueMin="1.05" pageId="4" pageNumber="337" unit="mm" value="1.51" valueMax="1.97" valueMin="1.05">1.051.97 mm</quantity>
wide; pterostigma present, large and oval; vein Sc elongated, ending opposite 3/4 the length of Rs; cross-vein sc-r before the end of Sc; R1 ending at the level of R2+3+4 midlength; r-r (R2) atrophied; R2+3+4 elongated and sinuous; cell r2 very wide; d-cell very broad and large, closed; M3 longer than d-cell and longer than half the length of R5; cross-vein m-cu in proximal 1/3 of d-cell base; A1 and A2 elongated, slightly subsinuous.
</paragraph>
<paragraph id="8BC136F0FFE5FFE6FF3BA33C3FCBBC66" blockId="4.[151,1436,151,699]" box="[199,923,343,368]" pageId="4" pageNumber="337">Male hypopygium unknown, female ovipositor long and narrow.</paragraph>
</subSubSection>
<subSubSection id="C364657BFFE5FFE6FF3BA3103D5CBFBF" pageId="4" pageNumber="337" type="discussion">
<paragraph id="8BC136F0FFE5FFE6FF3BA3103EC9BF2D" blockId="4.[151,1436,151,699]" pageId="4" pageNumber="337">
<emphasis id="B90AEAE2FFE5FFE6FF3BA3103D6BBC9D" bold="true" box="[199,315,379,404]" pageId="4" pageNumber="337">Remarks.</emphasis>
The species distinctly differ from
<taxonomicName id="4C7E4D73FFE5FFE6FD41A317384BBC9A" authority="Krzeminski, 1993" authorityName="Krzeminski" authorityYear="1993" box="[701,1051,379,404]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE5FFE6FD41A3173F69BC9A" box="[701,825,380,403]" italics="true" pageId="4" pageNumber="337">H. minutus</emphasis>
(
<bibRefCitation id="EFEF4B01FFE5FFE6FCBBA3103842BC9A" author="Krzeminski" box="[839,1042,379,404]" pageId="4" pageNumber="337" refString="Krzeminski, W. (1993) Fossil Tipulomorpha (Diptera, Nematocera) from Baltic amber (Upper Eocene). - Revision of the genus Helius Lepeletier et Serville (Limoniidae). Acta Zoologica Cracoviensia, 35, 597 - 601." type="journal article" year="1993">Krzemiński, 1993</bibRefCitation>
)
</taxonomicName>
in ration of the lengths of rostrum and the head. In
<taxonomicName id="4C7E4D73FFE5FFE6FEACA3CB3D89BCBE" box="[336,473,415,439]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B90AEAE2FFE5FFE6FEACA3CB3D89BCBE" box="[336,473,415,439]" italics="true" pageId="4" pageNumber="337">H. formosus</emphasis>
</taxonomicName>
rostrum is slightly less than twice as long as the head, in
<taxonomicName id="4C7E4D73FFE5FFE6FBA7A3CB3886BCBE" box="[1115,1238,416,439]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE5FFE6FBA7A3CB3886BCBE" box="[1115,1238,416,439]" italics="true" pageId="4" pageNumber="337">H. minutus</emphasis>
</taxonomicName>
rostrum is longer than head only approximately 0.2x. In
<taxonomicName id="4C7E4D73FFE5FFE6FDB1A3AF3E99BCD2" box="[589,713,452,475]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE5FFE6FDB1A3AF3E99BCD2" box="[589,713,452,475]" italics="true" pageId="4" pageNumber="337">H. minutus</emphasis>
</taxonomicName>
antenna is more than twice longer than rostrum, in
<taxonomicName id="4C7E4D73FFE5FFE6FAEEA3AF39CCBCD2" box="[1298,1436,451,475]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B90AEAE2FFE5FFE6FAEEA3AF39CCBCD2" box="[1298,1436,451,475]" italics="true" pageId="4" pageNumber="337">H. formosus</emphasis>
</taxonomicName>
antenna is only 1.2x the length of the rostrum. In
<taxonomicName id="4C7E4D73FFE5FFE6FD35A3833F15BCF6" box="[713,837,488,511]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE5FFE6FD35A3833F15BCF6" box="[713,837,488,511]" italics="true" pageId="4" pageNumber="337">H. minutus</emphasis>
</taxonomicName>
the last palpal segment is about equal two preceding segments together, in
<taxonomicName id="4C7E4D73FFE5FFE6FE77A0673E44BF2A" box="[395,532,523,547]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B90AEAE2FFE5FFE6FE77A0673E44BF2A" box="[395,532,523,547]" italics="true" pageId="4" pageNumber="337">H. formosus</emphasis>
</taxonomicName>
it is longer.
</paragraph>
<paragraph id="8BC136F0FFE5FFE6FF3BA0443D5CBFBF" blockId="4.[151,1436,151,699]" pageId="4" pageNumber="337">
These two species from Baltic amber also differ by wing venation: in
<taxonomicName id="4C7E4D73FFE5FFE6FC2EA05B380BBF4E" box="[978,1115,559,583]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B90AEAE2FFE5FFE6FC2EA05B380BBF4E" box="[978,1115,559,583]" italics="true" pageId="4" pageNumber="337">H. formosus</emphasis>
</taxonomicName>
vein Sc is elongated, ending opposite 3/4 the length of Rs, in
<taxonomicName id="4C7E4D73FFE5FFE6FDF6A03F3ED6BF62" box="[522,646,596,619]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE5FFE6FDF6A03F3ED6BF62" box="[522,646,596,619]" italics="true" pageId="4" pageNumber="337">H. minutus</emphasis>
</taxonomicName>
this vein is shorter, ending opposite 1/3 of Rs. Cross-vein sc-r ending before the end of Sc in
<taxonomicName id="4C7E4D73FFE5FFE6FE65A0133E71BF86" box="[409,545,631,655]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B90AEAE2FFE5FFE6FE65A0133E71BF86" box="[409,545,631,655]" italics="true" pageId="4" pageNumber="337">H. formosus</emphasis>
</taxonomicName>
, in
<taxonomicName id="4C7E4D73FFE5FFE6FDB0A0133E96BF86" box="[588,710,632,655]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B90AEAE2FFE5FFE6FDB0A0133E96BF86" box="[588,710,632,655]" italics="true" pageId="4" pageNumber="337">H. minutus</emphasis>
</taxonomicName>
at the end. In
<taxonomicName id="4C7E4D73FFE5FFE6FC98A0133FBDBF86" box="[868,1005,631,655]" class="Insecta" family="Limoniidae" genus="Helius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="337" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B90AEAE2FFE5FFE6FC98A0133FBDBF86" box="[868,1005,631,655]" italics="true" pageId="4" pageNumber="337">H. formosus</emphasis>
</taxonomicName>
R1 ending opposite 1/2 not opposite 1/ 4 of R2+3+4.
</paragraph>
</subSubSection>
</treatment>
</document>