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<document id="C76F440C09958C0E44E9C31F72895F18" ID-CLB-Dataset="51322" ID-DOI="10.5281/zenodo.156243" ID-GBIF-Dataset="7b470930-22b9-4a4c-af32-33db81b72fdc" ID-ISSN="1175­5326" ID-Zenodo-Dep="156243" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459615472219" checkinUser="plazi" docAuthor="Flowers, R. Wills" docDate="2003" docId="03FF01389A794349FEDFFBA6FAACF97F" docLanguage="en" docName="zt00221.pdf" docOrigin="Zootaxa 221" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Thysanomeros ulateae Flowers, new species" docType="treatment" docVersion="11" lastPageNumber="10" masterDocId="FFC679409A7F4340FFD7FFC2FFA6FFF0" masterDocTitle="A new genus and a new Costa Rican species of Endocephalites (Chrysomelidae: Eumolpinae: Eumolpini)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="7" updateTime="1698204370072" updateUser="ExternalLinkService">
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<mods:title id="43FFFB4D73256021273DD5490B01E18F">A new genus and a new Costa Rican species of Endocephalites (Chrysomelidae: Eumolpinae: Eumolpini)</mods:title>
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<mods:namePart id="95C4F52E7D32354E76065C658B3C92CE">Flowers, R. Wills</mods:namePart>
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<mods:date id="0E0A2CE4F5E01241BD32770F6A20E1ED">2003</mods:date>
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<mods:classification id="ED4098EC022C4946E3C7DA5A672788B8">journal article</mods:classification>
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<treatment id="03FF01389A794349FEDFFBA6FAACF97F" ID-DOI="http://doi.org/10.5281/zenodo.6276813" ID-GBIF-Taxon="119331675" ID-Zenodo-Dep="6276813" LSID="urn:lsid:plazi:treatment:03FF01389A794349FEDFFBA6FAACF97F" httpUri="http://treatment.plazi.org/id/03FF01389A794349FEDFFBA6FAACF97F" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7">
<subSubSection id="C34CE3A59A794346FEDFFBA6FDD5FB51" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BE9B02E9A794346FEDFFBA6FC87FB8E" blockId="6.[264,801,1124,1185]" box="[264,801,1124,1150]" pageId="6" pageNumber="7">
<heading id="D0A107429A794346FEDFFBA6FC87FB8E" bold="true" box="[264,801,1124,1150]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B9226C3C9A794346FEDFFBA6FC87FB8E" bold="true" box="[264,801,1124,1150]" pageId="6" pageNumber="7">
<taxonomicName id="4C56CBAD9A794346FEDFFBA6FD23FB8E" authority="Flowers" authorityName="Flowers" box="[264,645,1124,1150]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ulateae" status="sp. nov.">
<emphasis id="B9226C3C9A794346FEDFFBA6FDBEFB8E" bold="true" box="[264,536,1124,1150]" italics="true" pageId="6" pageNumber="7">Thysanomeros ulateae</emphasis>
Flowers
</taxonomicName>
,
<taxonomicNameLabel id="A211D1479A794346FD45FBA6FC87FB8E" box="[658,801,1124,1150]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE9B02E9A794346FEDFFB45FDD5FB51" blockId="6.[264,801,1124,1185]" box="[264,627,1159,1185]" pageId="6" pageNumber="7">
(
<figureCitation id="136DACAB9A794346FEC6FB45FEC4FB51" box="[273,354,1159,1185]" captionStart="FIGURE 1" captionStartId="1.[264,366,1485,1509]" captionText="FIGURE 1. Thysanomeros ulateae, female, dorsal view." pageId="6" pageNumber="7">Figs. 1</figureCitation>
,
<figureCitation id="136DACAB9A794346FEA7FB45FE3BFB51" box="[368,413,1159,1185]" captionStart="FIGURES 2 ­ 7" captionStartId="3.[264,382,1698,1722]" captionTargetBox="[303,1285,1089,1667]" captionTargetId="figure@3.[301,1287,1089,1672]" captionTargetPageId="3" captionText="FIGURES 2 ­ 7. External characters of Thysanomeros. 2 3. Hind femur of male. 2. T. jacobyi. 3. T. ulateae. 4 5. Lateral margin of pronotum. 4. T. jacobyi, 5. T. ulateae. 6. Fore tarsi of T. jacobyi: left, male; right, female. 7. Pygidium of T. jacobyi." httpUri="https://zenodo.org/record/156244/files/figure.png" pageId="6" pageNumber="7">3, 5</figureCitation>
,
<figureCitation id="136DACAB9A794346FE7CFB45FDA3FB51" box="[427,517,1159,1185]" captionStart="FIGURES 8 ­ 12" captionStartId="5.[264,382,1216,1240]" captionTargetBox="[326,1260,409,1187]" captionTargetId="figure@5.[325,1263,409,1189]" captionTargetPageId="5" captionText="FIGURES 8 ­ 12. Male genitalia of Thysanomeros. 8, 10. Median lobe. 8 T. jacobyi, showing endophalus and apical sclerite. 10. T. ulateae. 9, 12. Apex of median lobe. 9. T. jacobyi. 12. T. ulateae. 11. Apical sclerite, T. ulateae." httpUri="https://zenodo.org/record/156245/files/figure.png" pageId="6" pageNumber="7">10 12</figureCitation>
,
<figureCitation id="136DACAB9A794346FDC6FB45FDCCFB51" box="[529,618,1159,1185]" captionStart-0="FIGURES 13 ­ 17" captionStart-1="FIGURE 18" captionStartId-0="7.[264,382,1219,1243]" captionStartId-1="9.[264,368,1218,1242]" captionTargetBox-0="[317,1269,409,1191]" captionTargetBox-1="[264,1306,315,1209]" captionTargetId-0="figure@7.[316,1272,409,1193]" captionTargetId-1="figure@9.[264,1322,315,1215]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIGURES 13 ­ 17. Female characters of Thysanomeros. 13, 16 ­ 17. ovipositors. 13. T. jacobyi, ventral. 16. T. ulateae, ventral. 17. T. ulateae, dorsal. 14 ­ 15. spermatheca. 14. T. jacobyi. 15. T. ulateae." captionText-1="FIGURE 18. Distribution of Thysanomeros. T. jacobyi, closed circles. T. ulateae, closed squares." httpUri-0="https://zenodo.org/record/156246/files/figure.png" httpUri-1="https://zenodo.org/record/156247/files/figure.png" pageId="6" pageNumber="7">15 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34CE3A59A794347FEDFFB14FBCFF8AB" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BE9B02E9A794347FEDFFB14FD7BFEAE" blockId="6.[264,1323,1238,1905]" lastBlockId="7.[264,1322,284,350]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B9226C3C9A794346FEDFFB14FE62FB00" bold="true" box="[264,452,1238,1264]" pageId="6" pageNumber="7">
<typeStatus id="54ED0E8C9A794346FEDFFB14FEDCFB00" box="[264,378,1238,1264]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
Male
</emphasis>
. Body dorsally convex; length
<quantity id="4CAE1DCB9A794346FC92FB15FC03FB01" box="[837,933,1239,1265]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="7" unit="mm" value="6.6">6.6 mm</quantity>
. Head and pronotum yellowish brown, elytra shining violet; antenna with scape yellowish brown, washed with black, segments 3­11 black. Underside yellowish brown with femora yellowish brown, tibia brown to black, darkened apically, tarsi black. Head with apical margin of clypeus emarginate, antennal calli moderately swollen. Labrum emarginate, with 6 dorsal setae. Prothorax distinctly wider than long, L/W = 0.6, pronotum with lateral margin (
<figureCitation id="136DACAB9A794346FBEBFA5DFB25FA49" box="[1084,1155,1439,1465]" captionStart="FIGURES 2 ­ 7" captionStartId="3.[264,382,1698,1722]" captionTargetBox="[303,1285,1089,1667]" captionTargetId="figure@3.[301,1287,1089,1672]" captionTargetPageId="3" captionText="FIGURES 2 ­ 7. External characters of Thysanomeros. 2 3. Hind femur of male. 2. T. jacobyi. 3. T. ulateae. 4 5. Lateral margin of pronotum. 4. T. jacobyi, 5. T. ulateae. 6. Fore tarsi of T. jacobyi: left, male; right, female. 7. Pygidium of T. jacobyi." httpUri="https://zenodo.org/record/156244/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
) broad, undulate, greatest width of pronotum just behind middle; disc with scattered deep punctures, separated by distances greater than their diameters. Prosternum longitudinally weakly inclined behind coxae, posterior margin of intercoxal process truncate, width of intercoxal process 0.75 x diameter of procoxa. Mesosternum narrower than prosternum, flat between coxae, surface finely punctate, with numerous erect yellow setae. Metasternum with a median area of closely spaced sort recurved setae. Femora with a few setae on ventral margin in basal fourth, metafemur (
<figureCitation id="136DACAB9A794346FD66F975FD5EF921" box="[689,760,1719,1745]" captionStart="FIGURES 2 ­ 7" captionStartId="3.[264,382,1698,1722]" captionTargetBox="[303,1285,1089,1667]" captionTargetId="figure@3.[301,1287,1089,1672]" captionTargetPageId="3" captionText="FIGURES 2 ­ 7. External characters of Thysanomeros. 2 3. Hind femur of male. 2. T. jacobyi. 3. T. ulateae. 4 5. Lateral margin of pronotum. 4. T. jacobyi, 5. T. ulateae. 6. Fore tarsi of T. jacobyi: left, male; right, female. 7. Pygidium of T. jacobyi." httpUri="https://zenodo.org/record/156244/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
) with long ventral setae in basal third. Scutellum V­shaped. Abdomen with erect yellow setae on apical half of each segment. Male sternum VII broadly emarginate, pygidial surface with long, erect setae on apical half. Median lobe of aedeagus in lateral view strongly curved (
<figureCitation id="136DACAB9A794346FC7EF8EDFBA6F8B9" box="[937,1024,1839,1865]" captionStart="FIGURES 8 ­ 12" captionStartId="5.[264,382,1216,1240]" captionTargetBox="[326,1260,409,1187]" captionTargetId="figure@5.[325,1263,409,1189]" captionTargetPageId="5" captionText="FIGURES 8 ­ 12. Male genitalia of Thysanomeros. 8, 10. Median lobe. 8 T. jacobyi, showing endophalus and apical sclerite. 10. T. ulateae. 9, 12. Apex of median lobe. 9. T. jacobyi. 12. T. ulateae. 11. Apical sclerite, T. ulateae." httpUri="https://zenodo.org/record/156245/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
); basal hood long, lightly sclerotized, with apodemes indistinct at lateral margins of hood. Apex of median lobe mucronate in apical view (
<figureCitation id="136DACAB9A784347FD92FEDEFD38FEC6" box="[581,670,284,310]" captionStart="FIGURES 8 ­ 12" captionStartId="5.[264,382,1216,1240]" captionTargetBox="[326,1260,409,1187]" captionTargetId="figure@5.[325,1263,409,1189]" captionTargetPageId="5" captionText="FIGURES 8 ­ 12. Male genitalia of Thysanomeros. 8, 10. Median lobe. 8 T. jacobyi, showing endophalus and apical sclerite. 10. T. ulateae. 9, 12. Apex of median lobe. 9. T. jacobyi. 12. T. ulateae. 11. Apical sclerite, T. ulateae." httpUri="https://zenodo.org/record/156245/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
). Endophallus with apical sclerite straight, with apical and basal curved flaps present (
<figureCitation id="136DACAB9A784347FDAEFE86FD69FEAE" box="[633,719,324,350]" captionStart="FIGURES 8 ­ 12" captionStartId="5.[264,382,1216,1240]" captionTargetBox="[326,1260,409,1187]" captionTargetId="figure@5.[325,1263,409,1189]" captionTargetPageId="5" captionText="FIGURES 8 ­ 12. Male genitalia of Thysanomeros. 8, 10. Median lobe. 8 T. jacobyi, showing endophalus and apical sclerite. 10. T. ulateae. 9, 12. Apex of median lobe. 9. T. jacobyi. 12. T. ulateae. 11. Apical sclerite, T. ulateae." httpUri="https://zenodo.org/record/156245/files/figure.png" pageId="7" pageNumber="8">Fig. 11</figureCitation>
).
</paragraph>
<caption id="DF29E0A69A784347FEDFFB01FA8DFAF2" httpUri="https://zenodo.org/record/156246/files/figure.png" pageId="7" pageNumber="8" targetBox="[317,1269,409,1191]" targetPageId="7">
<paragraph id="8BE9B02E9A784347FEDFFB01FA8DFAF2" blockId="7.[264,1323,1219,1282]" pageId="7" pageNumber="8">
<emphasis id="B9226C3C9A784347FEDFFB01FE77FB2B" bold="true" box="[264,465,1219,1243]" pageId="7" pageNumber="8">FIGURES 13­ 17.</emphasis>
Female characters of
<taxonomicName id="4C56CBAD9A784347FD6AFB06FCF1FB2B" box="[701,855,1220,1243]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A784347FD6AFB06FCF1FB2B" box="[701,855,1220,1243]" italics="true" pageId="7" pageNumber="8">Thysanomeros</emphasis>
</taxonomicName>
. 13, 16­17. ovipositors. 13.
<taxonomicName id="4C56CBAD9A784347FB52FB06FB4BFB2B" box="[1157,1261,1220,1243]" class="Insecta" family="Chrysomelidae" genus="Prionodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="jacobyi">
<emphasis id="B9226C3C9A784347FB52FB06FB4BFB2B" box="[1157,1261,1220,1243]" italics="true" pageId="7" pageNumber="8">T. jacobyi</emphasis>
</taxonomicName>
, ventral. 16.
<taxonomicName id="4C56CBAD9A784347FEB6FB29FE61FAF2" box="[353,455,1259,1282]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ulateae">
<emphasis id="B9226C3C9A784347FEB6FB29FE61FAF2" box="[353,455,1259,1282]" italics="true" pageId="7" pageNumber="8">T. ulateae</emphasis>
</taxonomicName>
, ventral. 17.
<taxonomicName id="4C56CBAD9A784347FD82FB29FD1DFAF2" box="[597,699,1259,1282]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ulateae">
<emphasis id="B9226C3C9A784347FD82FB29FD1DFAF2" box="[597,699,1259,1282]" italics="true" pageId="7" pageNumber="8">T. ulateae</emphasis>
</taxonomicName>
, dorsal. 14­15. spermatheca. 14.
<taxonomicName id="4C56CBAD9A784347FBC9FB29FB22FAF2" box="[1054,1156,1259,1282]" class="Insecta" family="Chrysomelidae" genus="Prionodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="jacobyi">
<emphasis id="B9226C3C9A784347FBC9FB29FB22FAF2" box="[1054,1156,1259,1282]" italics="true" pageId="7" pageNumber="8">T. jacobyi</emphasis>
</taxonomicName>
. 15.
<taxonomicName id="4C56CBAD9A784347FB6CFB29FA85FAF2" box="[1211,1315,1259,1282]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ulateae">
<emphasis id="B9226C3C9A784347FB6CFB29FA85FAF2" box="[1211,1315,1259,1282]" italics="true" pageId="7" pageNumber="8">T. ulateae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE9B02E9A784347FEEFFAFBFBCFF8AB" blockId="7.[264,1323,1337,1883]" pageId="7" pageNumber="8">
<emphasis id="B9226C3C9A784347FEEFFAFBFDA3FAA3" bold="true" box="[312,517,1337,1363]" pageId="7" pageNumber="8">Allotype Female</emphasis>
. Body oval; length
<quantity id="4CAE1DCB9A784347FD2BFAFBFCFDFAA3" box="[764,859,1337,1363]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="7" pageNumber="8" unit="mm" value="7.8">7.8 mm</quantity>
; head, and pronotum as in male, elytra dark blue or blue­green, underside and antennae as in male; legs with femora and tarsi as in male, tibiae ranging from entirely yellowish brown to entirely black. Head with labrum, frons, clypeus, eyes and antennae similar to male; mouthparts similar to male. Prothorax distinctly wider than long, L/W = 0.67; pronotum with one to three teeth on lateral margin; punctation on disc as in male. Prosternum similar to male, but with width of intercoxal process 0.68.x diameter of procoxa. Mesosternum similar to male. Metasternum similar to male but with fewer setae in median area. Legs similar to male but fewer setae on ventral margins of femora. Scutellum U­shaped. abdominal sterna covered with long setae, 2­3 longer setae on each side of midline. Sternum VII with apical margin weakly emarginate, and with a transverse row of long setae. Abdominal segments VIII­XI forming short ovipositor (
<figureCitation id="136DACAB9A784347FEBDF933FE48F8FB" box="[362,494,1777,1803]" captionStart="FIGURES 13 ­ 17" captionStartId="7.[264,382,1219,1243]" captionTargetBox="[317,1269,409,1191]" captionTargetId="figure@7.[316,1272,409,1193]" captionTargetPageId="7" captionText="FIGURES 13 ­ 17. Female characters of Thysanomeros. 13, 16 ­ 17. ovipositors. 13. T. jacobyi, ventral. 16. T. ulateae, ventral. 17. T. ulateae, dorsal. 14 ­ 15. spermatheca. 14. T. jacobyi. 15. T. ulateae." httpUri="https://zenodo.org/record/156246/files/figure.png" pageId="7" pageNumber="8">Fig. 16, 17</figureCitation>
). Baculum distinct, apical, shorter than gonocoxae. Gonocoxae short, robust, with long setae in apical half, strongly incurved medially. Spermatheca as in
<figureCitation id="136DACAB9A784347FB2BF8DBFE81F8AB" captionStart="FIGURES 13 ­ 17" captionStartId="7.[264,382,1219,1243]" captionTargetBox="[317,1269,409,1191]" captionTargetId="figure@7.[316,1272,409,1193]" captionTargetPageId="7" captionText="FIGURES 13 ­ 17. Female characters of Thysanomeros. 13, 16 ­ 17. ovipositors. 13. T. jacobyi, ventral. 16. T. ulateae, ventral. 17. T. ulateae, dorsal. 14 ­ 15. spermatheca. 14. T. jacobyi. 15. T. ulateae." httpUri="https://zenodo.org/record/156246/files/figure.png" pageId="7" pageNumber="8">Fig. 15</figureCitation>
, receptacle tapered; spermathecal duct thin, sclerotized, loosely coiled.
</paragraph>
</subSubSection>
<subSubSection id="C34CE3A59A774348FEEFFED9FCD4FB56" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BE9B02E9A774348FEEFFED9FB1EFEAE" blockId="8.[264,1323,283,1830]" pageId="8" pageNumber="9">
<emphasis id="B9226C3C9A774348FEEFFED9FE6EFEC5" bold="true" box="[312,456,283,309]" pageId="8" pageNumber="9">Etymology:</emphasis>
This species is named for Elena Ulate Arce, in recognition of her talented and dedicated service as technician in the
<taxonomicName id="4C56CBAD9A774348FCEDFE86FC1CFEAE" box="[826,954,324,350]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
collections of INBio.
</paragraph>
<paragraph id="8BE9B02E9A774348FEEFFEA9FCD4FB56" blockId="8.[264,1323,283,1830]" pageId="8" pageNumber="9">
<emphasis id="B9226C3C9A774348FEEFFEA9FE63FE75" bold="true" box="[312,453,363,389]" pageId="8" pageNumber="9">Specimens:</emphasis>
(3 ɗ and 8 Ψ, all deposited in INBio) Male
<typeStatus id="54ED0E8C9A774348FC09FEAEFBDFFE76" box="[990,1145,364,390]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
labeled Sector San Ramón, Prov. Alaju,
<collectingCountry id="F341F0BE9A774348FD82FE56FCABFE5E" box="[597,781,404,430]" name="Costa Rica" pageId="8" pageNumber="9">COSTA RICA</collectingCountry>
.
<quantity id="4CAE1DCB9A774348FCF7FE56FCC0FE5E" box="[800,870,404,430]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="9" unit="m" value="620.0">620m</quantity>
,
<date id="FFE896EE9A774348FCADFE56FBEAFE5E" box="[890,1100,404,430]" pageId="8" pageNumber="9" value="1994-03-13" valueMax="1994-03-28" valueMin="1994-03-13">13­28 Mar 1994</date>
, D. Garcia, L N 318100_381900 # 2766” (INBIOCRI001738811). Female ALLOTYPE labeled “Valle la Estrella, R.B. Hitoy Cerere, Prov. Limón,
<collectingCountry id="F341F0BE9A774348FD2DFE26FC0DFE0E" box="[762,939,484,510]" name="Costa Rica" pageId="8" pageNumber="9">COSTA RICA</collectingCountry>
.
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<quantity id="4CAE1DCB9A774348FC61FE26FBA2FE0E" box="[950,1028,484,510]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="m" value="100.0">100 m</quantity>
.
</specimenCount>
<date id="FFE896EE9A774348FBC6FE26FBDEFE0E" box="[1041,1144,484,510]" pageId="8" pageNumber="9" value="1994-07">Jul 1994</date>
, J. F. Quesada, L S 398100_572800 # 3174” (INBIOCRI002004360).
<typeStatus id="54ED0E8C9A774348FC4DFDCEFB9AFDD6" box="[922,1084,524,550]" pageId="8" pageNumber="9" type="paratype">PARATYPES</typeStatus>
(2 ɗ, 7 Ψ):
<collectingCountry id="F341F0BE9A774348FB1AFDCEFEF5FDBE" name="Costa Rica" pageId="8" pageNumber="9">COSTA RICA</collectingCountry>
: Alajuela Province: 1 ɗ (INBIOCRI002250566), 1 Ψ (INBIOCRI002250565), Sect. San Ramón de Dos Rios,
<specimenCount id="9D507BA79A774348FD91FD9EFD37FD86" box="[582,657,604,630]" pageId="8" pageNumber="9" type="generic">
<quantity id="4CAE1DCB9A774348FD91FD9EFD2AFD86" box="[582,652,604,630]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="9" unit="m" value="620.0">620m</quantity>
.
</specimenCount>
<date id="FFE896EE9A774348FD4CFD9EFCC7FD86" box="[667,865,604,630]" pageId="8" pageNumber="9" value="1995-04-03" valueMax="1995-04-24" valueMin="1995-04-03">3­24 ABR 1995</date>
. M. Chinchilla, L_N_318100_381900 #5328; 1 Ψ (INBIOCRI002146264), same locality,
<date id="FFE896EE9A774348FC5EFD46FB14FD6E" box="[905,1202,644,670]" pageId="8" pageNumber="9" value="1995-04-27" valueMax="1995-05-11" valueMin="1995-04-27">27 ABR­11 MAY 1995</date>
. C. Cano #5276; 1 Ψ (INBIOCRI001874584), Est. San Ramón, P. N. Guanacaste,
<specimenCount id="9D507BA79A774348FB8EFD6EFB0CFD36" box="[1113,1194,684,710]" pageId="8" pageNumber="9" type="generic">
<quantity id="4CAE1DCB9A774348FB8EFD6EFB01FD36" box="[1113,1191,684,710]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="9" unit="m" value="620.0">620 m</quantity>
.
</specimenCount>
<date id="FFE896EE9A774348FB66FD6EFEE3FD1E" pageId="8" pageNumber="9" value="1994-04-03" valueMax="1994-04-19" valueMin="1994-04-03">03­19 Abr 1994</date>
, Fam. Hurtado Garcia, L N 318100_381900 # 3004; 1 Ψ (INBIOCRI001776894), Est. San Ramón Oeste,
<specimenCount id="9D507BA79A774348FDC8FD3EFDD7FCE6" box="[543,625,764,790]" pageId="8" pageNumber="9" type="generic">
<quantity id="4CAE1DCB9A774348FDC8FD3EFDCBFCE6" box="[543,621,764,790]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="9" unit="m" value="620.0">620 m</quantity>
.
</specimenCount>
<date id="FFE896EE9A774348FDAFFD3EFC91FCE6" box="[632,823,764,790]" pageId="8" pageNumber="9" value="1994-04-03" valueMax="1994-04-19" valueMin="1994-04-03">03­19 Abr 1994</date>
, F. Quesada, L N 318100_381900 # 2817. Guanacaste Province: 1 Ψ (INBIOCRI000698667), Est. Pitilla,
<quantity id="4CAE1DCB9A774348FC26FCE6FB91FCCE" box="[1009,1079,804,830]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="9" unit="m" value="700.0">700m</quantity>
,
<quantity id="4CAE1DCB9A774348FB95FCE6FBD8FCCE" box="[1090,1150,804,830]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="9" unit="km" value="9.0">9 km</quantity>
S Sta. Cecilia, P. N. Guanacaste, C. Moraga,
<date id="FFE896EE9A774348FD45FC8EFC18FC96" box="[658,958,844,870]" pageId="8" pageNumber="9" value="1992-03-31" valueMax="1992-04-15" valueMin="1992-03-31">31 mar. ­ 15 abr. 1992</date>
, L­ N 330200_380200; 1 Ψ (INBIOCRI002336380), same locality
<date id="FFE896EE9A774348FD01FCB6FCEEFC7E" box="[726,840,884,910]" pageId="8" pageNumber="9" value="1995-04">abr. 1995</date>
. P. Rios, L_N_329950_380450 #4814: 1 Ψ (INBIOCRI000422572), Río San Lorenzo,
<quantity id="4CAE1DCB9A774348FCFFFC5EFC23FC46" box="[808,901,924,950]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="9" unit="m" value="1050.0">1050 m</quantity>
., Tierras Morenas, Z. P. Tenorio, M. Segura,
<date id="FFE896EE9A774348FEA3FC06FD1EFC2E" box="[372,696,964,990]" pageId="8" pageNumber="9" value="1992-03-23" valueMax="1992-04-21" valueMin="1992-03-23">23 mar. a 21 abr. 1992</date>
, L_N_287800_427600. Heredia Province: 1 ɗ (INBIOCRI002729432), Est. Biol La Selva.
<quantity id="4CAE1DCB9A774348FCC9FC2EFC2DFBF6" box="[798,907,1004,1030]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="9" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
50­
<specimenCount id="9D507BA79A774348FC92FC2EFC2DFBF6" box="[837,907,1004,1030]" pageId="8" pageNumber="9" type="generic">100m</specimenCount>
</quantity>
.
<date id="FFE896EE9A774348FC4FFC2EFB87FBF6" box="[920,1057,1004,1030]" pageId="8" pageNumber="9" value="2000-05">MAY 2000</date>
. INBio­OET. Manual. L_N_268800_535300 #71434. Limón Province: 1 Ψ (INBIOCRI001954677), Río Sardinas, R.N.F.S. Barra del Colorado,
<specimenCount id="9D507BA79A774348FD40FBFEFD7FFBA6" box="[663,729,1084,1110]" pageId="8" pageNumber="9" type="generic">
<quantity id="4CAE1DCB9A774348FD40FBFEFD70FBA6" box="[663,726,1084,1110]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="m" value="10.0">10 m</quantity>
.
</specimenCount>
<date id="FFE896EE9A774348FD37FBFEFC3EFBA6" box="[736,920,1084,1110]" pageId="8" pageNumber="9" value="1993-06-18" valueMax="1993-06-27" valueMin="1993-06-18">18­27 Jun 1993</date>
, F. V. Araya, L N 291500_564700 # 2182; 1 Ψ (INBIOCRI000709586), Est. Hitoy Cerere,
<quantity id="4CAE1DCB9A774348FC4BFBA6FC44FB8E" box="[924,994,1124,1150]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="m" value="100.0">100m</quantity>
, R. Cerere, Res. Biol. Hitoy Cerere,
<date id="FFE896EE9A774348FEB2FB4EFE6DFB56" box="[357,459,1164,1190]" pageId="8" pageNumber="9" value="1992-07">Jul 1992</date>
, G. Carballo, L­ N 184200_643300.
</paragraph>
</subSubSection>
<subSubSection id="C34CE3A59A774349FEEFFB76FAACF97F" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BE9B02E9A774348FEEFFB76FDFFF8D6" blockId="8.[264,1323,283,1830]" pageId="8" pageNumber="9">
<emphasis id="B9226C3C9A774348FEEFFB76FE65FB3E" bold="true" box="[312,451,1204,1230]" pageId="8" pageNumber="9">Discussion.</emphasis>
The series Endocephalites was defined by
<bibRefCitation id="EFC7CDDF9A774348FC16FB76FBD0FB3E" author="Chapuis" box="[961,1142,1204,1230]" pageId="8" pageNumber="11" refString="Chapuis, F. (1874). In Lacordaire, Histoire naturelle des insectes. Genera de coleopteres. 10, 455 pp." type="book" year="1874">Chapuis (1874)</bibRefCitation>
as
<taxonomicName id="4C56CBAD9A774348FB4AFB76FA8DFB3E" box="[1181,1323,1204,1230]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subFamily" subFamily="Eumolpinae">Eumolpinae</taxonomicName>
having convex anterior edges on the lateral arms of the prosternum (the proepisternum in older literature). In Bechynés (1953) catalog this group was treated as a subtribe, but is here retained as an informal group pending broader study of the Eumolpini. Endocephalites contains the Neotropical genera
<taxonomicName id="4C56CBAD9A774348FD68FA96FC11FA9E" authority="Laporte" authorityName="Laporte" box="[703,951,1364,1390]" class="Insecta" family="Chrysomelidae" genus="Colaspoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FD68FA96FCF6FA9D" box="[703,848,1364,1389]" italics="true" pageId="8" pageNumber="9">Colaspoides</emphasis>
Laporte
</taxonomicName>
,
<taxonomicName id="4C56CBAD9A774348FC14FA96FB72FA9E" authority="Chapuis" authorityName="Chapuis" box="[963,1236,1364,1390]" class="Insecta" family="Chrysomelidae" genus="Endocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FC14FA96FBCFFA9D" box="[963,1129,1364,1389]" italics="true" pageId="8" pageNumber="9">Endocephalus</emphasis>
Chapuis
</taxonomicName>
,
<taxonomicName id="4C56CBAD9A774348FB36FA96FE35FA66" authority="Lefevre" authorityName="Lefevre" class="Insecta" family="Chrysomelidae" genus="Erotenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FB36FA96FE8BFA65" italics="true" pageId="8" pageNumber="9">Erotenia</emphasis>
Lefèvre
</taxonomicName>
,
<taxonomicName id="4C56CBAD9A774348FE48FABEFDD1FA66" authority="Jacoby" authorityName="Jacoby" box="[415,631,1404,1430]" class="Insecta" family="Chrysomelidae" genus="Melinodea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FE48FABEFDBAFA65" box="[415,540,1404,1429]" italics="true" pageId="8" pageNumber="9">Melinodea</emphasis>
Jacoby
</taxonomicName>
,
<taxonomicName id="4C56CBAD9A774348FD55FABEFCFEFA66" authority="Lefevre" authorityName="Lefevre" box="[642,856,1404,1430]" class="Insecta" family="Chrysomelidae" genus="Phanaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FD55FABEFD54FA65" box="[642,754,1404,1429]" italics="true" pageId="8" pageNumber="9">Phanaeta</emphasis>
Lefévre
</taxonomicName>
,
<taxonomicName id="4C56CBAD9A774348FCB3FABEFBE4FA66" authority="Bechyne" authorityName="Bechyne" box="[868,1090,1404,1430]" class="Insecta" family="Pteromalidae" genus="Susteraia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FCB3FABEFC75FA65" box="[868,979,1404,1429]" italics="true" pageId="8" pageNumber="9">Susteraia</emphasis>
Bechyné
</taxonomicName>
and
<taxonomicName id="4C56CBAD9A774348FBACFABEFA8CFA65" authority="Lefevre." authorityName="Lefevre." box="[1147,1322,1404,1430]" class="Insecta" family="Chrysomelidae" genus="Thyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FBACFABEFB67FA65" box="[1147,1217,1404,1429]" italics="true" pageId="8" pageNumber="9">Thyra</emphasis>
Lefèvre.
</taxonomicName>
<taxonomicName id="4C56CBAD9A774348FEDFFA66FEDEFA4D" box="[264,376,1444,1469]" class="Insecta" family="Chrysomelidae" genus="Phanaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FEDFFA66FEDEFA4D" box="[264,376,1444,1469]" italics="true" pageId="8" pageNumber="9">Phanaeta</emphasis>
</taxonomicName>
was later removed to the subtribe
<taxonomicName id="4C56CBAD9A774348FCC5FA66FBF7FA4E" authority="Bechyne 1958" authorityName="Bechyne" authorityYear="1958" box="[786,1105,1444,1470]" class="Insecta" family="Chrysomelidae" genus="Colaspina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
Colaspina (
<bibRefCitation id="EFC7CDDF9A774348FC4FFA66FBE1FA4E" author="Bechyne" box="[920,1095,1444,1470]" pageId="8" pageNumber="11" refString="Bechyne, J. (1958) Notizen zu den neotropischen Chrysomeloidea (Col. Phytophaga). Entomologische Arbeiten. aus dem Museum G. Frey, 9, 478 - 706" type="journal article" year="1958">Bechyné 1958</bibRefCitation>
)
</taxonomicName>
. Although
<taxonomicName id="4C56CBAD9A774348FB0CFA66FECDFA15" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FB0CFA66FECDFA15" italics="true" pageId="8" pageNumber="9">Thysanomeros</emphasis>
</taxonomicName>
does not have convex anterior margins on the prosternal arms, it shares with
<taxonomicName id="4C56CBAD9A774348FEDFFA36FE3FF9FD" box="[264,409,1524,1549]" class="Insecta" family="Chrysomelidae" genus="Colaspoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FEDFFA36FE3FF9FD" box="[264,409,1524,1549]" italics="true" pageId="8" pageNumber="9">Colaspoides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C56CBAD9A774348FE04FA36FDDFF9FD" box="[467,633,1524,1549]" class="Insecta" family="Chrysomelidae" genus="Endocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FE04FA36FDDFF9FD" box="[467,633,1524,1549]" italics="true" pageId="8" pageNumber="9">Endocephalus</emphasis>
</taxonomicName>
(the two largest New World genera of the Endocephalites) several other significant characters which together serve to define this subtribe: all three genera have the anterior margin of the prosternum broadly concave for reception of the head, completely lack an elytral locking groove on the pygidium and have a constriction at the junction of the attachment of the basal hood to the median lobe of the aedeagus.
<bibRefCitation id="EFC7CDDF9A774348FB33F956FE26F926" author="Flowers" pageId="8" pageNumber="11" refString="Flowers, R. W. (1996) La Subfamilia Eumolpinae (Coleoptera: Chrysomelidae) en America Central. Publicacion especial No. 2 de la Revista de Biologia Tropical, 59 pp." type="book" year="1996">Flowers (1996)</bibRefCitation>
noted that some species currently placed in
<taxonomicName id="4C56CBAD9A774348FC5BF97EFBBBF925" box="[908,1053,1724,1749]" class="Insecta" family="Chrysomelidae" genus="Colaspoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FC5BF97EFBBBF925" box="[908,1053,1724,1749]" italics="true" pageId="8" pageNumber="9">Colaspoides</emphasis>
</taxonomicName>
have grooved pygidia, as do some
<taxonomicName id="4C56CBAD9A774348FE7FF926FDE8F90D" box="[424,590,1764,1789]" class="Insecta" family="Chrysomelidae" genus="Endocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A774348FE7FF926FDE8F90D" box="[424,590,1764,1789]" italics="true" pageId="8" pageNumber="9">Endocephalus</emphasis>
</taxonomicName>
, but the
<typeStatus id="54ED0E8C9A774348FD1CF926FD58F90E" box="[715,766,1764,1790]" pageId="8" pageNumber="9">type</typeStatus>
species of both genera fit the definition of Endocephalites given above.
</paragraph>
<caption id="DF29E0A69A764349FEDFFB00FAB0FB29" box="[264,1302,1218,1242]" httpUri="https://zenodo.org/record/156247/files/figure.png" pageId="9" pageNumber="10" targetBox="[264,1306,315,1209]" targetPageId="9">
<paragraph id="8BE9B02E9A764349FEDFFB00FAB0FB29" blockId="9.[264,1302,1218,1242]" box="[264,1302,1218,1242]" pageId="9" pageNumber="10">
<emphasis id="B9226C3C9A764349FEDFFB00FE3EFB2A" bold="true" box="[264,408,1218,1242]" pageId="9" pageNumber="10">FIGURE 18.</emphasis>
Distribution of
<taxonomicName id="4C56CBAD9A764349FD94FB00FD7BFB29" box="[579,733,1218,1241]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A764349FD94FB00FD7BFB29" box="[579,733,1218,1241]" italics="true" pageId="9" pageNumber="10">Thysanomeros</emphasis>
</taxonomicName>
.
<taxonomicName id="4C56CBAD9A764349FD3CFB00FCF7FB29" box="[747,849,1218,1241]" class="Insecta" family="Chrysomelidae" genus="Prionodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jacobyi">
<emphasis id="B9226C3C9A764349FD3CFB00FCF7FB29" box="[747,849,1218,1241]" italics="true" pageId="9" pageNumber="10">T. jacobyi</emphasis>
</taxonomicName>
, closed circles.
<taxonomicName id="4C56CBAD9A764349FC29FB00FBC1FB29" box="[1022,1127,1218,1241]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ulateae">
<emphasis id="B9226C3C9A764349FC29FB00FBC1FB29" box="[1022,1127,1218,1241]" italics="true" pageId="9" pageNumber="10">T. ulateae</emphasis>
</taxonomicName>
, closed squares.
</paragraph>
</caption>
<paragraph id="8BE9B02E9A764349FEEFFAF7FAACF97F" blockId="9.[264,1323,1333,1679]" pageId="9" pageNumber="10">
The ranges of the two species in
<collectingCountry id="F341F0BE9A764349FD6AFAF7FC98FABF" box="[701,830,1333,1359]" name="Costa Rica" pageId="9" pageNumber="10">Costa Rica</collectingCountry>
(
<figureCitation id="136DACAB9A764349FC99FAF7FC3AFABF" box="[846,924,1333,1359]" captionStart="FIGURE 18" captionStartId="9.[264,368,1218,1242]" captionTargetBox="[264,1306,315,1209]" captionTargetId="figure@9.[264,1322,315,1215]" captionTargetPageId="9" captionText="FIGURE 18. Distribution of Thysanomeros. T. jacobyi, closed circles. T. ulateae, closed squares." httpUri="https://zenodo.org/record/156247/files/figure.png" pageId="9" pageNumber="10">Fig 18</figureCitation>
) show a largely disjunct distribution with most specimens of
<taxonomicName id="4C56CBAD9A764349FDB4FA9FFD73FA86" box="[611,725,1373,1398]" class="Insecta" family="Chrysomelidae" genus="Prionodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jacobyi">
<emphasis id="B9226C3C9A764349FDB4FA9FFD73FA86" box="[611,725,1373,1398]" italics="true" pageId="9" pageNumber="10">T. jacobyi</emphasis>
</taxonomicName>
coming from the Osa Peninsula on the southwest Pacific side and with
<taxonomicName id="4C56CBAD9A764349FDD5FA47FDD3FA6E" box="[514,629,1413,1438]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ulateae">
<emphasis id="B9226C3C9A764349FDD5FA47FDD3FA6E" box="[514,629,1413,1438]" italics="true" pageId="9" pageNumber="10">T. ulateae</emphasis>
</taxonomicName>
largely confined to the Atlantic slope of
<collectingCountry id="F341F0BE9A764349FB86FA47FB72FA6F" box="[1105,1236,1413,1439]" name="Costa Rica" pageId="9" pageNumber="10">Costa Rica</collectingCountry>
. There is an area of overlap around the volcanos of the Guanacaste Conservation Area in northwestern
<collectingCountry id="F341F0BE9A764349FEBDFA17FE48FA1F" box="[362,494,1493,1519]" name="Costa Rica" pageId="9" pageNumber="10">Costa Rica</collectingCountry>
. A single specimen of
<taxonomicName id="4C56CBAD9A764349FD29FA17FCC9FA1E" box="[766,879,1493,1518]" class="Insecta" family="Chrysomelidae" genus="Prionodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jacobyi">
<emphasis id="B9226C3C9A764349FD29FA17FCC9FA1E" box="[766,879,1493,1518]" italics="true" pageId="9" pageNumber="10">T. jacobyi</emphasis>
</taxonomicName>
is known from the San Ramón site in western
<collectingCountry id="F341F0BE9A764349FEBDFA3FFE4BF9E7" box="[362,493,1533,1559]" name="Costa Rica" pageId="9" pageNumber="10">Costa Rica</collectingCountry>
, but on the Atlantic slope, and one
<taxonomicName id="4C56CBAD9A764349FC5DFA3FFC5DF9E6" box="[906,1019,1533,1558]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ulateae">
<emphasis id="B9226C3C9A764349FC5DFA3FFC5DF9E6" box="[906,1019,1533,1558]" italics="true" pageId="9" pageNumber="10">T. ulateae</emphasis>
</taxonomicName>
has been collected on the western side of Volcán Tenorio. Both species of
<taxonomicName id="4C56CBAD9A764349FC99F9E7FC5EF9CE" box="[846,1016,1573,1598]" class="Insecta" family="Chrysomelidae" genus="Thysanomeros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B9226C3C9A764349FC99F9E7FC5EF9CE" box="[846,1016,1573,1598]" italics="true" pageId="9" pageNumber="10">Thysanomeros</emphasis>
</taxonomicName>
are currently known only from
<collectingCountry id="F341F0BE9A764349FE9EF98FFE6AF997" box="[329,460,1613,1639]" name="Costa Rica" pageId="9" pageNumber="10">Costa Rica</collectingCountry>
, but given that the ranges of both span the length of the country, it would not be surprising if these species were eventually found in both
<collectingCountry id="F341F0BE9A764349FC24F9B7FBCAF97F" box="[1011,1132,1653,1679]" name="Nicaragua" pageId="9" pageNumber="10">Nicaragua</collectingCountry>
and
<collectingCountry id="F341F0BE9A764349FB72F9B7FAA0F97F" box="[1189,1286,1653,1679]" name="Panama" pageId="9" pageNumber="10">Panamá</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>