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<document id="6AE64CA5FCB3AFC175827835553F8E3D" ID-DOI="10.5281/zenodo.5300381" ID-ISSN="0374-1036" ID-Zenodo-Dep="5300381" ID-ZooBank="FFD37B73-2DB4-4538-8BBE-323AA68B13C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630096633526" checkinUser="marcus" docAuthor="Diaz, Florentina, Torreias, Sharlene R. S., Spinelli, Gustavo R., Ronderos, María M. &amp; Unlp" docDate="2014" docId="03F7B30FFF803F4F82742FE1FBB7E60C" docLanguage="en" docName="ActaEntMusNatPra.2014.54.2.715-728.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Dasyhelea pseudopollinosa Diaz &amp; Torreias &amp; Spinelli &amp; Ronderos &amp; Unlp 2014, sp. nov." docType="treatment" docVersion="3" lastPageNumber="722" masterDocId="FFCECB77FF813F4883442B14FF88E47F" masterDocTitle="A new species of Dasyhelea from Brazilian Amazonas and the description of the male of D. paulistana (Diptera: Ceratopogonidae)" masterLastPageNumber="728" masterPageNumber="715" pageNumber="716" updateTime="1699143756584" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8940B74E334F8946742647823E86069B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4C6A2C26E59870DFF7D930428872D3C5">
<mods:title id="3345DD56DF53B7A11AFEB2E2AA0E701E">A new species of Dasyhelea from Brazilian Amazonas and the description of the male of D. paulistana (Diptera: Ceratopogonidae)</mods:title>
</mods:titleInfo>
<mods:name id="7AEFBA11A0B1A25FDBB10D90A1BEE579" type="personal">
<mods:role id="E04523321B23EC2C5DB9138358300A8F">
<mods:roleTerm id="6222B16EA9965E2F95F693CC0A1ADF16">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="69B1B222CC2F2CE1F4797F8F2FD8C842">Diaz, Florentina</mods:namePart>
<mods:affiliation id="56E180F53D955FB6A207D81280A1F1D1">División Entomología, Museo de La Plata, UNLP, Paseo del Bosque s / n, B 1900 WA La Plata, Buenos Aires, Argentina; e-mail: mfdiaz @ fcnym. unlp. edu. ar, spinelli @ fcnym. unlp. edu. ar &amp; corresponding author</mods:affiliation>
</mods:name>
<mods:name id="6642AADFE135589010D85743EC53AC25" type="personal">
<mods:role id="84DE153EDB032883F3D1B7860C0E7C11">
<mods:roleTerm id="EDC63A14DAAABFCB2A937A0F9710B550">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C71E8A1452D2EF9D4140D02DEA1E4DB">Torreias, Sharlene R. S.</mods:namePart>
<mods:affiliation id="3628DFE3716C1FA684A63D1930EF048E">Av. Torquato Tapajós, João Bosco, Bloco 8 Flores, CEP. 69048 - 660, Manaus, Amazonas, Brazil; e-mail: rtorreias @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="4C1406B79D1363518F96F0A848246DC6" type="personal">
<mods:role id="BEADFBA198D3F7286FD6441C982EC4F8">
<mods:roleTerm id="1A9D34806C0323D09E4A425E62F2A46C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE5EECA95906708F8FB99DB8F7E0A088">Spinelli, Gustavo R.</mods:namePart>
<mods:affiliation id="771FAE30BCFDF94A2774002269285C69">División Entomología, Museo de La Plata, UNLP, Paseo del Bosque s / n, B 1900 WA La Plata, Buenos Aires, Argentina; e-mail: mfdiaz @ fcnym. unlp. edu. ar, spinelli @ fcnym. unlp. edu. ar</mods:affiliation>
</mods:name>
<mods:name id="CAAE595C3EB85751A72FC4184E03C9B4" type="personal">
<mods:role id="938B2B97A1F1B4E507B22950DED75AD4">
<mods:roleTerm id="EB4D6D3E5D29D3E5B1456309D51EE312">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF001C39C1D0C825BE082FBFFE561C44">Ronderos, María M.</mods:namePart>
<mods:affiliation id="90341F3A1F81E6494B9F046E5E7A4DF7">División Entomología, Museo de La Plata, UNLP, Paseo del Bosque s / n, B 1900 WA La Plata, Buenos Aires, Argentina; e-mail: mfdiaz @ fcnym. unlp. edu. ar, spinelli @ fcnym. unlp. edu. ar</mods:affiliation>
</mods:name>
<mods:name id="34CF4C01CC74F033E4D837CC13A35E53" type="personal">
<mods:role id="9EE6D10E6683D854B7420415E9F1A920">
<mods:roleTerm id="1531619CF1538D741BCBA846B3D555A8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0BDCD8D2AF8820C7F4E26BF9084A885E">Unlp</mods:namePart>
</mods:name>
<mods:typeOfResource id="BCB9E777B60F405887B8CB37FDADB986">text</mods:typeOfResource>
<mods:relatedItem id="B531C5AB41E17026C19999E0D531AE47" type="host">
<mods:titleInfo id="2F6CD061C9D8A4784D75809D502E0DDE">
<mods:title id="737B70A09F75E04E7918CF6395B950E6">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="8C9DA8D0B29F42EBACA444012450D4E1">
<mods:date id="55D62A8EB5CB7FD8224C0D51487D3276">2014</mods:date>
<mods:detail id="4071E5CC711D14EF9F6D05ECF6C7911C" type="pubDate">
<mods:number id="3E2C98478C56B23B2B72DB85B44E971B">2014-12-15</mods:number>
</mods:detail>
<mods:detail id="E81E027F78B52FC824319A4E6EDEC118" type="volume">
<mods:number id="52A50DD46D1E1DAE9AA4DE0EC27560A3">54</mods:number>
</mods:detail>
<mods:detail id="FDD85D64982072E99630BCBC2F623D75" type="issue">
<mods:number id="F95679D3C4BA1FD8B7D958CD318FD94A">2</mods:number>
</mods:detail>
<mods:extent id="A177061DAFF45165BA1FEB97FEA2A297" unit="page">
<mods:start id="531F320636A2023E75560264EDF4F680">715</mods:start>
<mods:end id="12D2ECEF53FC0F9A63BA9B1A328DAECA">728</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6ABB31FBA2CD67711B99C79D9CBA1CB5">journal article</mods:classification>
<mods:identifier id="B2C524E1B4660E28D94381488DD3BDF2" type="DOI">10.5281/zenodo.5300381</mods:identifier>
<mods:identifier id="8D059CDB19AAAA0AFDB06025264AAD2A" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="9FFA297B6BC4FB2959F6FBBDFB28CBBE" type="Zenodo-Dep">5300381</mods:identifier>
<mods:identifier id="7F7A9363FB6D26623B0CD5B04D82E6C3" type="ZooBank">FFD37B73-2DB4-4538-8BBE-323AA68B13C1</mods:identifier>
</mods:mods>
<treatment id="03F7B30FFF803F4F82742FE1FBB7E60C" LSID="urn:lsid:plazi:treatment:03F7B30FFF803F4F82742FE1FBB7E60C" httpUri="http://treatment.plazi.org/id/03F7B30FFF803F4F82742FE1FBB7E60C" lastPageId="7" lastPageNumber="722" pageId="1" pageNumber="716">
<subSubSection id="C3445192FF803F4982742FE1FC0AE172" box="[304,898,1269,1293]" pageId="1" pageNumber="716" type="nomenclature">
<paragraph id="8BE10219FF803F4982742FE1FC0AE172" blockId="1.[304,898,1269,1293]" box="[304,898,1269,1293]" pageId="1" pageNumber="716">
<heading id="D0A9B575FF803F4982742FE1FC0AE172" bold="true" box="[304,898,1269,1293]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="716" reason="3">
<emphasis id="B92ADE0BFF803F4982742FE1FC0AE172" bold="true" box="[304,898,1269,1293]" pageId="1" pageNumber="716">
<taxonomicName id="4C5E799AFF803F4982742FE1FCAEE172" ID-CoL="9HY7N" authority="Diaz &amp; Ronderos" authorityName="Diaz &amp; Torreias &amp; Spinelli &amp; Ronderos &amp; Unlp" authorityYear="2014" box="[304,806,1269,1293]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="716" phylum="Arthropoda" rank="species" species="pseudopollinosa" status="sp. nov.">
<emphasis id="B92ADE0BFF803F4982742FE1FDD3E172" bold="true" box="[304,603,1269,1293]" italics="true" pageId="1" pageNumber="716">Dasyhelea pseudopollinosa</emphasis>
Díaz &amp; Ronderos
</taxonomicName>
<taxonomicNameLabel id="A2196370FF803F4980682FE2FC0AE172" box="[812,898,1270,1293]" pageId="1" pageNumber="716" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3445192FF803F4981532E00FD12E158" box="[535,666,1300,1319]" pageId="1" pageNumber="716" type="description">
<paragraph id="8BE10219FF803F4981532E00FD12E158" blockId="1.[113,1090,1300,1468]" box="[535,666,1300,1319]" pageId="1" pageNumber="716">
(
<figureCitation id="13651E9CFF803F4981592E00FDFDE158" box="[541,629,1300,1319]" captionStart-0="Figs 1–11" captionStart-1="Figs 12–20" captionStart-2="Figs 21–32" captionStartId-0="4.[151,188,1483,1502]" captionStartId-1="5.[113,149,1441,1460]" captionStartId-2="6.[151,188,1413,1432]" captionTargetBox-0="[182,1095,219,1464]" captionTargetBox-1="[162,1039,219,1413]" captionTargetBox-2="[231,1051,223,1379]" captionTargetId-0="figure-10@4.[182,1096,219,1464]" captionTargetId-1="figure-15@5.[162,1040,219,1414]" captionTargetId-2="figure-10@6.[226,1052,219,1380]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." captionText-1="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." captionText-2="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi-0="http://doi.org/10.5281/zenodo.5300383" figureDoi-1="http://doi.org/10.5281/zenodo.5300385" figureDoi-2="http://doi.org/10.5281/zenodo.5300387" httpUri-0="https://zenodo.org/record/5300383/files/figure.png" httpUri-1="https://zenodo.org/record/5300385/files/figure.png" httpUri-2="https://zenodo.org/record/5300387/files/figure.png" pageId="1" pageNumber="716">Figs 1–32</figureCitation>
, 62)
</paragraph>
</subSubSection>
<subSubSection id="C3445192FF803F4983352E2AFBCAE1C4" pageId="1" pageNumber="716" type="materials_examined">
<paragraph id="8BE10219FF803F4983352E2AFBCAE1C4" blockId="1.[113,1090,1300,1468]" pageId="1" pageNumber="716">
<emphasis id="B92ADE0BFF803F4983352E2AFF7AE12E" bold="true" box="[113,242,1342,1361]" pageId="1" pageNumber="716">Type material</emphasis>
.
<materialsCitation id="3B360844FF803F4983BA2E2AFD29E1F9" collectingDate="2011-05-17" collectionCode="INPA" country="Brazil" county="Iranduba" location="Lago Grande" municipality="Ilha da Marchantaria" pageId="1" pageNumber="716" specimenCount="2" stateProvince="Amazonas" typeStatus="holotype">
<typeStatus id="54E5BCBBFF803F4983BA2E2AFED9E130" box="[254,337,1342,1361]" pageId="1" pageNumber="716" type="holotype">HOLOTYPE</typeStatus>
: J (with ‘ex pupa’) on microscope slide, labelled: “
<typeStatus id="54E5BCBBFF803F49805F2E2AFC06E12E" box="[795,910,1342,1361]" pageId="1" pageNumber="716" type="holotype">HOLOTYPE</typeStatus>
”
<taxonomicName id="4C5E799AFF803F4980D82E29FEE0E114" authority="Diaz &amp; Ronderos." authorityName="Diaz &amp; Torreias &amp; Spinelli &amp; Ronderos &amp; Unlp" authorityYear="2014" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="716" phylum="Arthropoda" rank="species" species="pseudopollinosa">
<emphasis id="B92ADE0BFF803F4980D82E29FF48E114" italics="true" pageId="1" pageNumber="716">Dasyhelea pseudopollinosa</emphasis>
Díaz &amp; Ronderos.
</taxonomicName>
<emphasis id="B92ADE0BFF803F4982292E4CFDB8E114" bold="true" box="[365,560,1368,1387]" pageId="1" pageNumber="716">
<collectingCountry id="F3494289FF803F4982292E4CFE4AE114" box="[365,450,1368,1387]" name="Brazil" pageId="1" pageNumber="716">BRAZIL</collectingCountry>
:
<collectingRegion id="499ACCFBFF803F4982892E4CFDA1E115" box="[461,553,1368,1387]" country="Brazil" name="Amazonas" pageId="1" pageNumber="716">AMAZONAS</collectingRegion>
:
</emphasis>
<collectingCounty id="62807A95FF803F4981732E4CFD0DE114" box="[567,645,1368,1387]" pageId="1" pageNumber="716">Iranduba</collectingCounty>
,
<collectingMunicipality id="6B859863FF803F4981D42E4CFCCFE114" box="[656,839,1368,1387]" pageId="1" pageNumber="716">Ilha da Marchantaria</collectingMunicipality>
,
<location id="8E8154C2FF803F4980162E4CFC4EE114" LSID="urn:lsid:plazi:treatment:03F7B30FFF803F4F82742FE1FBB7E60C:8E8154C2FF803F4980162E4CFC4EE114" box="[850,966,1368,1387]" country="Brazil" county="Iranduba" municipality="Ilha da Marchantaria" name="Lago Grande" pageId="1" pageNumber="716" stateProvince="Amazonas">Lago Grande</location>
, 03°14މ43.9ފS 59°58މ54.2ފ W,
<date id="FFE024D9FF803F4983BD2E67FEC6E1F9" box="[249,334,1395,1414]" pageId="1" pageNumber="716" value="2011-05-17">
<collectingDate id="EFA4DD31FF803F4983BD2E67FEC6E1F9" box="[249,334,1395,1414]" pageId="1" pageNumber="716" value="2011-05-17">17.v.2011</collectingDate>
</date>
, Torreias-Ferreira-Keppler lgt., (
<collectionCode id="ED4F9ADCFF803F4981222E67FD13E1F9" box="[614,667,1395,1414]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="716">INPA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B360844FF803F4981EC2E67FF21E1DE" collectingDate="2011-05-17" collectionCode="INPA" country="Brazil" county="Iranduba" location="Lago Grande" municipality="Ilha da Marchantaria" pageId="1" pageNumber="716" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas" typeStatus="allotype">
<typeStatus id="54E5BCBBFF803F4981EC2E67FD71E1FB" box="[680,761,1395,1414]" pageId="1" pageNumber="716" type="allotype">ALLOTYPE</typeStatus>
:
<specimenCount id="9D58C990FF803F4980412E7BFC9BE1F9" box="[773,787,1391,1414]" pageId="1" pageNumber="716" type="female">♀</specimenCount>
(with ‘ex pupa’), same data as holotype
</materialsCitation>
.
<materialsCitation id="3B360844FF803F4983F52E9AFF34E1C4" collectionCode="INPA, MLPA" country="Brazil" pageId="1" pageNumber="716" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas" typeStatus="paratype">
<typeStatus id="54E5BCBBFF803F4983F52E9AFE80E1E0" box="[177,264,1422,1441]" pageId="1" pageNumber="716" type="paratype">PARATYPES</typeStatus>
: 5 JJ(with ‘ex pupa’) (
<quantity id="4CA6AFFCFF803F4982992E9AFE77E1DE" box="[477,511,1422,1441]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="1" pageNumber="716" unit="in" value="4.0">4 in</quantity>
<collectionCode id="ED4F9ADCFF803F4981472E9AFDBDE1DE" box="[515,565,1422,1441]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="716">INPA</collectionCode>
,
<quantity id="4CA6AFFCFF803F49817A2E9AFDD6E1DE" box="[574,606,1422,1441]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="716" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED4F9ADCFF803F4981272E9AFD16E1DE" box="[611,670,1422,1441]" pageId="1" pageNumber="716">MLPA</collectionCode>
) and
<specimenCount id="9D58C990FF803F4981882E9AFD62E1DE" box="[716,746,1418,1441]" pageId="1" pageNumber="716" type="female">1 ♀</specimenCount>
(with ‘ex pupa’), same date as
<typeStatus id="54E5BCBBFF803F4980B22E9AFBC9E1DE" box="[1014,1089,1422,1441]" pageId="1" pageNumber="716" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED4F9ADCFF803F4983332EBCFF3EE1C4" box="[119,182,1448,1467]" pageId="1" pageNumber="716">MLPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B360844FF803F4983872EBCFC9BE1C4" box="[195,787,1445,1468]" collectionCode="INPA, MLPA" country="Brazil" pageId="1" pageNumber="716" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">1 J (with ‘ex pupa’), same data except: Lago Camaleão, 03°13މ14.9ފS</materialsCitation>
;
<materialsCitation id="3B360844FF803F4980582EBCFBB6E1C4" box="[796,1086,1448,1467]" collectingDate="2011-05-20" collectionCode="MLPA" country="Brazil" pageId="1" pageNumber="716" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
59°56މ52.6ފW,
<date id="FFE024D9FF803F4980D82EBCFC67E1C4" box="[924,1007,1448,1467]" pageId="1" pageNumber="716" value="2011-05-20">
<collectingDate id="EFA4DD31FF803F4980D82EBCFC67E1C4" box="[924,1007,1448,1467]" pageId="1" pageNumber="716" value="2011-05-20">20.v.2011</collectingDate>
</date>
(
<collectionCode id="ED4F9ADCFF803F4980BE2EBCFBB1E1C4" box="[1018,1081,1448,1467]" pageId="1" pageNumber="716">MLPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3445192FF803F4A83352EC0FE08E52C" lastPageId="2" lastPageNumber="717" pageId="1" pageNumber="716" type="diagnosis">
<paragraph id="8BE10219FF803F4A83352EC0FE08E52C" blockId="1.[113,1089,1492,1547]" lastBlockId="2.[151,1128,220,1555]" lastPageId="2" lastPageNumber="717" pageId="1" pageNumber="716">
<emphasis id="B92ADE0BFF803F4983352EC0FF57E194" bold="true" box="[113,223,1492,1515]" pageId="1" pageNumber="716">Diagnosis</emphasis>
. Only Neotropical species of the
<emphasis id="B92ADE0BFF803F4981012EC0FD00E194" box="[581,648,1492,1515]" italics="true" pageId="1" pageNumber="716">grisea</emphasis>
species-group with the following combination of characters: male with posterolateral arms of aedeagus bearing a subapical, outer, pointed process; female with legs uniformly pale brown and subgenital plate represented by a slender transverse band. Pupa with respiratory organ thick at the base and distal half tapering to a sharp apex, with 18–20 apical pores, and terminal processes as long as length of body of segment 9.
</paragraph>
</subSubSection>
<subSubSection id="C3445192FF833F4B83D32A48FDC4E06C" lastPageId="3" lastPageNumber="718" pageId="2" pageNumber="717" type="description">
<paragraph id="8BE10219FF833F4A83D32A48FC96E76C" blockId="2.[151,1128,220,1555]" pageId="2" pageNumber="717">
<emphasis id="B92ADE0BFF833F4A83D32A48FEEAE50C" bold="true" box="[151,354,347,371]" pageId="2" pageNumber="717">
Description.
<emphasis id="B92ADE0BFF833F4A826E2A4FFEEAE50C" bold="true" box="[298,354,347,371]" italics="true" pageId="2" pageNumber="717">Male</emphasis>
</emphasis>
(
<figureCitation id="13651E9CFF833F4A82352A48FE5BE50C" box="[369,467,348,371]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Figs 1–4</figureCitation>
,
<figureCitation id="13651E9CFF833F4A829B2A48FDAEE50C" box="[479,550,348,371]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">21–24</figureCitation>
). Similar to female with the usual sexual differences. Flagellum as in
<figureCitation id="13651E9CFF833F4A82072A68FE0AE5EC" box="[323,386,380,403]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Fig. 1</figureCitation>
. Palpus (
<figureCitation id="13651E9CFF833F4A82A12A68FDADE5EC" box="[485,549,380,403]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Fig. 2</figureCitation>
) with segment 3 bearing scattered sensilla on surface;
<collectingRegion id="499ACCFBFF833F4A83D32A88FF30E5CC" box="[151,184,412,435]" country="Brazil" name="Para" pageId="2" pageNumber="717">PR</collectingRegion>
2.00–2.60 (2.45, n = 6). Scutellum with 6–7 strong, 2 thinner setae. Wing (
<figureCitation id="13651E9CFF833F4A80922A88FB9CE5CC" box="[982,1044,412,435]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Fig. 3</figureCitation>
) length 0.93–0.99 (0.95, n = 4) mm, width 0.24–0.30 (0.27, n = 4) mm, CR 0.47–0.51 (0.48, n = 4). Genitalia (
<figureCitation id="13651E9CFF833F4A82432AC8FEC3E58C" box="[263,331,476,499]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Figs 4</figureCitation>
,
<figureCitation id="13651E9CFF833F4A82132AC8FEFDE58C" box="[343,373,476,499]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">21</figureCitation>
): tergite 9 tapering distally, distal margin rounded; apicolateral process short, stout, with apical seta; cercus small, with 1–2 setae; sternite 9 (
<figureCitation id="13651E9CFF833F4A80372AE8FC3FE66C" box="[883,951,508,531]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Figs 4</figureCitation>
,
<figureCitation id="13651E9CFF833F4A80862AE8FC56E66C" box="[962,990,508,531]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">22</figureCitation>
) 0.3× longer than greatest width, distal margin slightly convex. Gonocoxite stout, 1.5× longer than greatest width, with blunt anteromedian process; gonostylus 0.8× longer than gonocoxite, narrow base, nearly straight, tip pointed, spur-like. Paramere and gonocoxal apodemes forming an asymmetrical structure (
<figureCitation id="13651E9CFF833F4A82D82968FE6AE6EC" box="[412,482,636,659]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Figs 4</figureCitation>
,
<figureCitation id="13651E9CFF833F4A82AB2968FD85E6EC" box="[495,525,636,659]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">23</figureCitation>
); gonocoxal apodeme stout, curved, not in contact with paramere, latter elongate, straight, tapering to slender blunt apex. Aedeagus (
<figureCitation id="13651E9CFF833F4A80F42988FC7BE6CC" box="[944,1011,668,691]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Figs 4</figureCitation>
,
<figureCitation id="13651E9CFF833F4A80B82988FB90E6CC" box="[1020,1048,668,691]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">24</figureCitation>
) nearly as long as greatest width, basal arch slightly concave, low, extending to 0.15 of total length; basal arms stout, directed anterolaterally; posterolateral arms stout, slightly convergent, each tapering to pointed tip with subapical, outer pointed process.
</paragraph>
<paragraph id="8BE10219FF833F4A83F1280FFDF7E7CC" blockId="2.[151,1128,220,1555]" pageId="2" pageNumber="717">
<emphasis id="B92ADE0BFF833F4A83F1280FFE80E74C" bold="true" box="[181,264,795,819]" italics="true" pageId="2" pageNumber="717">Female</emphasis>
(
<figureCitation id="13651E9CFF833F4A82522808FE09E74C" box="[278,385,796,819]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Figs 5–11</figureCitation>
,
<figureCitation id="13651E9CFF833F4A82CF2808FE5AE74C" box="[395,466,796,819]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">25–27</figureCitation>
). Head dark brown (
<figureCitation id="13651E9CFF833F4A81EC2808FD6FE74C" box="[680,743,796,819]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Fig. 5</figureCitation>
) Eyes contiguous by width of 2 ommatidia. Antenna with flagellum (
<figureCitation id="13651E9CFF833F4A82B82828FDB3E72C" box="[508,571,828,851]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Fig. 5</figureCitation>
) brown; AR 0.94–0.97 (0.95, n = 2). Frontal sclerite semicircular (
<figureCitation id="13651E9CFF833F4A826D2848FEF0E70C" box="[297,376,860,883]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">Fig. 25</figureCitation>
), with long, slender, posteromesally directed projection. Clypeus (
<figureCitation id="13651E9CFF833F4A877F2848FF2DE7EC" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Fig. 6</figureCitation>
) with three pairs of setae. Palpus (
<figureCitation id="13651E9CFF833F4A814F2868FDEEE7EC" box="[523,614,892,915]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Figs 5, 7</figureCitation>
) whitish, segment 5 slightly infuscated; segment 3 with 6–7 capitate sensillae,
<collectingRegion id="499ACCFBFF833F4A82972888FE7CE7CC" box="[467,500,924,947]" country="Brazil" name="Para" pageId="2" pageNumber="717">PR</collectingRegion>
1.86 (n = 2).
</paragraph>
<paragraph id="8BE10219FF833F4A83F128A8FD03E00C" blockId="2.[151,1128,220,1555]" pageId="2" pageNumber="717">
Thorax (
<figureCitation id="13651E9CFF833F4A825528A8FEDAE7AC" box="[273,338,956,979]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Fig. 8</figureCitation>
). Scutum brown, scutellum pale, with 6–7 strong, 1–2 thinner setae. Legs (
<figureCitation id="13651E9CFF833F4A83DB28C8FF69E78C" box="[159,225,988,1011]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Fig. 9</figureCitation>
) uniformly pale brown, tarsomeres 5 infuscated; apex of hind tibia with 5 spines; foreleg TR 2.37–2.50 (2.43, n = 2), midleg TR 2.41–2.44 (2.42, n = 2), hind leg TR 2.31–2.36 (2.33, n = 2). Wing (
<figureCitation id="13651E9CFF833F4A822A2F08FE34E04C" box="[366,444,1052,1075]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Fig. 10</figureCitation>
), length 0.96 (n = 2) mm, width 0.36–0.39 (0.37, n = 2) mm, CR 0.50–0.56 (0.53, n = 2); membrane hyaline, densely covered with macrotrichia; cubital fork at level of ¿rst radial cell. Halter knob whitish.
</paragraph>
<paragraph id="8BE10219FF833F4A83F12F68FC92E0AC" blockId="2.[151,1128,220,1555]" pageId="2" pageNumber="717">
Abdomen. Pale brown. Subgenital plate (
<figureCitation id="13651E9CFF833F4A81312F68FD40E0EC" box="[629,712,1148,1171]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Figs 11</figureCitation>
,
<figureCitation id="13651E9CFF833F4A81922F68FD7AE0EC" box="[726,754,1148,1171]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">26</figureCitation>
) represented by slender tranversal band, slightly arised anteromesally. Spermatheca ovoid (
<figureCitation id="13651E9CFF833F4A80442F88FCDBE0CC" box="[768,851,1180,1203]" captionStart="Figs 1–11" captionStartId="4.[151,188,1483,1502]" captionTargetBox="[182,1095,219,1464]" captionTargetId="figure-10@4.[182,1096,219,1464]" captionTargetPageId="4" captionText="Figs 1–11. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="2" pageNumber="717">Figs 11</figureCitation>
,
<figureCitation id="13651E9CFF833F4A80262F88FC08E0CC" box="[866,896,1180,1203]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">27</figureCitation>
), heavily sclerotized, diameter 252 µm, neck short, sclerotized, measuring 12 µm.
</paragraph>
<paragraph id="8BE10219FF833F4B83F12FCFFBCAE76C" blockId="2.[151,1128,220,1555]" lastBlockId="3.[113,1090,220,1555]" lastPageId="3" lastPageNumber="718" pageId="2" pageNumber="717">
<emphasis id="B92ADE0BFF833F4A83F12FCFFEC9E08C" bold="true" box="[181,321,1243,1267]" italics="true" pageId="2" pageNumber="717">Female pupa</emphasis>
(
<figureCitation id="13651E9CFF833F4A82092FC8FE4BE08C" box="[333,451,1244,1267]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="2" pageNumber="717">Figs 13–19</figureCitation>
,
<figureCitation id="13651E9CFF833F4A82882FC8FD9AE08C" box="[460,530,1244,1267]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">28–32</figureCitation>
). Total length
<quantity id="4CA6AFFCFF833F4A81E42FC8FD76E08C" box="[672,766,1244,1267]" metricMagnitude="-3" metricUnit="m" metricValue="2.88" pageId="2" pageNumber="717" unit="mm" value="2.88">2.88 mm</quantity>
. General coloration of exuviae pale brown. Dorsal apotome (
<figureCitation id="13651E9CFF833F4A82E52FE8FE66E16C" box="[417,494,1276,1299]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="2" pageNumber="717">Fig. 13</figureCitation>
) 2.11× broader than long, surface smooth with wrinkles on margins, anterior margin slightly concave, posterior margin rounded; dorsal apotome sensilla with one short, thin seta, one campaniform sensillum (
<figureCitation id="13651E9CFF833F4A81A22E28FCBEE12C" box="[742,822,1340,1363]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="2" pageNumber="717">Fig. 13</figureCitation>
); DAL 0.10–0.12 (0.11, n = 2) mm; DAW 0.240 (n = 2) mm; DAW/DAL 2.00–2.22 (2.11, n = 2). Cephalothorax surface with very small rounded tubercles, length 0.99–1.02 (1.00, n = 2) mm, width 0.69–0.75 (0.72, n = 2) mm. Cephalothoracic sensilla as follows: three dorsolateral cephalic sensilla (
<figureCitation id="13651E9CFF833F4A87542E88FBEAE1CC" box="[1040,1122,1436,1459]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="2" pageNumber="717">Figs 14</figureCitation>
,
<figureCitation id="13651E9CFF833F4A83D32EA8FF3CE1AC" box="[151,180,1468,1491]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">28</figureCitation>
), two minute setae, one campaniform sensilla; three anterolateral sensilla (
<figureCitation id="13651E9CFF833F4A80982EA8FBB8E1AC" box="[988,1072,1468,1491]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="2" pageNumber="717">Figs 14</figureCitation>
,
<figureCitation id="13651E9CFF833F4A87782EA8FBD1E1AC" box="[1084,1113,1468,1491]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">28</figureCitation>
), one long, thin seta, one medium-sized seta, one coeloconica sensillum; two anteromedial sensilla, one medium-sized seta, one short seta (
<figureCitation id="13651E9CFF833F4A81DB2EE8FD7BE26C" box="[671,755,1532,1555]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="2" pageNumber="717">Figs 14</figureCitation>
,
<figureCitation id="13651E9CFF833F4A81BB2EE8FC95E26C" box="[767,797,1532,1555]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">28</figureCitation>
); dorsals (
<figureCitation id="13651E9CFF833F4A80C82EE8FC68E26C" box="[908,992,1532,1555]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="2" pageNumber="717">Figs 15</figureCitation>
,
<figureCitation id="13651E9CFF833F4A80A82EE8FB83E26C" box="[1004,1035,1532,1555]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="2" pageNumber="717">29</figureCitation>
): D-1-T, D-2-T, D-4-T absent, D-3-T campaniform sensillum, D-5-T short, thin seta, supraalar (SA- 2-T) campaniform sensillum. Respiratory organ (
<figureCitation id="13651E9CFF823F4B813C2BE8FD4CE56C" box="[632,708,252,275]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Fig. 16</figureCitation>
) thick at base, distal half tapering to a sharp apex, 4.75× longer than broad, surface smooth, with 18–20 apical pores; RO length 0.23–0.25 (0.24, n = 2) mm, RO width 0.05 (n = 2) mm; pedicel (P) pale brown, smooth, short, pedicel length 0.024 (n = 2) mm, P/RO
<geoCoordinate id="EE6A64DEFF823F4B81102A48FD18E50C" box="[596,656,348,371]" degrees="0.100" direction="north" orientation="latitude" pageId="3" pageNumber="718" precision="55" value="0.1">0.100</geoCoordinate>
<geoCoordinate id="EE6A64DEFF823F4B81D42A48FD51E50C" box="[656,729,348,371]" degrees="0.105" direction="west" orientation="longitude" pageId="3" pageNumber="718" precision="55" value="-0.105">–0.105</geoCoordinate>
(0.1025, n = 2) mm. Two clypeal/ labral sensilla (
<figureCitation id="13651E9CFF823F4B82572A68FEEFE5EC" box="[275,359,380,403]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Figs 17</figureCitation>
,
<figureCitation id="13651E9CFF823F4B82362A68FE18E5EC" box="[370,400,380,403]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="3" pageNumber="718">30</figureCitation>
), CL-1-H long, thin seta, CL-2-H short seta; two ocular sensilla, one long, thin seta, one campaniform sensillum (
<figureCitation id="13651E9CFF823F4B81332A88FD42E5CC" box="[631,714,412,435]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Figs 17</figureCitation>
,
<figureCitation id="13651E9CFF823F4B81912A88FD7BE5CC" box="[725,755,412,435]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="3" pageNumber="718">30</figureCitation>
). Abdominal segments covered with small spinules. Abdominal segment 1 (
<figureCitation id="13651E9CFF823F4B81072AA8FD1EE5AC" box="[579,662,444,467]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Figs 18</figureCitation>
,
<figureCitation id="13651E9CFF823F4B81E52AA8FD34E5AC" box="[673,700,444,467]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="3" pageNumber="718">31</figureCitation>
) with sensilla as follows: D-2-I peg; D-4-I, D-7-I pores, D-8-I short seta; 3 lateral sensilla, L-1-I, L-3-I short setae, L-2-I minute seta. Second abdominal segment similar to ¿rst. Segment 4 with sensillar pattern (
<figureCitation id="13651E9CFF823F4B80952AE8FB95E66C" box="[977,1053,508,531]" captionStart="Figs 21–32" captionStartId="6.[151,188,1413,1432]" captionTargetBox="[231,1051,223,1379]" captionTargetId="figure-10@6.[226,1052,219,1380]" captionTargetPageId="6" captionText="Figs 21–32. Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov. 21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; D-3-T, D-5-T – dorsal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV – segment 4 chaetotaxy; O-1-H, O-2-H – ocular sensilla; SA-2-T – supraalar." figureDoi="http://doi.org/10.5281/zenodo.5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="3" pageNumber="718">Fig. 32</figureCitation>
) as follows: D-2-IV peg; D-4-IV, D-7-IV, D-8-IV without setae, all located on flattened tubercles; L-1-IV medium-sized, thin seta on triangular tubercle, L-2-IV campaniform sensillum, L-3-IV, L-4-IV minute setae, all located on triangular tubercles; two ventral sensilla, V-5-IV without seta, V-6-IV short, stout seta, both located on bi¿d tubercles. Segment 9 (
<figureCitation id="13651E9CFF823F4B80272968FC26E6EC" box="[867,942,636,659]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Fig. 19</figureCitation>
) 1.45× longer than width, ventral surface with many spinules; length
<geoCoordinate id="EE6A64DEFF823F4B81F52988FD64E6CC" box="[689,748,668,691]" degrees="0.348" direction="north" orientation="latitude" pageId="3" pageNumber="718" precision="55" value="0.348">0.348</geoCoordinate>
<geoCoordinate id="EE6A64DEFF823F4B81A82988FCBEE6CC" box="[748,822,668,691]" degrees="0.360" direction="west" orientation="longitude" pageId="3" pageNumber="718" precision="55" value="-0.36">–0.360</geoCoordinate>
(0.354, n = 2) mm, width
<geoCoordinate id="EE6A64DEFF823F4B833529A8FF25E6AC" box="[113,173,700,723]" degrees="0.204" direction="north" orientation="latitude" pageId="3" pageNumber="718" precision="55" value="0.204">0.204</geoCoordinate>
<geoCoordinate id="EE6A64DEFF823F4B83E929A8FF7FE6AC" box="[173,247,700,723]" degrees="0.240" direction="west" orientation="longitude" pageId="3" pageNumber="718" precision="55" value="-0.24">–0.240</geoCoordinate>
(0.222, n = 2) mm. Terminal process (
<figureCitation id="13651E9CFF823F4B81C329A8FD5AE6AC" box="[647,722,700,723]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Fig. 19</figureCitation>
) straight, as long as length of body of segment 9, tip pointed, base broad with two setae, one long, thin, hyaline seta on small rounded base, other medium-sized, stout seta on rounded tubercle; length 0.180 (n = 2) mm.
</paragraph>
<paragraph id="8BE10219FF823F4B83D4280FFDC4E06C" blockId="3.[113,1090,220,1555]" pageId="3" pageNumber="718">
<emphasis id="B92ADE0BFF823F4B83D4280FFE80E74C" bold="true" box="[144,264,795,819]" italics="true" pageId="3" pageNumber="718">Male pupa</emphasis>
(
<figureCitation id="13651E9CFF823F4B825C2808FE10E74C" box="[280,408,796,819]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Figs 12, 20</figureCitation>
). Similar to female with usual sexual differences: Total length 2.52–2.64 (2.61, n = 6) mm. Exuviae pale brown. Dorsal apotome with DAL 0.12–0.14 (0.130, n = 6) mm; DAW 0.24–0.26 (0.25, n = 6) mm, DAW/DAL 1.85–2.10 (1.98, n = 6). Respiratory organ (
<figureCitation id="13651E9CFF823F4B83AA2868FEB4E7EC" box="[238,316,892,915]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Fig. 12</figureCitation>
), RO length 0.22–0.24 (0.23, n = 7) mm, RO width 0.05 (n = 7); pedicel length
<geoCoordinate id="EE6A64DEFF823F4B83FF2888FF7FE7CC" box="[187,247,924,947]" degrees="0.024" direction="north" orientation="latitude" pageId="3" pageNumber="718" precision="55" value="0.024">0.024</geoCoordinate>
<geoCoordinate id="EE6A64DEFF823F4B83B32888FEC8E7CC" box="[247,320,924,947]" degrees="0.030" direction="west" orientation="longitude" pageId="3" pageNumber="718" precision="55" value="-0.03">–0.030</geoCoordinate>
(0.025, n = 7) mm, P/RO 0.10–0.13 (0.107, n = 7). Cephalothorax (
<figureCitation id="13651E9CFF823F4B87522888FF04E7AC" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Fig. 12</figureCitation>
) length 0.99–1.07 (1.03, n = 6) mm, width 0.66–0.74 (0.70, n = 6) mm. Segment 9 (
<figureCitation id="13651E9CFF823F4B875128A8FF05E78C" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Fig. 20</figureCitation>
) length 0.34–0.39 (0.35, n = 7) mm, width 0.19–0.24 (0.22, n = 7) mm; terminal process (
<figureCitation id="13651E9CFF823F4B833D28E8FF4FE06C" box="[121,199,1020,1043]" captionStart="Figs 12–20" captionStartId="5.[113,149,1441,1460]" captionTargetBox="[162,1039,219,1413]" captionTargetId="figure-15@5.[162,1040,219,1414]" captionTargetPageId="5" captionText="Figs 12–20.Dasyhelea pseudopollinosa Díaz &amp; Ronderos sp. nov., pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations: AL-1-T, AL-2-T, AL-3-T íanterolateral sensilla; AM-1-T, AM-2-T – anteromedial sensilla; CL-1-H, CL-2-H – clypeal/labral sensilla; DA – dorsal apotome; DA-1-H, DA-2-H – dorsal apotomal sensilla; DL-1-H, DL-2-H, DL-3-H – dorsolateral cephalic sclerite sensilla; D-3-T, D-5-T – dorsal sensilla; D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I – abdominal segment 1 chaetotaxy; GL – genital lobe; O-1-H, O-2-H – ocular sensilla; P – pedicel; RO – respiratory organ; SA-2-T – supraalar; TP – terminal process." figureDoi="http://doi.org/10.5281/zenodo.5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="3" pageNumber="718">Fig. 20</figureCitation>
) length 0.17–0.24 (0.19, n = 7) mm.
</paragraph>
</subSubSection>
<subSubSection id="C3445192FF823F4B83352F08FD21E02C" pageId="3" pageNumber="718" type="etymology">
<paragraph id="8BE10219FF823F4B83352F08FD21E02C" blockId="3.[113,1090,220,1555]" pageId="3" pageNumber="718">
<emphasis id="B92ADE0BFF823F4B83352F08FF7AE04C" bold="true" box="[113,242,1052,1075]" pageId="3" pageNumber="718">Etymology.</emphasis>
The species name
<taxonomicName id="4C5E799AFF823F4B82802F0FFD64E04C" authorityName="Diaz &amp; Torreias &amp; Spinelli &amp; Ronderos &amp; Unlp" authorityYear="2014" box="[452,748,1051,1075]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="718" phylum="Arthropoda" rank="species" species="pseudopollinosa">
<emphasis id="B92ADE0BFF823F4B82802F0FFD64E04C" box="[452,748,1051,1075]" italics="true" pageId="3" pageNumber="718">Dasyhelea pseudopollinosa</emphasis>
</taxonomicName>
refers to the similarity with its congener, Nearctic
<taxonomicName id="4C5E799AFF823F4B82042F2FFD2DE02C" authority="Wirth, 1952" authorityName="Wirth" authorityYear="1952" box="[320,677,1083,1107]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="718" phylum="Arthropoda" rank="species" species="pollinosa">
<emphasis id="B92ADE0BFF823F4B82042F2FFD93E02C" box="[320,539,1083,1107]" italics="true" pageId="3" pageNumber="718">Dasyhelea pollinosa</emphasis>
Wirth, 1952
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3445192FF823F4B83352F48FE54E00C" box="[113,476,1116,1139]" pageId="3" pageNumber="718" type="distribution">
<paragraph id="8BE10219FF823F4B83352F48FE54E00C" blockId="3.[113,1090,220,1555]" box="[113,476,1116,1139]" pageId="3" pageNumber="718">
<emphasis id="B92ADE0BFF823F4B83352F48FE8CE00C" bold="true" box="[113,260,1116,1139]" pageId="3" pageNumber="718">Distribution.</emphasis>
<collectingCountry id="F3494289FF823F4B824E2F48FEC3E00C" box="[266,331,1116,1139]" name="Brazil" pageId="3" pageNumber="718">Brazil</collectingCountry>
(
<collectingRegion id="499ACCFBFF823F4B821D2F48FE58E00C" box="[345,464,1116,1139]" country="Brazil" name="Amazonas" pageId="3" pageNumber="718">Amazonas</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3445192FF823F4F83352F68FBB7E60C" lastPageId="7" lastPageNumber="722" pageId="3" pageNumber="718" type="discussion">
<paragraph id="8BE10219FF823F4B83352F68FE35E16C" blockId="3.[113,1090,220,1555]" pageId="3" pageNumber="718">
<emphasis id="B92ADE0BFF823F4B83352F68FF78E0EC" bold="true" box="[113,240,1148,1171]" pageId="3" pageNumber="718">Discussion.</emphasis>
<bibRefCitation id="EFCF7FE8FF823F4B83B32F68FE75E0EC" author="WAUGH W. T. &amp; WIRTH W. W." box="[247,509,1148,1171]" pageId="3" pageNumber="718" pagination="219 - 247" refId="ref7471" refString="WAUGH W. T. &amp; WIRTH W. W. 1976: A revision of the genus Dasyhelea Kieffer of the eastern United States north of Florida (Diptera: Ceratopogonidae). Annals of the Entomological Society of America 69: 219 - 247." type="journal article" year="1976">WAUGH &amp; WIRTH (1976)</bibRefCitation>
proposed the
<taxonomicName id="4C5E799AFF823F4B81D22F68FD5FE0EC" baseAuthorityName="Coquillett" baseAuthorityYear="1901" box="[662,727,1148,1171]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="718" phylum="Arthropoda" rank="species" species="cincta">
<emphasis id="B92ADE0BFF823F4B81D22F68FD5FE0EC" box="[662,727,1148,1171]" italics="true" pageId="3" pageNumber="718">cincta</emphasis>
</taxonomicName>
,
<emphasis id="B92ADE0BFF823F4B81A22F68FCA1E0EC" box="[742,809,1148,1171]" italics="true" pageId="3" pageNumber="718">grisea</emphasis>
,
<emphasis id="B92ADE0BFF823F4B80732F6FFC45E0EC" box="[823,973,1147,1171]" italics="true" pageId="3" pageNumber="718">leptobranchia</emphasis>
and
<emphasis id="B92ADE0BFF823F4B87462F68FF17E0CC" italics="true" pageId="3" pageNumber="718">mutabilis</emphasis>
species-groups for the Nearctic
<taxonomicName id="4C5E799AFF823F4B82BD2F8FFD49E0CC" box="[505,705,1179,1203]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="718" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92ADE0BFF823F4B82BD2F8FFDE2E0CC" box="[505,618,1179,1203]" italics="true" pageId="3" pageNumber="718">Dasyhelea</emphasis>
species
</taxonomicName>
, working in an exclusively phenetic framework. Most of the Neotropical species ¿t within these groups or are in the
<emphasis id="B92ADE0BFF823F4B80962FAFFBC8E0AC" box="[978,1088,1211,1235]" italics="true" pageId="3" pageNumber="718">brevicosta</emphasis>
species-group proposed by
<bibRefCitation id="EFCF7FE8FF823F4B82DB2FC8FD4FE08C" author="GROGAN W. L. &amp; WIENERS J. A." box="[415,711,1244,1267]" pageId="3" pageNumber="718" pagination="467 - 473" refId="ref7426" refString="GROGAN W. L. &amp; WIENERS J. A. 2006. A new species of the biting midges genus Dasyhelea Kieffer (Diptera: Ceratopogonidae) from the Bahamas. Proceedings of the Entomological Society of Washington 108: 467 - 473." type="journal article" year="2006">GROGAN &amp; WIENERS (2006)</bibRefCitation>
and the
<taxonomicName id="4C5E799AFF823F4B806F2FC8FC2BE08C" authorityName="Ingram &amp; Macfie" authorityYear="1931" box="[811,931,1244,1267]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="718" phylum="Arthropoda" rank="species" species="patagonica">
<emphasis id="B92ADE0BFF823F4B806F2FC8FC2BE08C" box="[811,931,1244,1267]" italics="true" pageId="3" pageNumber="718">patagonica</emphasis>
</taxonomicName>
species-group proposed by
<bibRefCitation id="EFCF7FE8FF823F4B83BF2FE8FE31E16C" author="DIAZ F. &amp; SPINELLI G. R. &amp; RONDEROS M. M." box="[251,441,1276,1299]" pageId="3" pageNumber="718" pagination="2825 - 2849" refId="ref7342" refString="DIAZ F., SPINELLI G. R. &amp; RONDEROS M. M. 2010: A revision of the species allied to Dasyhelea patagonica Ingram &amp; Mac? e (Diptera: Ceratopogonidae). Journal of Natural History 44: 2825 - 2849." type="journal article" year="2010">DÍAZ et al. (2010)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE10219FF823F4B83D42E08FC1EE1CC" blockId="3.[113,1090,220,1555]" pageId="3" pageNumber="718">
We provisionally place this new species within the
<emphasis id="B92ADE0BFF823F4B81F52E08FD7CE14C" box="[689,756,1308,1331]" italics="true" pageId="3" pageNumber="718">grisea</emphasis>
group by virtue of the well developed frontal sclerite and sinuate paramere. On the other hand, the uniformly pale brown legs, the female subgenital plate represented by a transversal sclerotized band without anterior triangular projection and the posterolateral arms of aedeagus bearing an outer subapical pointed process, are not characters otherwise present in species of the
<emphasis id="B92ADE0BFF823F4B80432E88FCC2E1CC" box="[775,842,1436,1459]" italics="true" pageId="3" pageNumber="718">grisea</emphasis>
group.
</paragraph>
<paragraph id="8BE10219FF823F4F83D42EA8FC3FE54C" blockId="3.[113,1090,220,1555]" lastBlockId="7.[113,1089,220,627]" lastPageId="7" lastPageNumber="722" pageId="3" pageNumber="718">
The pupa of
<taxonomicName id="4C5E799AFF823F4B82562EAFFE71E1AC" authorityName="Forattini &amp; Rabello" authorityYear="1957" box="[274,505,1467,1491]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="718" phylum="Arthropoda" rank="species" species="paulistana">
<emphasis id="B92ADE0BFF823F4B82562EAFFE71E1AC" box="[274,505,1467,1491]" italics="true" pageId="3" pageNumber="718">Dasyhelea paulistana</emphasis>
</taxonomicName>
is very similar to this new species by virtue of the respiratory organ thick at base with its distal half tapering to a sharp apex, and by the slender and elongate terminal processes of the segment 9 but in
<taxonomicName id="4C5E799AFF823F4B81D32EE8FCA0E26C" authorityName="Forattini &amp; Rabello" authorityYear="1957" box="[663,808,1531,1555]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="718" phylum="Arthropoda" rank="species" species="paulistana">
<emphasis id="B92ADE0BFF823F4B81D32EE8FD22E26C" box="[663,682,1532,1555]" italics="true" pageId="3" pageNumber="718">D</emphasis>
.
<emphasis id="B92ADE0BFF823F4B81F32EEFFCA0E26C" box="[695,808,1531,1555]" italics="true" pageId="3" pageNumber="718">paulistana</emphasis>
</taxonomicName>
the terminal processes are very much longer, twice the body length of segment 9. The adult of
<taxonomicName id="4C5E799AFF863F4F80022BC8FC50E48B" authorityName="Forattini &amp; Rabello" authorityYear="1957" box="[838,984,220,244]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="722" phylum="Arthropoda" rank="species" species="paulistana">
<emphasis id="B92ADE0BFF863F4F80022BC8FCD1E48C" box="[838,857,220,243]" italics="true" pageId="7" pageNumber="722">D</emphasis>
.
<emphasis id="B92ADE0BFF863F4F80232BC8FC50E48B" box="[871,984,220,244]" italics="true" pageId="7" pageNumber="722">paulistana</emphasis>
</taxonomicName>
is readily distinguished from the new species by the ovoid female subgenital plate, the elongate and sinuate paramere hooked at tip, and by the aedeagus with V-shaped basal arch.
</paragraph>
<caption id="DF215291FF853F4C83D32EDFFDE0E26C" ID-DOI="http://doi.org/10.5281/zenodo.5300383" ID-Zenodo-Dep="5300383" httpUri="https://zenodo.org/record/5300383/files/figure.png" pageId="4" pageNumber="719" startId="4.[151,188,1483,1502]" targetBox="[182,1095,219,1464]" targetPageId="4">
<paragraph id="8BE10219FF853F4C83D32EDFFDE0E26C" blockId="4.[151,1127,1483,1555]" pageId="4" pageNumber="719">
Figs 1–11.
<taxonomicName id="4C5E799AFF853F4C83B22EDFFDF3E1A1" authority="Diaz &amp; Ronderos" authorityName="Diaz &amp; Torreias &amp; Spinelli &amp; Ronderos &amp; Unlp" authorityYear="2014" box="[246,635,1483,1502]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="719" phylum="Arthropoda" rank="species" species="pseudopollinosa" status="sp. nov.">
<emphasis id="B92ADE0BFF853F4C83B22EDFFE69E1A1" box="[246,481,1483,1502]" italics="true" pageId="4" pageNumber="719">Dasyhelea pseudopollinosa</emphasis>
Díaz &amp; Ronderos
</taxonomicName>
<taxonomicNameLabel id="A2196370FF853F4C81C42EDFFD4DE1A1" box="[640,709,1483,1502]" pageId="4" pageNumber="719" rank="species">sp. nov.</taxonomicNameLabel>
, adult. 1–4 – holotype, male; 5–11 – allotype, female. 1, 5 – flagellum, anterior view; 2, 7 – palpus; 3, 10 – wing; 4 – genitalia, ventral view; 6 – clypeus; 8 – thorax; 9 – legs; 11 – abdominal segments 8–10, ventral view.
</paragraph>
</caption>
<caption id="DF215291FF843F4D83352EB5FC22E211" ID-DOI="http://doi.org/10.5281/zenodo.5300385" ID-Zenodo-Dep="5300385" httpUri="https://zenodo.org/record/5300385/files/figure.png" pageId="5" pageNumber="720" startId="5.[113,149,1441,1460]" targetBox="[162,1039,219,1413]" targetPageId="5">
<paragraph id="8BE10219FF843F4D83352EB5FC22E211" blockId="5.[113,1090,1440,1646]" pageId="5" pageNumber="720">
Figs 12–20.
<taxonomicName id="4C5E799AFF843F4D83932EB4FDDBE1CB" authority="Diaz &amp; Ronderos" authorityName="Diaz &amp; Torreias &amp; Spinelli &amp; Ronderos &amp; Unlp" authorityYear="2014" box="[215,595,1440,1460]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="720" phylum="Arthropoda" rank="species" species="pseudopollinosa" status="sp. nov.">
<emphasis id="B92ADE0BFF843F4D83932EB4FE36E1CC" box="[215,446,1440,1459]" italics="true" pageId="5" pageNumber="720">Dasyhelea pseudopollinosa</emphasis>
Díaz &amp; Ronderos
</taxonomicName>
<taxonomicNameLabel id="A2196370FF843F4D81122EB5FD11E1CB" box="[598,665,1441,1460]" pageId="5" pageNumber="720" rank="species">sp. nov.</taxonomicNameLabel>
, pupa.12, 20 – male pupa; 13–19 – female pupa.12 – entire pupa; 13 – dorsal apotome; 14 – cephalothoracic chaetotaxy (dorsal view); 15 – dorsal sensilla and supraalar; 16 – respiratory organ; 17 – clypeal/labral sensilla and ocular sensilla; 18 – abdominal segment 1 chaetotaxy; 19–20 – segment 9. Abbreviations:
<emphasis id="B92ADE0BFF843F4D822D2EE5FDC0E27B" bold="true" box="[361,584,1521,1540]" pageId="5" pageNumber="720">AL-1-T, AL-2-T, AL-3-T</emphasis>
íanterolateral sensilla;
<emphasis id="B92ADE0BFF843F4D80592EE5FC33E27B" bold="true" box="[797,955,1521,1540]" pageId="5" pageNumber="720">AM-1-T, AM-2-T</emphasis>
– anteromedial sensilla;
<emphasis id="B92ADE0BFF843F4D83FA2D1FFED0E261" bold="true" box="[190,344,1547,1566]" pageId="5" pageNumber="720">CL-1-H, CL-2-H</emphasis>
– clypeal/labral sensilla;
<emphasis id="B92ADE0BFF843F4D817D2D1FFDD0E261" bold="true" box="[569,600,1547,1566]" pageId="5" pageNumber="720">DA</emphasis>
– dorsal apotome;
<emphasis id="B92ADE0BFF843F4D80452D1FFC16E261" bold="true" box="[769,926,1547,1566]" pageId="5" pageNumber="720">DA-1-H, DA-2-H</emphasis>
– dorsal apotomal sensilla;
<emphasis id="B92ADE0BFF843F4D83F82D32FE2EE246" bold="true" box="[188,422,1574,1593]" pageId="5" pageNumber="720">DL-1-H, DL-2-H, DL-3-H</emphasis>
– dorsolateral cephalic sclerite sensilla;
<emphasis id="B92ADE0BFF843F4D81B92D32FCFAE246" bold="true" box="[765,882,1574,1593]" pageId="5" pageNumber="720">D-3-T, D-5-T</emphasis>
– dorsal sensilla;
<emphasis id="B92ADE0BFF843F4D874F2D32FE43E22B" bold="true" pageId="5" pageNumber="720">D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I</emphasis>
– abdominal segment 1 chaetotaxy;
<emphasis id="B92ADE0BFF843F4D804D2D55FCA0E22B" bold="true" box="[777,808,1601,1620]" pageId="5" pageNumber="720">GL</emphasis>
– genital lobe;
<emphasis id="B92ADE0BFF843F4D80F42D55FBA7E22B" bold="true" box="[944,1071,1601,1620]" pageId="5" pageNumber="720">O-1-H, O-2-H</emphasis>
– ocular sensilla;
<emphasis id="B92ADE0BFF843F4D83B32D4FFE8CE211" bold="true" box="[247,260,1627,1646]" pageId="5" pageNumber="720">P</emphasis>
– pedicel;
<emphasis id="B92ADE0BFF843F4D82262D4FFE0AE211" bold="true" box="[354,386,1627,1646]" pageId="5" pageNumber="720">RO</emphasis>
– respiratory organ;
<emphasis id="B92ADE0BFF843F4D81712D4FFDFFE211" bold="true" box="[565,631,1627,1646]" pageId="5" pageNumber="720">SA-2-T</emphasis>
– supraalar;
<emphasis id="B92ADE0BFF843F4D81A22D4FFC89E211" bold="true" box="[742,769,1627,1646]" pageId="5" pageNumber="720">TP</emphasis>
– terminal process.
</paragraph>
</caption>
<caption id="DF215291FF873F4E83D32E91FCFDE212" ID-DOI="http://doi.org/10.5281/zenodo.5300387" ID-Zenodo-Dep="5300387" httpUri="https://zenodo.org/record/5300387/files/figure.png" pageId="6" pageNumber="721" startId="6.[151,188,1413,1432]" targetBox="[231,1051,223,1379]" targetPageId="6">
<paragraph id="8BE10219FF873F4E83D32E91FCFDE212" blockId="6.[151,1127,1412,1645]" pageId="6" pageNumber="721">
Figs 21–32.
<taxonomicName id="4C5E799AFF873F4E83BA2E90FDF4E1E7" authority="Diaz &amp; Ronderos" authorityName="Diaz &amp; Torreias &amp; Spinelli &amp; Ronderos &amp; Unlp" authorityYear="2014" box="[254,636,1412,1432]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="721" phylum="Arthropoda" rank="species" species="pseudopollinosa" status="sp. nov.">
<emphasis id="B92ADE0BFF873F4E83BA2E90FE6EE1E8" box="[254,486,1412,1431]" italics="true" pageId="6" pageNumber="721">Dasyhelea pseudopollinosa</emphasis>
Díaz &amp; Ronderos
</taxonomicName>
<taxonomicNameLabel id="A2196370FF873F4E81C42E91FD49E1E7" box="[640,705,1413,1432]" pageId="6" pageNumber="721" rank="species">sp. nov.</taxonomicNameLabel>
21–24 – holotype, male; 25–27 – allotype, female; 28–32 – female pupa. 21 – genitalia; 22 – sternite 9; 23 – paramere and gonocoxal apodemes; 24 – aedeagus; 25 – frontal sclerite; 26 – subgenital plate; 27 – spermatheca; 28 – cephalothoracic chaetotaxy (dorsal view); 29 – dorsal sensilla and supraalar; 30 – clypeal/labral sensilla and ocular sensilla; 31 – abdominal segment 1 chaetotaxy; 32 – segment 4 chaetotaxy. Scale bars = 0.05 mm. Abbreviations:
<emphasis id="B92ADE0BFF873F4E81FC2EFBFC1FE27D" bold="true" box="[696,919,1519,1538]" pageId="6" pageNumber="721">AL-1-T, AL-2-T, AL-3-T</emphasis>
íanterolateral sensilla;
<emphasis id="B92ADE0BFF873F4E83D32D1EFEBCE262" bold="true" box="[151,308,1546,1565]" pageId="6" pageNumber="721">AM-1-T, AM-2-T</emphasis>
– anteromedial sensilla;
<emphasis id="B92ADE0BFF873F4E814E2D1EFD2BE262" bold="true" box="[522,675,1546,1565]" pageId="6" pageNumber="721">CL-1-H, CL-2-H</emphasis>
– clypeal/labral sensilla;
<emphasis id="B92ADE0BFF873F4E80392D1EFC7BE262" bold="true" box="[893,1011,1546,1565]" pageId="6" pageNumber="721">D-3-T, D-5-T</emphasis>
– dorsal sensilla;
<emphasis id="B92ADE0BFF873F4E83822D31FE39E247" bold="true" box="[198,433,1573,1592]" pageId="6" pageNumber="721">DL-1-H, DL-2-H, DL-3-H</emphasis>
– dorsolateral cephalic sclerite sensilla;
<emphasis id="B92ADE0BFF873F4E804E2D31FF5DE22D" bold="true" pageId="6" pageNumber="721">D-2-I, D-4-I, D-7-I, D-8-I, L-1-I, L-2-I, L-3-I –</emphasis>
abdominal segment 1 chaetotaxy;
<emphasis id="B92ADE0BFF873F4E82B32D2BFF6EE212" bold="true" pageId="6" pageNumber="721">D-2-IV, D-4-IV, D-7-IV, D-8-IV, L-1-IV, L-2-IV, L-3-IV, L-4-IV, V-5-IV, V-6-IV –</emphasis>
segment 4 chaetotaxy;
<emphasis id="B92ADE0BFF873F4E82EB2D4EFDA5E212" bold="true" box="[431,557,1626,1645]" pageId="6" pageNumber="721">O-1-H, O-2-H</emphasis>
– ocular sensilla;
<emphasis id="B92ADE0BFF873F4E818D2D4EFC83E212" bold="true" box="[713,779,1626,1645]" pageId="6" pageNumber="721">SA-2-T</emphasis>
– supraalar.
</paragraph>
</caption>
<paragraph id="8BE10219FF863F4F83D42A28FBB7E60C" blockId="7.[113,1089,220,627]" pageId="7" pageNumber="722">
The pupae of the Nearctic species
<taxonomicName id="4C5E799AFF863F4F82B22A28FDF0E52B" authorityName="Wirth" authorityYear="1952" box="[502,632,316,340]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="722" phylum="Arthropoda" rank="species" species="pollinosa">
<emphasis id="B92ADE0BFF863F4F82B22A28FDF0E52B" box="[502,632,316,340]" italics="true" pageId="7" pageNumber="722">D. pollinosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5E799AFF863F4F81ED2A28FC43E52C" authority="Thomsen, 1935" authorityName="Thomsen" authorityYear="1935" box="[681,971,316,339]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="722" phylum="Arthropoda" rank="species" species="traverae">
<emphasis id="B92ADE0BFF863F4F81ED2A28FD34E52C" box="[681,700,316,339]" italics="true" pageId="7" pageNumber="722">D</emphasis>
.
<emphasis id="B92ADE0BFF863F4F818C2A28FCAAE52C" box="[712,802,316,339]" italics="true" pageId="7" pageNumber="722">traverae</emphasis>
Thomsen, 1935
</taxonomicName>
are similar to this new species. However, both species can be distinguished from
<taxonomicName id="4C5E799AFF863F4F80152A48FBA8E50B" authorityName="Diaz &amp; Torreias &amp; Spinelli &amp; Ronderos &amp; Unlp" authorityYear="2014" box="[849,1056,348,372]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="722" phylum="Arthropoda" rank="species" species="pseudopollinosa">
<emphasis id="B92ADE0BFF863F4F80152A48FCECE50C" box="[849,868,348,371]" italics="true" pageId="7" pageNumber="722">D</emphasis>
.
<emphasis id="B92ADE0BFF863F4F80352A48FBA8E50B" box="[881,1056,348,372]" italics="true" pageId="7" pageNumber="722">pseudopollinosa</emphasis>
</taxonomicName>
by several characters offered by the adults, as follows: in
<taxonomicName id="4C5E799AFF863F4F81872A68FCC0E5EB" authorityName="Wirth" authorityYear="1952" box="[707,840,380,404]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="722" phylum="Arthropoda" rank="species" species="pollinosa">
<emphasis id="B92ADE0BFF863F4F81872A68FD5EE5EC" box="[707,726,380,403]" italics="true" pageId="7" pageNumber="722">D</emphasis>
.
<emphasis id="B92ADE0BFF863F4F81A12A68FCC0E5EB" box="[741,840,380,404]" italics="true" pageId="7" pageNumber="722">pollinosa</emphasis>
</taxonomicName>
the legs are somewhat banded, the gonocoxal apodemes are fused with the paramere, and the female subgenital plate exhibits a conspicuous anterior, triangular projection.
<taxonomicName id="4C5E799AFF863F4F81EA2AA8FCF7E5AC" authorityName="Thomsen" authorityYear="1935" box="[686,895,444,468]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="722" phylum="Arthropoda" rank="species" species="traverae">
<emphasis id="B92ADE0BFF863F4F81EA2AA8FCF7E5AC" box="[686,895,444,468]" italics="true" pageId="7" pageNumber="722">Dasyhelea traverae</emphasis>
</taxonomicName>
is a larger species (female wing length
<quantity id="4CA6AFFCFF863F4F82172AC8FE79E58C" box="[339,497,476,499]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" metricValueMax="1.27" metricValueMin="1.21" pageId="7" pageNumber="722" unit="mm" value="1.24" valueMax="1.27" valueMin="1.21">1.21–1.27 mm</quantity>
) with a dark brown scutellum, male tergite 9 rounded, posterior margin of male sternite 9 nearly straight, paramere J-shaped and not in contact with the gonocoxal apodemes, aedeagus with the basal arch narrowed mesally and with straight lateral arms, each one bearing a slender, inner, posteriorly directed process with mesal tooth, and the female subgenital plate exhibits an elongate, slender, anteriorly directed projection.
</paragraph>
</subSubSection>
</treatment>
</document>
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