treatments-xml/data/03/F3/1D/03F31D2BFFCCBC3DFF68F9AEFB79F976.xml
2024-06-21 12:22:17 +02:00

716 lines
107 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="8B834895D9676FD7E1D190F52AE714B1" ID-CLB-Dataset="21915" ID-DOI="10.11646/zootaxa.4780.1.3" ID-GBIF-Dataset="02ba62cc-666f-490d-81d6-485a751257d3" ID-ISSN="1175-5326" ID-Zenodo-Dep="3839620" ID-ZooBank="B247849D-2204-436B-A3FC-2E00F0EFF208" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590138347486" checkinUser="plazi" docAuthor="Anker, Arthur, Al-Kandari, Manal &amp; Grave, Sammy De" docDate="2020" docId="03F31D2BFFCCBC3DFF68F9AEFB79F976" docLanguage="en" docName="zootaxa.4780.1.3.pdf" docOrigin="Zootaxa 4780 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Salmoneus gracilipes , Miya 1972" docType="treatment" docVersion="13" lastPageNumber="90" masterDocId="FFCA6553FFCBBC30FFFFFFBBFFBEFFED" masterDocTitle="On two species of the alpheid shrimp genus Salmoneus Holthuis, 1955 from Kuwait one of them new to science (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="92" masterPageNumber="77" pageNumber="84" updateTime="1698840891509" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6AD76D7E93251758561F91085C215E18" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6917990F3D4E8D5647F2EC03FA62C96A">
<mods:title id="DFB333EAC980D5F38CF2A3A0761861B2">On two species of the alpheid shrimp genus Salmoneus Holthuis, 1955 from Kuwait one of them new to science (Malacostraca: Decapoda: Caridea)</mods:title>
</mods:titleInfo>
<mods:name id="F6B8B33049B68A4D8EEF9209C2FA8A28" type="personal">
<mods:role id="7158D28FEEBA7CABD2280BBC84CFDE52">
<mods:roleTerm id="2D9E81D9829816D5FD33A09EFE50E816">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5CE5DDD708A00525F691CDF92B629566">Anker, Arthur</mods:namePart>
<mods:affiliation id="E1D0026B05CEC13347720DA89D181B85">Universidade Federal de Goiás, Campus Samambaia, Instituto de Ciências Biológicas-ICB- 5. Av. Esperança, s / n., 74690 - 900, Goiânia, Goiás, Brazil. arthuranker 7 @ gmail. com; http: // orcid. org / 0000 - 0002 - 5350 - 4267</mods:affiliation>
</mods:name>
<mods:name id="4FD98233B5C45718B63C50BB434B020B" type="personal">
<mods:role id="BFB4D63D199450B3F6D39E454CAF6422">
<mods:roleTerm id="0B1D26825F0D0A6A61B84129ABA612A9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09D162379859D54BDE82AA179B4F2632">Al-Kandari, Manal</mods:namePart>
<mods:affiliation id="18C0BD241D0932979140ED0BB588BC0F">Ecosystem-Based Management of Marine Resources, Environment and Life Sciences Research Center, Kuwait Institute of Scientific Research, Hamad Al-Mubarak Street, Building 900004, Area 1, Raas Salmiya, Kuwait. http: // orcid. org / 0000 - 0003 - 0073 - 7929 Oxford University Museum of Natural History, Parks Road, Oxford, OX 1 3 PW, United Kingdom.</mods:affiliation>
</mods:name>
<mods:name id="8BC54D9736E7250565C28BF9C7D40DD4" type="personal">
<mods:role id="A967B4A7EE290C5B46E6128C9A72CF15">
<mods:roleTerm id="777070CEE1DFC025AB63E10E4A0EC13E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EE6FB91F1ABDA69B24A666E4A9CEB9C2">Grave, Sammy De</mods:namePart>
</mods:name>
<mods:typeOfResource id="2152858FAF865519C7BEF73F60B3EB0E">text</mods:typeOfResource>
<mods:relatedItem id="85C9B722E3D2DF999B7954E11A871CF5" type="host">
<mods:titleInfo id="D11B28F422E85D2DB2180C1831D9A58B">
<mods:title id="CC86D163D9DE4F643853B456ADB35F91">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1B7600D7BD4E8B028E7E3D044B128FFD">
<mods:date id="2D1439F6783EF89674197B6F8F51A833">2020</mods:date>
<mods:detail id="79FF153C1B0C03C4691549F98E0916FD" type="pubDate">
<mods:number id="BE352035E28ED84EAB8DAAE72217E662">2020-05-22</mods:number>
</mods:detail>
<mods:detail id="1675AEF0189E9C30A3DB8B7BD6C8B8CE" type="volume">
<mods:number id="C7A8244CE0D1AE6E2426F625A360D44D">4780</mods:number>
</mods:detail>
<mods:detail id="FE184C44BD17924E9BE2BD0DE8804EB5" type="issue">
<mods:number id="01810163776C972542E5E838D41C1A06">1</mods:number>
</mods:detail>
<mods:extent id="32437CEBE6EF7CFF4AF97D016DF5A45E" unit="page">
<mods:start id="A3405F74B7FB4EB45A1B86BD13E6FB46">77</mods:start>
<mods:end id="4B35B5FFF81A6A43EACBA2AD1A6BE63B">92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9790FDC0A7F0387AB31FB3A49B2038B0">journal article</mods:classification>
<mods:identifier id="A2106F29FE756549DE74CC02383CAFEE" type="CLB-Dataset">21915</mods:identifier>
<mods:identifier id="69C0A6E95A118247B8FECEDDC12E0D9A" type="DOI">10.11646/zootaxa.4780.1.3</mods:identifier>
<mods:identifier id="370819F0158BDD426786C38D31F2C3B4" type="GBIF-Dataset">02ba62cc-666f-490d-81d6-485a751257d3</mods:identifier>
<mods:identifier id="C1F19FAF97C2FCEE691B58E57BBB735F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C8CFC422FEAEA5F9EB879FD4E2FEB3D3" type="Zenodo-Dep">3839620</mods:identifier>
<mods:identifier id="84F78154B72783226EC6973B8418132C" type="ZooBank">B247849D-2204-436B-A3FC-2E00F0EFF208</mods:identifier>
</mods:mods>
<treatment id="03F31D2BFFCCBC3DFF68F9AEFB79F976" ID-DOI="http://doi.org/10.5281/zenodo.3853021" ID-GBIF-Taxon="164248826" ID-Zenodo-Dep="3853021" LSID="urn:lsid:plazi:treatment:03F31D2BFFCCBC3DFF68F9AEFB79F976" httpUri="http://treatment.plazi.org/id/03F31D2BFFCCBC3DFF68F9AEFB79F976" lastPageId="13" lastPageNumber="90" pageId="7" pageNumber="84">
<subSubSection id="C340FFB6FFCCBC37FF68F9AEFD92F9C2" box="[151,556,1557,1584]" pageId="7" pageNumber="84" type="nomenclature">
<paragraph id="8BE5AC3DFFCCBC37FF68F9AEFD92F9C2" blockId="7.[151,556,1557,1619]" box="[151,556,1557,1584]" pageId="7" pageNumber="84">
<heading id="D0AD1B51FFCCBC37FF68F9AEFD92F9C2" bold="true" box="[151,556,1557,1584]" fontSize="11" level="1" pageId="7" pageNumber="84" reason="1">
<taxonomicName id="4C5AD7BEFFCCBC37FF68F9AEFD92F9C2" ID-CoL="854PP" authority="Miya, 1972" authorityName=", Miya" authorityYear="1972" box="[151,556,1557,1584]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFCCBC37FF68F9AEFD92F9C2" bold="true" box="[151,556,1557,1584]" pageId="7" pageNumber="84">
<emphasis id="B92E702FFFCCBC37FF68F9AEFE27F9DD" bold="true" box="[151,409,1557,1584]" italics="true" pageId="7" pageNumber="84">Salmoneus gracilipes</emphasis>
<bibRefCitation id="EFCBD1CCFFCCBC37FE5EF9AEFD92F9C2" author="Miya, Y." box="[417,556,1557,1583]" pageId="7" pageNumber="84" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya, 1972</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340FFB6FFCCBC37FF68F982FEBEF9BE" box="[151,256,1592,1619]" pageId="7" pageNumber="84" type="description">
<paragraph id="8BE5AC3DFFCCBC37FF68F982FEBEF9BE" blockId="7.[151,556,1557,1619]" box="[151,256,1592,1619]" pageId="7" pageNumber="84">
<figureCitation id="1361B0B8FFCCBC37FF68F982FEBEF9BE" box="[151,256,1592,1619]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="8.[151,250,1940,1965]" captionStartId-1="9.[151,250,1907,1932]" captionStartId-2="10.[151,250,1768,1793]" captionStartId-3="11.[151,250,880,905]" captionTargetBox-0="[166,1400,530,1908]" captionTargetBox-1="[151,1436,181,1882]" captionTargetBox-2="[151,1436,181,1742]" captionTargetBox-3="[151,1436,181,855]" captionTargetId-0="figure@8.[161,1406,525,1915]" captionTargetId-1="figure@9.[151,1436,181,1882]" captionTargetId-2="figure@10.[151,1436,181,1742]" captionTargetId-3="figure@11.[151,1436,181,855]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="FIGURE 5. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.5 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.048):A, frontal region, dorsal; B, anterior region of carapace and left eye, lateral; C, telson, dorsal; D, right (major) cheliped, lateral; E, same, chela, lateral; F, left (minor) cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral." captionText-1="FIGURE 6. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.9 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.047), living individual (A, B); ovigerous specimen (cl 4.5 mm) from the same locality (OUMNH.ZC. 2020.01.048), preserved individual (C): A, dorsal; B, lateral; C, frontal region with focus on left eye, lateral.All photographs by the first author (AA), photograph in (C) taken through a Leica stereomicroscope." captionText-2="FIGURE 7. Salmoneus gracilipes Miya, 1972, non-ovigerous specimen (cl 3.7 mm) from Failaka Island, Kuwait (OUMNH. ZC. 2020.01.046), living individual (A, B) and its presumed facultative host, the echiuran Listriolobus cf. brevirostris Chen &amp; Yeh 1958 (C, not to scale, size of the specimen approximately 60 mm with the proboscis fully extended): A, dorsal; B, lateral; C, host, lateral. Photographs by the first author (AA)." captionText-3="FIGURE 8. Salmoneus gracilipes Miya, 1972, 1 non-ovigerous specimen (cl 4.9 mm) from Tateyama, Honshu, Japan (CBM- ZC 14185), individual post-mortem, lateral. Photograph courtesy of Dr. Tomoyuki Komai." figureDoi-0="http://doi.org/10.5281/zenodo.3841684" figureDoi-1="http://doi.org/10.5281/zenodo.3841686" figureDoi-2="http://doi.org/10.5281/zenodo.3841688" figureDoi-3="http://doi.org/10.5281/zenodo.3841690" httpUri-0="https://zenodo.org/record/3841684/files/figure.png" httpUri-1="https://zenodo.org/record/3841686/files/figure.png" httpUri-2="https://zenodo.org/record/3841688/files/figure.png" httpUri-3="https://zenodo.org/record/3841690/files/figure.png" pageId="7" pageNumber="84">Figs. 58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C340FFB6FFCCBC37FF68F9C6FB10F93E" pageId="7" pageNumber="84" type="reference_group">
<paragraph id="8BE5AC3DFFCCBC37FF68F9C6FB10F93E" blockId="7.[151,1437,1661,1747]" pageId="7" pageNumber="84">
<treatmentCitationGroup id="AB4A8B13FFCCBC37FF68F9C6FB10F93E" pageId="7" pageNumber="84">
<treatmentCitation id="0AFB8A2CFFCCBC37FF68F9C6FDB3F978" author="Miya, Y." box="[151,525,1661,1686]" page="38" pageId="7" pageNumber="84" year="1972">
<taxonomicName id="4C5AD7BEFFCCBC37FF68F9C6FDB3F978" ID-CoL="854PP" authority="Miya 1972: 38" authorityName=", Miya" authorityPageNumber="38" authorityYear="1972" box="[151,525,1661,1686]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFCCBC37FF68F9C6FECEF978" box="[151,368,1661,1685]" italics="true" pageId="7" pageNumber="84">Salmoneus gracilipes</emphasis>
<bibRefCitation id="EFCBD1CCFFCCBC37FE89F9C6FDB3F978" author="Miya, Y." box="[374,525,1661,1686]" pageId="7" pageNumber="84" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972: 38</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 3;
<treatmentCitation id="0AFB8A2CFFCCBC37FDADF9C6FD56F97B" author="Miya, Y." box="[594,744,1661,1686]" page="86" pageId="7" pageNumber="84" year="1984">
<bibRefCitation id="EFCBD1CCFFCCBC37FDADF9C6FD56F97B" author="Miya, Y." box="[594,744,1661,1686]" pageId="7" pageNumber="84" pagination="67 - 100" refId="ref9147" refString="Miya, Y. (1984) Alpheid shrimps from the Truks, Ponape and Majuro Atolls (Crustacea Decapoda). Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 27, 67 - 100." type="journal article" year="1984">Miya 1984: 86</bibRefCitation>
</treatmentCitation>
, figs. 4ad;
<treatmentCitation id="0AFB8A2CFFCCBC37FC9CF9C6FBB6F978" author="Miya, Y." box="[867,1032,1661,1686]" page="287" pageId="7" pageNumber="84" year="1995">
<bibRefCitation id="EFCBD1CCFFCCBC37FC9CF9C6FBB6F978" author="Miya, Y." box="[867,1032,1661,1686]" pageId="7" pageNumber="84" pagination="271 - 288" refId="ref9186" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">Miya 1995: 287</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFB8A2CFFCCBC37FBEDF9C6FB6DF97B" author="Hayashi, K. - I." box="[1042,1235,1661,1686]" page="522" pageId="7" pageNumber="84" year="1995">
<bibRefCitation id="EFCBD1CCFFCCBC37FBEDF9C6FB6DF97B" author="Hayashi, K. - I." box="[1042,1235,1661,1686]" pageId="7" pageNumber="84" pagination="520 - 524" refId="ref8593" refString="Hayashi, K. - I. (1995) Prawns, shrimps and lobsters from Japan (85). Family Alpheidae - genera Automate and Salmoneus. Aquabiology, 101 (17), 520 - 524." type="journal article" year="1995">Hayashi 1995: 522</bibRefCitation>
</treatmentCitation>
, figs. 287b, 288cf;
<bibRefCitation id="EFCBD1CCFFCCBC37FF38F920FEFDF959" author="Anker, A." box="[199,323,1691,1716]" pageId="7" pageNumber="84" refId="ref7566" refString="Anker, A. (2001) Taxonomie et evolution des Alpheidae (Crustacea, Decapoda). Vols. 1 &amp; 2. PhD Thesis, Museum National d'Histoire Naturelle, Paris, 547 + 331 pp." type="book" year="2001">Anker 2001</bibRefCitation>
: (vol. 2) 272;
<bibRefCitation id="EFCBD1CCFFCCBC37FE2BF927FD35F959" author="Koo, H. &amp; Kim, W." box="[468,651,1691,1716]" pageId="7" pageNumber="84" pagination="43 - 48" refId="ref8939" refString="Koo, H. &amp; Kim, W. (2003) First report of the alpheid shrimp Salmoneus gracilipes (Decapoda: Caridea: Alpheidae) from Korea. The Korean Journal of Systematic Zoology, 19, 43 - 48." type="journal article" year="2003">Koo &amp; Kim 2003</bibRefCitation>
: fig. 1;
<treatmentCitation id="0AFB8A2CFFCCBC37FD28F920FC3DF959" author="Anker, A." box="[727,899,1691,1716]" page="114" pageId="7" pageNumber="84" year="2003">
<bibRefCitation id="EFCBD1CCFFCCBC37FD28F920FC3DF959" author="Anker, A." box="[727,899,1691,1716]" pageId="7" pageNumber="84" pagination="101 - 109" refId="ref7605" refString="Anker, A. (2003) New records of Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Alpheidae) from northern Australia, with description of one new species and remarks on S. serratidigitus (Coutiere, 1896). Beagle-Records of the Museum and Art Gallery of the Northern Territory, 19, 101 - 109." type="journal article" year="2003">Anker 2003: 114</bibRefCitation>
</treatmentCitation>
(key);
<treatmentCitation id="0AFB8A2CFFCCBC37FC32F920FB47F959" ID-DOI="http://doi.org/10.5281/zenodo.5255527" ID-Zenodo-Dep="5255527" author="Naderloo, R. &amp; Turkay, M." box="[973,1273,1691,1716]" httpUri="http://treatment.plazi.org/id/E177585FC60A1B3FFF3CFFB1FE1AFE13" page="13" pageId="7" pageNumber="84" year="2012">
<bibRefCitation id="EFCBD1CCFFCCBC37FC32F920FB47F959" author="Naderloo, R. &amp; Turkay, M." box="[973,1273,1691,1716]" pageId="7" pageNumber="84" pagination="1 - 67" refId="ref8982" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012: 13</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFB8A2CFFCCBC37FAFBF920FE8EF93E" author="Naderloo, R. &amp; Turkay, M. &amp; Sari, A." page="448" pageId="7" pageNumber="84" year="2013">
<bibRefCitation id="EFCBD1CCFFCCBC37FAFBF920FE8EF93E" author="Naderloo, R. &amp; Turkay, M. &amp; Sari, A." pageId="7" pageNumber="84" pagination="445 - 462" refId="ref9038" refString="Naderloo, R., Turkay, M. &amp; Sari, A. (2013) Intertidal habitats and decapod (Crustacea) diversity of Qeshm Island, a biodiversity hotspot within the Persian Gulf. Marine Biodiversity, 43, 445 - 462. https: // doi. org / 10.1007 / s 12526 - 013 - 0174 - 3" type="journal article" year="2013">
Naderloo
<emphasis id="B92E702FFFCCBC37FA96F926FA23F959" box="[1385,1437,1692,1716]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
2013: 448
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFB8A2CFFCCBC37FEC6F901FD9EF93E" ID-DOI="http://doi.org/10.5281/zenodo.5073383" ID-Zenodo-Dep="5073383" author="Anker, A. &amp; Pratama, I. S. &amp; Firdaus, M. &amp; Rahayu, D. L." box="[313,544,1722,1747]" httpUri="http://treatment.plazi.org/id/AD437D0CF16DAD6694CE1061F4381F59" page="326" pageId="7" pageNumber="84" year="2015">
<bibRefCitation id="EFCBD1CCFFCCBC37FEC6F901FD9EF93E" author="Anker, A. &amp; Pratama, I. S. &amp; Firdaus, M. &amp; Rahayu, D. L." box="[313,544,1722,1747]" pageId="7" pageNumber="84" pagination="301 - 342" refId="ref8017" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans (Alpheidae, Laomediidae, Strahlaxiidae) from Lombok, Indonesia. Zootaxa, 3911 (3), 301 - 342. https: // doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker
<emphasis id="B92E702FFFCCBC37FE80F900FE0CF93E" box="[383,434,1723,1747]" italics="true" pageId="7" pageNumber="84">et al.</emphasis>
2015: 326
</bibRefCitation>
</treatmentCitation>
, fig. 19;
<treatmentCitation id="0AFB8A2CFFCCBC37FD85F901FCCDF93E" author="Duris, Z. &amp; Horka, I." box="[634,883,1722,1747]" page="12" pageId="7" pageNumber="84" year="2016">
<bibRefCitation id="EFCBD1CCFFCCBC37FD85F901FCCDF93E" author="Duris, Z. &amp; Horka, I." box="[634,883,1722,1747]" pageId="7" pageNumber="84" pagination="773 - 793" refId="ref8441" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016: 12</bibRefCitation>
</treatmentCitation>
, fig. 11e [see Remarks below].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C340FFB6FFCCBC38FF68F8BBFC05FF3A" lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84" type="materials_examined">
<paragraph id="8BE5AC3DFFCCBC38FF68F8BBFC05FF3A" blockId="7.[151,1436,1792,2035]" lastBlockId="8.[151,1436,152,503]" lastPageId="8" lastPageNumber="85" pageId="7" pageNumber="84">
<emphasis id="B92E702FFFCCBC37FF68F8BBFEC5F8F6" bold="true" box="[151,379,1792,1819]" pageId="7" pageNumber="84">Material examined</emphasis>
.
<materialsCitation id="3B32A660FFCCBC37FE75F8BAFDD9F81E" ID-GBIF-Occurrence="2622605303" collectingDate="2019-09-26" collectingDateMax="2019-10-03" collectingDateMin="2019-09-26" collectionCode="OUMNH, ZC" collectorName="A. Anker &amp; S. De Grave" country="Kuwait" elevation="3" latitude="29.068419" location="Kubbar Island" longLatPrecision="1" longitude="48.48941" municipality="Failaka Island" pageId="7" pageNumber="84" specimenCount="1">
<collectingCountry id="F34DECADFFCCBC37FE75F8BAFE62F8F6" box="[394,476,1793,1819]" name="Kuwait" pageId="7" pageNumber="84">Kuwait</collectingCountry>
: 1 non-ovigerous specimen (cl
<quantity id="4CA201D8FFCCBC37FCB9F8BAFC1CF8F6" box="[838,930,1793,1819]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="84" unit="mm" value="3.7">3.7 mm</quantity>
),
<collectionCode id="ED4B34F8FFCCBC37FC49F8BAFB98F8F6" box="[950,1062,1793,1819]" pageId="7" pageNumber="84">OUMNH</collectionCode>
.
<collectionCode id="ED4B34F8FFCCBC37FBD6F8BAFBF0F8F6" box="[1065,1102,1793,1819]" country="Austria" name="Zoological Collection, University of Vienna" pageId="7" pageNumber="84" type="University or college">ZC</collectionCode>
. 2020.01.046,
<collectingMunicipality id="6B813647FFCCBC37FB09F8BAFA26F8F6" box="[1270,1432,1793,1819]" pageId="7" pageNumber="84">Failaka Island</collectingMunicipality>
, eastern side,
<geoCoordinate id="EE6ECAFAFFCCBC37FED5F89EFE02F8D2" box="[298,444,1829,1855]" degrees="29" direction="north" minutes="23.409" orientation="latitude" pageId="7" pageNumber="84" precision="1" value="29.39015">29°23.409N</geoCoordinate>
<geoCoordinate id="EE6ECAFAFFCCBC37FE3BF89EFDE3F8D2" box="[452,605,1829,1855]" degrees="48" direction="east" minutes="23.927" orientation="longitude" pageId="7" pageNumber="84" precision="1" value="48.398785">48° 23.927E</geoCoordinate>
, mixed rocky-muddy intertidal, muddy patches between rocks, depth less than
<quantity id="4CA201D8FFCCBC37FF32F8F2FEB0F88E" box="[205,270,1865,1891]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="84" unit="m" value="0.3">
<elevation id="00774B0EFFCCBC37FF32F8F2FEB0F88E" box="[205,270,1865,1891]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="84" unit="m" value="0.3">0.3 m</elevation>
</quantity>
, suction (yabby) pump, in burrow of echiuran,
<taxonomicName id="4C5AD7BEFFCCBC37FCEAF8F2FBF8F88E" baseAuthorityName="Edmondson" baseAuthorityYear="1930" box="[789,1094,1865,1891]" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="7" pageNumber="84" phylum="Echiura" rank="species" species="brevirostris">
<emphasis id="B92E702FFFCCBC37FCEAF8F2FC22F88E" box="[789,924,1865,1891]" italics="true" pageId="7" pageNumber="84">Listriolobus</emphasis>
cf.
<emphasis id="B92E702FFFCCBC37FC3AF8F2FBF8F88E" box="[965,1094,1865,1891]" italics="true" pageId="7" pageNumber="84">brevirostris</emphasis>
</taxonomicName>
, leg.
<collectorName id="26AFC9EBFFCCBC37FB80F8F2FB59F88E" box="[1151,1255,1865,1891]" pageId="7" pageNumber="84">A. Anker</collectorName>
,
<date id="FFE48AFDFFCCBC37FB0FF8F2FA22F88E" box="[1264,1436,1865,1891]" pageId="7" pageNumber="84" value="2019-10-03">
<collectingDate id="EFA07315FFCCBC37FB0FF8F2FA22F88E" box="[1264,1436,1865,1891]" pageId="7" pageNumber="84" value="2019-10-03">3 October 2019</collectingDate>
</date>
[fcn KUW-070]; 1 ovigerous specimen (cl
<quantity id="4CA201D8FFCCBC37FD79F8D6FD5DF86A" box="[646,739,1901,1927]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="84" unit="mm" value="4.9">4.9 mm</quantity>
),
<collectionCode id="ED4B34F8FFCCBC37FD07F8D6FCD6F86A" box="[760,872,1901,1927]" pageId="7" pageNumber="84">OUMNH</collectionCode>
.
<collectionCode id="ED4B34F8FFCCBC37FC94F8D6FC2EF86A" box="[875,912,1901,1927]" country="Austria" name="Zoological Collection, University of Vienna" pageId="7" pageNumber="84" type="University or college">ZC</collectionCode>
. 2020.01.047,
<location id="8E85FAE6FFCCBC37FBC5F8D6FB60F86A" LSID="urn:lsid:plazi:treatment:03F31D2BFFCCBC3DFF68F9AEFB79F976:8E85FAE6FFCCBC37FBC5F8D6FB60F86A" box="[1082,1246,1901,1927]" country="Kuwait" latitude="29.068419" longLatPrecision="1" longitude="48.48941" municipality="Failaka Island" name="Kubbar Island" pageId="7" pageNumber="84">Kubbar Island</location>
,
<geoCoordinate id="EE6ECAFAFFCCBC37FB13F8D6FA22F86A" box="[1260,1436,1901,1927]" degrees="29" direction="north" minutes="04" orientation="latitude" pageId="7" pageNumber="84" precision="1" seconds="06.30" value="29.068419">29°0406.30N</geoCoordinate>
<geoCoordinate id="EE6ECAFAFFCCBC37FF68F82AFEF9F846" box="[151,327,1937,1963]" degrees="48" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="84" precision="1" seconds="21.88" value="48.48941">48°2921.88E</geoCoordinate>
, depth
<quantity id="4CA201D8FFCCBC37FE64F82AFE77F846" box="[411,457,1937,1963]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="84" unit="m" value="7.0">
<elevation id="00774B0EFFCCBC37FE64F82AFE77F846" box="[411,457,1937,1963]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="84" unit="m" value="7.0">7 m</elevation>
</quantity>
, sand slope with dead coral “bommies”, under rocks and rubble, leg.
<collectorName id="26AFC9EBFFCCBC37FB1BF82AFACDF846" box="[1252,1395,1937,1963]" pageId="7" pageNumber="84">S. De Grave</collectorName>
,
<date id="FFE48AFDFFCCBC37FA7FF82AFEF1F822" pageId="7" pageNumber="84" value="2019-09-26">
<collectingDate id="EFA07315FFCCBC37FA7FF82AFEF1F822" pageId="7" pageNumber="84" value="2019-09-26">26 September 2019</collectingDate>
</date>
[fcn KUW-015]; 1 ovigerous specimen (cl
<quantity id="4CA201D8FFCCBC37FCC1F80FFC27F822" box="[830,921,1972,1999]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="84" unit="mm" value="4.5">4.5 mm</quantity>
),
<collectionCode id="ED4B34F8FFCCBC37FC52F80EFBA3F822" box="[941,1053,1973,1999]" pageId="7" pageNumber="84">OUMNH</collectionCode>
.
<collectionCode id="ED4B34F8FFCCBC37FBDFF80EFBFBF822" box="[1056,1093,1973,1999]" country="Austria" name="Zoological Collection, University of Vienna" pageId="7" pageNumber="84" type="University or college">ZC</collectionCode>
. 2020.01.048, same collection as for previous specimen [fcn KUW-016]
</materialsCitation>
.
<materialsCitation id="3B32A660FFCCBC38FD8EF862FC09FF3A" ID-GBIF-Occurrence="2622605301" collectingDate="2015-04-07" collectionCode="T" collectorName="Komai &amp; Dr. Tomoyuki Komai" country="Japan" county="Honshu" lastPageId="8" lastPageNumber="85" latitude="34.988" location="Okinoshima" longLatPrecision="11" longitude="139.8225" municipality="Tateyama" pageId="7" pageNumber="84" specimenCount="1" stateProvince="Chiba">
<collectingCountry id="F34DECADFFCCBC37FD8EF862FD0FF81E" box="[625,689,2009,2035]" name="Japan" pageId="7" pageNumber="84">Japan</collectingCountry>
: 1 non-ovigerous specimen (cl
<quantity id="4CA201D8FFCCBC37FBF1F862FBD9F81E" box="[1038,1127,2009,2035]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="84" unit="mm" value="4.9">4.9 mm</quantity>
), CBM-ZC 14185,
<collectingCounty id="6284D4B1FFCCBC37FAC1F862FA26F81E" box="[1342,1432,2009,2035]" pageId="7" pageNumber="84">Honshu</collectingCounty>
,
<collectingRegion id="499E62DFFFC3BC38FF68FF22FEE6FF5E" box="[151,344,153,179]" country="Japan" name="Chiba" pageId="8" pageNumber="85">Chiba Prefecture</collectingRegion>
,
<collectingMunicipality id="6B813647FFC3BC38FE9BFF22FE6DFF5E" box="[356,467,153,179]" pageId="8" pageNumber="85">Tateyama</collectingMunicipality>
,
<location id="8E85FAE6FFC3BC38FE1FFF22FDD2FF5E" LSID="urn:lsid:plazi:treatment:03F31D2BFFCCBC3DFF68F9AEFB79F976:8E85FAE6FFC3BC38FE1FFF22FDD2FF5E" box="[480,620,153,179]" country="Japan" county="Honshu" latitude="34.988" longLatPrecision="11" longitude="139.8225" municipality="Tateyama" name="Okinoshima" pageId="8" pageNumber="85" stateProvince="Chiba">Okinoshima</location>
,
<geoCoordinate id="EE6ECAFAFFC3BC38FD87FF23FD42FF5E" box="[632,764,152,179]" degrees="34" direction="north" minutes="59.28" orientation="latitude" pageId="8" pageNumber="85" precision="9" value="34.988">34°59.28N</geoCoordinate>
<geoCoordinate id="EE6ECAFAFFC3BC38FCFAFF23FC28FF5E" box="[773,918,152,179]" degrees="139" direction="east" minutes="49.35" orientation="longitude" pageId="8" pageNumber="85" precision="9" value="139.8225">139°49.35E</geoCoordinate>
, intertidal, sand, under rock, leg.
<collectionCode id="ED4B34F8FFC3BC38FAEBFF22FA98FF5E" box="[1300,1318,153,179]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="85">T</collectionCode>
.
<collectorName id="26AFC9EBFFC3BC38FACCFF22FA3CFF5E" box="[1331,1410,153,179]" pageId="8" pageNumber="85">Komai</collectorName>
,
<date id="FFE48AFDFFC3BC38FA71FF22FEAFFF3A" pageId="8" pageNumber="85" value="2015-04-07">
<collectingDate id="EFA07315FFC3BC38FA71FF22FEAFFF3A" pageId="8" pageNumber="85" value="2015-04-07">7 April 2015</collectingDate>
</date>
[material examined at our request by
<collectorName id="26AFC9EBFFC3BC38FD44FF06FC0FFF3A" box="[699,945,188,215]" pageId="8" pageNumber="85">Dr. Tomoyuki Komai</collectorName>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C340FFB6FFC3BC38FF38FF5BFDB3FE1A" pageId="8" pageNumber="85" type="description">
<paragraph id="8BE5AC3DFFC3BC38FF38FF5BFC59FEF2" blockId="8.[151,1436,152,503]" pageId="8" pageNumber="85">
<emphasis id="B92E702FFFC3BC38FF38FF5BFEEFFF16" bold="true" box="[199,337,224,251]" pageId="8" pageNumber="85">Description</emphasis>
. See
<bibRefCitation id="EFCBD1CCFFC3BC38FE6EFF5AFD9FFF16" author="Miya, Y." box="[401,545,225,251]" pageId="8" pageNumber="85" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya (1972)</bibRefCitation>
,
<bibRefCitation id="EFCBD1CCFFC3BC38FDD1FF5AFD6BFF16" author="Hayashi, K. - I." box="[558,725,224,251]" pageId="8" pageNumber="85" pagination="520 - 524" refId="ref8593" refString="Hayashi, K. - I. (1995) Prawns, shrimps and lobsters from Japan (85). Family Alpheidae - genera Automate and Salmoneus. Aquabiology, 101 (17), 520 - 524." type="journal article" year="1995">Hayashi (1995</bibRefCitation>
, in Japanese) and
<bibRefCitation id="EFCBD1CCFFC3BC38FC5AFF5AFB3CFF16" author="Koo, H. &amp; Kim, W." box="[933,1154,225,251]" pageId="8" pageNumber="85" pagination="43 - 48" refId="ref8939" refString="Koo, H. &amp; Kim, W. (2003) First report of the alpheid shrimp Salmoneus gracilipes (Decapoda: Caridea: Alpheidae) from Korea. The Korean Journal of Systematic Zoology, 19, 43 - 48." type="journal article" year="2003">Koo &amp; Kim (2003)</bibRefCitation>
; taxonomically most important morphologically features of the Kuwaiti specimens shown in
<figureCitation id="1361B0B8FFC3BC38FC60FEBEFC5CFEF2" box="[927,994,260,287]" captionStart="FIGURE 5" captionStartId="8.[151,250,1940,1965]" captionTargetBox="[166,1400,530,1908]" captionTargetId="figure@8.[161,1406,525,1915]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.5 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.048):A, frontal region, dorsal; B, anterior region of carapace and left eye, lateral; C, telson, dorsal; D, right (major) cheliped, lateral; E, same, chela, lateral; F, left (minor) cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral." figureDoi="http://doi.org/10.5281/zenodo.3841684" httpUri="https://zenodo.org/record/3841684/files/figure.png" pageId="8" pageNumber="85">Fig. 5</figureCitation>
.
</paragraph>
<paragraph id="8BE5AC3DFFC3BC38FF38FE93FDB3FE1A" blockId="8.[151,1436,152,503]" pageId="8" pageNumber="85">
<emphasis id="B92E702FFFC3BC38FF38FE93FEE6FEAE" bold="true" box="[199,344,296,323]" pageId="8" pageNumber="85">Colouration</emphasis>
. Originally described as entirely transparent, with dull reddish orange cornea and yellowish orange eggs (
<bibRefCitation id="EFCBD1CCFFC3BC38FEE6FEF6FE24FE8A" author="Miya, Y." box="[281,410,333,359]" pageId="8" pageNumber="85" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
), generally corresponding well to the colour of the Kuwaiti specimens, especially the two ovigerous specimens from Kubbar Island (
<figureCitation id="1361B0B8FFC3BC38FD8CFECAFD53FE66" box="[627,749,369,395]" captionStart="FIGURE 6" captionStartId="9.[151,250,1907,1932]" captionTargetBox="[151,1436,181,1882]" captionTargetId="figure@9.[151,1436,181,1882]" captionTargetPageId="9" captionText="FIGURE 6. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.9 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.047), living individual (A, B); ovigerous specimen (cl 4.5 mm) from the same locality (OUMNH.ZC. 2020.01.048), preserved individual (C): A, dorsal; B, lateral; C, frontal region with focus on left eye, lateral.All photographs by the first author (AA), photograph in (C) taken through a Leica stereomicroscope." figureDoi="http://doi.org/10.5281/zenodo.3841686" httpUri="https://zenodo.org/record/3841686/files/figure.png" pageId="8" pageNumber="85">Fig. 6A, B</figureCitation>
), as well as the material from Tateyama,
<collectingCountry id="F34DECADFFC3BC38FB48FECAFB49FE66" box="[1207,1271,369,395]" name="Japan" pageId="8" pageNumber="85">Japan</collectingCountry>
(
<figureCitation id="1361B0B8FFC3BC38FAF9FECAFAF7FE66" box="[1286,1353,369,395]" captionStart="FIGURE 8" captionStartId="11.[151,250,880,905]" captionTargetBox="[151,1436,181,855]" captionTargetId="figure@11.[151,1436,181,855]" captionTargetPageId="11" captionText="FIGURE 8. Salmoneus gracilipes Miya, 1972, 1 non-ovigerous specimen (cl 4.9 mm) from Tateyama, Honshu, Japan (CBM- ZC 14185), individual post-mortem, lateral. Photograph courtesy of Dr. Tomoyuki Komai." figureDoi="http://doi.org/10.5281/zenodo.3841690" httpUri="https://zenodo.org/record/3841690/files/figure.png" pageId="8" pageNumber="85">Fig. 8</figureCitation>
),
<collectingCountry id="F34DECADFFC3BC38FAA0FECAFF6FFE42" name="Indonesia" pageId="8" pageNumber="85">Indonesia</collectingCountry>
(
<bibRefCitation id="EFCBD1CCFFC3BC38FF21FE2EFE18FE42" author="Anker, A. &amp; Pratama, I. S. &amp; Firdaus, M. &amp; Rahayu, D. L." box="[222,422,404,431]" pageId="8" pageNumber="85" pagination="301 - 342" refId="ref8017" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans (Alpheidae, Laomediidae, Strahlaxiidae) from Lombok, Indonesia. Zootaxa, 3911 (3), 301 - 342. https: // doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker
<emphasis id="B92E702FFFC3BC38FED2FE2DFEDAFE42" box="[301,356,405,431]" italics="true" pageId="8" pageNumber="85">et al.</emphasis>
2015
</bibRefCitation>
: fig. 19) and
<collectingCountry id="F34DECADFFC3BC38FDC8FE2EFCB3FE42" box="[567,781,405,431]" name="Papua New Guinea" pageId="8" pageNumber="85">Papua New Guinea</collectingCountry>
(
<bibRefCitation id="EFCBD1CCFFC3BC38FCE4FE2FFBB8FE42" author="Duris, Z. &amp; Horka, I." box="[795,1030,404,431]" pageId="8" pageNumber="85" pagination="773 - 793" refId="ref8441" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016</bibRefCitation>
: fig. 11e). In the Kuwaiti specimens, the background colour varies from white to pale yellowish and eggs in ovigerous specimens are bright orange rather than yellow (
<figureCitation id="1361B0B8FFC3BC38FED5FE66FE10FE1A" box="[298,430,477,503]" captionStart="FIGURE 6" captionStartId="9.[151,250,1907,1932]" captionTargetBox="[151,1436,181,1882]" captionTargetId="figure@9.[151,1436,181,1882]" captionTargetPageId="9" captionText="FIGURE 6. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.9 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.047), living individual (A, B); ovigerous specimen (cl 4.5 mm) from the same locality (OUMNH.ZC. 2020.01.048), preserved individual (C): A, dorsal; B, lateral; C, frontal region with focus on left eye, lateral.All photographs by the first author (AA), photograph in (C) taken through a Leica stereomicroscope." figureDoi="http://doi.org/10.5281/zenodo.3841686" httpUri="https://zenodo.org/record/3841686/files/figure.png" pageId="8" pageNumber="85">Figs. 6A, B</figureCitation>
,
<figureCitation id="1361B0B8FFC3BC38FE45FE66FE41FE1A" box="[442,511,477,503]" captionStart="FIGURE 7" captionStartId="10.[151,250,1768,1793]" captionTargetBox="[151,1436,181,1742]" captionTargetId="figure@10.[151,1436,181,1742]" captionTargetPageId="10" captionText="FIGURE 7. Salmoneus gracilipes Miya, 1972, non-ovigerous specimen (cl 3.7 mm) from Failaka Island, Kuwait (OUMNH. ZC. 2020.01.046), living individual (A, B) and its presumed facultative host, the echiuran Listriolobus cf. brevirostris Chen &amp; Yeh 1958 (C, not to scale, size of the specimen approximately 60 mm with the proboscis fully extended): A, dorsal; B, lateral; C, host, lateral. Photographs by the first author (AA)." figureDoi="http://doi.org/10.5281/zenodo.3841688" httpUri="https://zenodo.org/record/3841688/files/figure.png" pageId="8" pageNumber="85">7A, B</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF25FCB5FFC3BC38FF68F82FFA35F818" ID-DOI="http://doi.org/10.5281/zenodo.3841684" ID-Zenodo-Dep="3841684" httpUri="https://zenodo.org/record/3841684/files/figure.png" pageId="8" pageNumber="85" startId="8.[151,250,1940,1965]" targetBox="[166,1400,530,1908]" targetPageId="8">
<paragraph id="8BE5AC3DFFC3BC38FF68F82FFA35F818" blockId="8.[151,1437,1940,2037]" pageId="8" pageNumber="85">
<emphasis id="B92E702FFFC3BC38FF68F82FFEB1F841" bold="true" box="[151,271,1940,1965]" pageId="8" pageNumber="85">FIGURE 5</emphasis>
.
<taxonomicName id="4C5AD7BEFFC3BC38FEE2F82FFDC7F841" authority="Miya, 1972" authorityName=", Miya" authorityYear="1972" box="[285,633,1940,1965]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC3BC38FEE2F82FFE46F841" box="[285,504,1940,1964]" italics="true" pageId="8" pageNumber="85">Salmoneus gracilipes</emphasis>
<bibRefCitation id="EFCBD1CCFFC3BC38FDFFF82FFDC7F841" author="Miya, Y." box="[512,633,1940,1965]" pageId="8" pageNumber="85" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya, 1972</bibRefCitation>
</taxonomicName>
, ovigerous specimen (cl 4.5 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.048): A, frontal region, dorsal; B, anterior region of carapace and left eye, lateral; C, telson, dorsal; D, right (major) cheliped, lateral; E, same, chela, lateral; F, left (minor) cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral.
</paragraph>
</caption>
<caption id="DF25FCB5FFC2BC39FF68F8C8FC8EF815" ID-DOI="http://doi.org/10.5281/zenodo.3841686" ID-Zenodo-Dep="3841686" httpUri="https://zenodo.org/record/3841686/files/figure.png" pageId="9" pageNumber="86" startId="9.[151,250,1907,1932]" targetBox="[151,1436,181,1882]" targetPageId="9">
<paragraph id="8BE5AC3DFFC2BC39FF68F8C8FC8EF815" blockId="9.[151,1437,1907,2040]" pageId="9" pageNumber="86">
<emphasis id="B92E702FFFC2BC39FF68F8C8FEB1F861" bold="true" box="[151,271,1907,1932]" pageId="9" pageNumber="86">FIGURE 6</emphasis>
.
<taxonomicName id="4C5AD7BEFFC2BC39FEE2F8CFFDC7F861" authority="Miya" authorityName="Miya" authorityYear="1972" box="[285,633,1907,1932]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="86" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC2BC39FEE2F8CFFE35F861" box="[285,395,1908,1932]" italics="true" pageId="9" pageNumber="86">Salmoneus</emphasis>
<emphasis id="B92E702FFFC2BC39FE6CF8CFFE46F861" box="[403,504,1908,1932]" italics="true" pageId="9" pageNumber="86">gracilipes</emphasis>
<bibRefCitation id="EFCBD1CCFFC2BC39FDFFF8C8FDC7F861" author="Miya, Y." box="[512,633,1907,1932]" pageId="9" pageNumber="86" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya, 1972</bibRefCitation>
</taxonomicName>
, ovigerous specimen (cl 4.9 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.047), living individual (A, B); ovigerous specimen (cl 4.5 mm) from the same locality (OUMNH.ZC. 2020.01.048), preserved individual (C): A, dorsal; B, lateral; C, frontal region with focus on left eye, lateral.All photographs by the first author (AA), photograph in (C) taken through a Leica stereomicroscope.
</paragraph>
</caption>
<caption id="DF25FCB5FFC1BC3AFF68F953FD0AF880" ID-DOI="http://doi.org/10.5281/zenodo.3841688" ID-Zenodo-Dep="3841688" httpUri="https://zenodo.org/record/3841688/files/figure.png" pageId="10" pageNumber="87" startId="10.[151,250,1768,1793]" targetBox="[151,1436,181,1742]" targetPageId="10">
<paragraph id="8BE5AC3DFFC1BC3AFF68F953FD0AF880" blockId="10.[151,1437,1768,1901]" pageId="10" pageNumber="87">
<emphasis id="B92E702FFFC1BC3AFF68F953FEB0F8EC" bold="true" box="[151,270,1768,1793]" pageId="10" pageNumber="87">FIGURE 7</emphasis>
.
<taxonomicName id="4C5AD7BEFFC1BC3AFEE3F952FDC9F8EC" authority="Miya, 1972" authorityName=", Miya" authorityYear="1972" box="[284,631,1768,1793]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="87" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC1BC3AFEE3F952FE34F8EC" box="[284,394,1769,1793]" italics="true" pageId="10" pageNumber="87">Salmoneus</emphasis>
<emphasis id="B92E702FFFC1BC3AFE6EF952FE48F8EC" box="[401,502,1769,1793]" italics="true" pageId="10" pageNumber="87">gracilipes</emphasis>
<bibRefCitation id="EFCBD1CCFFC1BC3AFE01F953FDC9F8EC" author="Miya, Y." box="[510,631,1768,1793]" pageId="10" pageNumber="87" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya, 1972</bibRefCitation>
</taxonomicName>
, non-ovigerous specimen (cl 3.7 mm) from Failaka Island, Kuwait (OUMNH. ZC. 2020.01.046), living individual (A, B) and its presumed facultative host, the echiuran
<taxonomicName id="4C5AD7BEFFC1BC3AFBD0F8B6FF47F8A4" authority="Chen &amp; Yeh 1958" authorityName="Chen &amp; Yeh" authorityYear="1958" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="10" pageNumber="87" phylum="Echiura" rank="species" species="brevirostris">
<emphasis id="B92E702FFFC1BC3AFBD0F8B6FB17F8C8" box="[1071,1193,1805,1829]" italics="true" pageId="10" pageNumber="87">Listriolobus</emphasis>
cf.
<emphasis id="B92E702FFFC1BC3AFB2EF8B6FAFBF8C8" box="[1233,1349,1805,1829]" italics="true" pageId="10" pageNumber="87">brevirostris</emphasis>
<bibRefCitation id="EFCBD1CCFFC1BC3AFAB3F8B7FF47F8A4" author="Chen, Y. &amp; Yeh, C." pageId="10" pageNumber="87" pagination="265 - 278" refId="ref8176" refString="Chen, Y. &amp; Yeh, C. (1958) Notes on some Gephyrea of China with description of four new species. Acta Zoologica Sinica, 10, 265 - 278." type="journal article" year="1958">Chen &amp; Yeh 1958</bibRefCitation>
</taxonomicName>
(C, not to scale, size of the specimen approximately 60 mm with the proboscis fully extended): A, dorsal; B, lateral; C, host, lateral. Photographs by the first author (AA).
</paragraph>
</caption>
<caption id="DF25FCB5FFC0BC3BFF68FCCBFB95FC40" ID-DOI="http://doi.org/10.5281/zenodo.3841690" ID-Zenodo-Dep="3841690" httpUri="https://zenodo.org/record/3841690/files/figure.png" pageId="11" pageNumber="88" startId="11.[151,250,880,905]" targetBox="[151,1436,181,855]" targetPageId="11">
<paragraph id="8BE5AC3DFFC0BC3BFF68FCCBFB95FC40" blockId="11.[151,1436,880,941]" pageId="11" pageNumber="88">
<emphasis id="B92E702FFFC0BC3BFF68FCCBFEB0FC64" bold="true" box="[151,270,880,905]" pageId="11" pageNumber="88">FIGURE 8</emphasis>
.
<taxonomicName id="4C5AD7BEFFC0BC3BFEE5FCCAFDCDFC64" authority="Miya, 1972" authorityName=", Miya" authorityYear="1972" box="[282,627,880,905]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC0BC3BFEE5FCCAFE36FC64" box="[282,392,881,905]" italics="true" pageId="11" pageNumber="88">Salmoneus</emphasis>
<emphasis id="B92E702FFFC0BC3BFE70FCCAFE4AFC64" box="[399,500,881,905]" italics="true" pageId="11" pageNumber="88">gracilipes</emphasis>
<bibRefCitation id="EFCBD1CCFFC0BC3BFE04FCCBFDCDFC64" author="Miya, Y." box="[507,627,880,905]" pageId="11" pageNumber="88" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya, 1972</bibRefCitation>
</taxonomicName>
, 1 non-ovigerous specimen (cl 4.9 mm) from Tateyama, Honshu, Japan (CBM- ZC 14185), individual
<emphasis id="B92E702FFFC0BC3BFE80FC2DFE41FC40" box="[383,511,918,941]" italics="true" pageId="11" pageNumber="88">post-mortem</emphasis>
, lateral. Photograph courtesy of Dr. Tomoyuki Komai.
</paragraph>
</caption>
<subSubSection id="C340FFB6FFC0BC3BFF38FC61FC21FB68" pageId="11" pageNumber="88" type="distribution">
<paragraph id="8BE5AC3DFFC0BC3BFF38FC61FC21FB68" blockId="11.[151,1437,986,2022]" pageId="11" pageNumber="88">
<emphasis id="B92E702FFFC0BC3BFF38FC61FEE7FC18" bold="true" box="[199,345,986,1013]" pageId="11" pageNumber="88">Distribution</emphasis>
. Indo-West Pacific:
<collectingCountry id="F34DECADFFC0BC3BFDBAFC67FD3BFC1B" box="[581,645,988,1014]" name="Japan" pageId="11" pageNumber="88">Japan</collectingCountry>
(e.g., Shimo-shima, Genkai-nada,
<collectingRegion id="499E62DFFFC0BC3BFBECFC60FBE8FC18" box="[1043,1110,987,1013]" country="Japan" name="Chiba" pageId="11" pageNumber="88">Chiba</collectingRegion>
Peninsula, Ryukyu Islands),
<collectingCountry id="F34DECADFFC0BC3BFF68FBBBFF65FBF7" box="[151,219,1024,1050]" name="South Korea" pageId="11" pageNumber="88">Korea</collectingCountry>
(
<collectingRegion id="499E62DFFFC0BC3BFF10FC44FEA0FBF4" box="[239,286,1023,1049]" country="South Korea" name="Jeju" pageId="11" pageNumber="88">Jeju</collectingRegion>
Island),
<collectingCountry id="F34DECADFFC0BC3BFE75FC44FDB6FBF4" box="[394,520,1023,1049]" name="Micronesia" pageId="11" pageNumber="88">Micronesia</collectingCountry>
(Majuro Atoll),
<collectingCountry id="F34DECADFFC0BC3BFD30FC44FC82FBF4" box="[719,828,1023,1049]" name="Indonesia" pageId="11" pageNumber="88">Indonesia</collectingCountry>
(Lombok),
<collectingCountry id="F34DECADFFC0BC3BFC33FBBBFB10FBF4" box="[972,1198,1023,1050]" name="Papua New Guinea" pageId="11" pageNumber="88">Papua New Guinea</collectingCountry>
(
<collectingRegion id="499E62DFFFC0BC3BFB3EFC44FAE6FBF7" box="[1217,1368,1023,1050]" country="Papua New Guinea" name="New Ireland" pageId="11" pageNumber="88">New Ireland</collectingRegion>
),
<collectingCountry id="F34DECADFFC0BC3BFA90FBBBFA22FBF7" box="[1391,1436,1024,1050]" name="Iran" pageId="11" pageNumber="88">Iran</collectingCountry>
(
<collectingCountry id="F34DECADFFC0BC3BFF60FB98FEFEFBD3" box="[159,320,1059,1086]" name="Qeshm Island" pageId="11" pageNumber="88">Qeshm Island</collectingCountry>
) and
<collectingCountry id="F34DECADFFC0BC3BFE7BFB98FE68FBD0" box="[388,470,1059,1085]" name="Kuwait" pageId="11" pageNumber="88">Kuwait</collectingCountry>
(Failaka and Kubbar Islands) (
<bibRefCitation id="EFCBD1CCFFC0BC3BFCC0FB98FC7FFBD0" author="Miya, Y." box="[831,961,1059,1085]" pageId="11" pageNumber="88" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
,
<bibRefCitation id="EFCBD1CCFFC0BC3BFC30FB98FBB4FBD0" author="Miya, Y." box="[975,1034,1059,1085]" pageId="11" pageNumber="88" pagination="67 - 100" refId="ref9147" refString="Miya, Y. (1984) Alpheid shrimps from the Truks, Ponape and Majuro Atolls (Crustacea Decapoda). Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 27, 67 - 100." type="journal article" year="1984">1984</bibRefCitation>
,
<bibRefCitation id="EFCBD1CCFFC0BC3BFBE7FB99FBEDFBD0" author="Miya, Y." box="[1048,1107,1058,1085]" pageId="11" pageNumber="88" pagination="271 - 288" refId="ref9186" refString="Miya, Y. (1995) Four species of Alpheus from intertidal and shallow water mudflats in the Sea of Ariake, Kyushu, Japan (Crustacea, Decapoda, Alpheidae). Bulletin of the Faculty of Liberal Arts, Nagasaki University, Natural Science, 35, 271 - 288." type="journal article" year="1995">1995</bibRefCitation>
;
<bibRefCitation id="EFCBD1CCFFC0BC3BFB9EFB98FABCFBD0" author="Hayashi, K. - I." box="[1121,1282,1058,1085]" pageId="11" pageNumber="88" pagination="520 - 524" refId="ref8593" refString="Hayashi, K. - I. (1995) Prawns, shrimps and lobsters from Japan (85). Family Alpheidae - genera Automate and Salmoneus. Aquabiology, 101 (17), 520 - 524." type="journal article" year="1995">Hayashi 1995</bibRefCitation>
;
<bibRefCitation id="EFCBD1CCFFC0BC3BFAEEFB9FFF6CFB8C" author="Koo, H. &amp; Kim, W." pageId="11" pageNumber="88" pagination="43 - 48" refId="ref8939" refString="Koo, H. &amp; Kim, W. (2003) First report of the alpheid shrimp Salmoneus gracilipes (Decapoda: Caridea: Alpheidae) from Korea. The Korean Journal of Systematic Zoology, 19, 43 - 48." type="journal article" year="2003">Koo &amp; Kim 2003</bibRefCitation>
;
<bibRefCitation id="EFCBD1CCFFC0BC3BFF21FBFCFE43FB8C" author="Naderloo, R. &amp; Turkay, M." box="[222,509,1095,1121]" pageId="11" pageNumber="88" pagination="1 - 67" refId="ref8982" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012</bibRefCitation>
;
<bibRefCitation id="EFCBD1CCFFC0BC3BFDF9FBFCFD73FB8C" author="Anker, A. &amp; Pratama, I. S. &amp; Firdaus, M. &amp; Rahayu, D. L." box="[518,717,1094,1121]" pageId="11" pageNumber="88" pagination="301 - 342" refId="ref8017" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans (Alpheidae, Laomediidae, Strahlaxiidae) from Lombok, Indonesia. Zootaxa, 3911 (3), 301 - 342. https: // doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker
<emphasis id="B92E702FFFC0BC3BFDACFBF3FD35FB8C" box="[595,651,1095,1121]" italics="true" pageId="11" pageNumber="88">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="EFCBD1CCFFC0BC3BFD26FBFDFC7AFB8C" author="Duris, Z. &amp; Horka, I." box="[729,964,1094,1121]" pageId="11" pageNumber="88" pagination="773 - 793" refId="ref8441" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016</bibRefCitation>
; present study); additional possible record from northern
<collectingCountry id="F34DECADFFC0BC3BFEC6FBD0FE1EFB68" box="[313,416,1131,1157]" name="Australia" pageId="11" pageNumber="88">Australia</collectingCountry>
(
<bibRefCitation id="EFCBD1CCFFC0BC3BFE51FBD0FD87FB68" author="Anker, A." box="[430,569,1131,1157]" pageId="11" pageNumber="88" refId="ref7566" refString="Anker, A. (2001) Taxonomie et evolution des Alpheidae (Crustacea, Decapoda). Vols. 1 &amp; 2. PhD Thesis, Museum National d'Histoire Naturelle, Paris, 547 + 331 pp." type="book" year="2001">Anker 2001</bibRefCitation>
). However, see Remarks below.
</paragraph>
</subSubSection>
<subSubSection id="C340FFB6FFC0BC3BFF38FB34FE93F9FF" pageId="11" pageNumber="88" type="biology_ecology">
<paragraph id="8BE5AC3DFFC0BC3BFF38FB34FE93F9FF" blockId="11.[151,1437,986,2022]" pageId="11" pageNumber="88">
<emphasis id="B92E702FFFC0BC3BFF38FB34FE98FB44" bold="true" box="[199,294,1167,1193]" pageId="11" pageNumber="88">Ecology</emphasis>
.
<taxonomicName id="4C5AD7BEFFC0BC3BFECAFB34FD99FB44" authorityName=", Miya" authorityYear="1972" box="[309,551,1167,1193]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC0BC3BFECAFB34FD99FB44" box="[309,551,1167,1193]" italics="true" pageId="11" pageNumber="88">Salmoneus gracilipes</emphasis>
</taxonomicName>
appears to be ecologically rather variable, perhaps indicating the existence of a species complex (see Remarks below). The Kuwaiti specimens were collected either by flipping coral rocks on a heavily degraded reef, at a depth of
<quantity id="4CA201D8FFC0BC3BFDB9FB63FDCCFB1F" box="[582,626,1240,1266]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="88" unit="m" value="7.0">7 m</quantity>
(Kubbar Island), or extracted from a burrow of an echiuran worm,
<taxonomicName id="4C5AD7BEFFC0BC3BFA8EFB63FCE0FAFB" authority="Chen &amp; Yeh, 1958" authorityName="Chen &amp; Yeh" authorityYear="1958" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="11" pageNumber="88" phylum="Echiura" rank="species" species="brevirostris">
<emphasis id="B92E702FFFC0BC3BFA8EFB63FF42FAF8" italics="true" pageId="11" pageNumber="88">Listriolobus</emphasis>
cf.
<emphasis id="B92E702FFFC0BC3BFED4FB40FE12FAF8" box="[299,428,1275,1301]" italics="true" pageId="11" pageNumber="88">brevirostris</emphasis>
<bibRefCitation id="EFCBD1CCFFC0BC3BFE4BFB40FD37FAF8" author="Chen, Y. &amp; Yeh, C." box="[436,649,1274,1301]" pageId="11" pageNumber="88" pagination="265 - 278" refId="ref8176" refString="Chen, Y. &amp; Yeh, C. (1958) Notes on some Gephyrea of China with description of four new species. Acta Zoologica Sinica, 10, 265 - 278." type="journal article" year="1958">Chen &amp; Yeh, 1958</bibRefCitation>
(Thalassematidae)
</taxonomicName>
(
<figureCitation id="1361B0B8FFC0BC3BFC8FFB40FC77FAF8" box="[880,969,1275,1301]" captionStart="FIGURE 7" captionStartId="10.[151,250,1768,1793]" captionTargetBox="[151,1436,181,1742]" captionTargetId="figure@10.[151,1436,181,1742]" captionTargetPageId="10" captionText="FIGURE 7. Salmoneus gracilipes Miya, 1972, non-ovigerous specimen (cl 3.7 mm) from Failaka Island, Kuwait (OUMNH. ZC. 2020.01.046), living individual (A, B) and its presumed facultative host, the echiuran Listriolobus cf. brevirostris Chen &amp; Yeh 1958 (C, not to scale, size of the specimen approximately 60 mm with the proboscis fully extended): A, dorsal; B, lateral; C, host, lateral. Photographs by the first author (AA)." figureDoi="http://doi.org/10.5281/zenodo.3841688" httpUri="https://zenodo.org/record/3841688/files/figure.png" pageId="11" pageNumber="88">Fig. 7C</figureCitation>
), on a shallow subtidal flat characterised by murky water and a fine sandy-muddy bottom, near large rock platforms (Failaka Island). The
<typeStatus id="54E1129FFFC0BC3BFB10FA9BFAA1FAD7" box="[1263,1311,1312,1338]" pageId="11" pageNumber="88">type</typeStatus>
series was collected in both the intertidal, under boulders on sandy-mud flats at low tide (as was the specimen from Tateyama), and at a depth of
<quantity id="4CA201D8FFC0BC3BFEA1FADCFE26FA6F" box="[350,408,1383,1410]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="88" unit="m" value="10.0">10 m</quantity>
(
<bibRefCitation id="EFCBD1CCFFC0BC3BFE57FADCFD8EFA6C" author="Miya, Y." box="[424,560,1383,1409]" pageId="11" pageNumber="88" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya, 1972</bibRefCitation>
). In contrast, the Majuro Atoll specimen was found under a coral rock buried deep in the sand at the low tide level of an open ocean-side reef flat (
<bibRefCitation id="EFCBD1CCFFC0BC3BFC4DFA30FB8DFA48" author="Miya, Y." box="[946,1075,1419,1445]" pageId="11" pageNumber="88" pagination="67 - 100" refId="ref9147" refString="Miya, Y. (1984) Alpheid shrimps from the Truks, Ponape and Majuro Atolls (Crustacea Decapoda). Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 27, 67 - 100." type="journal article" year="1984">Miya 1984</bibRefCitation>
), whilst the Lombok specimens were extracted from burrows on an intertidal / shallow subtidal sea grass flat with abundant coral rubble (
<bibRefCitation id="EFCBD1CCFFC0BC3BFAC6FA14FF47FA00" author="Anker, A. &amp; Pratama, I. S. &amp; Firdaus, M. &amp; Rahayu, D. L." pageId="11" pageNumber="88" pagination="301 - 342" refId="ref8017" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans (Alpheidae, Laomediidae, Strahlaxiidae) from Lombok, Indonesia. Zootaxa, 3911 (3), 301 - 342. https: // doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker
<emphasis id="B92E702FFFC0BC3BFA77FA0BFF13FA00" italics="true" pageId="11" pageNumber="88">et al</emphasis>
. 2015
</bibRefCitation>
). The habitat of the
<collectingRegion id="499E62DFFFC0BC3BFE24FA6FFDD8FA00" box="[475,614,1491,1518]" country="Papua New Guinea" name="New Ireland" pageId="11" pageNumber="88">New Ireland</collectingRegion>
(Kavieng) specimen was noted as “
<quantity id="4CA201D8FFC0BC3BFC03FA68FBE2FA03" box="[1020,1116,1491,1518]" metricMagnitude="0" metricUnit="m" metricValue="8.0" metricValueMax="12.0" metricValueMin="4.0" pageId="11" pageNumber="88" unit="m" value="8.0" valueMax="12.0" valueMin="4.0">4 12 m</quantity>
, sand, rubble, silt” (
<bibRefCitation id="EFCBD1CCFFC0BC3BFAC0FA69FE9FF9FF" author="Duris, Z. &amp; Horka, I." pageId="11" pageNumber="88" pagination="773 - 793" refId="ref8441" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C340FFB6FFC0BC3DFF38F9A0FB79F976" lastPageId="13" lastPageNumber="90" pageId="11" pageNumber="88" type="discussion">
<paragraph id="8BE5AC3DFFC0BC3BFF38F9A0FDA0F92B" blockId="11.[151,1437,986,2022]" pageId="11" pageNumber="88">
<emphasis id="B92E702FFFC0BC3BFF38F9A0FE8AF9D8" bold="true" box="[199,308,1563,1589]" pageId="11" pageNumber="88">Remarks</emphasis>
. The present material is morphologically similar to
<taxonomicName id="4C5AD7BEFFC0BC3BFC7DF9A7FBB1F9D8" authorityName=", Miya" authorityYear="1972" box="[898,1039,1563,1590]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC0BC3BFC7DF9A7FBB1F9D8" box="[898,1039,1563,1590]" italics="true" pageId="11" pageNumber="88">S. gracilipes</emphasis>
</taxonomicName>
from
<collectingCountry id="F34DECADFFC0BC3BFBA9F9A7FB27F9DB" box="[1110,1177,1564,1590]" name="Japan" pageId="11" pageNumber="88">Japan</collectingCountry>
, especially in the main diagnostic features of the rostro-orbital region, tail fan and major and minor chelipeds, including the presence of the characteristic smooth ridges on the ventral surface of the major chela palm (
<bibRefCitation id="EFCBD1CCFFC0BC3BFC1BF9D8FBDAF990" author="Miya, Y." box="[996,1124,1635,1661]" pageId="11" pageNumber="88" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
: pl. 3, figs. C, D). However, there are also several important discrepancies between the Kuwaiti material and the
<typeStatus id="54E1129FFFC0BC3BFBC5F933FBD4F94F" box="[1082,1130,1672,1698]" pageId="11" pageNumber="88">type</typeStatus>
specimens, as well as other material identified as
<taxonomicName id="4C5AD7BEFFC0BC3BFE74F917FDA9F928" authorityName=", Miya" authorityYear="1972" box="[395,535,1707,1734]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC0BC3BFE74F917FDA9F928" box="[395,535,1707,1734]" italics="true" pageId="11" pageNumber="88">S. gracilipes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE5AC3DFFC0BC3CFF38F974FCF9FF3A" blockId="11.[151,1437,986,2022]" lastBlockId="12.[151,1436,152,215]" lastPageId="12" lastPageNumber="89" pageId="11" pageNumber="88">
According to the description and illustration of
<bibRefCitation id="EFCBD1CCFFC0BC3BFD21F974FCD0F904" author="Miya, Y." box="[734,878,1743,1769]" pageId="11" pageNumber="88" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya (1972)</bibRefCitation>
, the
<typeStatus id="54E1129FFFC0BC3BFC5CF974FBBBF904" box="[931,1029,1743,1769]" pageId="11" pageNumber="88" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5AD7BEFFC0BC3BFBD4F96BFB06F904" authorityName=", Miya" authorityYear="1972" box="[1067,1208,1743,1770]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="88" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC0BC3BFBD4F96BFB06F904" box="[1067,1208,1743,1770]" italics="true" pageId="11" pageNumber="88">S. gracilipes</emphasis>
</taxonomicName>
was lacking a small subdistal tooth on the ventral margin of the rostrum (
<bibRefCitation id="EFCBD1CCFFC0BC3BFD1CF948FCDCF8E0" author="Miya, Y." box="[739,866,1779,1805]" pageId="11" pageNumber="88" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
: pl. 3, fig. A). Later,
<bibRefCitation id="EFCBD1CCFFC0BC3BFBB6F948FB68F8E0" author="Miya, Y." box="[1097,1238,1779,1805]" pageId="11" pageNumber="88" pagination="67 - 100" refId="ref9147" refString="Miya, Y. (1984) Alpheid shrimps from the Truks, Ponape and Majuro Atolls (Crustacea Decapoda). Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 27, 67 - 100." type="journal article" year="1984">Miya (1984)</bibRefCitation>
reported an ovigerous specimen from Majuro Atoll (
<collectingCountry id="F34DECADFFC0BC3BFDD8F8ACFD15F8DF" box="[551,683,1815,1842]" name="Micronesia" pageId="11" pageNumber="88">Micronesia</collectingCountry>
), in which a small subdistal tooth was present on the ventral margin of the rostrum (
<bibRefCitation id="EFCBD1CCFFC0BC3BFEB5F880FE75F8B8" author="Miya, Y." box="[330,459,1851,1877]" pageId="11" pageNumber="88" pagination="67 - 100" refId="ref9147" refString="Miya, Y. (1984) Alpheid shrimps from the Truks, Ponape and Majuro Atolls (Crustacea Decapoda). Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 27, 67 - 100." type="journal article" year="1984">Miya 1984</bibRefCitation>
: fig. 4A), further stating that because of this small difference and a different ecology, the Majuro specimen could belong to a different taxon. In the specimen from Tateyama, the rostrum is devoid of a subdistal ventral tooth (T. Komai, pers. comm.). In contrast, in all three Kuwaiti specimens, the rostrum possesses a more or less developed subdistal ventral tooth (
<figureCitation id="1361B0B8FFC0BC3BFD57F81CFCBEF82C" box="[680,768,1958,1985]" captionStart="FIGURE 5" captionStartId="8.[151,250,1940,1965]" captionTargetBox="[166,1400,530,1908]" captionTargetId="figure@8.[161,1406,525,1915]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.5 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.048):A, frontal region, dorsal; B, anterior region of carapace and left eye, lateral; C, telson, dorsal; D, right (major) cheliped, lateral; E, same, chela, lateral; F, left (minor) cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral." figureDoi="http://doi.org/10.5281/zenodo.3841684" httpUri="https://zenodo.org/record/3841684/files/figure.png" pageId="11" pageNumber="88">Fig. 5B</figureCitation>
), which raises the same taxonomic doubts as
<bibRefCitation id="EFCBD1CCFFC0BC3BFB02F81CFA22F82C" author="Miya, Y." box="[1277,1436,1959,1985]" pageId="11" pageNumber="88" pagination="67 - 100" refId="ref9147" refString="Miya, Y. (1984) Alpheid shrimps from the Truks, Ponape and Majuro Atolls (Crustacea Decapoda). Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 27, 67 - 100." type="journal article" year="1984">Miyas (1984)</bibRefCitation>
for the specimen from Majuro Atoll. Furthermore, the Kuwaiti specimens also have a small (in one specimen ob- solete) post-rostral tubercle (
<figureCitation id="1361B0B8FFC7BC3CFE20FF22FD87FF5E" box="[479,569,152,179]" captionStart="FIGURE 5" captionStartId="8.[151,250,1940,1965]" captionTargetBox="[166,1400,530,1908]" captionTargetId="figure@8.[161,1406,525,1915]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.5 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.048):A, frontal region, dorsal; B, anterior region of carapace and left eye, lateral; C, telson, dorsal; D, right (major) cheliped, lateral; E, same, chela, lateral; F, left (minor) cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral." figureDoi="http://doi.org/10.5281/zenodo.3841684" httpUri="https://zenodo.org/record/3841684/files/figure.png" pageId="12" pageNumber="89">Fig. 5B</figureCitation>
), which seems to be absent in the type specimens, at least based on Miyas descriptive text and illustrations (
<bibRefCitation id="EFCBD1CCFFC7BC3CFDF3FF06FD32FF3A" author="Miya, Y." box="[524,652,189,215]" pageId="12" pageNumber="89" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
: pl. 3, fig. A, B).
</paragraph>
<caption id="DF25FCB5FFC7BC3CFF68F946FDC2F84B" ID-DOI="http://doi.org/10.5281/zenodo.3841692" ID-Zenodo-Dep="3841692" httpUri="https://zenodo.org/record/3841692/files/figure.png" pageId="12" pageNumber="89" startId="12.[151,250,1789,1814]" targetBox="[151,1436,255,1765]" targetPageId="12">
<paragraph id="8BE5AC3DFFC7BC3CFF68F946FDC2F84B" blockId="12.[151,1437,1789,1959]" pageId="12" pageNumber="89">
<emphasis id="B92E702FFFC7BC3CFF68F946FEB0F8FB" bold="true" box="[151,270,1789,1814]" pageId="12" pageNumber="89">FIGURE 9</emphasis>
. Habitats of
<taxonomicName id="4C5AD7BEFFC7BC3CFE6CF945FDBFF8FB" authorityName="Holthuis" authorityYear="1955" box="[403,513,1790,1814]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B92E702FFFC7BC3CFE6CF945FDBFF8FB" box="[403,513,1790,1814]" italics="true" pageId="12" pageNumber="89">Salmoneus</emphasis>
</taxonomicName>
spp. in Kuwait: A, rocky-muddy intertidal flat on the eastern coast of Failaka Island, type locality of
<taxonomicName id="4C5AD7BEFFC7BC3CFEFAF899FE09F8D7" authority="Anker &amp; Al-Kandari &amp; Grave, 2020" authorityName="Anker &amp; Al-Kandari &amp; Grave" authorityYear="2020" box="[261,439,1826,1850]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="89" phylum="Arthropoda" rank="species" species="ikaros" status="sp. nov.">
<emphasis id="B92E702FFFC7BC3CFEFAF899FE09F8D7" box="[261,439,1826,1850]" italics="true" pageId="12" pageNumber="89">Salmoneus ikaros</emphasis>
</taxonomicName>
<emphasis id="B92E702FFFC7BC3CFE41F899FDB0F8D7" bold="true" box="[446,526,1826,1850]" pageId="12" pageNumber="89">
<taxonomicNameLabel id="A21DCD54FFC7BC3CFE41F899FDB0F8D7" box="[446,526,1826,1850]" pageId="12" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
associated with
<taxonomicName id="4C5AD7BEFFC7BC3CFD48F899FBE9F8D7" authority="(Tirmizi 1970)" baseAuthorityName="Tirmizi" baseAuthorityYear="1970" box="[695,1111,1825,1850]" class="Malacostraca" family="Callianassidae" genus="Balsscallichirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="89" phylum="Arthropoda" rank="species" species="masoomi">
<emphasis id="B92E702FFFC7BC3CFD48F899FCE4F8D7" box="[695,858,1826,1850]" italics="true" pageId="12" pageNumber="89">Balsscallichirus</emphasis>
<emphasis id="B92E702FFFC7BC3CFC9FF898FC05F8D7" box="[864,955,1827,1850]" italics="true" pageId="12" pageNumber="89">masoomi</emphasis>
(
<bibRefCitation id="EFCBD1CCFFC7BC3CFC36F89AFBEEF8D7" author="Tirmizi, N. M." box="[969,1104,1825,1850]" pageId="12" pageNumber="89" pagination="245 - 250" refId="ref9296" refString="Tirmizi, N. M. (1970) A new species of Callianassa (Decapoda, Thalassinidea) from West Pakistan. Crustaceana, 19, 245 - 250. https: // doi. org / 10.1163 / 156854070 X 00329" type="journal article" year="1970">Tirmizi 1970</bibRefCitation>
)
</taxonomicName>
and habitat of one individual of
<taxonomicName id="4C5AD7BEFFC7BC3CFF68F8FCFEABF8B3" authorityName=", Miya" authorityYear="1972" box="[151,277,1862,1886]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="89" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC7BC3CFF68F8FCFEABF8B3" box="[151,277,1862,1886]" italics="true" pageId="12" pageNumber="89">S. gracilipes</emphasis>
</taxonomicName>
associated with
<taxonomicName id="4C5AD7BEFFC7BC3CFE42F8FDFC2EF8B3" authority="Chen" authorityName="Chen" authorityYear="1958" box="[445,912,1861,1886]" class="Polychaeta" family="Echiuridae" genus="Listriolobus" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="12" pageNumber="89" phylum="Echiura" rank="species" species="brevirostris">
<emphasis id="B92E702FFFC7BC3CFE42F8FDFD89F8B3" box="[445,567,1862,1886]" italics="true" pageId="12" pageNumber="89">Listriolobus</emphasis>
cf.
<emphasis id="B92E702FFFC7BC3CFDA2F8FDFD6FF8B3" box="[605,721,1862,1886]" italics="true" pageId="12" pageNumber="89">brevirostris</emphasis>
<bibRefCitation id="EFCBD1CCFFC7BC3CFD27F8FEFC2EF8B3" author="Chen, Y. &amp; Yeh, C." box="[728,912,1861,1886]" pageId="12" pageNumber="89" pagination="265 - 278" refId="ref8176" refString="Chen, Y. &amp; Yeh, C. (1958) Notes on some Gephyrea of China with description of four new species. Acta Zoologica Sinica, 10, 265 - 278." type="journal article" year="1958">Chen &amp; Yeh 1958</bibRefCitation>
</taxonomicName>
; B, degraded reef at Kubbar Island, about 7 m deep, habitat of two possibly free-living individuals of
<taxonomicName id="4C5AD7BEFFC7BC3CFD79F8D0FCBAF86F" authorityName=", Miya" authorityYear="1972" box="[646,772,1898,1922]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="89" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC7BC3CFD79F8D0FCBAF86F" box="[646,772,1898,1922]" italics="true" pageId="12" pageNumber="89">S. gracilipes</emphasis>
</taxonomicName>
, collected under coral rocks. Photographs: (A) by the third author (SDG); (B) courtesy of Dr. Shakar Al- Hazeem.
</paragraph>
</caption>
<paragraph id="8BE5AC3DFFC6BC3DFF38FF22FA28FD46" blockId="13.[151,1437,153,1691]" pageId="13" pageNumber="90">
In the original description of
<taxonomicName id="4C5AD7BEFFC6BC3DFDE9FF22FCFCFF5E" authority=", Miya (1972)" authorityName=", Miya" authorityYear="1972" box="[534,834,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFDE9FF22FD1AFF5E" box="[534,676,153,179]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
,
<bibRefCitation id="EFCBD1CCFFC6BC3DFD4CFF22FCFCFF5E" author="Miya, Y." box="[691,834,153,179]" pageId="13" pageNumber="90" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya (1972)</bibRefCitation>
</taxonomicName>
did not mention nor illustrate the stout cuspidate setae on the ischia of both the major and minor cheliped, as well as on the ischium of the second pereiopod (
<bibRefCitation id="EFCBD1CCFFC6BC3DFAA0FF06FF6DFF16" author="Miya, Y." pageId="13" pageNumber="90" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
: pl. 3, figs. A, C). It is important to note that in the specimens subsequently identified as
<taxonomicName id="4C5AD7BEFFC6BC3DFB2EFF5AFAE1FF16" authorityName=", Miya" authorityYear="1972" box="[1233,1375,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFB2EFF5AFAE1FF16" box="[1233,1375,225,251]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
or
<taxonomicName id="4C5AD7BEFFC6BC3DFA78FF5AFE8AFEF2" authorityName=", Miya" authorityYear="1972" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFA78FF5AFA22FF16" box="[1415,1436,225,251]" italics="true" pageId="13" pageNumber="90">S.</emphasis>
aff.
<emphasis id="B92E702FFFC6BC3DFF3BFEBEFE8AFEF2" box="[196,308,261,287]" italics="true" pageId="13" pageNumber="90">gracilipes</emphasis>
</taxonomicName>
, for which illustrations of some morphological parts were provided, the cheliped ischia were shown as devoid of stout cuspidate setae (e.g.,
<bibRefCitation id="EFCBD1CCFFC6BC3DFDAAFE92FD6DFEAE" author="Miya, Y." box="[597,723,297,323]" pageId="13" pageNumber="90" pagination="67 - 100" refId="ref9147" refString="Miya, Y. (1984) Alpheid shrimps from the Truks, Ponape and Majuro Atolls (Crustacea Decapoda). Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 27, 67 - 100." type="journal article" year="1984">Miya 1984</bibRefCitation>
: fig. 4C;
<bibRefCitation id="EFCBD1CCFFC6BC3DFCC5FE92FC67FEAE" author="Hayashi, K. - I." box="[826,985,296,323]" pageId="13" pageNumber="90" pagination="520 - 524" refId="ref8593" refString="Hayashi, K. - I. (1995) Prawns, shrimps and lobsters from Japan (85). Family Alpheidae - genera Automate and Salmoneus. Aquabiology, 101 (17), 520 - 524." type="journal article" year="1995">Hayashi 1995</bibRefCitation>
: fig. 288i;
<bibRefCitation id="EFCBD1CCFFC6BC3DFBADFE92FAA5FEAE" author="Koo, H. &amp; Kim, W." box="[1106,1307,297,323]" pageId="13" pageNumber="90" pagination="43 - 48" refId="ref8939" refString="Koo, H. &amp; Kim, W. (2003) First report of the alpheid shrimp Salmoneus gracilipes (Decapoda: Caridea: Alpheidae) from Korea. The Korean Journal of Systematic Zoology, 19, 43 - 48." type="journal article" year="2003">Koo &amp; Kim 2003</bibRefCitation>
: fig. 1D, E;
<bibRefCitation id="EFCBD1CCFFC6BC3DFF68FEF7FEC1FE8A" author="Duris, Z. &amp; Horka, I." box="[151,383,332,359]" pageId="13" pageNumber="90" pagination="773 - 793" refId="ref8441" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016</bibRefCitation>
: fig. 12c, g). In the specimen from Kavieng reported as
<taxonomicName id="4C5AD7BEFFC6BC3DFC19FEF6FBCFFE8A" authorityName=", Miya" authorityYear="1972" box="[998,1137,333,359]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFC19FEF6FBCFFE8A" box="[998,1137,333,359]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
and the specimen from the Red Sea reported as
<taxonomicName id="4C5AD7BEFFC6BC3DFE82FECAFD88FE66" authorityName=", Miya" authorityYear="1972" box="[381,566,369,395]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFE82FECAFE2CFE66" box="[381,402,369,395]" italics="true" pageId="13" pageNumber="90">S.</emphasis>
aff.
<emphasis id="B92E702FFFC6BC3DFE39FECAFD88FE66" box="[454,566,369,395]" italics="true" pageId="13" pageNumber="90">gracilipes</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFCBD1CCFFC6BC3DFD9FFECBFCE2FE61" author="Duris, Z. &amp; Horka, I." box="[608,860,368,396]" pageId="13" pageNumber="90" pagination="773 - 793" refId="ref8441" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká (2016)</bibRefCitation>
, the second pereiopod ischium was armed with one cuspidate seta (
<emphasis id="B92E702FFFC6BC3DFEBAFE2EFEC5FE42" box="[325,379,405,431]" italics="true" pageId="13" pageNumber="90">idem</emphasis>
: fig. 12h). In the specimen from Tateyama, the major cheliped ischium is unarmed, the minor cheliped bears one small cuspidate seta and the second pereiopod ischia are each armed with one small cuspidate seta (T. Komai, pers. comm.). Regarding the Kuwaiti material, in the two specimens from Kubbar Island, the major and minor cheliped ischia, as well as the second pereiopod ischia, are each armed with one stout, rather conspicuous cuspidate seta (
<figureCitation id="1361B0B8FFC6BC3DFEBCFD9EFE65FDD2" box="[323,475,548,575]" captionStart="FIGURE 5" captionStartId="8.[151,250,1940,1965]" captionTargetBox="[166,1400,530,1908]" captionTargetId="figure@8.[161,1406,525,1915]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.5 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.048):A, frontal region, dorsal; B, anterior region of carapace and left eye, lateral; C, telson, dorsal; D, right (major) cheliped, lateral; E, same, chela, lateral; F, left (minor) cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral." figureDoi="http://doi.org/10.5281/zenodo.3841684" httpUri="https://zenodo.org/record/3841684/files/figure.png" pageId="13" pageNumber="90">Fig. 5D, F, G</figureCitation>
). In the specimen from Failaka Island, only the minor cheliped ischium is armed with a cuspidate seta, whilst the major cheliped ischium is unarmed, as in the specimen from Tateyama. Whether
<bibRefCitation id="EFCBD1CCFFC6BC3DFA9EFDF2FF5CFD6A" author="Miya, Y." pageId="13" pageNumber="90" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya (1972)</bibRefCitation>
overlooked the ischial armature on both the major and the minor chelipeds, as well as on the second pereiopods, or whether it was actually absent on all these appendages in the
<typeStatus id="54E1129FFFC6BC3DFC58FD2AFC69FD46" box="[935,983,657,683]" pageId="13" pageNumber="90">type</typeStatus>
specimens currently remains unknown.
</paragraph>
<paragraph id="8BE5AC3DFFC6BC3DFF38FD0EFEB1FC02" blockId="13.[151,1437,153,1691]" pageId="13" pageNumber="90">
The longitudinal groove on the dorsal (“outer”) surface of the major chela palm was considered to be one of the diagnostic features of
<taxonomicName id="4C5AD7BEFFC6BC3DFE48FD62FDFCFD1E" authorityName=", Miya" authorityYear="1972" box="[439,578,729,755]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFE48FD62FDFCFD1E" box="[439,578,729,755]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCBD1CCFFC6BC3DFDAFFD62FD71FD1E" author="Miya, Y." box="[592,719,729,755]" pageId="13" pageNumber="90" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
: pl. 3, fig. 3C, D;
<bibRefCitation id="EFCBD1CCFFC6BC3DFC69FD63FB3EFD1E" author="Duris, Z. &amp; Horka, I." box="[918,1152,728,755]" pageId="13" pageNumber="90" pagination="773 - 793" refId="ref8441" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016</bibRefCitation>
; shown for
<taxonomicName id="4C5AD7BEFFC6BC3DFAFEFD62FF02FCFA" authorityName=", Miya" authorityYear="1972" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFAFEFD62FAB1FD1E" box="[1281,1295,729,755]" italics="true" pageId="13" pageNumber="90">S</emphasis>
. aff.
<emphasis id="B92E702FFFC6BC3DFAB7FD62FF02FCFA" italics="true" pageId="13" pageNumber="90">gracilipes</emphasis>
</taxonomicName>
in their fig.
<quantity id="4CA201D8FFC6BC3DFEB4FD46FEC6FCFA" box="[331,376,765,791]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="13" pageNumber="90" unit="g" value="12.0">12g</quantity>
). In the Kuwaiti specimens, this groove is either very shallow and barely noticeable, as in the specimens from Kubbar Island (
<figureCitation id="1361B0B8FFC6BC3DFE04FC9AFDEAFCD6" box="[507,596,800,827]" captionStart="FIGURE 5" captionStartId="8.[151,250,1940,1965]" captionTargetBox="[166,1400,530,1908]" captionTargetId="figure@8.[161,1406,525,1915]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.5 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.048):A, frontal region, dorsal; B, anterior region of carapace and left eye, lateral; C, telson, dorsal; D, right (major) cheliped, lateral; E, same, chela, lateral; F, left (minor) cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral." figureDoi="http://doi.org/10.5281/zenodo.3841684" httpUri="https://zenodo.org/record/3841684/files/figure.png" pageId="13" pageNumber="90">Fig. 5D</figureCitation>
), or very conspicuous and similar to that of the
<typeStatus id="54E1129FFFC6BC3DFB9DFC9AFB79FCD6" box="[1122,1223,801,827]" pageId="13" pageNumber="90" type="holotype">holotype</typeStatus>
, as in the specimen from Failaka Island. Noteworthy,
<bibRefCitation id="EFCBD1CCFFC6BC3DFDE9FCFEFD70FCB2" author="Coutiere, H." box="[534,718,837,863]" pageId="13" pageNumber="90" pagination="380 - 386" refId="ref8211" refString="Coutiere, H. (1897 a) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sous-famille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1897">Coutière (1897a</bibRefCitation>
,
<bibRefCitation id="EFCBD1CCFFC6BC3DFD24FCFEFC9CFC8D" author="Coutiere, H." box="[731,802,837,864]" pageId="13" pageNumber="90" pagination="233 - 236" refId="ref8247" refString="Coutiere, H. (1897 b) Notes sur quelques Alpheides nouveaux ou peu connus rapportes de Djibouti (Afrique Orientale). Bulletin du Museum d'Histoire Naturelle, 3, 233 - 236. https: // doi. org / 10.5962 / bhl. part. 19259" type="journal article" year="1897">1897b</bibRefCitation>
) mentioned the presence of a deep groove on the dorsal (“superior”) surface of the major chela in
<taxonomicName id="4C5AD7BEFFC6BC3DFD91FCD2FC9FFC6E" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[622,801,873,899]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="B92E702FFFC6BC3DFD91FCD2FC9FFC6E" box="[622,801,873,899]" italics="true" pageId="13" pageNumber="90">S. serratidigitus</emphasis>
</taxonomicName>
, illustrating it in his famous monograph (
<bibRefCitation id="EFCBD1CCFFC6BC3DFB0CFCD2FA26FC6E" author="Coutiere, H." box="[1267,1432,873,899]" pageId="13" pageNumber="90" pagination="1 - 560" refId="ref8297" refString="Coutiere, H. (1899) Les &quot; Alpheidae &quot;, morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie, Serie 8, 9, 1 - 560, pls. 1 - 6. [Masson, Paris] https: // doi. org / 10.5962 / bhl. title. 13143" type="journal article" year="1899">Coutière 1899</bibRefCitation>
: fig. 215). Another species that has a shallow longitudinal depression on the dorsal surface of the major chela is
<taxonomicName id="4C5AD7BEFFC6BC3DFA78FC36FE9AFC26" authorityName="Anker" authorityYear="2010" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="camaroncito">
<emphasis id="B92E702FFFC6BC3DFA78FC36FE9AFC26" italics="true" pageId="13" pageNumber="90">S. camaroncito</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCBD1CCFFC6BC3DFECCFC0AFE00FC26" author="Anker, A." box="[307,446,945,971]" pageId="13" pageNumber="90" pagination="177 - 205" refId="ref7666" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372 (1), 177 - 205. https: // doi. org / 10.11646 / zootaxa. 2839.1.3" type="journal article" year="2010">Anker 2010</bibRefCitation>
: fig. 6a), indicating that it may be present in other species of
<taxonomicName id="4C5AD7BEFFC6BC3DFB8FFC0AFB57FC26" authorityName="Holthuis" authorityYear="1955" box="[1136,1257,945,971]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="genus">
<emphasis id="B92E702FFFC6BC3DFB8FFC0AFB57FC26" box="[1136,1257,945,971]" italics="true" pageId="13" pageNumber="90">Salmoneus</emphasis>
</taxonomicName>
, but not always illustrated.
</paragraph>
<paragraph id="8BE5AC3DFFC6BC3DFF38FC42FAF7FB49" blockId="13.[151,1437,153,1691]" pageId="13" pageNumber="90">
The armature of the major chela fingers also appears to be variable in the material identified as
<taxonomicName id="4C5AD7BEFFC6BC3DFAF7FC42FA2BFBFE" authorityName=", Miya" authorityYear="1972" box="[1288,1429,1017,1043]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFAF7FC42FA2BFBFE" box="[1288,1429,1017,1043]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
, ranging from about 14 teeth in the Japanese
<typeStatus id="54E1129FFFC6BC3DFD79FBA6FD08FBDA" box="[646,694,1053,1079]" pageId="13" pageNumber="90">type</typeStatus>
specimens (
<bibRefCitation id="EFCBD1CCFFC6BC3DFCBFFBA6FC01FBDA" author="Miya, Y." box="[832,959,1053,1079]" pageId="13" pageNumber="90" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya 1972</bibRefCitation>
: pl. 3, fig. D), as well as in the Korean and Micronesian specimens (
<bibRefCitation id="EFCBD1CCFFC6BC3DFE4FFBFAFD8FFBB6" author="Miya, Y." box="[432,561,1089,1115]" pageId="13" pageNumber="90" pagination="67 - 100" refId="ref9147" refString="Miya, Y. (1984) Alpheid shrimps from the Truks, Ponape and Majuro Atolls (Crustacea Decapoda). Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 27, 67 - 100." type="journal article" year="1984">Miya 1984</bibRefCitation>
: fig. 4D;
<bibRefCitation id="EFCBD1CCFFC6BC3DFD62FBFAFCD7FBB6" author="Koo, H. &amp; Kim, W." box="[669,873,1089,1115]" pageId="13" pageNumber="90" pagination="43 - 48" refId="ref8939" refString="Koo, H. &amp; Kim, W. (2003) First report of the alpheid shrimp Salmoneus gracilipes (Decapoda: Caridea: Alpheidae) from Korea. The Korean Journal of Systematic Zoology, 19, 43 - 48." type="journal article" year="2003">Koo &amp; Kim 2003</bibRefCitation>
: fig. 1D), to only
<quantity id="4CA201D8FFC6BC3DFBC9FBFAFBCEFBB6" box="[1078,1136,1089,1115]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="90" unit="in" value="10.0">10 in</quantity>
<taxonomicName id="4C5AD7BEFFC6BC3DFB87FBFAFA8DFBB6" authorityName=", Miya" authorityYear="1972" box="[1144,1331,1089,1115]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFB87FBFAFB33FBB6" box="[1144,1165,1089,1115]" italics="true" pageId="13" pageNumber="90">S.</emphasis>
aff.
<emphasis id="B92E702FFFC6BC3DFB3CFBFAFA8DFBB6" box="[1219,1331,1089,1115]" italics="true" pageId="13" pageNumber="90">gracilipes</emphasis>
</taxonomicName>
from the Red Sea (
<bibRefCitation id="EFCBD1CCFFC6BC3DFEFCFBDFFE4EFB92" author="Duris, Z. &amp; Horka, I." box="[259,496,1124,1151]" pageId="13" pageNumber="90" pagination="773 - 793" refId="ref8441" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016</bibRefCitation>
: fig.
<quantity id="4CA201D8FFC6BC3DFDD7FBDEFDE8FB92" box="[552,598,1125,1151]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="13" pageNumber="90" unit="g" value="12.0">12g</quantity>
). In the Kuwaiti specimens, the mean number is around 15 teeth (
<figureCitation id="1361B0B8FFC6BC3DFAC7FBDEFA31FB92" box="[1336,1423,1124,1151]" captionStart="FIGURE 5" captionStartId="8.[151,250,1940,1965]" captionTargetBox="[166,1400,530,1908]" captionTargetId="figure@8.[161,1406,525,1915]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.5 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.048):A, frontal region, dorsal; B, anterior region of carapace and left eye, lateral; C, telson, dorsal; D, right (major) cheliped, lateral; E, same, chela, lateral; F, left (minor) cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral." figureDoi="http://doi.org/10.5281/zenodo.3841684" httpUri="https://zenodo.org/record/3841684/files/figure.png" pageId="13" pageNumber="90">Fig. 5E</figureCitation>
), thus corresponding more to
<taxonomicName id="4C5AD7BEFFC6BC3DFE2EFB32FDE3FB4E" authorityName=", Miya" authorityYear="1972" box="[465,605,1161,1187]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFE2EFB32FDE3FB4E" box="[465,605,1161,1187]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
of
<bibRefCitation id="EFCBD1CCFFC6BC3DFD7CFB32FCB1FB4E" author="Miya, Y." box="[643,783,1161,1187]" pageId="13" pageNumber="90" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya (1972)</bibRefCitation>
than to
<taxonomicName id="4C5AD7BEFFC6BC3DFC95FB32FB9DFB4E" authorityName=", Miya" authorityYear="1972" box="[874,1059,1161,1187]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFC95FB32FC3EFB4E" box="[874,896,1161,1187]" italics="true" pageId="13" pageNumber="90">S.</emphasis>
aff.
<emphasis id="B92E702FFFC6BC3DFC4CFB32FB9DFB4E" box="[947,1059,1161,1187]" italics="true" pageId="13" pageNumber="90">gracilipes</emphasis>
</taxonomicName>
of
<bibRefCitation id="EFCBD1CCFFC6BC3DFBB7FB33FAFBFB49" author="Duris, Z. &amp; Horka, I." box="[1096,1349,1160,1188]" pageId="13" pageNumber="90" pagination="773 - 793" refId="ref8441" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká (2016)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE5AC3DFFC6BC3DFF38FB16FE03FA91" blockId="13.[151,1437,153,1691]" pageId="13" pageNumber="90">
It is also interesting to note that in the Kuwaiti and Tateyama specimens of
<taxonomicName id="4C5AD7BEFFC6BC3DFBEDFB16FB20FB2A" authorityName=", Miya" authorityYear="1972" box="[1042,1182,1197,1223]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFBEDFB16FB20FB2A" box="[1042,1182,1197,1223]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
, the pigmented part of the eye cornea is divided into two distinct areas: a smaller, more basal, black area and a larger, globular, more distal, brown area (
<figureCitation id="1361B0B8FFC6BC3DFEDBFB4EFE3BFAE2" box="[292,389,1268,1295]" captionStart="FIGURE 5" captionStartId="8.[151,250,1940,1965]" captionTargetBox="[166,1400,530,1908]" captionTargetId="figure@8.[161,1406,525,1915]" captionTargetPageId="8" captionText="FIGURE 5. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.5 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.048):A, frontal region, dorsal; B, anterior region of carapace and left eye, lateral; C, telson, dorsal; D, right (major) cheliped, lateral; E, same, chela, lateral; F, left (minor) cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral." figureDoi="http://doi.org/10.5281/zenodo.3841684" httpUri="https://zenodo.org/record/3841684/files/figure.png" pageId="13" pageNumber="90">Figs. 5B</figureCitation>
,
<figureCitation id="1361B0B8FFC6BC3DFE6EFB4EFE0BFAE2" box="[401,437,1269,1295]" captionStart="FIGURE 6" captionStartId="9.[151,250,1907,1932]" captionTargetBox="[151,1436,181,1882]" captionTargetId="figure@9.[151,1436,181,1882]" captionTargetPageId="9" captionText="FIGURE 6. Salmoneus gracilipes Miya, 1972, ovigerous specimen (cl 4.9 mm) from Kubbar Island, Kuwait (OUMNH.ZC. 2020.01.047), living individual (A, B); ovigerous specimen (cl 4.5 mm) from the same locality (OUMNH.ZC. 2020.01.048), preserved individual (C): A, dorsal; B, lateral; C, frontal region with focus on left eye, lateral.All photographs by the first author (AA), photograph in (C) taken through a Leica stereomicroscope." figureDoi="http://doi.org/10.5281/zenodo.3841686" httpUri="https://zenodo.org/record/3841686/files/figure.png" pageId="13" pageNumber="90">6C</figureCitation>
; also visible in
<figureCitation id="1361B0B8FFC6BC3DFD9CFB4EFD19FAE2" box="[611,679,1269,1295]" captionStart="FIGURE 8" captionStartId="11.[151,250,880,905]" captionTargetBox="[151,1436,181,855]" captionTargetId="figure@11.[151,1436,181,855]" captionTargetPageId="11" captionText="FIGURE 8. Salmoneus gracilipes Miya, 1972, 1 non-ovigerous specimen (cl 4.9 mm) from Tateyama, Honshu, Japan (CBM- ZC 14185), individual post-mortem, lateral. Photograph courtesy of Dr. Tomoyuki Komai." figureDoi="http://doi.org/10.5281/zenodo.3841690" httpUri="https://zenodo.org/record/3841690/files/figure.png" pageId="13" pageNumber="90">Fig. 8</figureCitation>
). Some unusual corneal pigmentation is also seen in the Indonesian specimens reported as
<taxonomicName id="4C5AD7BEFFC6BC3DFE69FAA2FD9CFADE" authorityName=", Miya" authorityYear="1972" box="[406,546,1305,1331]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFE69FAA2FD9CFADE" box="[406,546,1305,1331]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFCBD1CCFFC6BC3DFDB4FAA2FCA2FADE" author="Anker, A. &amp; Pratama, I. S. &amp; Firdaus, M. &amp; Rahayu, D. L." box="[587,796,1304,1331]" pageId="13" pageNumber="90" pagination="301 - 342" refId="ref8017" refString="Anker, A., Pratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans (Alpheidae, Laomediidae, Strahlaxiidae) from Lombok, Indonesia. Zootaxa, 3911 (3), 301 - 342. https: // doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker
<emphasis id="B92E702FFFC6BC3DFD67FAA1FD6EFADE" box="[664,720,1305,1331]" italics="true" pageId="13" pageNumber="90">et al.</emphasis>
(2015
</bibRefCitation>
: fig. 19A). Nevertheless, the general configuration of the eyes and the black and brown areas in
<taxonomicName id="4C5AD7BEFFC6BC3DFDB3FA86FD66FABA" authorityName=", Miya" authorityYear="1972" box="[588,728,1341,1367]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFDB3FA86FD66FABA" box="[588,728,1341,1367]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
is very different from that of the above-described
<taxonomicName id="4C5AD7BEFFC6BC3DFAEDFA86FACDFABA" authority="Anker &amp; Al-Kandari &amp; Grave, 2020" authorityName="Anker &amp; Al-Kandari &amp; Grave" authorityYear="2020" box="[1298,1395,1341,1367]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="ikaros" status="sp. nov.">
<emphasis id="B92E702FFFC6BC3DFAEDFA86FACDFABA" box="[1298,1395,1341,1367]" italics="true" pageId="13" pageNumber="90">S. ikaros</emphasis>
</taxonomicName>
<emphasis id="B92E702FFFC6BC3DFA84FA86FF76FA96" bold="true" pageId="13" pageNumber="90">
<taxonomicNameLabel id="A21DCD54FFC6BC3DFA84FA86FF76FA96" pageId="13" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(cf.
<figureCitation id="1361B0B8FFC6BC3DFF03FADAFEF7FA96" box="[252,329,1377,1403]" captionStart="FIGURE 4" captionStartId="6.[151,250,1412,1437]" captionTargetBox="[151,1436,957,1387]" captionTargetId="figure@6.[151,1436,957,1387]" captionTargetPageId="6" captionText="FIGURE 4. Salmoneus ikaros sp. nov., holotype, ovigerous specimen (cl 4.2 mm) from Failaka Island, Kuwait (OUMNH.ZC. 2020.01.045), preserved individual:A, left eyestalk, lateral; B, left and right eyestalks, anterolateral; C, same, anterior (frontal). Photographs taken by the first author (AA) through a Leica stereomicroscope." figureDoi="http://doi.org/10.5281/zenodo.3841682" httpUri="https://zenodo.org/record/3841682/files/figure.png" pageId="13" pageNumber="90">Figs. 4</figureCitation>
AC, 6C).
</paragraph>
<paragraph id="8BE5AC3DFFC6BC3DFF38FA3EFB79F976" blockId="13.[151,1437,153,1691]" pageId="13" pageNumber="90">
<bibRefCitation id="EFCBD1CCFFC6BC3DFF38FA3EFE4BFA72" author="Naderloo, R. &amp; Turkay, M." box="[199,501,1413,1439]" pageId="13" pageNumber="90" pagination="1 - 67" refId="ref8982" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay (2012)</bibRefCitation>
listed
<taxonomicName id="4C5AD7BEFFC6BC3DFDC1FA3EFD74FA72" authorityName=", Miya" authorityYear="1972" box="[574,714,1413,1439]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFDC1FA3EFD74FA72" box="[574,714,1413,1439]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
from the Iranian side of the Gulf (
<collectingCountry id="F34DECADFFC6BC3DFBB6FA3EFB59FA4D" box="[1097,1255,1413,1440]" name="Qeshm Island" pageId="13" pageNumber="90">Qeshm Island</collectingCountry>
) based on a single specimen reported in an unpublished M.Sc. thesis of
<bibRefCitation id="EFCBD1CCFFC6BC3DFCEDFA12FBA1FA2E" author="Sheibani-Tezerji, R." box="[786,1055,1449,1475]" pageId="13" pageNumber="90" refId="ref9240" refString="Sheibani-Tezerji, R. (2008) Taxonomy and zoogeography of littoral caridean shrimps (Crustacea: Decapoda: Caridea) of the Persian Gulf and Gulf of Oman. Unpublished M. Sc. thesis, University of Tehran, Tehran, 144 pp. [in Farsi with English abstract, not seen]" type="book" year="2008">Sheibani-Tezerji (2008)</bibRefCitation>
, stating that her specimen “agrees well with the original description by
<bibRefCitation id="EFCBD1CCFFC6BC3DFDCAFA76FD01FA0A" author="Miya, Y." box="[565,703,1485,1511]" pageId="13" pageNumber="90" pagination="23 - 101" refId="ref9101" refString="Miya, Y. (1972) The Alpheidae (Crustacea Decapoda) of Japan and its adjacent waters. Part I. Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 3, 23 - 101, pls. 1 - 14." type="journal article" year="1972">Miya (1972</bibRefCitation>
….)”. However, this species was listed as “
<taxonomicName id="4C5AD7BEFFC6BC3DFB65FA76FAF4FA0A" authorityName=", Miya" authorityYear="1972" box="[1178,1354,1485,1511]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFB65FA76FB11FA0A" box="[1178,1199,1485,1511]" italics="true" pageId="13" pageNumber="90">S.</emphasis>
cf.
<emphasis id="B92E702FFFC6BC3DFB25FA76FAF4FA0A" box="[1242,1354,1485,1511]" italics="true" pageId="13" pageNumber="90">gracilipes</emphasis>
</taxonomicName>
” in the abstract of
<bibRefCitation id="EFCBD1CCFFC6BC3DFEEDFA4AFDFEF9E6" author="Naderloo, R. &amp; Turkay, M." box="[274,576,1521,1547]" pageId="13" pageNumber="90" pagination="1 - 67" refId="ref8982" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay (2012)</bibRefCitation>
, without any further comments. Due to the uncertainties in the previous identifications of
<taxonomicName id="4C5AD7BEFFC6BC3DFED1F9AEFE04F9C2" authorityName=", Miya" authorityYear="1972" box="[302,442,1557,1583]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFED1F9AEFE04F9C2" box="[302,442,1557,1583]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
, including by the descriptor of the species itself (
<bibRefCitation id="EFCBD1CCFFC6BC3DFC17F9AEFBD7F9C2" author="Miya, Y." box="[1000,1129,1557,1583]" pageId="13" pageNumber="90" pagination="67 - 100" refId="ref9147" refString="Miya, Y. (1984) Alpheid shrimps from the Truks, Ponape and Majuro Atolls (Crustacea Decapoda). Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 27, 67 - 100." type="journal article" year="1984">Miya 1984</bibRefCitation>
), the identity of the Iranian material needs confirmation. More generally, it seems likely that all material previously reported as
<taxonomicName id="4C5AD7BEFFC6BC3DFAF0F982FA22F9BE" authorityName=", Miya" authorityYear="1972" box="[1295,1436,1593,1619]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B92E702FFFC6BC3DFAF0F982FA22F9BE" box="[1295,1436,1593,1619]" italics="true" pageId="13" pageNumber="90">S. gracilipes</emphasis>
</taxonomicName>
(e.g., some material from
<collectingCountry id="F34DECADFFC6BC3DFE44F9E6FE45F99A" box="[443,507,1629,1655]" name="Japan" pageId="13" pageNumber="90">Japan</collectingCountry>
+ material from
<collectingCountry id="F34DECADFFC6BC3DFD44F9E6FC82F99A" box="[699,828,1629,1655]" name="Micronesia" pageId="13" pageNumber="90">Micronesia</collectingCountry>
,
<collectingCountry id="F34DECADFFC6BC3DFCB7F9E6FC0BF99A" box="[840,949,1629,1655]" name="Indonesia" pageId="13" pageNumber="90">Indonesia</collectingCountry>
and
<collectingCountry id="F34DECADFFC6BC3DFC13F9E6FB74F995" box="[1004,1226,1629,1656]" name="Papua New Guinea" pageId="13" pageNumber="90">Papua New Guinea</collectingCountry>
), together with the present material from
<collectingCountry id="F34DECADFFC6BC3DFE70F93AFE5AF976" box="[399,484,1665,1691]" name="Kuwait" pageId="13" pageNumber="90">Kuwait</collectingCountry>
, in reality represents a species complex, which needs unravelling.
</paragraph>
</subSubSection>
</treatment>
</document>