372 lines
54 KiB
XML
372 lines
54 KiB
XML
<document id="AAC7E76C54D305416F87E00A51501D8F" ID-CLB-Dataset="25304" ID-DOI="10.5852/ejt.2019.561" ID-GBIF-Dataset="f0779c03-8e65-424d-b680-ff8f219fd345" ID-Zenodo-Dep="3474359" ID-ZooBank="6C81FC6F-2CDD-46B8-9825-6D09DF9475F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570396803884" checkinUser="plazi" docAuthor="Prokofiev, Artem M." docDate="2019" docId="03E72F184007FFF7FD80516FFD71FB95" docLanguage="en" docName="ejt-561_prokofiev.pdf.imf" docOrigin="European Journal of Taxonomy 561" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Engertia allolepis Prokofiev 2019, sp. nov." docType="treatment" docUuid="3E17E5EE-4309-4836-9A06-4AB5994E3F32" docUuidSource="ZooBank" docVersion="8" lastPageNumber="6" masterDocId="FFDE57604005FFF2FFFE5345FF9AFFE5" masterDocTitle="Two new species of Engertia Dalla Torre, 1913 and Philacelota Heller, 1900 from Indonesia and the Philippines with a revised identification key (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="3" updateTime="1698757900835" updateUser="ExternalLinkService">
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<mods:mods id="C3EEE2FFF395F1863367FEBC10945E8D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A2807DA161F09D58C8F5B8B181115A0F">
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<mods:title id="CCDDD20283027EF41B05F1E66DCA3FDC">Two new species of Engertia Dalla Torre, 1913 and Philacelota Heller, 1900 from Indonesia and the Philippines with a revised identification key (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F28C472C0A39328F8323AF0D70D8A3E5" type="personal">
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<mods:roleTerm id="04C5DFD8042A5BE0294DC89201C607A9">Author</mods:roleTerm>
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<mods:namePart id="6E0300121B5B00135925DD060FEF2EFB">Prokofiev, Artem M.</mods:namePart>
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<mods:typeOfResource id="4A7234ACF2626724CDF3BCC2207B9681">text</mods:typeOfResource>
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<mods:title id="A7CAFBC86DCC60D6134D56D1EE2AC7A9">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="B331EFE7F28EBB118B963685D96CE3CA">
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<mods:date id="9FD061CAD9165177657D6C9AA7814550">2019</mods:date>
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<mods:number id="B99E72D9B8DD18E093486BA3982176AF">2019-10-03</mods:number>
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<mods:number id="7C7AE578744D3A90F7AC181707FFBD2E">561</mods:number>
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<mods:classification id="BBDBBDB2987E90F2013241F8D798D48F">journal article</mods:classification>
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<mods:identifier id="C75D1CA130E5D2EC7C85E941C9C8D0B3" type="DOI">10.5852/ejt.2019.561</mods:identifier>
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<treatment id="03E72F184007FFF7FD80516FFD71FB95" ID-DOI="http://doi.org/10.5281/zenodo.3475275" ID-GBIF-Taxon="159647021" ID-Zenodo-Dep="3475275" LSID="urn:lsid:zoobank.org:act:3E17E5EE-4309-4836-9A06-4AB5994E3F32" httpUri="http://treatment.plazi.org/id/03E72F184007FFF7FD80516FFD71FB95" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
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<subSubSection id="C354CD854007FFF0FD80516FFB27FD8D" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BF19E0E4007FFF0FD80516FFC2CFDA1" blockId="2.[375,1213,553,652]" box="[638,950,553,580]" pageId="2" pageNumber="3">
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<heading id="D0B929624007FFF0FD80516FFC2CFDA1" box="[638,950,553,580]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
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<taxonomicName id="4C4EE58D4007FFF0FD80516FFCCEFDA1" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[638,852,553,580]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="allolepis" status="sp. nov.">
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<emphasis id="B93A421C4007FFF0FD80516FFCCEFDA1" bold="true" box="[638,852,553,580]" italics="true" pageId="2" pageNumber="3">Engertia allolepis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A209FF674007FFF0FCA5516FFC2CFDA1" box="[859,950,554,580]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BF19E0E4007FFF0FE895108FB27FD8D" blockId="2.[375,1213,553,652]" box="[375,1213,589,616]" pageId="2" pageNumber="3">
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<uri id="FFDF920C4007FFF0FE895108FB27FD8D" box="[375,1213,589,616]" pageId="2" pageNumber="3">
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urn:lsid:zoobank.org:act:
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<uuid id="FFE8A4DB4007FFF0FD695108FB27FD8D" box="[663,1213,589,616]" pageId="2" pageNumber="3">3E17E5EE-4309-4836-9A06-4AB5994E3F32</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C354CD854007FFF0FD345134FCF3FD69" box="[714,873,625,652]" pageId="2" pageNumber="3" type="description">
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<paragraph id="8BF19E0E4007FFF0FD345134FCF3FD69" blockId="2.[375,1213,553,652]" box="[714,873,625,652]" pageId="2" pageNumber="3">
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<figureCitation id="1375828B4007FFF0FD345134FC89FD69" box="[714,787,625,652]" captionStart="Fig" captionStartId="3.[189,232,2002,2028]" captionTargetBox="[637,953,1294,1977]" captionTargetId="figure@3.[631,961,1290,1987]" captionTargetPageId="3" captionText="Fig. 1. Engertia allolepis sp. nov., habitus, holotype, ♂ (MNHN). Scale bar = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474361" httpUri="https://zenodo.org/record/3474361/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
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,
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<figureCitation id="1375828B4007FFF0FCDF5137FCB5FD69" box="[801,815,626,652]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="2" pageNumber="3">2</figureCitation>
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A–G
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</paragraph>
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</subSubSection>
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<subSubSection id="C354CD854007FFF0FF4351F6FA9EFC69" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="8BF19E0E4007FFF0FF4351F6FEACFD28" blockId="2.[189,310,691,717]" box="[189,310,691,717]" pageId="2" pageNumber="3">
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<heading id="D0B929624007FFF0FF4351F6FEACFD28" bold="true" box="[189,310,691,717]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="B93A421C4007FFF0FF4351F6FEACFD28" bold="true" box="[189,310,691,717]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF19E0E4007FFF0FF4351A1FA9EFC69" blockId="2.[189,1399,739,909]" pageId="2" pageNumber="3">
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<emphasis id="B93A421C4007FFF0FF4351A1FEB1FD18" box="[189,299,740,765]" italics="true" pageId="2" pageNumber="3">
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<taxonomicName id="4C4EE58D4007FFF0FF4351A1FEBCFD18" authorityName="Dalla Torre" authorityYear="1913" box="[189,294,740,765]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Engertia</taxonomicName>
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,
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</emphasis>
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with five lamellae in antennal club of males. Disc of pronotum with irregular callose areas. Elytra with glabrous costae and scale pattern forming longitudinal bands on interstices; scales on elytra lanceolate in 1
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<superScript id="7C3B33464007FFF0FE96506CFEE9FCD2" attach="left" box="[360,371,809,823]" fontSize="6" pageId="2" pageNumber="3">st</superScript>
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to 3
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<superScript id="7C3B33464007FFF0FE56506CFE2CFCD2" attach="left" box="[424,438,809,823]" fontSize="6" pageId="2" pageNumber="3">rd</superScript>
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interstices, but intermixed with setiform scales on 4
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<superScript id="7C3B33464007FFF0FBE2506CFBB3FCD2" attach="left" box="[1052,1065,809,823]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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and 5
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<superScript id="7C3B33464007FFF0FB8F506CFBE4FCD2" attach="left" box="[1137,1150,809,823]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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interstices and along apical margin of elytra. Propygidium with dense long setae; pygidium fully scaled. Abdominal ventrites 2 to 4 with isolated round patches of scales along midline. Aedeagus strongly arcuate, robust.
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</paragraph>
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</subSubSection>
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<subSubSection id="C354CD854007FFF0FF4350F0FCEDFBC7" pageId="2" pageNumber="3" type="etymology">
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<paragraph id="8BF19E0E4007FFF0FF4350F0FEDEFC2A" blockId="2.[189,324,949,975]" box="[189,324,949,975]" pageId="2" pageNumber="3">
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<heading id="D0B929624007FFF0FF4350F0FEDEFC2A" bold="true" box="[189,324,949,975]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="B93A421C4007FFF0FF4350F0FEDEFC2A" bold="true" box="[189,324,949,975]" pageId="2" pageNumber="3">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF19E0E4007FFF0FF4350A1FCEDFBC7" blockId="2.[189,1398,996,1058]" pageId="2" pageNumber="3">
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The species epithet is based on the Greek words ‘
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<emphasis id="B93A421C4007FFF0FCD550A1FCFEFC1B" box="[811,868,996,1022]" italics="true" pageId="2" pageNumber="3">allos</emphasis>
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’, meaning ‘different’, and ‘
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<emphasis id="B93A421C4007FFF0FB4350A1FB6EFC1B" box="[1213,1268,996,1022]" italics="true" pageId="2" pageNumber="3">lepis</emphasis>
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’, meaning ‘scale’, due to the heterogeneous scales covering the elytra.
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</paragraph>
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</subSubSection>
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<subSubSection id="C354CD854007FFF0FF43570FFBECFB52" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="8BF19E0E4007FFF0FF43570FFEF6FB81" blockId="2.[189,364,1098,1124]" box="[189,364,1098,1124]" pageId="2" pageNumber="3">
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<heading id="D0B929624007FFF0FF43570FFEF6FB81" bold="true" box="[189,364,1098,1124]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="B93A421C4007FFF0FF43570FFEF6FB81" bold="true" box="[189,364,1098,1124]" pageId="2" pageNumber="3">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF19E0E4007FFF0FF14573CFEC6FB76" blockId="2.[189,1142,1145,1207]" box="[234,348,1145,1171]" pageId="2" pageNumber="3">
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<emphasis id="B93A421C4007FFF0FF14573CFEC6FB76" bold="true" box="[234,348,1145,1171]" pageId="2" pageNumber="3">
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<typeStatus id="54F520AC4007FFF0FF14573CFEC6FB76" box="[234,348,1145,1171]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BF19E0E4007FFF0FF4357D8FBECFB52" blockId="2.[189,1142,1145,1207]" box="[189,1142,1181,1207]" pageId="2" pageNumber="3">
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<materialsCitation id="3B2694534007FFF0FF4357D8FBECFB52" ID-GBIF-Occurrence="2424121667" box="[189,1142,1181,1207]" collectionCode="MNHN" country="Indonesia" location="INDONESIA" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<collectingCountry id="F359DE9E4007FFF0FF4357D8FEC1FB52" box="[189,347,1181,1207]" name="Indonesia" pageId="2" pageNumber="3">INDONESIA</collectingCountry>
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•
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<specimenCount id="9D4855874007FFF0FE8A57D8FE17FB52" box="[372,397,1181,1207]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
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; “Molucc. Ambon / Has / Ex Musaeo van Lansberge”;
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<collectionCode id="ED5F06CB4007FFF0FBEA57D8FBE8FB52" box="[1044,1138,1181,1207]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C354CD854007FFF7FF43579AFEA3FEAC" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" type="description">
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<paragraph id="8BF19E0E4007FFF0FF43579AFE4AFB1F" blockId="2.[189,464,1247,1274]" box="[189,464,1247,1274]" pageId="2" pageNumber="3">
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<emphasis id="B93A421C4007FFF0FF43579AFED4FB1C" bold="true" box="[189,334,1247,1273]" pageId="2" pageNumber="3">Description</emphasis>
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(
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<typeStatus id="54F520AC4007FFF0FEA3579AFE5DFB1F" box="[349,455,1247,1274]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
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)
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</paragraph>
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<paragraph id="8BF19E0E4007FFF0FF14564BFEB0FACD" blockId="2.[189,1398,1294,1393]" box="[234,298,1294,1320]" pageId="2" pageNumber="3">
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<emphasis id="B93A421C4007FFF0FF14564BFEB0FACD" bold="true" box="[234,298,1294,1320]" pageId="2" pageNumber="3">Male</emphasis>
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</paragraph>
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<paragraph id="8BF19E0E4007FFF0FF435676FB03FA95" blockId="2.[189,1398,1294,1393]" pageId="2" pageNumber="3">
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BODY (
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<figureCitation id="1375828B4007FFF0FEEA5677FEC4FAA9" box="[276,350,1330,1357]" captionStart="Fig" captionStartId="3.[189,232,2002,2028]" captionTargetBox="[637,953,1294,1977]" captionTargetId="figure@3.[631,961,1290,1987]" captionTargetPageId="3" captionText="Fig. 1. Engertia allolepis sp. nov., habitus, holotype, ♂ (MNHN). Scale bar = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474361" httpUri="https://zenodo.org/record/3474361/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
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). Length
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<quantity id="4CB633EB4007FFF0FE215677FDCAFAA8" box="[479,592,1330,1357]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="2" pageNumber="3" unit="mm" value="18.5">18.5 mm</quantity>
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, greatest width
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<quantity id="4CB633EB4007FFF0FCE55677FCE4FAA8" box="[795,894,1330,1357]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="3" unit="mm" value="7.5">7.5 mm</quantity>
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. Integument dark reddish-brown; vertex, pronotum and tarsal claws more infuscated; scales and pilosity yellowish to whitish.
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</paragraph>
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<paragraph id="8BF19E0E4007FFF0FF4356DCFE08F8D8" blockId="2.[189,1399,1432,1853]" pageId="2" pageNumber="3">
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HEAD. Clypeus pentagonal, with anterior margin weakly bisinuate in frontal view, slightly produced in middle and conspicuously reflexed; surface rugo-punctate; punctures moderately deep, bearing narrowly lanceolate to spiniform scales becoming much shorter and thinner along midline in posterior half of clypeus. Fronto-clypeal suture conspicuous, weakly concave. Frons rugo-punctate; punctures much denser along inner margins of eyes, bearing lanceolate scales replaced by moderately long erect setae on lateral thirds of anterior half-length of frons; vertex glabrous. Antennae consist of ten antennomeres, with five antennomeres in club (
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<figureCitation id="1375828B4007FFF0FDBA552AFD3CF96F" box="[580,678,1647,1674]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="2" pageNumber="3">Fig. 2A</figureCitation>
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); all antennomeres of club of equal length; 5
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<superScript id="7C3B33464007FFF0FB36552BFB4FF999" attach="left" box="[1224,1237,1646,1660]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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antennomere half as long as 4
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<superScript id="7C3B33464007FFF0FE8355D7FE10F945" attach="none" box="[381,394,1682,1696]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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, 3
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<superScript id="7C3B33464007FFF0FE5655D7FE2CF945" attach="left" box="[424,438,1682,1696]" fontSize="6" pageId="2" pageNumber="3">rd</superScript>
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and 4
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<superScript id="7C3B33464007FFF0FE0155D7FD96F945" attach="left" box="[511,524,1682,1696]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
|
||
antennomeres of equal length; antennomeres of club strongly shagreened, antennomeres of funicle glabrous. Last maxillary palpomere elongate fusiform, with narrow, but deep impression on its outer side. Labrum weakly bilobed, covered by moderately long setae intermixed at sides with narrowly lanceolate scales; mentum deeply concave, bearing long setae along its lateral and posterior margins.
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4007FFF1FF435423FC72FC7A" blockId="2.[189,1398,1893,2027]" lastBlockId="3.[189,1399,267,928]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
THORAX AND ABDOMEN. Pronotum narrower at base than base of elytra, 1.5 times as broad as long, broadest just before middle, strongly and almost equally convergent anteriad and posteriad; sides straight anteriad, weakly concave posteriad; anterior and posterior angles obtuse; anterior margin bisinuate; posterior margin strongly convex before scutellar shield; all sides of pronotum unbeaded. Pronotum roughly, but not deeply and very unevenly punctured, punctures slightly rugose with interspaces between punctures on disc somewhat callose; glabrous and callose medial longitudinal stripe present; scales on pronotum mostly lanceolate, but intermixed on disc with few setiform scales; moderately long setae present along anterior margin and near anterior angles of pronotum only. Propleura setose, with dense patch of lanceolate scales in their anterior halves. Scutellar shield broadly rounded apically, only punctured along lateral sides; punctures bearing narrowly lanceolate scales. Elytra with five conspicuous glabrous costae; base of elytron somewhat swollen; humeral and apical umbones rather weakly expressed. Interstices shallowly, but densely punctured; punctures of 1
|
||
<superScript id="7C3B33464006FFF1FCFC5145FC97FDEB" attach="left" box="[770,781,512,526]" fontSize="6" pageId="3" pageNumber="4">st</superScript>
|
||
to 3
|
||
<superScript id="7C3B33464006FFF1FCBB5145FCC9FDEB" attach="left" box="[837,851,512,526]" fontSize="6" pageId="3" pageNumber="4">rd</superScript>
|
||
interstices bearing lanceolate scales, partially replaced by setiform scales in 4
|
||
<superScript id="7C3B33464006FFF1FDBD5166FDCAFDD4" attach="left" box="[579,592,547,561]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
and 5
|
||
<superScript id="7C3B33464006FFF1FD5E5166FD37FDD4" attach="left" box="[672,685,547,561]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
interstices and along apical margin of elytra (
|
||
<figureCitation id="1375828B4006FFF1FB185160FAAAFDDA" box="[1254,1328,549,575]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
B–C). Propygidium finely and densely punctured; punctures setigerous, bearing long setae; distal margin of propygidium with few isolated lanceolate scales (
|
||
<figureCitation id="1375828B4006FFF1FCE8512EFCEDFD60" box="[790,887,619,645]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="3" pageNumber="4">Fig. 2C</figureCitation>
|
||
). Pygidium densely covered by lanceolate scales (
|
||
<figureCitation id="1375828B4006FFF1FEE851CBFEECFD4C" box="[278,374,654,681]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="3" pageNumber="4">Fig. 2D</figureCitation>
|
||
), setose along lateral and apical margins. Prosternum, mesoventrite and metaventrite densely and finely rugo-punctate, densely covered with long setae; disc of metaventrite, metepimeron and metepisternum bearing some scales. First abdominal ventrite flat in middle; 1
|
||
<superScript id="7C3B33464006FFF1FB625191FB3DFD07" attach="left" box="[1180,1191,724,738]" fontSize="6" pageId="3" pageNumber="4">st</superScript>
|
||
and 2
|
||
<superScript id="7C3B33464006FFF1FB085196FA9DFD04" attach="left" box="[1270,1287,723,737]" fontSize="6" pageId="3" pageNumber="4">nd</superScript>
|
||
ventrites setose, remaining ventrites with isolated setae in their lateral thirds only, except last visible ventrite bearing rather dense setae of moderate length along its distal margin; all abdominal ventrites covered by ovate (lanceolate in uppermost parts) scales, more densely aggregated in uppermost reaches of each ventrite, forming small round patches of scales on 2
|
||
<superScript id="7C3B33464006FFF1FCEF5025FCB8FC8B" attach="left" box="[785,802,864,878]" fontSize="6" pageId="3" pageNumber="4">nd</superScript>
|
||
to 4
|
||
<superScript id="7C3B33464006FFF1FCAD5025FCFAFC8B" attach="left" box="[851,864,864,878]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
ventrites along midline of abdomen (
|
||
<figureCitation id="1375828B4006FFF1FAF15027FAF3FC99" box="[1295,1385,866,892]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="3" pageNumber="4">Fig. 2E</figureCitation>
|
||
); sutures between 2
|
||
<superScript id="7C3B33464006FFF1FE7150C6FE3AFC74" attach="left" box="[399,416,899,913]" fontSize="6" pageId="3" pageNumber="4">nd</superScript>
|
||
and 5
|
||
<superScript id="7C3B33464006FFF1FE1650C6FE6FFC74" attach="left" box="[488,501,899,913]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
abdominal ventrites obliterated in middle.
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4006FFF1FF435082FA85FB88" blockId="3.[189,1398,966,1134]" pageId="3" pageNumber="4">
|
||
LEGS. Protibiae unidentate, with spur short and thick; middle and hind tibiae gracile; apex of hind tibiae with 12 separated bristles. Femora setose; lanceolate scales present on hind femora only; tibiae with sparse narrow scales and spiniform setae; tarsi with setae only. Tarsomeres 1
|
||
<emphasis id="B93A421C4006FFF1FBA95748FBFCFBC2" bold="true" box="[1111,1126,1037,1063]" pageId="3" pageNumber="4">–</emphasis>
|
||
4 shortened, especially of fore tarsi; 1
|
||
<superScript id="7C3B33464006FFF1FE98576AFEEBFBD8" attach="left" box="[358,369,1071,1085]" fontSize="6" pageId="3" pageNumber="4">st</superScript>
|
||
tarsomere of hind tarsi equal to 2
|
||
<superScript id="7C3B33464006FFF1FCFF576BFC88FBD9" attach="left" box="[769,786,1070,1084]" fontSize="6" pageId="3" pageNumber="4">nd</superScript>
|
||
in length. Claws bifurcate, with lower lobe situated slightly closer to base of claw, parallel to upper one, distinctly shorter, but as broad as the latter.
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4006FFF1FF4357D0FBE2FB10" blockId="3.[189,1399,1173,1270]" pageId="3" pageNumber="4">
|
||
GENITALIA. Aedeagus (
|
||
<figureCitation id="1375828B4006FFF1FE3957D0FDD2FB4A" box="[455,584,1173,1199]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="3" pageNumber="4">Fig. 2E, G</figureCitation>
|
||
) strongly arcuate, robust; phallobase remarkably deepened distally; parameres considerably narrowed toward apex, their ventral contour very weakly arcuate and dorsal contour almost straight; apical extensions of parameres small, oval, not upturned.
|
||
</paragraph>
|
||
<caption id="DF31CE864006FFF1FF435497FB2EF808" ID-DOI="http://doi.org/10.5281/zenodo.3474361" ID-Zenodo-Dep="3474361" box="[189,1204,2002,2029]" httpUri="https://zenodo.org/record/3474361/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,2002,2028]" targetBox="[637,953,1294,1977]" targetPageId="3">
|
||
<paragraph id="8BF19E0E4006FFF1FF435497FB2EF808" blockId="3.[189,1204,2002,2029]" box="[189,1204,2002,2029]" pageId="3" pageNumber="4">
|
||
<emphasis id="B93A421C4006FFF1FF435497FE91F809" bold="true" box="[189,267,2002,2028]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
|
||
<taxonomicName id="4C4EE58D4006FFF1FEEC5496FE7FF809" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[274,485,2002,2028]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="allolepis" status="sp. nov.">
|
||
<emphasis id="B93A421C4006FFF1FEEC5496FE7FF809" box="[274,485,2002,2028]" italics="true" pageId="3" pageNumber="4">Engertia allolepis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF674006FFF1FE125496FDD3F808" box="[492,585,2003,2029]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, habitus, holotype, ♂ (MNHN). Scale bar = 2.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF31CE864001FFF6FF43553BFC05F889" ID-DOI="http://doi.org/10.5281/zenodo.3474363" ID-Zenodo-Dep="3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1662,1688]" targetBox="[201,1389,302,1606]" targetPageId="4">
|
||
<paragraph id="8BF19E0E4001FFF6FF43553BFC05F889" blockId="4.[189,1399,1662,1900]" pageId="4" pageNumber="5">
|
||
<emphasis id="B93A421C4001FFF6FF43553BFED3F97D" bold="true" box="[189,329,1662,1688]" pageId="4" pageNumber="5">Fig. 2. A–G</emphasis>
|
||
.
|
||
<taxonomicName id="4C4EE58D4001FFF6FEAB553AFDBCF97D" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[341,550,1662,1688]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="allolepis" status="sp. nov.">
|
||
<emphasis id="B93A421C4001FFF6FEAB553AFDBCF97D" box="[341,550,1662,1688]" italics="true" pageId="4" pageNumber="5">Engertia allolepis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF674001FFF6FDD2553AFD1DF97C" box="[556,647,1663,1689]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (MNHN).
|
||
<emphasis id="B93A421C4001FFF6FC67553BFC34F97D" bold="true" box="[921,942,1662,1688]" pageId="4" pageNumber="5">A</emphasis>
|
||
. Antenna.
|
||
<emphasis id="B93A421C4001FFF6FBD7553AFBA7F97C" bold="true" box="[1065,1085,1663,1689]" pageId="4" pageNumber="5">B</emphasis>
|
||
. Lateral part of the elytron.
|
||
<emphasis id="B93A421C4001FFF6FF4355E4FF48F95E" bold="true" box="[189,210,1697,1723]" pageId="4" pageNumber="5">C</emphasis>
|
||
. Apical margin of elytra and propygidium.
|
||
<emphasis id="B93A421C4001FFF6FD2955E7FD76F959" bold="true" box="[727,748,1698,1724]" pageId="4" pageNumber="5">D</emphasis>
|
||
. Pygidium.
|
||
<emphasis id="B93A421C4001FFF6FC8355E7FC0BF959" bold="true" box="[893,913,1698,1724]" pageId="4" pageNumber="5">E</emphasis>
|
||
. Abdominal ventrites 2–4.
|
||
<emphasis id="B93A421C4001FFF6FB2D55E7FB7FF959" bold="true" box="[1235,1253,1698,1724]" pageId="4" pageNumber="5">F</emphasis>
|
||
. Parameres, dorsal view.
|
||
<emphasis id="B93A421C4001FFF6FEA65580FEF5F93A" bold="true" box="[344,367,1733,1759]" pageId="4" pageNumber="5">G</emphasis>
|
||
. Aedeagus, lateral view. —
|
||
<emphasis id="B93A421C4001FFF6FD395580FD61F93A" bold="true" box="[711,763,1733,1759]" pageId="4" pageNumber="5">H–J</emphasis>
|
||
.
|
||
<taxonomicName id="4C4EE58D4001FFF6FCF35583FBD7F93A" authority="(Moser, 1913)" authorityName="Moser" authorityYear="1913" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[781,1101,1733,1759]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis id="B93A421C4001FFF6FCF35583FC0DF93A" box="[781,919,1734,1759]" italics="true" pageId="4" pageNumber="5">E. papuana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF4001FFF6FC575580FBDEF93A" author="Moser" box="[937,1092,1733,1759]" firstAuthor="Moser" pageId="4" pageNumber="5" pagination="271 - 297" refId="ref5971" refString="Moser J. 1913. Beitrag zur Kenntnis der Melolonthiden I. Deutsche entomologische Zeitschrift 3: 271 - 297. Available from https: // archive. org / details / biostor- 101429 [accessed 10 Sep. 2019]." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♂, from Sentani (cAP).
|
||
<emphasis id="B93A421C4001FFF6FF4355ADFF4EF8E7" bold="true" box="[189,212,1768,1794]" pageId="4" pageNumber="5">H</emphasis>
|
||
. Lateral part of elytron.
|
||
<emphasis id="B93A421C4001FFF6FDFE55ADFD91F8E7" bold="true" box="[512,523,1768,1794]" pageId="4" pageNumber="5">I</emphasis>
|
||
. Apical margin of elytra and propygidium.
|
||
<emphasis id="B93A421C4001FFF6FBE555ADFBB0F8E7" bold="true" box="[1051,1066,1768,1794]" pageId="4" pageNumber="5">J</emphasis>
|
||
. Aedeagus, lateral view. —
|
||
<emphasis id="B93A421C4001FFF6FF43544EFF4EF8C0" bold="true" box="[189,212,1803,1829]" pageId="4" pageNumber="5">K</emphasis>
|
||
.
|
||
<taxonomicName id="4C4EE58D4001FFF6FF1B5449FDD2F8C0" authority="(Brenske, 1897)" authorityName="Brenske" authorityYear="1897" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[229,584,1803,1829]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="amboinae">
|
||
<emphasis id="B93A421C4001FFF6FF1B5449FEE6F8C0" box="[229,380,1803,1829]" italics="true" pageId="4" pageNumber="5">E. amboinae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF4001FFF6FE70544EFDA5F8C0" author="Brenske" box="[398,575,1803,1829]" firstAuthor="Brenske" pageId="4" pageNumber="5" pagination="96 - 120" refId="ref5871" refString="Brenske E. 1897. Neue Gattungen und Arten der Melolonthiden aus Afrika und Asien. Stettiner entomologische Zeitung 58: 96 - 120." type="journal article" year="1897">Brenske, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♂, from Ambon (MNHN), lateral part of elytron. —
|
||
<emphasis id="B93A421C4001FFF6FB29544EFB71F8C0" bold="true" box="[1239,1259,1803,1829]" pageId="4" pageNumber="5">L</emphasis>
|
||
.
|
||
<taxonomicName id="4C4EE58D4001FFF6FB055449FEFCF8AC" authority="(Moser, 1913)" authorityName="Moser" authorityYear="1913" baseAuthorityName="Moser" baseAuthorityYear="1913" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="setifera">
|
||
<emphasis id="B93A421C4001FFF6FB055449FAECF8C0" box="[1275,1398,1803,1829]" italics="true" pageId="4" pageNumber="5">E. setifera</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF4001FFF6FF3B546AFEC7F8AC" author="Moser" box="[197,349,1839,1865]" firstAuthor="Moser" pageId="4" pageNumber="5" pagination="271 - 297" refId="ref5971" refString="Moser J. 1913. Beitrag zur Kenntnis der Melolonthiden I. Deutsche entomologische Zeitschrift 3: 271 - 297. Available from https: // archive. org / details / biostor- 101429 [accessed 10 Sep. 2019]." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF19E0E4000FFF7FF14524EFEDFFEC0" blockId="5.[189,325,267,329]" box="[234,325,267,293]" pageId="5" pageNumber="6">
|
||
<emphasis id="B93A421C4000FFF7FF14524EFEDFFEC0" bold="true" box="[234,325,267,293]" pageId="5" pageNumber="6">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FF43526AFEA3FEAC" blockId="5.[189,325,267,329]" box="[189,313,303,329]" pageId="5" pageNumber="6">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354CD854000FFF7FF435237FD71FB95" pageId="5" pageNumber="6" type="diagnosis">
|
||
<paragraph id="8BF19E0E4000FFF7FF435237FE53FE69" blockId="5.[189,457,370,396]" box="[189,457,370,396]" pageId="5" pageNumber="6">
|
||
<heading id="D0B929624000FFF7FF435237FE53FE69" bold="true" box="[189,457,370,396]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="B93A421C4000FFF7FF435237FE53FE69" bold="true" box="[189,457,370,396]" pageId="5" pageNumber="6">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FF4352E6FEA8FCC9" blockId="5.[189,1399,418,812]" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C4EE58D4000FFF7FF4352E6FE0AFE58" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[189,400,419,445]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="allolepis" status="sp. nov.">
|
||
<emphasis id="B93A421C4000FFF7FF4352E6FE0AFE58" box="[189,400,419,445]" italics="true" pageId="5" pageNumber="6">Engertia allolepis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF674000FFF7FE6652E6FE69FE58" box="[408,499,419,445]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is unique within the genus in the possession of both lanceolate and setiform scales in the outer elytral interstices (vs elytral scales more or less homogeneous in shape in all interstices in the other
|
||
<taxonomicName id="4C4EE58D4000FFF7FEAF52A8FE22FDE3" authorityName="Dalla Torre" authorityYear="1913" box="[337,440,493,518]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93A421C4000FFF7FEAF52A8FE22FDE3" box="[337,440,493,518]" italics="true" pageId="5" pageNumber="6">Engertia</emphasis>
|
||
</taxonomicName>
|
||
spp.; see
|
||
<figureCitation id="1375828B4000FFF7FDCC52A9FD14FDE3" box="[562,654,492,518]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="5" pageNumber="6">Fig. 2H</figureCitation>
|
||
for representation). In other respects this new species is most similar to
|
||
<taxonomicName id="4C4EE58D4000FFF7FECC5157FDFDFDCE" authority="(Moser, 1913)" authorityName="Moser" authorityYear="1913" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[306,615,529,555]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis id="B93A421C4000FFF7FECC5157FE22FDCE" box="[306,440,530,555]" italics="true" pageId="5" pageNumber="6">E. papuana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF4000FFF7FE395154FDC4FDCE" author="Moser" box="[455,606,529,555]" firstAuthor="Moser" pageId="5" pageNumber="6" pagination="271 - 297" refId="ref5971" refString="Moser J. 1913. Beitrag zur Kenntnis der Melolonthiden I. Deutsche entomologische Zeitschrift 3: 271 - 297. Available from https: // archive. org / details / biostor- 101429 [accessed 10 Sep. 2019]." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but differs from the latter in the much longer and denser pilosity of the propygidium (compare
|
||
<figureCitation id="1375828B4000FFF7FE095170FD01FDAA" box="[503,667,565,591]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="5" pageNumber="6">
|
||
Fig. 2C and
|
||
<date id="FFF0B8CE4000FFF7FD7C5170FD01FDAA" box="[642,667,565,591]" pageId="5" pageNumber="6">2I</date>
|
||
</figureCitation>
|
||
) and in the somewhat callose disc of the pronotum. The aedeagi of
|
||
<taxonomicName id="4C4EE58D4000FFF7FF20511EFEFFFD91" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[222,357,602,628]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="allolepis" status="sp. nov.">
|
||
<emphasis id="B93A421C4000FFF7FF20511EFEFFFD91" box="[222,357,602,628]" italics="true" pageId="5" pageNumber="6">E. allolepis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF674000FFF7FE90511EFE57FD90" box="[366,461,603,629]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C4EE58D4000FFF7FE24511EFCA0FD91" authority="(Brenske, 1897)" authorityName="Brenske" authorityYear="1897" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[474,826,602,628]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="amboinae">
|
||
<emphasis id="B93A421C4000FFF7FE24511EFDEAFD91" box="[474,624,602,628]" italics="true" pageId="5" pageNumber="6">E. amboinae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF4000FFF7FD7F511FFCAAFD91" author="Brenske" box="[641,816,602,628]" firstAuthor="Brenske" pageId="5" pageNumber="6" pagination="96 - 120" refId="ref5871" refString="Brenske E. 1897. Neue Gattungen und Arten der Melolonthiden aus Afrika und Asien. Stettiner entomologische Zeitung 58: 96 - 120." type="journal article" year="1897">Brenske, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EE58D4000FFF7FCB9511EFB34FD91" authority="Prokofiev, 2016" authorityName="Prokofiev" authorityYear="2016" box="[839,1198,602,628]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="germanica">
|
||
<emphasis id="B93A421C4000FFF7FCB9511EFC7CFD91" box="[839,998,603,628]" italics="true" pageId="5" pageNumber="6">E. germanica</emphasis>
|
||
<bibRefCitation id="EFDFE3FF4000FFF7FC11511FFB34FD91" author="Prokofiev A. M." box="[1007,1198,602,628]" journalOrPublisher="Euroasian Entomological Journal" pageId="5" pageNumber="6" pagination="68 - 74" part="15" refId="ref6048" refString="Prokofiev A. M. 2016 a. New genera and species of the Papuan Leucopholini and Melolonthini (Coleoptera, Scarabaeidae: Melolonthinae). Euroasian Entomological Journal 15: 68 - 74. [In Russian.]" title="New genera and species of the Papuan Leucopholini and Melolonthini (Coleoptera, Scarabaeidae: Melolonthinae)" type="journal article" year="2016">Prokofiev, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EE58D4000FFF7FB45511EFADEFD91" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[1211,1348,603,628]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis id="B93A421C4000FFF7FB45511EFADEFD91" box="[1211,1348,603,628]" italics="true" pageId="5" pageNumber="6">E. papuana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EE58D4000FFF7FF4351C5FE78FD7C" authority="(Moser, 1913)" authorityName="Moser" authorityYear="1913" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[189,482,639,665]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="setifera">
|
||
<emphasis id="B93A421C4000FFF7FF4351C5FEAFFD7C" box="[189,309,639,665]" italics="true" pageId="5" pageNumber="6">E. setifera</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF4000FFF7FEBA513AFE40FD7C" author="Moser" box="[324,474,639,665]" firstAuthor="Moser" pageId="5" pageNumber="6" pagination="271 - 297" refId="ref5971" refString="Moser J. 1913. Beitrag zur Kenntnis der Melolonthiden I. Deutsche entomologische Zeitschrift 3: 271 - 297. Available from https: // archive. org / details / biostor- 101429 [accessed 10 Sep. 2019]." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
are very similar to each other, but the aedeagus of
|
||
<taxonomicName id="4C4EE58D4000FFF7FBCD51C5FB22FD7C" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[1075,1208,639,665]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="allolepis" status="sp. nov.">
|
||
<emphasis id="B93A421C4000FFF7FBCD51C5FB22FD7C" box="[1075,1208,639,665]" italics="true" pageId="5" pageNumber="6">E. allolepis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF674000FFF7FB41513AFA83FD7C" box="[1215,1305,639,665]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is more arched in lateral view than that of the aforementioned species except
|
||
<taxonomicName id="4C4EE58D4000FFF7FC1C51E1FBE4FD58" authorityName="Prokofiev" authorityYear="2016" box="[994,1150,676,701]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="germanica">
|
||
<emphasis id="B93A421C4000FFF7FC1C51E1FBE4FD58" box="[994,1150,676,701]" italics="true" pageId="5" pageNumber="6">E. germanica</emphasis>
|
||
</taxonomicName>
|
||
, and it can be further distinguished from the aedeagi of all the other species mentioned above in the deeper and broader apical portion of the phallobase and in the more robust and massive parameres (compare
|
||
<figureCitation id="1375828B4000FFF7FB8151A8FB43FCE2" box="[1151,1241,749,775]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="5" pageNumber="6">Fig. 2G</figureCitation>
|
||
with 2J as an example).
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FF435010FD71FB95" blockId="5.[189,1399,852,1136]" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C4EE58D4000FFF7FF435010FE08FC8A" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[189,402,853,879]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="allolepis" status="sp. nov.">
|
||
<emphasis id="B93A421C4000FFF7FF435010FE08FC8A" box="[189,402,853,879]" italics="true" pageId="5" pageNumber="6">Engertia allolepis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF674000FFF7FE625010FE63FC8A" box="[412,505,853,879]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is known from
|
||
<collectingCountry id="F359DE9E4000FFF7FD475011FCFCFC8B" box="[697,870,852,878]" name="Ambon Island" pageId="5" pageNumber="6">Ambon Island</collectingCountry>
|
||
, where two other members of the genus are distributed:
|
||
<taxonomicName id="4C4EE58D4000FFF7FEB4503FFE44FC76" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[330,478,889,915]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="amboinae">
|
||
<emphasis id="B93A421C4000FFF7FEB4503FFE44FC76" box="[330,478,889,915]" italics="true" pageId="5" pageNumber="6">E. amboinae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EE58D4000FFF7FDE9503FFD15FC76" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[535,655,889,915]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="setifera">
|
||
<emphasis id="B93A421C4000FFF7FDE9503FFD15FC76" box="[535,655,889,915]" italics="true" pageId="5" pageNumber="6">E. setifera</emphasis>
|
||
</taxonomicName>
|
||
. Besides the heterogeneous vestiture of the elytra and the more robust aedeagus, the new species can be distinguished from
|
||
<taxonomicName id="4C4EE58D4000FFF7FC6950DAFBB4FC5D" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[919,1070,926,952]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="amboinae">
|
||
<emphasis id="B93A421C4000FFF7FC6950DAFBB4FC5D" box="[919,1070,926,952]" italics="true" pageId="5" pageNumber="6">E. amboinae</emphasis>
|
||
</taxonomicName>
|
||
by the scales in the elytral interstices being much narrower (lanceolate to setiform vs ovoid:
|
||
<figureCitation id="1375828B4000FFF7FC385086FBBEFC38" box="[966,1060,963,989]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="5" pageNumber="6">Fig. 2K</figureCitation>
|
||
), scales along midline of the abdominal ventrites aggregated into spots (vs not aggregated), propygidium with long (vs short) setae, and from
|
||
<taxonomicName id="4C4EE58D4000FFF7FECA5748FE35FBC3" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[308,431,1036,1062]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="setifera">
|
||
<emphasis id="B93A421C4000FFF7FECA5748FE35FBC3" box="[308,431,1036,1062]" italics="true" pageId="5" pageNumber="6">E. setifera</emphasis>
|
||
</taxonomicName>
|
||
by the pygidium being densely covered with scales (vs mainly setose:
|
||
<figureCitation id="1375828B4000FFF7FAF25749FAF2FBC3" box="[1292,1384,1036,1062]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="5" pageNumber="6">Fig. 2L</figureCitation>
|
||
), scales along midline of the abdominal ventrites aggregated into spots (vs not aggregated), and scales present on labrum and metaventrite (vs absent).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |