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<mods:title id="9C991E28BA5223CC09E9689ECCF52322">Overview of the bee genus Trachusa Panzer, 1804 (Hymenoptera: Apoidea Megachilidae: Anthidiini) from China with description of three new species</mods:title>
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<mods:namePart id="6EE93465A8EDA9ED6AE9F5006F6BD116">Niu, Ze-Qing</mods:namePart>
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<mods:namePart id="8A9D79F215F2C4A44DA2A9C6619131A5">Griswold, Terry</mods:namePart>
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<mods:namePart id="BC5D3DD7066EEC5A01B5FBA350A0F15A">Zhu, Chao-Dong</mods:namePart>
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<subSubSection id="C36565601558BC1DEC87B451A24EFA03" box="[151,856,1390,1417]" pageId="4" pageNumber="255" type="nomenclature">
<paragraph id="8BC036EB1558BC1DEC87B451A24EFA03" blockId="4.[151,856,1390,1453]" box="[151,856,1390,1417]" pageId="4" pageNumber="255">
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<taxonomicName id="4C7F4D681558BC1DEC87B451A24EFA03" ID-CoL="57PMB" authority="(Mavromoustakis, 1953)" baseAuthorityName="Mavromoustakis" baseAuthorityYear="1953" box="[151,856,1390,1417]" class="Insecta" family="Megachilidae" genus="Trachusa" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="maai" subGenus="Paraanthidium">
<emphasis id="B90BEAF91558BC1DEC87B451A24EFA03" bold="true" box="[151,856,1390,1417]" pageId="4" pageNumber="255">
<emphasis id="B90BEAF91558BC1DEC87B451A01CFA02" bold="true" box="[151,266,1390,1417]" italics="true" pageId="4" pageNumber="255">Trachusa</emphasis>
(
<emphasis id="B90BEAF91558BC1DED0BB451A0C5FA02" bold="true" box="[283,467,1390,1417]" italics="true" pageId="4" pageNumber="255">Paraanthidium</emphasis>
)
<emphasis id="B90BEAF91558BC1DEDF4B451A336FA03" bold="true" box="[484,544,1390,1416]" italics="true" pageId="4" pageNumber="255">maai</emphasis>
(
<bibRefCitation id="EFEE4B1A1558BC1DEE20B451A246FA03" author="Mavromoustakis, G. A." box="[560,848,1390,1416]" pageId="4" pageNumber="255" pagination="834 - 840" refId="ref11186" refString="Mavromoustakis, G. A. (1953) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VI. Annals and Magazine for Natural History, 12, Ser., 67, 834 - 840. https: // doi. org / 10.1080 / 00222935308654492." type="journal article" year="1953">Mavromoustakis, 1953</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36565601558BC1DEC87B4ADA090FA26" box="[151,390,1425,1453]" pageId="4" pageNumber="255" type="description">
<paragraph id="8BC036EB1558BC1DEC87B4ADA090FA26" blockId="4.[151,856,1390,1453]" box="[151,390,1425,1453]" pageId="4" pageNumber="255">
(
<figureCitation id="13442A6E1558BC1DECB0B4ADA1F3FA27" box="[160,229,1425,1452]" captionStart="FIGURE 5" captionStartId="9.[151,250,1959,1984]" captionTargetBox="[151,1433,212,1902]" captionTargetId="figure@9.[151,1436,206,1911]" captionTargetPageId="9" captionText="FIGURE 5. Trachusa (Paraanthidium) maai (Mavromoustakis, 1953), AE, female.A: Body in lateral view; B: Head in frontal view; C: Mesosoma in dorsal view; D: Fore wing in frontal view; E: Metasoma in dorsal view. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/3349435/files/figure.png" pageId="4" pageNumber="255">Fig. 5</figureCitation>
: AE, female)
</paragraph>
</subSubSection>
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<paragraph id="8BC036EB1558BC1DEC87B4E9A31BF9A7" blockId="4.[151,1437,1494,1641]" pageId="4" pageNumber="255">
<taxonomicName id="4C7F4D681558BC1DEC87B4E9A3DDFA64" authority="Mavromoustakis, 1953: 834 - 836" authorityName="Mavromoustakis" authorityPageNumber="834" authorityYear="1953" box="[151,715,1494,1519]" class="Insecta" family="Megachilidae" genus="Paraanthidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="maai">
<emphasis id="B90BEAF91558BC1DEC87B4E9A07BFA65" box="[151,365,1494,1518]" italics="true" pageId="4" pageNumber="255">Paraanthidium maai</emphasis>
<bibRefCitation id="EFEE4B1A1558BC1DED63B4E9A3DDFA64" author="Mavromoustakis, G. A." box="[371,715,1494,1519]" pageId="4" pageNumber="255" pagination="834 - 840" refId="ref11186" refString="Mavromoustakis, G. A. (1953) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VI. Annals and Magazine for Natural History, 12, Ser., 67, 834 - 840. https: // doi. org / 10.1080 / 00222935308654492." type="journal article" year="1953">Mavromoustakis, 1953: 834836</bibRefCitation>
</taxonomicName>
,
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.
<typeStatus id="54C488491558BC1DEEE9B4E9A24DFA64" box="[761,859,1494,1519]" pageId="4" pageNumber="255" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D79FD621558BC1DEF78B4E8A26DFA65" box="[872,891,1495,1518]" pageId="4" pageNumber="255" type="female"></specimenCount>
, “S.E.
<collectingCountry id="F368767B1558BC1DEFD9B4E9A51FFA64" box="[969,1033,1494,1519]" name="China" pageId="4" pageNumber="255">China</collectingCountry>
: Psingshanpu,
<specimenCount id="9D79FD621558BC1DE8B5B4E9A5E6FA65" box="[1189,1264,1494,1519]" pageId="4" pageNumber="255" type="generic">
<quantity id="4C879B0E1558BC1DE8B5B4E9A5FAFA65" box="[1189,1260,1494,1519]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="255" unit="m" value="600.0">600 m</quantity>
.
</specimenCount>
, Changtinghsien (Fukien Province), leg. Tsing-chao Maa” [Changdin Xian,
<collectingRegion id="49BBF8091558BC1DEF32B4CBA273F986" box="[802,869,1524,1549]" country="China" name="Fujian" pageId="4" pageNumber="255">Fujian</collectingRegion>
,
<collectingCountry id="F368767B1558BC1DEF60B4CBA2A7F986" box="[880,945,1524,1549]" name="China" pageId="4" pageNumber="255">China</collectingCountry>
], coll. of G. Mavromoustakis;
<bibRefCitation id="EFEE4B1A1558BC1DE8FCB4CBA039F9A7" author="Mavromoustakis, G. A." pageId="4" pageNumber="255" pagination="249 - 252" refId="ref11239" refString="Mavromoustakis, G. A. (1954) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VII. Annals and Magazine for Natural History, 12, Ser., 7, 249 - 252. https: // doi. org / 10.1080 / 00222935408651726." type="journal article" year="1954">Mavromoustakis, 1954: 250</bibRefCitation>
,
<specimenCount id="9D79FD621558BC1DED2AB72BA05BF9A0" box="[314,333,1556,1579]" pageId="4" pageNumber="255" type="male"></specimenCount>
(new description).
</paragraph>
<paragraph id="8BC036EB1558BC1DEC87B70DA578F9E2" blockId="4.[151,1437,1494,1641]" pageId="4" pageNumber="255">
<treatmentCitationGroup id="AB6F11C51558BC1DEC87B70DA578F9E2" pageId="4" pageNumber="255">
<taxonomicName id="4C7F4D681558BC1DEC87B70DA389F9C0" ID-CoL="57PMB" authority=": Pasteels, 1972: 82" authorityName="Pasteels" authorityPageNumber="82" authorityYear="1972" box="[151,671,1586,1611]" class="Insecta" family="Megachilidae" genus="Trachusa" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="maai" subGenus="Proanthidium">
<emphasis id="B90BEAF91558BC1DEC87B70DA1E3F9C1" box="[151,245,1586,1610]" italics="true" pageId="4" pageNumber="255">Trachusa</emphasis>
(
<emphasis id="B90BEAF91558BC1DED15B70DA087F9C1" box="[261,401,1586,1610]" italics="true" pageId="4" pageNumber="255">Proanthidium</emphasis>
)
<emphasis id="B90BEAF91558BC1DEDB1B70CA0C2F9C1" box="[417,468,1587,1610]" italics="true" pageId="4" pageNumber="255">maai</emphasis>
:
<treatmentCitation id="0ADE10FA1558BC1DEDF2B70DA389F9C0" author="Pasteels, J. J." box="[482,671,1586,1611]" page="82" pageId="4" pageNumber="255" year="1972">
<bibRefCitation id="EFEE4B1A1558BC1DEDF2B70DA389F9C0" author="Pasteels, J. J." box="[482,671,1586,1611]" pageId="4" pageNumber="255" pagination="72 - 128" refId="ref11663" refString="Pasteels, J. J. (1972) Revision des Anthidiinae (Hymenoptera, Apoidea) de la region Indo-Malaise. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 108, 72 - 128." type="journal article" year="1972">Pasteels, 1972: 82</bibRefCitation>
</treatmentCitation>
</taxonomicName>
,
<specimenCount id="9D79FD621558BC1DEEBAB70CA3ABF9C1" box="[682,701,1587,1610]" pageId="4" pageNumber="255" type="female"></specimenCount>
(redescription), Figs 2124 (assigned to
<emphasis id="B90BEAF91558BC1DE873B70DA5F9F9C1" box="[1123,1263,1586,1610]" italics="true" pageId="4" pageNumber="255">Proanthidium</emphasis>
, but listed under
<emphasis id="B90BEAF91558BC1DECD7B76EA04CF9E2" box="[199,346,1617,1641]" italics="true" pageId="4" pageNumber="255">Protanthidium</emphasis>
);
<treatmentCitation id="0ADE10FA1558BC1DED7FB76DA31EF9E2" author="Wu, Y. R." box="[367,520,1616,1641]" page="175" pageId="4" pageNumber="255" year="2006">
<bibRefCitation id="EFEE4B1A1558BC1DED7FB76DA31EF9E2" author="Wu, Y. R." box="[367,520,1616,1641]" pageId="4" pageNumber="255" refId="ref11919" refString="Wu, Y. R. (2006) Fauna Sinica Insecta Vol. 44. Hymenoptera. Megachilidae. Science Press, Beijing, xvi + 474 pp., 4 pls. [in Chinese]" type="book" year="2006">Wu, 2006: 175</bibRefCitation>
</treatmentCitation>
,
<specimenCount id="9D79FD621558BC1DEE02B76DA333F9E2" box="[530,549,1618,1641]" pageId="4" pageNumber="255" type="female"></specimenCount>
(description);
<treatmentCitation id="0ADE10FA1558BC1DEEAAB76FA298F9E2" author="Kasparek, M." box="[698,910,1616,1641]" page="118" pageId="4" pageNumber="255" year="2017">
<bibRefCitation id="EFEE4B1A1558BC1DEEAAB76FA298F9E2" author="Kasparek, M." box="[698,910,1616,1641]" pageId="4" pageNumber="255" pagination="1 - 151" refId="ref10897" refString="Kasparek, M. (2017) Resin bees of the anthidiine genus Trachusa: identification, taxonomy, distribution and biology of the old world species. Entomofauna, supplement 21, 1 - 151." type="journal article" year="2017">Kasparek, 2017: 118</bibRefCitation>
</treatmentCitation>
,
<specimenCount id="9D79FD621558BC1DEF89B76DA2BAF9E2" box="[921,940,1618,1641]" pageId="4" pageNumber="255" type="female"></specimenCount>
,
<specimenCount id="9D79FD621558BC1DEFA8B76DA2DDF9E2" box="[952,971,1618,1641]" pageId="4" pageNumber="255" type="male"></specimenCount>
(redescription).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36565601558BC1DEC87B7A8A59FF8ED" pageId="4" pageNumber="255" type="materials_examined">
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<emphasis id="B90BEAF91558BC1DEC87B7A8A092F939" bold="true" box="[151,388,1687,1714]" pageId="4" pageNumber="255">Material examined:</emphasis>
<materialsCitation id="3B173CB61558BC1DED9CB7A8A587F939" ID-GBIF-Occurrence="2294326321" box="[396,1169,1687,1714]" collectingDate="1983-07-12" country="China" latitude="25.7" location="Longsheng" longLatPrecision="1245" longitude="110.0" pageId="4" pageNumber="255" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi">
<collectingCountry id="F368767B1558BC1DED9CB7A8A0C3F939" box="[396,469,1687,1714]" name="China" pageId="4" pageNumber="255">
<emphasis id="B90BEAF91558BC1DED9CB7A8A0C3F939" bold="true" box="[396,469,1687,1714]" pageId="4" pageNumber="255">China</emphasis>
</collectingCountry>
:
<collectingRegion id="49BBF8091558BC1DEDF6B7A8A35FF93A" box="[486,585,1687,1713]" country="China" name="Guangxi" pageId="4" pageNumber="255">Guangxi</collectingRegion>
,
<location id="8EA060301558BC1DEE45B7A8A3C7F93A" LSID="urn:lsid:plazi:treatment:03D687FD1558BC1DEC87B451A59FF8ED:8EA060301558BC1DEE45B7A8A3C7F93A" box="[597,721,1687,1713]" country="China" latitude="25.7" longLatPrecision="1245" longitude="110.0" name="Longsheng" pageId="4" pageNumber="255" stateProvince="Guangxi">Longsheng</location>
(
<geoCoordinate id="EE4B502C1558BC1DEEF2B7A8A257F939" box="[738,833,1687,1714]" degrees="25" direction="north" minutes="42" orientation="latitude" pageId="4" pageNumber="255" precision="925" value="25.7">25º42N</geoCoordinate>
,
<geoCoordinate id="EE4B502C1558BC1DEF5EB7A8A2A1F939" box="[846,951,1687,1714]" degrees="110" direction="east" minutes="00" orientation="longitude" pageId="4" pageNumber="255" precision="925" value="110.0">110º00E</geoCoordinate>
),
<specimenCount id="9D79FD621558BC1DEFDBB7A7A2E0F939" box="[971,1014,1688,1714]" pageId="4" pageNumber="255" type="female">1 ♀</specimenCount>
,
<collectingDate id="EF85E9C31558BC1DE816B7A7A587F939" box="[1030,1169,1688,1714]" pageId="4" pageNumber="255" value="1983-07-12">
<date id="FFC1102B1558BC1DE816B7A7A587F939" box="[1030,1169,1688,1714]" pageId="4" pageNumber="255" value="1983-07-12">12.VII.1983</date>
</collectingDate>
</materialsCitation>
, leg. no information on label;
<materialsCitation id="3B173CB61558BC1DECCDB784A454F95D" ID-GBIF-Occurrence="2294326318" box="[221,1346,1723,1750]" collectingDate="1988-07-05" collectionCode="YANG" collectorName="Long-Long" country="China" elevation="1000" latitude="26.333334" location="Taojiang" longLatPrecision="1242" longitude="108.15" municipality="Leishan" pageId="4" pageNumber="255" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou">
<collectingRegion id="49BBF8091558BC1DECCDB784A056F95E" box="[221,320,1723,1749]" country="China" name="Guizhou" pageId="4" pageNumber="255">Guizhou</collectingRegion>
,
<collectingMunicipality id="6BA4AC911558BC1DED5DB784A0BFF95E" box="[333,425,1723,1749]" pageId="4" pageNumber="255">Leishan</collectingMunicipality>
,
<location id="8EA060301558BC1DEDA4B784A300F95E" LSID="urn:lsid:plazi:treatment:03D687FD1558BC1DEC87B451A59FF8ED:8EA060301558BC1DEDA4B784A300F95E" box="[436,534,1723,1749]" country="China" latitude="26.333334" longLatPrecision="1242" longitude="108.15" municipality="Leishan" name="Taojiang" pageId="4" pageNumber="255" stateProvince="Guizhou">Taojiang</location>
(
<geoCoordinate id="EE4B502C1558BC1DEE36B784A392F95D" box="[550,644,1723,1750]" degrees="26" direction="north" minutes="20" orientation="latitude" pageId="4" pageNumber="255" precision="925" value="26.333334">26º20N</geoCoordinate>
,
<geoCoordinate id="EE4B502C1558BC1DEE81B784A3ECF95D" box="[657,762,1723,1750]" degrees="108" direction="east" minutes="09" orientation="longitude" pageId="4" pageNumber="255" precision="925" value="108.15">108º09E</geoCoordinate>
),
<elevation id="0052D1D81558BC1DEF1FB783A271F95D" box="[783,871,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="255" unit="m" value="1000.0">
<quantity id="4C879B0E1558BC1DEF1FB783A271F95D" box="[783,871,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="255" unit="m" value="1000.0">1000 m</quantity>
</elevation>
,
<specimenCount id="9D79FD621558BC1DEF64B783A289F95D" box="[884,927,1724,1750]" pageId="4" pageNumber="255" type="female">1 ♀</specimenCount>
,
<collectingDate id="EF85E9C31558BC1DEFBEB784A53AF95D" box="[942,1068,1723,1750]" pageId="4" pageNumber="255" value="1988-07-05">
<date id="FFC1102B1558BC1DEFBEB784A53AF95D" box="[942,1068,1723,1750]" pageId="4" pageNumber="255" value="1988-07-05">5.VII.1988</date>
</collectingDate>
, leg.
<collectorName id="268A533D1558BC1DE879B783A5FFF95D" box="[1129,1257,1724,1750]" pageId="4" pageNumber="255">Long-Long</collectorName>
<collectionCode id="ED6EAE2E1558BC1DE8E0B783A454F95D" box="[1264,1346,1724,1750]" pageId="4" pageNumber="255">YANG</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B173CB61558BC1DE95EB784A53AF971" ID-GBIF-Occurrence="2294326319" collectingDate="1991-07-03" collectionCode="WANG" collectorName="Xing-Jian" country="China" elevation="490" latitude="26.7" location="Jiangle Xian" longLatPrecision="1240" longitude="117.4" pageId="4" pageNumber="255" specimenCount="1" specimenCount-female="1" stateProvince="Fujian">
<collectingRegion id="49BBF8091558BC1DE95EB784A48FF95E" box="[1358,1433,1723,1749]" country="China" name="Fujian" pageId="4" pageNumber="255">Fujian</collectingRegion>
,
<location id="8EA060301558BC1DEC87B7E0A032F972" LSID="urn:lsid:plazi:treatment:03D687FD1558BC1DEC87B451A59FF8ED:8EA060301558BC1DEC87B7E0A032F972" box="[151,292,1759,1785]" country="China" latitude="26.7" longLatPrecision="1240" longitude="117.4" name="Jiangle Xian" pageId="4" pageNumber="255" stateProvince="Fujian">Jiangle Xian</location>
(
<geoCoordinate id="EE4B502C1558BC1DED23B7E0A084F971" box="[307,402,1759,1786]" degrees="26" direction="north" minutes="42" orientation="latitude" pageId="4" pageNumber="255" precision="925" value="26.7">26º42N</geoCoordinate>
,
<geoCoordinate id="EE4B502C1558BC1DED8DB7E0A310F971" box="[413,518,1759,1786]" degrees="117" direction="east" minutes="24" orientation="longitude" pageId="4" pageNumber="255" precision="925" value="117.4">117º24E</geoCoordinate>
),
<elevation id="0052D1D81558BC1DEE09B7DFA374F971" box="[537,610,1760,1786]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="255" unit="m" value="490.0">
<quantity id="4C879B0E1558BC1DEE09B7DFA374F971" box="[537,610,1760,1786]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="255" unit="m" value="490.0">490 m</quantity>
</elevation>
,
<specimenCount id="9D79FD621558BC1DEE7EB7DFA38EF971" box="[622,664,1760,1786]" pageId="4" pageNumber="255" type="female">1 ♀</specimenCount>
,
<collectingDate id="EF85E9C31558BC1DEEB6B7DFA232F971" box="[678,804,1760,1786]" pageId="4" pageNumber="255" value="1991-07-03">
<date id="FFC1102B1558BC1DEEB6B7DFA232F971" box="[678,804,1760,1786]" pageId="4" pageNumber="255" value="1991-07-03">3.VII.1991</date>
</collectingDate>
, leg.
<collectorName id="268A533D1558BC1DEF4FB7E0A2DBF972" box="[863,973,1759,1785]" pageId="4" pageNumber="255">Xing-Jian</collectorName>
<collectionCode id="ED6EAE2E1558BC1DEFC4B7DFA53AF971" box="[980,1068,1760,1786]" pageId="4" pageNumber="255">WANG</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC036EB1558BC1DECD7B63CA2A2F895" blockId="4.[151,1436,1687,1894]" box="[199,948,1795,1822]" pageId="4" pageNumber="255">
<emphasis id="B90BEAF91558BC1DECD7B63CA0D0F895" bold="true" box="[199,454,1795,1822]" pageId="4" pageNumber="255">General distribution:</emphasis>
<materialsCitation id="3B173CB61558BC1DEDDDB63CA2A6F895" ID-GBIF-Occurrence="2294326332" box="[461,944,1795,1822]" country="China" location="Fujian" pageId="4" pageNumber="255" specimenCount="1" stateProvince="Fujian">
<collectingCountry id="F368767B1558BC1DEDDDB63CA306F896" box="[461,528,1795,1821]" name="China" pageId="4" pageNumber="255">China</collectingCountry>
(
<collectingRegion id="49BBF8091558BC1DEE0EB63CA37CF896" box="[542,618,1795,1821]" country="China" name="Fujian" pageId="4" pageNumber="255">Fujian</collectingRegion>
,
<collectingRegion id="49BBF8091558BC1DEE65B63CA3DFF896" box="[629,713,1795,1821]" country="China" name="Jiangxi" pageId="4" pageNumber="255">Jiangxi</collectingRegion>
,
<collectingRegion id="49BBF8091558BC1DEEC4B63CA221F896" box="[724,823,1795,1821]" country="China" name="Guangxi" pageId="4" pageNumber="255">Guangxi</collectingRegion>
,
<collectingRegion id="49BBF8091558BC1DEF53B63CA2BFF895" box="[835,937,1795,1822]" country="China" name="Guizhou" pageId="4" pageNumber="255">Guizhou</collectingRegion>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC036EB1558BC1DECD7B617A59FF8ED" blockId="4.[151,1436,1687,1894]" pageId="4" pageNumber="255">
<emphasis id="B90BEAF91558BC1DECD7B617A02BF8C9" bold="true" box="[199,317,1832,1858]" pageId="4" pageNumber="255">Remarks:</emphasis>
<bibRefCitation id="EFEE4B1A1558BC1DED54B617A0AAF8C9" author="Wu, Y. R." box="[324,444,1831,1858]" pageId="4" pageNumber="255" refId="ref11919" refString="Wu, Y. R. (2006) Fauna Sinica Insecta Vol. 44. Hymenoptera. Megachilidae. Science Press, Beijing, xvi + 474 pp., 4 pls. [in Chinese]" type="book" year="2006">Wu (2006)</bibRefCitation>
only described the female of the species, and thought the male is unknown. However, the male was described by
<bibRefCitation id="EFEE4B1A1558BC1DED8CB674A3BCF8ED" author="Mavromoustakis, G. A." box="[412,682,1867,1894]" pageId="4" pageNumber="255" pagination="249 - 252" refId="ref11239" refString="Mavromoustakis, G. A. (1954) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VII. Annals and Magazine for Natural History, 12, Ser., 7, 249 - 252. https: // doi. org / 10.1080 / 00222935408651726." type="journal article" year="1954">Mavromoustakis (1954)</bibRefCitation>
(see also redescription in
<bibRefCitation id="EFEE4B1A1558BC1DEFDFB674A56BF8ED" author="Kasparek, M." box="[975,1149,1867,1894]" pageId="4" pageNumber="255" pagination="1 - 151" refId="ref10897" refString="Kasparek, M. (2017) Resin bees of the anthidiine genus Trachusa: identification, taxonomy, distribution and biology of the old world species. Entomofauna, supplement 21, 1 - 151." type="journal article" year="2017">Kasparek 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>