363 lines
48 KiB
XML
363 lines
48 KiB
XML
<document id="FCA8C1B1FA02833F84CDFF7B7ACC4726" ID-CLB-Dataset="50633" ID-DOI="10.5281/zenodo.172131" ID-GBIF-Dataset="636f2b6e-7672-455b-b541-01c3657fa517" ID-ISSN="11755326" ID-Zenodo-Dep="172131" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459764802000" checkinUser="plazi" docAuthor="Karaman, Gordan S., Mato, Sanja Gottstein & Ec" docDate="2006" docId="03D687F0FFC10C5D3174FEB1AE82FBA3" docLanguage="en" docName="zt01150p068.pdf" docOrigin="Zootaxa 1150" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Niphargus echion Karaman, Mato & Ec, 2006, n. sp." docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="FFEFFF88FFC30C50307CFFADAC56FFA5" masterDocTitle="Niphargus echion, a new species of amphipod (Crustacea Amphipoda, Niphargidae) from Istra, Croatia" masterLastPageNumber="68" masterPageNumber="53" pageNumber="55" updateTime="1698222148265" updateUser="ExternalLinkService">
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<mods:titleInfo id="326FC7493587AFA80D15EFDF39849391">
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<mods:title id="E1E0478D613C1F8565D0E698FC38E3CB">Niphargus echion, a new species of amphipod (Crustacea Amphipoda, Niphargidae) from Istra, Croatia</mods:title>
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<mods:name id="C8844B6768352855F31BEE50FC153F46" type="personal">
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<mods:roleTerm id="E6F8E8D0A9A97596FEDD0725374A80E5">Author</mods:roleTerm>
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<mods:namePart id="520F05110EDA129F3DF7C165C849F0F8">Karaman, Gordan S.</mods:namePart>
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<mods:roleTerm id="69DC43151516F06BFCCF51FC8F03133D">Author</mods:roleTerm>
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<mods:namePart id="879E6CB7C6DCA3FC378623C7FD1600ED">Mato, Sanja Gottstein</mods:namePart>
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<mods:roleTerm id="5BBD9F40BF64F30AAE0C63718EDF9ED9">Author</mods:roleTerm>
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<mods:namePart id="C06610F65341CC837712D3ADBF1565D7">Ec</mods:namePart>
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<mods:title id="2BAB4C7A4159202FE44E80E20DE2971B">Zootaxa</mods:title>
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<mods:part id="C7789D0AACD0F8362D6724E91C3BC9C4">
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<mods:date id="ADD1E7DBB19AEE0C1B1D6C180D4AB553">2006</mods:date>
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<mods:number id="1FFA0BF57C30C935A38C1A055B4B7C9D">1150</mods:number>
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<mods:identifier id="217103A75A8EBA8920039B233E85116C" type="DOI">10.5281/zenodo.172131</mods:identifier>
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<treatment id="03D687F0FFC10C5D3174FEB1AE82FBA3" ID-DOI="http://doi.org/10.5281/zenodo.6263348" ID-GBIF-Taxon="119345766" ID-Zenodo-Dep="6263348" LSID="urn:lsid:plazi:treatment:03D687F0FFC10C5D3174FEB1AE82FBA3" httpUri="http://treatment.plazi.org/id/03D687F0FFC10C5D3174FEB1AE82FBA3" lastPageId="13" lastPageNumber="66" pageId="2" pageNumber="55">
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<subSubSection id="C365656DFFC10C523174FEB1ADD7FEFC" pageId="2" pageNumber="55" type="nomenclature">
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<paragraph id="8BC036E6FFC10C523174FEB1AE65FE90" blockId="2.[264,563,283,345]" box="[264,563,283,310]" pageId="2" pageNumber="55">
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<heading id="D088818AFFC10C523174FEB1AE65FE90" bold="true" box="[264,563,283,310]" fontSize="11" level="1" pageId="2" pageNumber="55" reason="1">
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<emphasis id="B90BEAF4FFC10C523174FEB1AE65FE90" bold="true" box="[264,563,283,310]" pageId="2" pageNumber="55">
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<taxonomicName id="4C7F4D65FFC10C523174FEB1ADB4FE93" ID-CoL="47FZJ" box="[264,482,284,310]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="echion" status="sp. nov.">
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<emphasis id="B90BEAF4FFC10C523174FEB1ADB4FE93" bold="true" box="[264,482,284,310]" italics="true" pageId="2" pageNumber="55">Niphargus echion</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A238578FFFC10C52318CFEB6AE65FE90" box="[496,563,283,309]" pageId="2" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC036E6FFC10C523174FE92ADD7FEFC" blockId="2.[264,563,283,345]" box="[264,385,319,345]" pageId="2" pageNumber="55">
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(
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<figureCitation id="13442A63FFC10C52316CFE92AD2EFEFC" box="[272,376,319,345]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="3.[264,368,1684,1708]" captionStartId-1="4.[264,368,1789,1813]" captionStartId-2="6.[264,367,1795,1819]" captionStartId-3="7.[264,368,1780,1804]" captionStartId-4="10.[264,368,1767,1791]" captionStartId-5="11.[264,367,1794,1818]" captionTargetBox-0="[296,1308,334,1646]" captionTargetBox-1="[294,1271,331,1787]" captionTargetBox-2="[295,1309,328,1754]" captionTargetBox-3="[310,1283,336,1744]" captionTargetBox-4="[288,1308,322,1729]" captionTargetBox-5="[326,1298,328,1753]" captionTargetId-0="figure@3.[267,1326,317,1678]" captionTargetId-1="figure@4.[264,1323,298,1790]" captionTargetId-2="figure@6.[264,1323,300,1771]" captionTargetId-3="figure@7.[273,1314,298,1770]" captionTargetId-4="figure@10.[264,1323,294,1760]" captionTargetId-5="figure@11.[273,1314,298,1770]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="10" captionTargetPageId-5="11" captionText-0="FIGURE 1. Niphargus echion, n. sp., holotype male, 11 mm, Pliškovićeva jama cave: A, head; B – C, antenna 1; D, antenna 2; E, maxilla 2; F – G, maxilliped. Scales bars: A, B, D, 1 mm; E, F, 0.25 mm." captionText-1="FIGURE 2. Niphargus echion, n. sp., holotype male, 11 mm, PliŠkovićeva jamacave: A – C, gnathopod 1; D – F, gnathopod 2. Scales bars: A, B, D, E, 1 mm." captionText-2="FIGURE 4. Niphargus echion, n. sp., holotype male, 11 mm, PliŠkovićeva jama cave: A – B, pereopod 5; C, pereopod 6; D – E, pereopod 7; F, epimeral plates 1 – 3; G, telson. Scales bars: A, C, D, F, 1 mm; G, 0.5 mm." captionText-3="FIGURE 5. Niphargus echion, n. sp., holotype male, 11 mm, PliŠkovićeva jama cave: A, labrum; B, labium; C – D, maxilla 1; E, mandibular palp, inner face; F, mandibular palp, outer face. Female 10.3 mm (paratype): G – J, coxae 1 – 4. Scales bars: A – C, 0.5 mm; E – F, 0.25 mm; G – J, 1 mm." captionText-4="FIGURE 6. Niphargus echion, n. sp., paratype female, 10.3 mm, PliŠkovićeva jama cave: A – B, gnathopod 1; C – D, gnathopod 2; E, urosome with uropods 1 – 2. Scales bars: A, C, 0.5 mm; E, 1 mm." captionText-5="FIGURE 7. Niphargus echion, n. sp., paratype female, 10.3 mm, PliŠkovićeva jama cave: A, inner plate of maxilliped; B, dactyl of maxilliped; C – D, pereopod 7; E, epimeral plates 2 – 3; F, uropod 3; G, telson. Scales bars: C, E, F, G, 1 mm." httpUri-0="https://zenodo.org/record/172132/files/figure.png" httpUri-1="https://zenodo.org/record/172133/files/figure.png" httpUri-2="https://zenodo.org/record/172134/files/figure.png" httpUri-3="https://zenodo.org/record/172135/files/figure.png" httpUri-4="https://zenodo.org/record/172136/files/figure.png" httpUri-5="https://zenodo.org/record/172137/files/figure.png" pageId="2" pageNumber="55">Figs 1–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C365656DFFC10C523174FE22A8D7FC29" pageId="2" pageNumber="55" type="materials_examined">
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<paragraph id="8BC036E6FFC10C523174FE22ADB0FE0D" blockId="2.[264,1323,399,588]" box="[264,486,399,424]" pageId="2" pageNumber="55">
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<emphasis id="B90BEAF4FFC10C523174FE22ADB0FE0D" box="[264,486,399,424]" italics="true" pageId="2" pageNumber="55">Material examined</emphasis>
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</paragraph>
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<paragraph id="8BC036E6FFC10C523144FE17AD34FDE9" blockId="2.[264,1323,399,588]" pageId="2" pageNumber="55">
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“Pliškovićeva jama” cave,Š ikuti village, St. Vinčenat, Istra (
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<collectingCountry id="F3687676FFC10C523478FE17A837FE71" box="[1028,1121,442,468]" name="Croatia" pageId="2" pageNumber="55">Croatia</collectingCountry>
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);
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<date id="FFC11026FFC10C523409FE17A970FE71" box="[1141,1318,442,468]" pageId="2" pageNumber="55" value="2000-08-06">6 August 2000</date>
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, coll. B. Jalżić,
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<specimenCount id="9D79FD6FFFC10C5231C9FE4FAE1FFE59" box="[437,585,482,508]" pageId="2" pageNumber="55" type="generic">2 specimens</specimenCount>
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.
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<typeStatus id="54C48844FFC10C52322FFE4FAE96FE59" box="[595,704,482,508]" pageId="2" pageNumber="55" type="holotype">Holotype</typeStatus>
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(coll. no. C1587, male 11.0 mm), and
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<typeStatus id="54C48844FFC10C5234FEFE4FA8B1FE59" box="[1154,1255,482,508]" pageId="2" pageNumber="55" type="paratype">paratype</typeStatus>
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(coll. no. C1588, female 10.3 mm) are deposited in the Croatian Natural History Museum, Zagreb.
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</paragraph>
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<paragraph id="8BC036E6FFC10C523174FD2FAD2BFD39" blockId="2.[264,1322,642,908]" box="[264,381,642,668]" pageId="2" pageNumber="55">Diagnosis</paragraph>
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<paragraph id="8BC036E6FFC10C523144FD07A8D7FC29" blockId="2.[264,1322,642,908]" pageId="2" pageNumber="55">
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Outer plate of maxilla 1 with 19–20 spines, each with 1 or 2 very slender lateral and elongated distal bifurcations. Dactylus of maxilliped with 2 nails. Pleopods with 2 or 3 retinaculae. Gnathopod 2 basis with 4 or 5 long outer setae near corner spine; dactyl with row of outer marginal setae. Dactyl of pereopods 3–7 with 1 strong spine on inner margin. Epimeral plates with angular or subacute posteroventral corner. Uropods 1 and 2 with subequal rami. Second segment of uropod
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<quantity id="4C879B03FFC10C523280FCDFAF7FFC29" box="[764,809,882,908]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="55" unit="in" value="3.0">3 in</quantity>
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males moderately elongated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C365656DFFC10C5C3174FC6EA8A3FA7B" lastPageId="12" lastPageNumber="65" pageId="2" pageNumber="55" type="description">
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<paragraph id="8BC036E6FFC10C523174FC6EADC4FC79" blockId="2.[264,1324,963,1828]" box="[264,402,963,988]" pageId="2" pageNumber="55">
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<emphasis id="B90BEAF4FFC10C523174FC6EADC4FC79" box="[264,402,963,988]" italics="true" pageId="2" pageNumber="55">Description</emphasis>
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</paragraph>
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<paragraph id="8BC036E6FFC10C523144FC47AEC7FBD9" blockId="2.[264,1324,963,1828]" pageId="2" pageNumber="55">
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ADULT MALE (
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<typeStatus id="54C48844FFC10C523278FC47AE27FBA1" box="[516,625,1002,1028]" pageId="2" pageNumber="55" type="holotype">holotype</typeStatus>
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). Body: moderately slender, metasomal segments 1–3 with 10–15 dorsoposterior short marginal setae [segment 3 with 15 setae] (fig. 4F). Epimeral plates 2–3 with poorly acute posteroventral corner, never produced, plates 2–3 with 2 ventrofacial spines each (fig. 4F).
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</paragraph>
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<paragraph id="8BC036E6FFC10C523144FB27AE31FB51" blockId="2.[264,1324,963,1828]" pageId="2" pageNumber="55">Urosomite 1 with one posteroventral spine (fig. 3F). Urosomite 1 on each dorsolateral side with 2 strong spines; urosomite 2 on each dorsolateral side with 2 strong spines. Urosomite 3 smooth (fig. 3F).</paragraph>
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<paragraph id="8BC036E6FFC10C523144FAAFA947FAE1" blockId="2.[264,1324,963,1828]" pageId="2" pageNumber="55">Head with slightly convex dorsal margin (in lateral view); rostrum short, lateral cephalic lobes short and subrounded, anteroventral sinus shallow (fig. 1A). Eyes absent.</paragraph>
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<paragraph id="8BC036E6FFC10C523144FAFFAD6DF9A9" blockId="2.[264,1324,963,1828]" pageId="2" pageNumber="55">Antenna 1: slightly shorter than half body length (4.7: 11); peduncular segments 1–3 relatively stout, progressively shorter; peduncular segment 3 slightly shorter than half peduncular segment 2; primary flagellum longer than peduncle, consisting of 22 articles bearing one short aesthetasc each (fig. 1B); accessory flagellum short, 2segmented (fig. 1C).</paragraph>
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<paragraph id="8BC036E6FFC10C523144F9B7AFA5F921" blockId="2.[264,1324,963,1828]" pageId="2" pageNumber="55">Antenna 2: much shorter than antenna 1; peduncular segment 4 slightly longer than segment 5 (4.3: 5.3); flagellum moderately slender, longer than last peduncular segment and consisting of 9 articles; antennal gland cone short (fig. 1D).</paragraph>
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<paragraph id="8BC036E6FFC10C523144F93FA896F971" blockId="2.[264,1324,963,1828]" pageId="2" pageNumber="55">Coxae: relatively short, coxa 1 slightly broader than long, with subrounded anteroventral corner; ventral margin with about 10 short marginal setae (fig. 2A).</paragraph>
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<paragraph id="8BC036E6FFC10C523144F94FAF2BF881" blockId="2.[264,1324,963,1828]" pageId="2" pageNumber="55">Coxae 2–4 slightly longer than broad, with up to 12 relatively short, marginal setae (figs 2D, 3A & E). Coxae 5–7 short (fig. 4A, C & D).</paragraph>
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<caption id="DF00666EFFC00C533174F939AD6FF954" httpUri="https://zenodo.org/record/172132/files/figure.png" pageId="3" pageNumber="56" targetBox="[296,1308,334,1646]" targetPageId="3">
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<paragraph id="8BC036E6FFC00C533174F939AD6FF954" blockId="3.[264,1324,1684,1777]" pageId="3" pageNumber="56">
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<emphasis id="B90BEAF4FFC00C533174F939ADDBF909" bold="true" box="[264,397,1684,1708]" pageId="3" pageNumber="56">FIGURE 1.</emphasis>
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<taxonomicName id="4C7F4D65FFC00C5331EBF938AE01F909" box="[407,599,1685,1708]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="echion" status="sp. nov.">
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<emphasis id="B90BEAF4FFC00C5331EBF938AE01F909" box="[407,599,1685,1708]" italics="true" pageId="3" pageNumber="56">Niphargus echion</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A238578FFFC00C53321BF939AEF3F909" box="[615,677,1684,1708]" pageId="3" pageNumber="56" rank="species">
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<emphasis id="B90BEAF4FFC00C53321BF939AEF3F909" bold="true" box="[615,677,1684,1708]" pageId="3" pageNumber="56">n. sp.</emphasis>
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</taxonomicNameLabel>
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, holotype male, 11 mm, Pliškovićeva jama cave: A, head; B–C, antenna 1; D, antenna 2; E, maxilla 2; F–G, maxilliped. Scales bars: A, B, D, 1 mm; E, F, 0.25 mm.
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</paragraph>
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</caption>
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<caption id="DF00666EFFC70C543174F950AFF9F892" httpUri="https://zenodo.org/record/172133/files/figure.png" pageId="4" pageNumber="57" targetBox="[294,1271,331,1787]" targetPageId="4">
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<paragraph id="8BC036E6FFC70C543174F950AFF9F892" blockId="4.[264,1323,1789,1847]" pageId="4" pageNumber="57">
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<emphasis id="B90BEAF4FFC70C543174F950ADC6F8B0" bold="true" box="[264,400,1789,1813]" pageId="4" pageNumber="57">FIGURE 2.</emphasis>
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<taxonomicName id="4C7F4D65FFC70C5431E1F950AE36F8B1" box="[413,608,1789,1812]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="echion" status="sp. nov.">
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<emphasis id="B90BEAF4FFC70C5431E1F950AE36F8B1" box="[413,608,1789,1812]" italics="true" pageId="4" pageNumber="57">Niphargus echion</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A238578FFFC70C54320FF950AEE3F8B0" box="[627,693,1789,1813]" pageId="4" pageNumber="57" rank="species">
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<emphasis id="B90BEAF4FFC70C54320FF950AEE3F8B0" bold="true" box="[627,693,1789,1813]" pageId="4" pageNumber="57">n. sp.</emphasis>
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</taxonomicNameLabel>
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, holotype male, 11 mm, PliŠkovićeva jamacave: A–C, gnathopod 1; D–F, gnathopod 2. Scales bars: A, B, D, E, 1 mm.
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</paragraph>
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</caption>
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<paragraph id="8BC036E6FFC60C553174F941AD97F8EC" pageId="5" pageNumber="58">
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<table id="F97FC446FFC6F3AF3174F941A97CF8EC" box="[264,1322,1772,1865]" gridcols="3" gridrows="3" pageId="5" pageNumber="58">
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<tr id="354F34A4FFC6F3AF3174F941A97CF8A1" box="[264,1322,1772,1796]" gridrow="0" pageId="5" pageNumber="58">
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<th id="769E5DD8FFC6F3AF3174F941AE30F8A1" box="[264,614,1772,1796]" gridcol="0" gridrow="0" pageId="5" pageNumber="58">
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<emphasis id="B90BEAF4FFC60C553174F941ADD9F8A1" bold="true" box="[264,399,1772,1796]" pageId="5" pageNumber="58">FIGURE 3.</emphasis>
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<taxonomicName id="4C7F4D65FFC60C5531E0F940AE09F8A1" box="[412,607,1773,1796]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="echion">
|
||
<emphasis id="B90BEAF4FFC60C5531E0F940AE09F8A1" box="[412,607,1773,1796]" italics="true" pageId="5" pageNumber="58">Niphargus echion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
</th>
|
||
<th id="769E5DD8FFC6F3AF3211F941AF3DF8A1" box="[621,875,1772,1796]" gridcol="1" gridrow="0" pageId="5" pageNumber="58">
|
||
<emphasis id="B90BEAF4FFC60C55320EF941AEE5F8A1" bold="true" box="[626,691,1772,1796]" pageId="5" pageNumber="58">n. sp.</emphasis>
|
||
, holotype male,
|
||
</th>
|
||
<th id="769E5DD8FFC6F3AF330EF941A97CF8A1" box="[882,1322,1772,1796]" gridcol="2" gridrow="0" pageId="5" pageNumber="58">11 mm, PliŠkovićeva jama cave: A–B,</th>
|
||
</tr>
|
||
<tr id="354F34A4FFC6F3AF3174F8A2A97CF883" box="[264,1322,1807,1830]" gridrow="1" pageId="5" pageNumber="58">
|
||
<th id="769E5DD8FFC6F3AF3174F8A2AE30F883" box="[264,614,1807,1830]" gridcol="0" gridrow="1" pageId="5" pageNumber="58">pereopod 3; C–D, pereopod 4; E,</th>
|
||
<td id="769E5DD8FFC6F3AF3211F8A2AF3DF883" box="[621,875,1807,1830]" gridcol="1" gridrow="1" pageId="5" pageNumber="58">coxa 4; F, urosome with</td>
|
||
<td id="769E5DD8FFC6F3AF330EF8A2A97CF883" box="[882,1322,1807,1830]" gridcol="2" gridrow="1" pageId="5" pageNumber="58">uropods 1–2; G, uropod 3. Scales bars: A,</td>
|
||
</tr>
|
||
<tr id="354F34A4FFC6F3AF3174F89FA97CF8EC" box="[264,1322,1842,1865]" gridrow="2" pageId="5" pageNumber="58" rowspan-1="1" rowspan-2="1">
|
||
<th id="769E5DD8FFC6F3AF3174F89FAE30F8EC" box="[264,614,1842,1865]" gridcol="0" gridrow="2" pageId="5" pageNumber="58">C, E, F, G, 1 mm.</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF00666EFFC50C563174F8AEAE52F8C5" httpUri="https://zenodo.org/record/172134/files/figure.png" pageId="6" pageNumber="59" targetBox="[295,1309,328,1754]" targetPageId="6">
|
||
<paragraph id="8BC036E6FFC50C563174F8AEAE52F8C5" blockId="6.[264,1323,1795,1888]" pageId="6" pageNumber="59">
|
||
<emphasis id="B90BEAF4FFC50C563174F8AEADD9F8BE" bold="true" box="[264,399,1795,1819]" pageId="6" pageNumber="59">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C7F4D65FFC50C5631E0F8AEAE09F8BF" box="[412,607,1795,1818]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="echion" status="sp. nov.">
|
||
<emphasis id="B90BEAF4FFC50C5631E0F8AEAE09F8BF" box="[412,607,1795,1818]" italics="true" pageId="6" pageNumber="59">Niphargus echion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A238578FFFC50C56320EF8AEAEE5F8BE" box="[626,691,1795,1819]" pageId="6" pageNumber="59" rank="species">
|
||
<emphasis id="B90BEAF4FFC50C56320EF8AEAEE5F8BE" bold="true" box="[626,691,1795,1819]" pageId="6" pageNumber="59">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male, 11 mm, PliŠkovićeva jama cave: A–B, pereopod 5; C, pereopod 6; D–E, pereopod 7; F, epimeral plates 1–3; G, telson. Scales bars: A, C, D, F, 1 mm; G, 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00666EFFC40C573174F959A889F8F4" httpUri="https://zenodo.org/record/172135/files/figure.png" pageId="7" pageNumber="60" targetBox="[310,1283,336,1744]" targetPageId="7">
|
||
<paragraph id="8BC036E6FFC40C573174F959A889F8F4" blockId="7.[264,1322,1780,1873]" pageId="7" pageNumber="60">
|
||
<emphasis id="B90BEAF4FFC40C573174F959ADDDF8A9" bold="true" box="[264,395,1780,1804]" pageId="7" pageNumber="60">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C7F4D65FFC40C5731EFF958AE07F8A9" box="[403,593,1781,1804]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="echion" status="sp. nov.">
|
||
<emphasis id="B90BEAF4FFC40C5731EFF958AE07F8A9" box="[403,593,1781,1804]" italics="true" pageId="7" pageNumber="60">Niphargus echion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A238578FFFC40C57321CF959AECAF8A9" box="[608,668,1780,1804]" pageId="7" pageNumber="60" rank="species">
|
||
<emphasis id="B90BEAF4FFC40C57321CF959AECAF8A9" bold="true" box="[608,668,1780,1804]" pageId="7" pageNumber="60">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male, 11 mm, PliŠkovićeva jama cave: A, labrum; B, labium; C–D, maxilla 1; E, mandibular palp, inner face; F, mandibular palp, outer face. Female 10.3 mm (paratype): G–J, coxae 1–4. Scales bars: A–C, 0.5 mm; E–F, 0.25 mm; G–J, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036E6FFCB0C583144FEB1AE10FEFB" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="61">Mouthparts: Labrum short, subrounded (fig. 5A). Labium normal, broader than long, outer lobes entire (fig. 5B).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C583144FEC1A976FE73" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="61">Maxilla 1: inner plate conical, with 5 distal setae; outer plate not dilated, but provided with 19 or 20 spines bearing 1 or 2 lateral long and slender teeth each. Palp short, 2 segmented, reaching distal margin of outer plate, and bearing 6 distal setae (fig. 5C & D).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C583144FE49A828FE5B" blockId="8.[264,1325,284,1870]" box="[312,1150,484,510]" pageId="8" pageNumber="61">Maxilla 2: normal, both lobes with numerous distal setae only (fig. 1E).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C583144FDA1A801FDD3" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="61">Maxilliped: inner plate with 4 or 5 distal chisellike spines intermixed with single setae; outer plate with row of spines along inner margin and distal tip, reaching nearly half of palp segment 2; dactyl (segment 4) of palp with 2 nails (fig. 1F & G).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C583144FD29AE28FCB3" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="61">Mandibular palp: 3segmented, first segment smooth, second segment with 13 strong, spinelike setae along posterior margin; palp segment 3 slightly longer than segment 2, bearing 22 Dsetae, 8 distal Esetae, on outer face one group of 5 setae, on inner face 5 groups of Bsetae (fig. 1E & F).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C583144FC89AEFCFBF3" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="61">Gnathopod 1: size moderate, slightly smaller than gnathopod 2; segment 3 with one posterior group of setae only; segment 5 markedly shorter than segment 6. Segment 6 longer than broad, poorly dilated distally, and with 9 groups of posterior marginal setae; palm slope oblique nearly 3/8 of posterior margin of segment 6, defined on outer face by short corner spine accompanied laterally by 3 slender toothed spines and 5 long outer setae; on inner face defined by one short subcorner spine; dactyl reaching posterior margin of segment 6, with row of 9 setae along outer margin, setae sitting in group of 1 or 2 only (1–1–2–1–1–1–2) (fig. 2A, B & C).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C583144FBC9AEF3FACB" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="61">Gnathopod 2: segment 3 with one posterior group of setae; segment 5 slightly shorter than segment 6. Segment 6 nearly as long as broad, with 13 groups of setae along posterior margin; palm slope oblique nearly 3/8 of posterior margin of segment 6, defined on outer face by 1 strong corner spine accompanied by 2 slender toothed spines placed laterally and by 4 long setae on outer surface; at inner face by 1 short submarginal spine; dactyl reaching posterior margin of segment 6, provided along outer margin with row of 9 setae (2–2–1–1–1–1–1) (fig. 2D, E & F).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C583144FAD1A8D7FA1B" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="61">Pereopods 3 and 4: subequal in length, relatively stout; dactyls short, with one strong spine at inner margin, nail poorly longer than peduncle (fig. 3A, B, C & D).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C583144FA61AD9AF973" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="61">Pereopods 5–7: relatively stout, pereopod 5 remarkably shorter than pereopod 7, all pereopods with segment 2 distinctly longer than broad, without anteroventral or posteroventral lobe, and with slightly convex posterior margin provided with row of short marginal setae (fig. 4A, C & D). Segments 3–6 stout, with spines along both margins. Dactyl of pereopod 5 slightly longer than peduncle, with one strong spine at inner margin (fig. 4B). Dactyl of pereopod 7 with nail as long as peduncle, and with one spine at inner margin (fig. 4E).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C583144F949AFB1F95B" blockId="8.[264,1325,284,1870]" box="[312,999,1764,1790]" pageId="8" pageNumber="61">Pleopods 1–3 with 2 or 3 retinaculae each (2–3–2, 2–2–2).</paragraph>
|
||
<paragraph id="8BC036E6FFCB0C593144F8A1AF72FE93" blockId="8.[264,1325,284,1870]" lastBlockId="9.[264,1325,284,1750]" lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61">Uropod 1: peduncle with row of dorsomedial and dorsolateral row of spines, rami shorter than peduncle, of subequal length, bearing single lateral and 4 or 5 distal short spines accompanied by several setae (fig. 3F).</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FEE9A975FE23" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">Uropod 2: rami subequal, outer ramus slightly recurved, and almost reaching tip of inner ramus, both rami provided with single lateral and 4 or 5 distal short spines (fig. 3F).</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FE39AF74FD83" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">Uropod 3: slightly elongated, peduncle much shorter than first segment of outer ramus; first segment at both margins with spines, single long plumose seta at its inner margin; second segment of outer ramus slightly exceeding 1/3 of first segment of outer ramus (fig. 3G). Inner ramus short, scalelike.</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FD99ADE6FD63" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">Telson: not elongated, slightly broader than long, incised nearly 2/3 of its length; each lobe with 3 distal and 2 facial spines, along outer margin with 2 single spines, along inner margin with 1 or 2 spines; a pair of short plumose setae appears near the middle of each lobe (fig. 4G).</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FD79AFCFFD4B" blockId="9.[264,1325,284,1750]" box="[312,921,724,750]" pageId="9" pageNumber="62">Coxal gills: 1–6 moderately long (figs 2D, 3A, 4A).</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FD51AFAEFC2B" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">
|
||
ADULT FEMALE (
|
||
<typeStatus id="54C48844FFCA0C59324AFD51AEF5FCB3" box="[566,675,764,790]" pageId="9" pageNumber="62" type="paratype">paratype</typeStatus>
|
||
): Body: with setose oostegites bearing
|
||
<specimenCount id="9D79FD6FFFCA0C5934DDFD51A952FCB3" box="[1185,1284,764,790]" pageId="9" pageNumber="62" type="egg">10 eggs</specimenCount>
|
||
in marsupium. Metasomal segments 1–3 with row of posterodorsal marginal setae as in male (fig.7E). Urosomite 1 with one posteroventral spine, as in male (fig. 6E). Urosomites 1 and 2 on each side with 2 spines each; urosomite 3 smooth (fig. 6E).
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FC31AF22FC7B" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">Epimeral plates: 2 or 3 acute, but not pointed, as in male; epimeral plate 2 with 2, and epimeral plate 3 with 3 ventrolateral spines (fig. 7E).</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FC41ADBAFBF3" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">Head: as in male, antenna 1 hardly longer than that of male (4.7: 10.3); primary flagellum consisting of 21 or 22 articles bearing one aesthetasc each. Antenna 2: flagellum with 8 or 9 articles.</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FBC9AEC7FBDB" blockId="9.[264,1325,284,1750]" box="[312,657,1124,1150]" pageId="9" pageNumber="62">Mouthparts: similar to males.</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FB21A941FB6B" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">Maxilla 1: inner plate with 3 distal setae only; outer plate with up to 20 slender spines, bearing 1 or 2 slender elongated lateral teeth each; palp 2segmented, with 6 distal setae.</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FB71AE05FABB" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">Maxilliped: as in males: inner plate with 4 or 5 distal chisellike spines (fig. 7A), dactyl with 2 nails (fig. 7B).</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FA81AD90FA33" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">Coxae: similar to males; coxa 1 broader than long (fig. 5G); coxae 2–4 longer than broad (fig. 5H, I & J), all with row of short to slightly longer marginal setae each. Coxae 5–7 as in males.</paragraph>
|
||
<paragraph id="8BC036E6FFCA0C593144FA09AEA2F973" blockId="9.[264,1325,284,1750]" pageId="9" pageNumber="62">Gnathopods: 1 or 2 of moderate size, gnathopod 2 moderately larger than gnathopod 1 (fig. 6A, 6C). Gnathopod 1: segment 3 with one posterior marginal group of setae; segment 5 shorter than 6; segment 6 hardly longer than broad, along posterior margin with 9 groups of marginal setae; palm slope oblique slightly less than half of posterior margin of segment 6; outer face with one strong corner spine accompanied by 6 lateral setae and by 3 slender toothed spines sitting laterally of main spine (fig. 6B); corner of inner face with one short submarginal spine; dactyl reaching posterior margin of segment 6, along outer margin with 8 single setae (fig. 6A).</paragraph>
|
||
<caption id="DF00666EFFC90C5A3174F94AAD6FF8E6" httpUri="https://zenodo.org/record/172136/files/figure.png" pageId="10" pageNumber="63" targetBox="[288,1308,322,1729]" targetPageId="10">
|
||
<paragraph id="8BC036E6FFC90C5A3174F94AAD6FF8E6" blockId="10.[264,1322,1767,1859]" pageId="10" pageNumber="63">
|
||
<emphasis id="B90BEAF4FFC90C5A3174F94AADDAF95A" bold="true" box="[264,396,1767,1791]" pageId="10" pageNumber="63">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C7F4D65FFC90C5A31EAF94AAE00F95B" box="[406,598,1767,1790]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="63" phylum="Arthropoda" rank="species" species="echion" status="sp. nov.">
|
||
<emphasis id="B90BEAF4FFC90C5A31EAF94AAE00F95B" box="[406,598,1767,1790]" italics="true" pageId="10" pageNumber="63">Niphargus echion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A238578FFFC90C5A3219F94AAEF2F95A" box="[613,676,1767,1791]" pageId="10" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEAF4FFC90C5A3219F94AAEF2F95A" bold="true" box="[613,676,1767,1791]" pageId="10" pageNumber="63">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype female, 10.3 mm, PliŠkovićeva jama cave: A–B, gnathopod 1; C–D, gnathopod 2; E, urosome with uropods 1–2. Scales bars: A, C, 0.5 mm; E, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF00666EFFC80C5B3174F8AFAEE7F8FA" httpUri="https://zenodo.org/record/172137/files/figure.png" pageId="11" pageNumber="64" targetBox="[326,1298,328,1753]" targetPageId="11">
|
||
<paragraph id="8BC036E6FFC80C5B3174F8AFAEE7F8FA" blockId="11.[264,1323,1794,1887]" pageId="11" pageNumber="64">
|
||
<emphasis id="B90BEAF4FFC80C5B3174F8AFADDCF8BF" bold="true" box="[264,394,1794,1818]" pageId="11" pageNumber="64">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C7F4D65FFC80C5B31EDF8AEAE19F8BF" box="[401,591,1795,1818]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="echion" status="sp. nov.">
|
||
<emphasis id="B90BEAF4FFC80C5B31EDF8AEAE19F8BF" box="[401,591,1795,1818]" italics="true" pageId="11" pageNumber="64">Niphargus echion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A238578FFFC80C5B3220F8AFAECEF8BF" box="[604,664,1794,1818]" pageId="11" pageNumber="64" rank="species">
|
||
<emphasis id="B90BEAF4FFC80C5B3220F8AFAECEF8BF" bold="true" box="[604,664,1794,1818]" pageId="11" pageNumber="64">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype female, 10.3 mm, PliŠkovićeva jama cave: A, inner plate of maxilliped; B, dactyl of maxilliped; C–D, pereopod 7; E, epimeral plates 2–3; F, uropod 3; G, telson. Scales bars: C, E, F, G, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC036E6FFCF0C5C3144FEB1AEE2FE5B" blockId="12.[264,1325,284,1150]" pageId="12" pageNumber="65">Gnathopod 2: segment 3 with one posterior marginal group of setae; segment 5 shorter than 6; segment 6 hardly broader than long, with palm slope oblique slightly over half of posterior margin of segment 6; palm defined on outer face by 1 strong corner spine accompanied by 5 long setae and by 2 slender toothed spines sitting lateral to main spine (fig. 6D); dactyl reaching posterior margin, and provided along outer margin with row of 11 setae (1–2–1–2–2–1–2) (fig. 6C).</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3144FDA1AE23FD3B" blockId="12.[264,1325,284,1150]" pageId="12" pageNumber="65">Pereopods 3–7: as in males. Pereopod 7 segment 2 longer than broad, without distinct anteroventral or posteroventral lobe; posterior margin with row of short marginal setae (fig. 7C). Dactyl of pereopod 7 with nail nearly as long as peduncle, and with one strong spine at inner margin (fig. 7D).</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3144FD01AFB8FD63" blockId="12.[264,1325,284,1150]" box="[312,1006,684,710]" pageId="12" pageNumber="65">Pleopods 1–3: with 2 or 3 retinaculae each (2–2–3, 2–2–3).</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3144FD79A8A7FCB3" blockId="12.[264,1325,284,1150]" pageId="12" pageNumber="65">Uropod 1: peduncle with dorsolateral row of spines and dorsomedial row of spinelike setae; rami nearly subequal, with single lateral and 4 or 5 distal short spines (fig. 6E).</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3144FC89AD6FFCC3" blockId="12.[264,1325,284,1150]" pageId="12" pageNumber="65">Uropod 2: rami subequal, long, with single lateral and 4 or 5 short distal spines (fig. 6E).</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3144FCD9AFE2FC7B" blockId="12.[264,1325,284,1150]" pageId="12" pageNumber="65">Uropod 3: with short second segment on outer ramus, bearing numerous distomarginal setae; first segment with spines on both margins and a row of long single plumose setae along inner margin (fig. 7F). Inner ramus short, scalelike.</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3144FC41ADE6FBDB" blockId="12.[264,1325,284,1150]" pageId="12" pageNumber="65">Telson: distinctly broader than long, incised slightly less than 2/3 of its length; each lobe with 3 distal and one group of 2 facial spines; along outer margin with 1 or 2 spines, along inner margin with 1 spine; a pair of plumose setae appears near the middle of each lobe (fig. 7G).</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3174FB02ADD2FB6D" blockId="12.[264,1323,1199,1502]" box="[264,388,1199,1224]" pageId="12" pageNumber="65">
|
||
<emphasis id="B90BEAF4FFCF0C5C3174FB02ADD2FB6D" box="[264,388,1199,1224]" italics="true" pageId="12" pageNumber="65">Variability</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3144FB79A8A3FA7B" blockId="12.[264,1323,1199,1502]" pageId="12" pageNumber="65">
|
||
Owing to the very scarce at hand, variability in taxonomic characters is difficult to assess. The number of distal spines on inner plate of maxilliped is rather variable (4 or 5), as well as the number of setae on inner plate of maxilla 1 (3–5). The number of retinaculae is 2 or 3, but not the all pleopods bear the same number. Thus, the variability in the retinacula remains problematic and must be resolved based on a additional material. Among the stable characters are the presence of two nails on each dactyl of the maxilliped, a character already observed and known for
|
||
<taxonomicName id="4C7F4D65FFCF0C5C336CFA69A8A7FA7B" authority="Schellelenberg, 1935" authorityName="Schellelenberg" authorityYear="1935" box="[784,1265,1476,1502]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="krameri">
|
||
<emphasis id="B90BEAF4FFCF0C5C336CFA69AFB9FA78" box="[784,1007,1476,1501]" italics="true" pageId="12" pageNumber="65">Niphargus krameri</emphasis>
|
||
Schellelenberg, 1935
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365656DFFCF0C5D3174F9A2A8E4FE5B" lastPageId="13" lastPageNumber="66" pageId="12" pageNumber="65" type="discussion">
|
||
<paragraph id="8BC036E6FFCF0C5C3174F9A2AD38F98D" blockId="12.[264,1324,1551,1899]" box="[264,366,1551,1576]" pageId="12" pageNumber="65">
|
||
<emphasis id="B90BEAF4FFCF0C5C3174F9A2AD38F98D" box="[264,366,1551,1576]" italics="true" pageId="12" pageNumber="65">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3144F99AAD3DF904" blockId="12.[264,1324,1551,1899]" pageId="12" pageNumber="65">
|
||
<taxonomicName id="4C7F4D65FFCF0C5C3144F99AAE5CF9F5" box="[312,522,1591,1616]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="echion" status="sp. nov.">
|
||
<emphasis id="B90BEAF4FFCF0C5C3144F99AAE5CF9F5" box="[312,522,1591,1616]" italics="true" pageId="12" pageNumber="65">Niphargus echion</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A238578FFFCF0C5C3260F99BAE37F9F5" box="[540,609,1590,1616]" pageId="12" pageNumber="65" rank="species">
|
||
<emphasis id="B90BEAF4FFCF0C5C3260F99BAE37F9F5" bold="true" box="[540,609,1590,1616]" pageId="12" pageNumber="65">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from all known species of the genus by the highest known number of distal spines (19 or 20) on the outer plate of maxilla
|
||
<quantity id="4C879B03FFCF0C5C3408F9F2A8F3F9DC" box="[1140,1189,1631,1657]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="65" unit="in" value="1.0">1 in</quantity>
|
||
males and females.
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCF0C5C3144F902ADC9F8CE" blockId="12.[264,1324,1551,1899]" pageId="12" pageNumber="65">
|
||
Most known
|
||
<taxonomicName id="4C7F4D65FFCF0C5C31AFF902AE18F96D" box="[467,590,1711,1736]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAF4FFCF0C5C31AFF902AE18F96D" box="[467,590,1711,1736]" italics="true" pageId="12" pageNumber="65">Niphargus</emphasis>
|
||
</taxonomicName>
|
||
species are provided with seven distal spines on outer plate of maxilla 1.
|
||
<taxonomicName id="4C7F4D65FFCF0C5C31F5F97AAF13F954" authority="G. Karaman, 1985" authorityName="G. Karaman" authorityYear="1985" box="[393,837,1751,1777]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="lunaris">
|
||
<emphasis id="B90BEAF4FFCF0C5C31F5F97AAE37F955" box="[393,609,1751,1776]" italics="true" pageId="12" pageNumber="65">Niphargus lunaris</emphasis>
|
||
<bibRefCitation id="EFEE4B17FFCF0C5C3216F97AAF13F954" author="Karaman" box="[618,837,1751,1777]" pageId="12" pageNumber="67" refString="Karaman, G. (1985) Contribution to the Knowledge of the Amphipoda 145. Two subterranean species of the Family Niphargidae, Niphargus lunaris, n. sp. and N. timavi S. Kar. 1954. Glasnik Republi c kog zavoda za za s titu prirode - Prirodnja c kog muzeja Titograd, 17, 29 - 46 (1984)." type="journal article" year="1985">G. Karaman, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3687676FFCF0C5C33EDF97AAFB3F954" box="[913,997,1751,1777]" name="Bosnia and Herzegovina" pageId="12" pageNumber="65">Bosnia</collectingCountry>
|
||
, however, is provided with eight distal spines;
|
||
<taxonomicName id="4C7F4D65FFCF0C5C3197F952AF14F8BC" authority="G. Karaman, 1984" authorityName="G. Karaman" authorityYear="1984" box="[491,834,1791,1817]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="sodalis">
|
||
<emphasis id="B90BEAF4FFCF0C5C3197F952AE37F8BD" box="[491,609,1791,1816]" italics="true" pageId="12" pageNumber="65">N. sodalis</emphasis>
|
||
G. Karaman, 1984
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="F3687676FFCF0C5C33ECF952AF91F8BC" box="[912,967,1791,1817]" name="Italy" pageId="12" pageNumber="65">Italy</collectingCountry>
|
||
, is provided with 10–12 distal spines; and
|
||
<taxonomicName id="4C7F4D65FFCF0C5C31E9F884AF67F8E6" authority="Vigna Taglianti, 1966" authorityName="Vigna Taglianti" authorityYear="1966" box="[405,817,1833,1859]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="65" phylum="Arthropoda" rank="species" species="pasquinii">
|
||
<emphasis id="B90BEAF4FFCF0C5C31E9F884AE70F8E7" box="[405,550,1833,1858]" italics="true" pageId="12" pageNumber="65">N. pasquinii</emphasis>
|
||
Vigna Taglianti, 1966
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="F3687676FFCF0C5C33FCF884AFE0F8E6" box="[896,950,1833,1859]" name="Italy" pageId="12" pageNumber="65">Italy</collectingCountry>
|
||
also, is provided with up to 15 distal spines.
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCE0C5D3144FEB1AF77FE73" blockId="13.[264,1322,284,510]" pageId="13" pageNumber="66">
|
||
The second very important character is the presence of two nails on the tip of maxilliped palp, a rare condition in
|
||
<taxonomicName id="4C7F4D65FFCE0C5D32C9FEE9AF66FEF8" box="[693,816,324,349]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAF4FFCE0C5D32C9FEE9AF66FEF8" box="[693,816,324,349]" italics="true" pageId="13" pageNumber="66">Niphargus</emphasis>
|
||
</taxonomicName>
|
||
. Only
|
||
<taxonomicName id="4C7F4D65FFCE0C5D33F8FEE9AD13FE23" authority="Schellellenberg, 1935" authorityName="Schellellenberg" authorityYear="1935" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="66" phylum="Arthropoda" rank="species" species="krameri">
|
||
<emphasis id="B90BEAF4FFCE0C5D33F8FEE9A832FEF8" box="[900,1124,324,349]" italics="true" pageId="13" pageNumber="66">Niphargus krameri</emphasis>
|
||
Schellellenberg, 1935
|
||
</taxonomicName>
|
||
, also from Istra and vicinity, is also provided usually with two nails (rarely with only one) on the dactyl of the maxilliped. Other species of
|
||
<taxonomicName id="4C7F4D65FFCE0C5D33E9FE39A846FE08" box="[917,1040,404,429]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAF4FFCE0C5D33E9FE39A846FE08" box="[917,1040,404,429]" italics="true" pageId="13" pageNumber="66">Niphargus</emphasis>
|
||
</taxonomicName>
|
||
are provided with only one distal nail on the dactyl of the maxilliped.
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCE0C5D3144FE49A8E4FE5B" blockId="13.[264,1322,284,510]" box="[312,1202,484,510]" pageId="13" pageNumber="66">
|
||
In other taxonomic characters
|
||
<taxonomicName id="4C7F4D65FFCE0C5D32E0FE49AF5AFE58" box="[668,780,484,509]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="66" phylum="Arthropoda" rank="species" species="echion">
|
||
<emphasis id="B90BEAF4FFCE0C5D32E0FE49AF5AFE58" box="[668,780,484,509]" italics="true" pageId="13" pageNumber="66">N. echion</emphasis>
|
||
</taxonomicName>
|
||
resembles other
|
||
<taxonomicName id="4C7F4D65FFCE0C5D33A8FE49A819FE58" box="[980,1103,484,509]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90BEAF4FFCE0C5D33A8FE49A819FE58" box="[980,1103,484,509]" italics="true" pageId="13" pageNumber="66">Niphargus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365656DFFCE0C5D3174FD99A808FC2B" pageId="13" pageNumber="66" type="etymology">
|
||
<paragraph id="8BC036E6FFCE0C5D3174FD99ADD3FDE8" blockId="13.[264,1325,564,790]" box="[264,389,564,589]" pageId="13" pageNumber="66">
|
||
<emphasis id="B90BEAF4FFCE0C5D3174FD99ADD3FDE8" box="[264,389,564,589]" italics="true" pageId="13" pageNumber="66">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCE0C5D3144FDF1AEDBFCB3" blockId="13.[264,1325,564,790]" pageId="13" pageNumber="66">
|
||
The new species,
|
||
<emphasis id="B90BEAF4FFCE0C5D3276FDF1AED5FDD0" box="[522,643,604,629]" italics="true" pageId="13" pageNumber="66">
|
||
<taxonomicName id="4C7F4D65FFCE0C5D3276FDF1AE28FDD0" box="[522,638,604,629]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="66" phylum="Arthropoda" rank="species" species="echion">N. echion</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
is named after
|
||
<taxonomicName id="4C7F4D65FFCE0C5D3347FDF1AFC7FDD3" box="[827,913,604,630]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="66" phylum="Arthropoda" rank="species" species="echion">Echion</taxonomicName>
|
||
, one of the five surviving warriors in Greek mythology, who was born out of the dragon’s teeth that Cadmus had sown when founding Thebes. The dragon’s triple row of teeth is reminiscent of the numerous distal spines on outer plate of the first maxilla of
|
||
<taxonomicName id="4C7F4D65FFCE0C5D337FFD79AF22FD48" box="[771,884,724,749]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="66" phylum="Arthropoda" rank="species" species="echion">
|
||
<emphasis id="B90BEAF4FFCE0C5D337FFD79AF22FD48" box="[771,884,724,749]" italics="true" pageId="13" pageNumber="66">N. echion</emphasis>
|
||
</taxonomicName>
|
||
, which distinguishes the new species from all other known Congeners.
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCE0C5D3174FCE1AD3EFCC0" blockId="13.[264,1118,844,910]" box="[264,360,844,869]" pageId="13" pageNumber="66">
|
||
<emphasis id="B90BEAF4FFCE0C5D3174FCE1AD3EFCC0" box="[264,360,844,869]" italics="true" pageId="13" pageNumber="66">Ecology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCE0C5D3144FCD9A808FC2B" blockId="13.[264,1118,844,910]" box="[312,1118,884,910]" pageId="13" pageNumber="66">The material was collected from a cave pool in the unsaturated zone.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C365656DFFCE0C5D3174FC69AE82FBA3" box="[264,724,964,1030]" pageId="13" pageNumber="66" type="distribution">
|
||
<paragraph id="8BC036E6FFCE0C5D3174FC69ADC1FC78" blockId="13.[264,724,964,1030]" box="[264,407,964,989]" pageId="13" pageNumber="66">
|
||
<emphasis id="B90BEAF4FFCE0C5D3174FC69ADC1FC78" box="[264,407,964,989]" italics="true" pageId="13" pageNumber="66">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC036E6FFCE0C5D3144FC41AE82FBA3" blockId="13.[264,724,964,1030]" box="[312,724,1004,1030]" pageId="13" pageNumber="66">
|
||
Known from the
|
||
<typeStatus id="54C48844FFCE0C5D327DFC41AE62FBA3" box="[513,564,1004,1030]" pageId="13" pageNumber="66">type</typeStatus>
|
||
locality only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |