841 lines
104 KiB
XML
841 lines
104 KiB
XML
<document id="1AF63C3A9A45563B3C011A5AE95B9BDB" ID-CLB-Dataset="27943" ID-DOI="10.11646/zootaxa.4523.1.1" ID-GBIF-Dataset="23903efe-ba7a-438c-a04f-d4b08680b640" ID-ISSN="1175-5326" ID-Zenodo-Dep="2610202" ID-ZooBank="96B7B9E3-9F49-4983-A46C-D29CD6B2EE49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553624654688" checkinUser="plazi" docAuthor="Mahony, Stephen, Kamei, Rachunliu G. & Teeling, Emma C." docDate="2018" docId="03D6878AFFE10270FF73FEF5FD74F92C" docLanguage="en" docName="zootaxa.4523.1.1.pdf" docOrigin="Zootaxa 4523 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Megophrys (Xenophrys) glandulosa Fei, Ye and Huang 1990" docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="FFEFFFF2FFDE0232FFE4FFD2FFDEFFCC" masterDocTitle="Cryptic diversity within the Megophrys major species group (Amphibia: Megophryidae) of the Asian Horned Frogs: Phylogenetic perspectives and a taxonomic revision of South Asian taxa, with descriptions of four new species" masterLastPageNumber="96" masterPageNumber="1" pageNumber="64" updateTime="1698713983112" updateUser="ExternalLinkService">
|
||
<mods:mods id="E381EA3EB6C8C47815B466407249AEFF" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="862C7805BE4B89CF5046FD813F877BA6">
|
||
<mods:title id="E86F72191E6DE42D045DE158A1BB79EA">Cryptic diversity within the Megophrys major species group (Amphibia: Megophryidae) of the Asian Horned Frogs: Phylogenetic perspectives and a taxonomic revision of South Asian taxa, with descriptions of four new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="ED47B9D8FBAFA952E1D6EACD12123999" type="personal">
|
||
<mods:role id="C0926EF369AD0C32FEDC2AB0398DFBA9">
|
||
<mods:roleTerm id="FEB41BC81E3661FDA688F4A1A28E9E30">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E4B4AE7160794895ABBF9B1D6FA665A4">Mahony, Stephen</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="504662BD22C45445D461E0D13394F682" type="personal">
|
||
<mods:role id="1A586360CA8E770BA81CA9BCAFF262B0">
|
||
<mods:roleTerm id="421E931327C0A7D04D5884D4D113861C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="71BCA06C8181EC4926183D4AB6F50E78">Kamei, Rachunliu G.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A0FE41AB6BFDCC768DD5C08CFC677227" type="personal">
|
||
<mods:role id="FDC737C34FD462E570984CA8B6AECA9C">
|
||
<mods:roleTerm id="CD18825C2942B13D72E208BAD950FED2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AAEE00960A249CB1092DDF79C2D8EAA0">Teeling, Emma C.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="6A4894C65761FA5C2D0CB215B8B137FF">text</mods:typeOfResource>
|
||
<mods:relatedItem id="DA10C37E8F6E5FA75534BFF201726F9A" type="host">
|
||
<mods:titleInfo id="DECCB08DA4E2FF44F4FF6E021E2D5E31">
|
||
<mods:title id="A4414ECE9D61B7510F286EBD5CA84CA4">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="256C91F6324399AE56D2A9F69D92B0C0">
|
||
<mods:date id="EC8B73B9A5CAA3B3F1242A13B308ED11">2018</mods:date>
|
||
<mods:detail id="B80EA7E8DCCB0EA52902C2722D186E58" type="pubDate">
|
||
<mods:number id="A4A5ED6D12139EC04DA618E7C308F56D">2018-11-19</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="97D0044E25F59A9D84303AAFC97C24AC" type="volume">
|
||
<mods:number id="E99D7304A1D0A1508B72EB444EEB7833">4523</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B7826FE40467698B507E69E488E1A881" type="issue">
|
||
<mods:number id="DD727563D16A9098B9C080C535F41758">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4DA342642B8703FF3DB54F3779914EE7" unit="page">
|
||
<mods:start id="62A849F775D73D26B188663B7FE6CB8C">1</mods:start>
|
||
<mods:end id="D2CAE01596997A443B117D4DE98F40CD">96</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="1ECCC35AFA685C6879153B6897E54A48">journal article</mods:classification>
|
||
<mods:identifier id="B185FDC0647DD684C3C34E80F3F901C4" type="CLB-Dataset">27943</mods:identifier>
|
||
<mods:identifier id="68EF78D7EDA825AADEBC51F133A5439A" type="DOI">10.11646/zootaxa.4523.1.1</mods:identifier>
|
||
<mods:identifier id="ECAD0C0A50F79EEF952ED2BA71428318" type="GBIF-Dataset">23903efe-ba7a-438c-a04f-d4b08680b640</mods:identifier>
|
||
<mods:identifier id="09113DD9CA577BA77FD41663A3E7F0B7" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="412E6B31589070E8A3D327F4A73E5C1E" type="Zenodo-Dep">2610202</mods:identifier>
|
||
<mods:identifier id="ED848A37296FCB90B0D814658B57AC58" type="ZooBank">96B7B9E3-9F49-4983-A46C-D29CD6B2EE49</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03D6878AFFE10270FF73FEF5FD74F92C" ID-DOI="http://doi.org/10.5281/zenodo.6490316" ID-GBIF-Taxon="154499352" ID-Zenodo-Dep="6490316" LSID="urn:lsid:plazi:treatment:03D6878AFFE10270FF73FEF5FD74F92C" httpUri="http://treatment.plazi.org/id/03D6878AFFE10270FF73FEF5FD74F92C" lastPageId="66" lastPageNumber="67" pageId="63" pageNumber="64">
|
||
<subSubSection id="C3656517FFE1020DFF73FEF5FC5EFE8E" box="[151,896,295,322]" pageId="63" pageNumber="64" type="nomenclature">
|
||
<paragraph id="8BC0369CFFE1020DFF73FEF5FC5EFE8E" blockId="63.[151,896,295,355]" box="[151,896,295,322]" pageId="63" pageNumber="64">
|
||
<heading id="D08881F0FFE1020DFF73FEF5FC5EFE8E" bold="true" box="[151,896,295,322]" fontSize="11" level="1" pageId="63" pageNumber="64" reason="1">
|
||
<taxonomicName id="4C7F4D1FFFE1020DFF73FEF5FC5EFE8E" ID-CoL="6QXMX" authority="Fei, Ye and Huang, 1990" authorityName="Fei, Ye and Huang" authorityYear="1990" box="[151,896,295,322]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="glandulosa" subGenus="Xenophrys">
|
||
<emphasis id="B90BEA8EFFE1020DFF73FEF5FC5EFE8E" bold="true" box="[151,896,295,322]" pageId="63" pageNumber="64">
|
||
<emphasis id="B90BEA8EFFE1020DFF73FEF5FEC0FE8D" bold="true" box="[151,286,295,321]" italics="true" pageId="63" pageNumber="64">Megophrys</emphasis>
|
||
(
|
||
<emphasis id="B90BEA8EFFE1020DFECBFEF5FE6FFE8D" bold="true" box="[303,433,295,321]" italics="true" pageId="63" pageNumber="64">Xenophrys</emphasis>
|
||
)
|
||
<emphasis id="B90BEA8EFFE1020DFE26FEF5FD96FE8D" bold="true" box="[450,584,295,321]" italics="true" pageId="63" pageNumber="64">glandulosa</emphasis>
|
||
<bibRefCitation id="EFEE4B6DFFE1020DFDABFEFAFC5EFE8E" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[591,896,296,322]" pageId="63" pageNumber="64" pagination="2" refId="ref52831" refString="Fei, L., Ye, C. - Y. & Huang, Y. - Z. (1990) Key to Chinese Amphibians. Chongqing Branch, Science and Technology Literature Publishing House, Chongqing, 364 + 2 + [iv] pp." type="journal article" year="1990">Fei, Ye and Huang, 1990</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FFE1020DFF73FE98FEA9FEAF" box="[151,375,330,355]" pageId="63" pageNumber="64" type="description">
|
||
<paragraph id="8BC0369CFFE1020DFF73FE98FEA9FEAF" blockId="63.[151,896,295,355]" box="[151,375,330,355]" pageId="63" pageNumber="64">
|
||
(
|
||
<figureCitation id="13442A19FFE1020DFF7AFE98FED1FEAF" box="[158,271,330,355]" captionStart="FIGURE 24" captionStartId="64.[151,250,1907,1929]" captionTargetBox="[151,1436,553,1885]" captionTargetId="figure@64.[151,1436,553,1885]" captionTargetPageId="64" captionText="FIGURE 24. Megophrys glandulosa holotype: adult male (CIB 873112: SVL 78.4 mm [Fei et al. 1992]) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610250/files/figure.png" pageId="63" pageNumber="64">Figure 24</figureCitation>
|
||
;
|
||
<tableCitation id="C6FD0327FFE1020DFEFFFE98FEB2FEAF" box="[283,364,330,355]" captionStart="TABLE 1" captionStartId="22.[147,225,525,545]" captionText="TABLE 1. Morphometric details for specimens new and known included in the Taxonomic review section. Refer to Methods and materials section for explanation of measurement abbreviations and museum acronyms. HT = holotype; PT = paratype; LT = lectotype; PLT = paralectotype; NT = neotype; RS = referred specimen; M = adult male; F = adult female; SA = subadult; J = juvenile [post metamorphic]; † = BMNH voucher numbers; # = SDBDU voucher numbers; ind = character limits insufficiently distinct to reliably measure." pageId="63" pageNumber="64">Table 1</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FFE1020DFF73FE40FC93FE0B" pageId="63" pageNumber="64" type="reference_group">
|
||
<paragraph id="8BC0369CFFE1020DFF73FE40FC93FE0B" blockId="63.[151,1436,402,455]" pageId="63" pageNumber="64">
|
||
<treatmentCitationGroup id="AB6F11B2FFE1020DFF73FE40FC93FE0B" pageId="63" pageNumber="64">
|
||
<taxonomicName id="4C7F4D1FFFE1020DFF73FE40FD7AFE64" ID-CoL="6QXMX" authority="Fei, Ye and Huang 1990: 99" authorityName="Fei, Ye and Huang" authorityPageNumber="99" authorityYear="1990" box="[151,676,402,424]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFFE1020DFF73FE40FE5FFE64" box="[151,385,402,424]" italics="true" pageId="63" pageNumber="64">Megophrys glandulosa</emphasis>
|
||
<treatmentCitation id="0ADE108DFFE1020DFE6CFE40FD7AFE64" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[392,676,402,424]" page="99" pageId="63" pageNumber="64" year="1990">
|
||
<bibRefCitation id="EFEE4B6DFFE1020DFE6CFE40FD58FE64" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[392,646,402,424]" pageId="63" pageNumber="64" pagination="2" refId="ref52831" refString="Fei, L., Ye, C. - Y. & Huang, Y. - Z. (1990) Key to Chinese Amphibians. Chongqing Branch, Science and Technology Literature Publishing House, Chongqing, 364 + 2 + [iv] pp." type="journal article" year="1990">Fei, Ye and Huang 1990</bibRefCitation>
|
||
:99
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, 273.
|
||
<emphasis id="B90BEA8EFFE1020DFD06FE41FD29FE64" box="[738,759,403,424]" italics="true" pageId="63" pageNumber="64">In</emphasis>
|
||
:
|
||
<emphasis id="B90BEA8EFFE1020DFCE2FE41FBF7FE64" box="[774,1065,402,424]" italics="true" pageId="63" pageNumber="64">Key to Chinese Amphibians.</emphasis>
|
||
Publishing House for Scientific and Technological Literature,
|
||
<collectingRegion id="49BBF87EFFE1020DFE28FE63FDE1FE0B" box="[460,575,433,455]" country="China" name="Chongqing" pageId="63" pageNumber="64">Chongqing</collectingRegion>
|
||
,
|
||
<collectingCountry id="F368760CFFE1020DFDAEFE63FD54FE0B" box="[586,650,433,455]" name="China" pageId="63" pageNumber="64">China</collectingCountry>
|
||
: 364 pp. + 2 + [iv].
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FFE1020DFF73FE26FF08FBC8" pageId="63" pageNumber="64" type="materials_examined">
|
||
<paragraph id="8BC0369CFFE1020DFF73FE26FC28FD98" blockId="63.[151,1437,500,2037]" pageId="63" pageNumber="64">
|
||
<materialsCitation id="3B173CC1FFE1020DFF73FE26FC28FD98" ID-GBIF-Occurrence="2028521210" collectingDate="1987-03-29" collectionCode="CIB" collectorName="Liang Fei" country="China" county="Jingdong Yi Autonomous County" elevation="1900" latitude="24.824167" location="Adult" longLatPrecision="21" longitude="100.44195" municipality="Pu'er Prefecture" pageId="63" pageNumber="64" specimenCode="CIB 873112" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
|
||
<emphasis id="B90BEA8EFFE1020DFF73FE26FED5FDC1" bold="true" box="[151,267,500,525]" pageId="63" pageNumber="64">
|
||
<typeStatus id="54C4883EFFE1020DFF73FE26FED8FDC1" box="[151,262,500,525]" pageId="63" pageNumber="64" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<location id="8EA06047FFE1020DFEF5FE26FE8FFDC1" LSID="urn:lsid:plazi:treatment:03D6878AFFE10270FF73FEF5FD74F92C:8EA06047FFE1020DFEF5FE26FE8FFDC1" box="[273,337,500,525]" country="China" county="Jingdong Yi Autonomous County" latitude="24.824167" longLatPrecision="21" longitude="100.44195" municipality="Pu'er Prefecture" name="Adult" pageId="63" pageNumber="64" stateProvince="Yunnan">Adult</location>
|
||
male (
|
||
<specimenCode id="DBD99EE7FFE1020DFE78FE26FDF4FDC0" box="[412,554,500,524]" collectionCode="CIB" country="China" httpUri="http://grbio.org/cool/vxkr-andy" name="Chengdu Institute of Biology" pageId="63" pageNumber="64">CIB 873112</specimenCode>
|
||
:
|
||
<figureCitation id="13442A19FFE1020DFDD0FE26FD7CFDC0" box="[564,674,500,525]" captionStart="FIGURE 24" captionStartId="64.[151,250,1907,1929]" captionTargetBox="[151,1436,553,1885]" captionTargetId="figure@64.[151,1436,553,1885]" captionTargetPageId="64" captionText="FIGURE 24. Megophrys glandulosa holotype: adult male (CIB 873112: SVL 78.4 mm [Fei et al. 1992]) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610250/files/figure.png" pageId="63" pageNumber="64">Figure 24</figureCitation>
|
||
), from “Wuliang shan, Jingdong,
|
||
<collectingRegion id="49BBF87EFFE1020DFBFEFE27FB03FDC1" box="[1050,1245,500,525]" country="China" name="Yunnan" pageId="63" pageNumber="64">Yunnan Province</collectingRegion>
|
||
, altitude
|
||
<elevation id="0052D1AFFFE1020DFAA6FE26FA4CFDC0" box="[1346,1426,500,524]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="63" pageNumber="64" unit="m" value="1900.0">
|
||
<quantity id="4C879B79FFE1020DFAA6FE26FA4CFDC0" box="[1346,1426,500,524]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="63" pageNumber="64" unit="m" value="1900.0">1900m</quantity>
|
||
</elevation>
|
||
” (=
|
||
<location id="8EA06047FFE1020DFF49FDC5FE59FDFC" LSID="urn:lsid:plazi:treatment:03D6878AFFE10270FF73FEF5FD74F92C:8EA06047FFE1020DFF49FDC5FE59FDFC" box="[173,391,535,560]" country="China" county="Jingdong Yi Autonomous County" latitude="24.824167" longLatPrecision="21" longitude="100.44195" municipality="Pu'er Prefecture" name="Wuliang Mountain" pageId="63" pageNumber="64" stateProvince="Yunnan">Wuliang Mountain</location>
|
||
[~
|
||
<geoCoordinate id="EE4B505BFFE1020DFE43FDCAFDEEFDFC" box="[423,560,536,560]" direction="north" orientation="latitude" pageId="63" pageNumber="64" precision="15" value="24.824167">24°49'27"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFFE1020DFDDBFDC5FD0DFDFC" box="[575,723,535,560]" direction="east" orientation="longitude" pageId="63" pageNumber="64" precision="15" value="100.44195">100°26'31"E</geoCoordinate>
|
||
],
|
||
<collectingCounty id="62A14E10FFE1020DFD0CFDC5FBB3FDFC" box="[744,1133,535,560]" pageId="63" pageNumber="64">Jingdong Yi Autonomous County</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BA4ACE6FFE1020DFB9FFDCAFAE6FDFC" box="[1147,1336,535,560]" pageId="63" pageNumber="64">Pu’er Prefecture</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49BBF87EFFE1020DFAA3FDCAFF20FD99" country="China" name="Yunnan" pageId="63" pageNumber="64">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F368760CFFE1020DFEECFDEEFE91FD99" box="[264,335,572,597]" name="China" pageId="63" pageNumber="64">China</collectingCountry>
|
||
), collected by
|
||
<collectorName id="268A534AFFE1020DFE16FDEEFD80FD99" box="[498,606,572,597]" pageId="63" pageNumber="64">Liang Fei</collectorName>
|
||
on
|
||
<collectingDate id="EF85E9B4FFE1020DFD6CFDEEFCEDFD98" box="[648,819,572,597]" pageId="63" pageNumber="64" value="1987-03-29">
|
||
<date id="FFC1105CFFE1020DFD6CFDEEFCEDFD98" box="[648,819,572,597]" pageId="63" pageNumber="64" value="1987-03-29">29 March 1987</date>
|
||
</collectingDate>
|
||
(
|
||
<bibRefCitation id="EFEE4B6DFFE1020DFCA6FDEEFC37FD98" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[834,1001,572,597]" pageId="63" pageNumber="64" pagination="5 - 12" refId="ref52880" refString="Fei, L., Ye, C. - Y. & Huang, Y. - Z. (1992) Two new species of the genus Megophrys, Pelobatidae (Amphibia, Anura) from China. Zoological Research, 13 (1), 5 - 12." type="journal article" year="1992">
|
||
Fei
|
||
<emphasis id="B90BEA8EFFE1020DFC8AFDEFFC78FD99" box="[878,934,572,597]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
|
||
1992
|
||
</bibRefCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE1020DFF23FD8DFBEAFD28" blockId="63.[151,1437,500,2037]" pageId="63" pageNumber="64">
|
||
<emphasis id="B90BEA8EFFE1020DFF23FD8DFE98FDB4" bold="true" box="[199,326,607,632]" pageId="63" pageNumber="64">
|
||
<typeStatus id="54C4883EFFE1020DFF23FD8DFE9FFDB4" box="[199,321,607,632]" pageId="63" pageNumber="64" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult female (“
|
||
<typeStatus id="54C4883EFFE1020DFDE5FD8DFDBFFDB4" box="[513,609,607,632]" pageId="63" pageNumber="64" type="allotype">allotype</typeStatus>
|
||
” CIB 873201), from the “same locality as that of the
|
||
<typeStatus id="54C4883EFFE1020DFB32FD8DFAE5FDB4" box="[1238,1339,607,632]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
, altitude
|
||
<quantity id="4C879B79FFE1020DFF73FD56FF37FD50" box="[151,233,644,668]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="63" pageNumber="64" unit="m" value="2100.0">2100m</quantity>
|
||
”, collector not mentioned, 0
|
||
<date id="FFC1105CFFE1020DFDCFFD56FD66FD51" box="[555,696,644,669]" pageId="63" pageNumber="64" value="1975-05-02">2 May 1975</date>
|
||
;
|
||
<specimenCount id="9D79FD15FFE1020DFD22FD56FCF5FD51" box="[710,811,644,669]" pageId="63" pageNumber="64" type="male">25 males</specimenCount>
|
||
and two females (specimen numbers not mentioned in
|
||
<bibRefCitation id="EFEE4B6DFFE1020DFF73FD75FE8CFD0C" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[151,338,679,704]" pageId="63" pageNumber="64" pagination="5 - 12" refId="ref52880" refString="Fei, L., Ye, C. - Y. & Huang, Y. - Z. (1992) Two new species of the genus Megophrys, Pelobatidae (Amphibia, Anura) from China. Zoological Research, 13 (1), 5 - 12." type="journal article" year="1992">
|
||
Fei
|
||
<emphasis id="B90BEA8EFFE1020DFF27FD7BFF22FD0C" box="[195,252,679,704]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
|
||
[1992]
|
||
</bibRefCitation>
|
||
), from “Wuliang shan, Jingdong,
|
||
<collectingRegion id="49BBF87EFFE1020DFD31FD7AFC44FD0C" box="[725,922,679,704]" country="China" name="Yunnan" pageId="63" pageNumber="64">Yunnan Province</collectingRegion>
|
||
, altitude
|
||
<quantity id="4C879B79FFE1020DFBE7FD7AFB45FD0C" box="[1027,1179,680,704]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="63" pageNumber="64" unit="m" value="2000.0" valueMax="2100.0" valueMin="1900.0">1900–2100m</quantity>
|
||
”, collectors Liang Fei, Younzhao Huang and Jiarui Luo, between
|
||
<date id="FFC1105CFFE1020DFD90FD1EFCACFD28" box="[628,882,716,741]" pageId="63" pageNumber="64" value="1987-03-29" valueMax="1987-03-30" valueMin="1987-03-29">29 and 30 March 1987</date>
|
||
(
|
||
<bibRefCitation id="EFEE4B6DFFE1020DFC64FD1EFBF9FD28" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[896,1063,716,741]" pageId="63" pageNumber="64" pagination="5 - 12" refId="ref52880" refString="Fei, L., Ye, C. - Y. & Huang, Y. - Z. (1992) Two new species of the genus Megophrys, Pelobatidae (Amphibia, Anura) from China. Zoological Research, 13 (1), 5 - 12." type="journal article" year="1992">
|
||
Fei
|
||
<emphasis id="B90BEA8EFFE1020DFC48FD1FFC3AFD29" box="[940,996,716,741]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
|
||
1992
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE1020DFF23FD3DFBDBFCC4" blockId="63.[151,1437,500,2037]" box="[199,1029,751,776]" pageId="63" pageNumber="64">
|
||
<emphasis id="B90BEA8EFFE1020DFF23FD3DFE16FCC4" bold="true" box="[199,456,751,776]" pageId="63" pageNumber="64">Examined specimens.</emphasis>
|
||
Adult male (CIB 873112––photos only),
|
||
<typeStatus id="54C4883EFFE1020DFC7FFD3DFBDFFCC4" box="[923,1025,751,776]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE1020DFF23FCC6FF08FBC8" blockId="63.[151,1437,500,2037]" pageId="63" pageNumber="64">
|
||
<emphasis id="B90BEA8EFFE1020DFF23FCC6FD3CFCE1" bold="true" box="[199,738,788,813]" pageId="63" pageNumber="64">Provisionally referred examined specimens.</emphasis>
|
||
Two adult males (CAS 221395, CAS 221484), from two localities on the road between Ahtonga and Babaw (
|
||
<geoCoordinate id="EE4B505BFFE1020DFD39FCE5FCA4FC9C" box="[733,890,823,848]" direction="north" orientation="latitude" pageId="63" pageNumber="64" precision="1" value="27.257555">27°15'27.2"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFFE1020DFC61FCE5FBC4FC9C" box="[901,1050,823,848]" direction="east" orientation="longitude" pageId="63" pageNumber="64" precision="1" value="97.84234">97°50'32.4"E</geoCoordinate>
|
||
and
|
||
<geoCoordinate id="EE4B505BFFE1020DFBB4FCEAFB32FC9C" box="[1104,1260,824,848]" direction="north" orientation="latitude" pageId="63" pageNumber="64" precision="1" value="27.290083">27°17'24.3"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFFE1020DFB12FCE5FA4EFC9C" box="[1270,1424,823,848]" direction="east" orientation="longitude" pageId="63" pageNumber="64" precision="1" value="97.864555">97°51'52.4"E</geoCoordinate>
|
||
), Machanbaw township, Putao district,
|
||
<collectingRegion id="49BBF87EFFE1020DFDD9FC8EFD50FCB9" box="[573,654,860,885]" country="Myanmar" name="Kachin" pageId="63" pageNumber="64">Kachin</collectingRegion>
|
||
state,
|
||
<collectingCountry id="F368760CFFE1020DFD31FC8FFC9AFCB9" box="[725,836,861,885]" name="Myanmar" pageId="63" pageNumber="64">Myanmar</collectingCountry>
|
||
, collected by H. Win and R. Shaung on 0
|
||
<date id="FFC1105CFFE1020DFAF5FC8EFF0CFC54" pageId="63" pageNumber="64" value="2001-09-05">5 September 2001</date>
|
||
, and J.B. Slowinski, D.-O. Rao, G.O.U. Wogan, H. Win, A.K. Shein and H. Tun on 0
|
||
<date id="FFC1105CFFE1020DFB7CFC52FAB6FC54" box="[1176,1384,895,920]" pageId="63" pageNumber="64" value="2001-09-07">7 September 2001</date>
|
||
;
|
||
<materialsCitation id="3B173CC1FFE1020DFA90FC52FF0DFBC8" ID-GBIF-Occurrence="2028521219" collectingDate="2001-09-07" collectionCode="CAS" collectorName="J. B. Slowinski & Slowinski, G. O. U & Wogan, H & A. K. Shein" country="Myanmar" latitude="27.293444" location="Naung Mon township" longLatPrecision="13" longitude="97.863884" municipality="Putao district" pageId="63" pageNumber="64" specimenCode="CAS 221442" specimenCount="1" stateProvince="Kachin">
|
||
one adult male (
|
||
<specimenCode id="DBD99EE7FFE1020DFEFEFC76FE6FFC70" box="[282,433,932,956]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="64" type="Museum">CAS 221442</specimenCode>
|
||
), from Aureinga Camp (
|
||
<geoCoordinate id="EE4B505BFFE1020DFD25FC76FC80FC71" box="[705,862,932,957]" direction="north" orientation="latitude" pageId="63" pageNumber="64" precision="1" value="27.293444">27°17'36.4"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFFE1020DFC8DFC76FC32FC71" box="[873,1004,932,957]" direction="east" orientation="longitude" pageId="63" pageNumber="64" precision="15" value="97.863884">97°51'50"E</geoCoordinate>
|
||
),
|
||
<location id="8EA06047FFE1020DFBE4FC77FB29FC71" LSID="urn:lsid:plazi:treatment:03D6878AFFE10270FF73FEF5FD74F92C:8EA06047FFE1020DFBE4FC77FB29FC71" box="[1024,1271,932,957]" country="Myanmar" latitude="27.293444" longLatPrecision="13" longitude="97.863884" municipality="Putao district" name="Naung Mon township" pageId="63" pageNumber="64" stateProvince="Kachin">Naung Mon township</location>
|
||
,
|
||
<collectingMunicipality id="6BA4ACE6FFE1020DFAE5FC77FA47FC71" box="[1281,1433,932,957]" pageId="63" pageNumber="64">Putao district</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49BBF87EFFE1020DFF73FC15FF36FC2C" box="[151,232,967,992]" country="Myanmar" name="Kachin" pageId="63" pageNumber="64">Kachin</collectingRegion>
|
||
state,
|
||
<collectingCountry id="F368760CFFE1020DFED3FC1AFE78FC2C" box="[311,422,968,992]" name="Myanmar" pageId="63" pageNumber="64">Myanmar</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268A534AFFE1020DFDAEFC1AFD2DFC2C" box="[586,755,967,992]" pageId="63" pageNumber="64">J.B. Slowinski</collectorName>
|
||
, G.O.U.
|
||
<collectorName id="268A534AFFE1020DFCBFFC1AFC0CFC13" box="[859,978,968,992]" pageId="63" pageNumber="64">Wogan, H</collectorName>
|
||
. Win and
|
||
<collectorName id="268A534AFFE1020DFBA9FC1AFB10FC2C" box="[1101,1230,967,992]" pageId="63" pageNumber="64">A.K. Shein</collectorName>
|
||
on 0
|
||
<date id="FFC1105CFFE1020DFAE9FC1AFF0DFBC8" pageId="63" pageNumber="64" value="2001-09-07">
|
||
<collectingDate id="EF85E9B4FFE1020DFAE9FC1AFF0DFBC8" pageId="63" pageNumber="64" value="2001-09-07">7 September 2001</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FFE10270FF23FBDDFB7DFE39" lastPageId="66" lastPageNumber="67" pageId="63" pageNumber="64" type="description">
|
||
<paragraph id="8BC0369CFFE1020DFF23FBDDFEDEFBBC" blockId="63.[151,1437,500,2037]" pageId="63" pageNumber="64">
|
||
<emphasis id="B90BEA8EFFE1020DFF23FBDDFE23FBE4" bold="true" box="[199,509,1039,1064]" pageId="63" pageNumber="64">Description of type series.</emphasis>
|
||
Refer to
|
||
<bibRefCitation id="EFEE4B6DFFE1020DFD8CFBDDFCC0FBE4" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[616,798,1039,1064]" pageId="63" pageNumber="64" pagination="5 - 12" refId="ref52880" refString="Fei, L., Ye, C. - Y. & Huang, Y. - Z. (1992) Two new species of the genus Megophrys, Pelobatidae (Amphibia, Anura) from China. Zoological Research, 13 (1), 5 - 12." type="journal article" year="1992">
|
||
Fei
|
||
<emphasis id="B90BEA8EFFE1020DFD77FBC3FD12FBE4" box="[659,716,1039,1064]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
|
||
(1992)
|
||
</bibRefCitation>
|
||
for the description in Chinese language text, and
|
||
<bibRefCitation id="EFEE4B6DFFE1020DFAB6FBC2FEFBFB80" author="Huang, C. - M. & Lathrop, A. & Murphy, R. W." pageId="63" pageNumber="64" pagination="1 - 20" refId="ref54291" refString="Huang, C. - M., Lathrop, A. & Murphy, R. W. (1998) Translations of " Two new species of amphibian in Tibet " Huang & Fei 1981 and " Description of two new species of the genus Megophrys " Fei, Ye, & Huang 1995 [1992]. Smithsonian Herpetological Information Service, 118, 1 - 20." type="journal article" year="1998">
|
||
Huang
|
||
<emphasis id="B90BEA8EFFE1020DFF73FBE7FF0FFB81" box="[151,209,1076,1101]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
|
||
(1998)
|
||
</bibRefCitation>
|
||
for the English translation of
|
||
<bibRefCitation id="EFEE4B6DFFE1020DFD66FBE6FCE1FB80" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[642,831,1076,1101]" pageId="63" pageNumber="64" pagination="5 - 12" refId="ref52880" refString="Fei, L., Ye, C. - Y. & Huang, Y. - Z. (1992) Two new species of the genus Megophrys, Pelobatidae (Amphibia, Anura) from China. Zoological Research, 13 (1), 5 - 12." type="journal article" year="1992">
|
||
Fei
|
||
<emphasis id="B90BEA8EFFE1020DFD4BFBE7FD37FB81" box="[687,745,1076,1101]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
|
||
(1992)
|
||
</bibRefCitation>
|
||
. See
|
||
<figureCitation id="13442A19FFE1020DFC9AFBE6FC33FB80" box="[894,1005,1076,1101]" captionStart="FIGURE 24" captionStartId="64.[151,250,1907,1929]" captionTargetBox="[151,1436,553,1885]" captionTargetId="figure@64.[151,1436,553,1885]" captionTargetPageId="64" captionText="FIGURE 24. Megophrys glandulosa holotype: adult male (CIB 873112: SVL 78.4 mm [Fei et al. 1992]) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610250/files/figure.png" pageId="63" pageNumber="64">Figure 24</figureCitation>
|
||
for colouration in preservative of the
|
||
<typeStatus id="54C4883EFFE1020DFF73FB85FF23FBBC" box="[151,253,1111,1136]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE1020DFF23FBAEFD10F9EC" blockId="63.[151,1437,500,2037]" pageId="63" pageNumber="64">
|
||
<emphasis id="B90BEA8EFFE1020DFF23FBAEFBFBFB59" bold="true" box="[199,1061,1148,1173]" pageId="63" pageNumber="64">Description of referred specimen CAS 22 1442 (measurements in mm).</emphasis>
|
||
Mature male (SVL 81.0). Head large (HW 31.3, HL 30.2, IFE 13.6, IBE 21.2), wider than long, snout broadly pointed in dorsal view, obtusely protruding beyond mandible in lateral view, without rostral appendage; loreal region acute, concave, with well developed canthus rostralis; dorsal surface of snout concave; eye diameter larger than maximum tympanum diameter, and shorter than snout (EL 7.9, TYD 5.1, SL 10.1); eye–tympanum distance (TYE 6.7) longer than tympanum diameter; tympanum distinctly oval, obliquely orientated, upper ~10% concealed by supratympanic ridge; pupil vertically elliptical; nostril positioned laterally, oval, obliquely oriented with raised posterior rim, closer to eye than to snout (EN 4.6, SN 6.2); upper eyelid width subequal to narrowest point between upper eyelids, and less than internarial distance (UEW 7.5, IUE 7.6, IN 9.3); pineal ocellus absent; vomerine ridges well developed, ovoid, positioned between choanae, extending posterior to choanae, slightly closer to choanae than to each other; vomerine teeth present, moderately long; maxillary teeth present; tongue moderately large, distinctly notched posteriorly, medial lingual process absent.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE1020DFF23F9FEFA5BF844" blockId="63.[151,1437,500,2037]" pageId="63" pageNumber="64">
|
||
Forelimbs moderately long, thick; forearms enlarged relative to upper forelimbs, and shorter than hand length (FAL 17.3, HAL 21.3); fingers moderately long, narrow, finger length formula IV<I=II<III (FIL 10.6, FIIL 10.6, FIIIL 13.1, FIVL 8.4); interdigital webbing, lateral fringes, subarticular and supernumerary tubercles absent; thenar tubercle distinct; metacarpal tubercles absent; digit tips slightly dilated, flattened, with subcircular pads, terminal grooves on pads absent. Hindlimbs relatively long, thin; thigh length slightly shorter than shanks, and longer than feet (TL 39.3, SHL 41.4, FOL 37.9); toes long, thin, relative toe lengths I<II<V<III<IV; digit tips not dilated but flattened, with subcircular pads, terminal grooves on pads absent; toes with thick basal webbing, rudimentary between digits I and II, remaining webbing formula as follows: II1.9–
|
||
<date id="FFC1105CFFE1020DFC3EF8FAFBDDF88C" box="[986,1027,1832,1856]" pageId="63" pageNumber="64">3III</date>
|
||
2.3–3.9IV3.8–
|
||
<date id="FFC1105CFFE1020DFB4DF8FAFB15F88C" box="[1193,1227,1832,1856]" pageId="63" pageNumber="64">2V</date>
|
||
; moderately wide lateral fringes present on all toes; outer metatarsal, subarticular and supernumerary tubercles all absent; inner metatarsal tubercle present but barely distinguishable; ridge of thickened skin absent on ventral surface of digits.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE10272FF23F846FB24FE39" blockId="63.[151,1437,500,2037]" lastBlockId="64.[151,1437,151,501]" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">Skin on dorsal surfaces of snout, head and back primarily smooth; throat, chest and limbs smooth; flanks with moderately large tubercles approximately forming two longitudinal rows, interspersed with densely scattered small granular tubercles; posterior thigh and cloacal region primarily smooth with small scattered weakly raised tubercles; tympanum mostly smooth, slightly concave; sides of head finely granular; palpebral horn absent, replaced by short fleshy bump; supratympanic ridge narrow anteriorly, extends from posterior orbital border, curves down along upper and posterior border of tympanum, considerably broadening posteriorly, terminating above shoulder; narrow, raised dorsolateral ridge extending posteriorly from behind supratympanic ridge to ~75% trunk length on both sides; weakly raised V-shaped parietoscapular ridge present, extending posteriorly from temporal region on each side, meeting medially at level of forelimb insertion; pectoral glands flat, level with axilla on chest; femoral glands slightly raised, on posterior surface of thighs, closer to knees than to cloaca; small black and white dermal asperities form a broad dense band circummarginally on gular region, moderately dense on tympanic regions between tympanum and eyes, and along upper lips, sparse on anterior dorsum of body restricted to dorsal ridges, increasing in density posteriorly across dorsum, absent from all remaining surfaces.</paragraph>
|
||
<caption id="DF006614FF9E0272FF73F8A1FB7BF86B" httpUri="https://zenodo.org/record/2610250/files/figure.png" pageId="64" pageNumber="65" startId="64.[151,250,1907,1929]" targetBox="[151,1436,553,1885]" targetPageId="64">
|
||
<paragraph id="8BC0369CFF9E0272FF73F8A1FB7BF86B" blockId="64.[151,1435,1907,1959]" pageId="64" pageNumber="65">
|
||
<emphasis id="B90BEA8EFF9E0272FF73F8A1FEFFF844" bold="true" box="[151,289,1907,1929]" pageId="64" pageNumber="65">FIGURE 24.</emphasis>
|
||
<taxonomicName id="4C7F4D1FFF9E0272FECCF8A1FDCDF845" authorityName="Fei, Ye and Huang" authorityYear="1990" box="[296,531,1907,1929]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9E0272FECCF8A1FDCDF845" box="[296,531,1907,1929]" italics="true" pageId="64" pageNumber="65">Megophrys glandulosa</emphasis>
|
||
</taxonomicName>
|
||
holotype: adult male (CIB 873112: SVL 78.4 mm [Fei
|
||
<emphasis id="B90BEA8EFF9E0272FBB1F8A6FB54F845" box="[1109,1162,1907,1929]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
|
||
1992]) in preservation:
|
||
<emphasis id="B90BEA8EFF9E0272FA67F8A1FA4BF844" bold="true" box="[1411,1429,1907,1928]" pageId="64" pageNumber="65">A</emphasis>
|
||
. dorsal view;
|
||
<emphasis id="B90BEA8EFF9E0272FEFEF843FEF5F86A" bold="true" box="[282,299,1937,1958]" pageId="64" pageNumber="65">B</emphasis>
|
||
. ventral view;
|
||
<emphasis id="B90BEA8EFF9E0272FE26F843FE0AF86B" bold="true" box="[450,468,1937,1959]" pageId="64" pageNumber="65">C</emphasis>
|
||
. profile view of head;
|
||
<emphasis id="B90BEA8EFF9E0272FD5FF843FD13F86A" bold="true" box="[699,717,1937,1958]" pageId="64" pageNumber="65">D</emphasis>
|
||
. ventral view of hand;
|
||
<emphasis id="B90BEA8EFF9E0272FC5CF843FC17F86A" bold="true" box="[952,969,1937,1958]" pageId="64" pageNumber="65">E</emphasis>
|
||
. ventral view of foot.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0369CFF9F0273FF23FF45FC04FCF4" blockId="65.[151,1437,151,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B90BEA8EFF9F0273FF23FF45FE8EFF7C" box="[199,336,151,176]" italics="true" pageId="65" pageNumber="66">Colouration</emphasis>
|
||
:
|
||
<emphasis id="B90BEA8EFF9F0273FEBBFF4BFDD8FF7C" box="[351,518,153,176]" italics="true" pageId="65" pageNumber="66">In preservative</emphasis>
|
||
: Dorsal and lateral surfaces of head and body primarily plain brown; light-edged, dark brown triangular marking between eyes; faint darker brown V-shaped marking on anterior dorsum surrounding parietoscapular ridge, narrow broken dark brown stripe follows lower edge of dorsolateral ridges; larger tubercles on flanks dark brown with light tip; front of snout and lateral surfaces below canthus rostralis dark brown; greyish-white stripe on upper lip extends from nostril to rear of upper jaw, interrupted by wide dark brown vertical bar between lower edge of orbits and edge of mouth; besides narrow brown oblique stripe extending from posterior edge of orbits posteriorly to join light upper lip stripe, remaining lateral surfaces of head below supratympanic ridges solid dark brown; outer edges of eyelids dark brown; lower half of supratympanic ridges dark brown, upper half brown anteriorly, light yellowish-cream posteriorly; dorsal surfaces of forelimbs and hindlimbs brown; two dark brown blotches on anterior lateral surface of forearms; dorsal surface of outer three fingers with dark brown blotches; dorsal surfaces of hindlimbs with distinct brown transverse crossbars; outer lateral surfaces of thighs and shanks with dark brown spots and blotches; throat with large light brown blotches, few small white spots along its edge, light-edged wide dark brown stripe extends from posterior edge of lower jaw to ventral surface of upper forelimb on each side; chest and abdomen with faint large brown blotches on an otherwise pale greyishwhite background; ventral surfaces of forelimbs pale greyish-white; ventral surfaces of thighs densely mottled light brown and greyish-white, posterior surfaces of thighs (including area surrounding cloaca) and ventral surfaces of shanks and tarsi primarily mid to dark brown with pale greyish-white scattered spots; some large dark brown spots and blotches laterally along abdomen on both sides; ventral surfaces of hands and feet greyish-brown; pectoral and femoral glands creamish-white.
|
||
<emphasis id="B90BEA8EFF9F0273FE1BFCF3FD9FFCF4" box="[511,577,799,824]" italics="true" pageId="65" pageNumber="66">In life</emphasis>
|
||
: Not documented for this specimen.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFF9F0273FF23FC96FEEEFB24" blockId="65.[151,1437,151,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B90BEA8EFF9F0273FF23FC96FEE1FC91" bold="true" box="[199,319,836,861]" pageId="65" pageNumber="66">Variation.</emphasis>
|
||
Refer to
|
||
<tableCitation id="C6FD0327FF9F0273FE51FC96FDD2FC90" box="[437,524,836,861]" captionStart="TABLE 1" captionStartId="22.[147,225,525,545]" captionText="TABLE 1. Morphometric details for specimens new and known included in the Taxonomic review section. Refer to Methods and materials section for explanation of measurement abbreviations and museum acronyms. HT = holotype; PT = paratype; LT = lectotype; PLT = paralectotype; NT = neotype; RS = referred specimen; M = adult male; F = adult female; SA = subadult; J = juvenile [post metamorphic]; † = BMNH voucher numbers; # = SDBDU voucher numbers; ind = character limits insufficiently distinct to reliably measure." pageId="65" pageNumber="66">Table 1</tableCitation>
|
||
for morphometric variation within the provisionally referred specimen series, consisting of three adult males. The two other referred specimens generally resemble the referred specimen described above for most morphological characters with the following exceptions: CAS 221484
|
||
<materialsCitation id="3B173CC1FF9F0273FACBFC5EFAB5FC04" ID-GBIF-Occurrence="2028521231" collectingDate="II 2" collectingDateMax="4 IV" collectingDateMin="II 2" collectionCode="IV" pageId="65" pageNumber="66" specimenCode="III2.8, 2.5, CAS 221484" specimenCount="1">
|
||
has more extensive webbing between Toes II to V (i.e., webbing formula:
|
||
<collectingDate id="EF85E9B4FF9F0273FC7EFC7DFC65FC04" box="[922,955,943,968]" pageId="65" pageNumber="66">
|
||
<date id="FFC1105CFF9F0273FC7EFC7DFC65FC04" box="[922,955,943,968]" pageId="65" pageNumber="66">II2</date>
|
||
</collectingDate>
|
||
–3.1
|
||
<specimenCode id="DBD99EE7FF9F0273FC08FC7DFBF5FC04" box="[1004,1067,943,968]" collectionCode="III" pageId="65" pageNumber="66">III2.8</specimenCode>
|
||
–
|
||
<collectingDate id="EF85E9B4FF9F0273FBDFFC7DFBB9FC04" box="[1083,1127,943,968]" pageId="65" pageNumber="66">
|
||
<date id="FFC1105CFF9F0273FBDFFC7DFBB9FC04" box="[1083,1127,943,968]" pageId="65" pageNumber="66">
|
||
4
|
||
<collectionCode id="ED6EAE59FF9F0273FBADFC7DFBB9FC04" box="[1097,1127,943,968]" pageId="65" pageNumber="66">IV</collectionCode>
|
||
</date>
|
||
</collectingDate>
|
||
4–
|
||
<specimenCode id="DBD99EE7FF9F0273FB61FC7DFB76FC04" box="[1157,1192,943,968]" collectionCode="IV" pageId="65" pageNumber="66">2.5</specimenCode>
|
||
V);
|
||
<specimenCode id="DBD99EE7FF9F0273FB31FC62FAB5FC04" box="[1237,1387,944,968]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS 221484</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B173CC1FF9F0273FA93FC7DFD66FC21" ID-GBIF-Occurrence="2028521229" collectionCode="IV, III" pageId="65" pageNumber="66" specimenCode="CAS 221395" specimenCount="1">
|
||
has relative finger lengths II=I<
|
||
<collectionCode id="ED6EAE59FF9F0273FE34FC07FE31FC20" box="[464,495,981,1004]" pageId="65" pageNumber="66">IV</collectionCode>
|
||
<
|
||
<collectionCode id="ED6EAE59FF9F0273FE1AFC07FDC5FC20" box="[510,539,981,1004]" pageId="65" pageNumber="66">III</collectionCode>
|
||
,
|
||
<specimenCode id="DBD99EE7FF9F0273FDC3FC06FD66FC21" box="[551,696,980,1005]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS 221395</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B173CC1FF9F0273FD5BFC06FE83FBDC" ID-GBIF-Occurrence="2028521205" collectionCode="IV, III" pageId="65" pageNumber="66" specimenCode="CAS 221484" specimenCount="1">
|
||
has
|
||
<collectionCode id="ED6EAE59FF9F0273FD08FC07FCD5FC20" box="[748,779,981,1004]" pageId="65" pageNumber="66">IV</collectionCode>
|
||
<II<I<
|
||
<collectionCode id="ED6EAE59FF9F0273FCB2FC07FCADFC20" box="[854,883,981,1004]" pageId="65" pageNumber="66">III</collectionCode>
|
||
; finger tips not dilated relative to finger width on
|
||
<specimenCode id="DBD99EE7FF9F0273FF73FC2AFEF5FBDC" box="[151,299,1016,1040]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS 221484</specimenCode>
|
||
and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B173CC1FF9F0273FE82FC2AFE20FBDC" ID-GBIF-Occurrence="2028521230" box="[358,510,1015,1040]" collectionCode="CAS" isTailEnumeration="true" pageId="65" pageNumber="66" specimenCode="CAS 221395" specimenCount="1">
|
||
<specimenCode id="DBD99EE7FF9F0273FE82FC2AFE20FBDC" box="[358,510,1015,1040]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS 221395</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B173CC1FF9F0273FDEFFC25FC12FBDC" ID-GBIF-Occurrence="2028521235" box="[523,972,1015,1040]" collectionCode="CAS" pageId="65" pageNumber="66" specimenCode="CAS 221484" specimenCount="1">
|
||
vomerine teeth absent on
|
||
<specimenCode id="DBD99EE7FF9F0273FCD0FC2AFC12FBDC" box="[820,972,1016,1040]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS 221484</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B173CC1FF9F0273FC3DFC25FE03FBF8" ID-GBIF-Occurrence="2028521243" collectionCode="CAS" pageId="65" pageNumber="66" specimenCode="CAS 221484" specimenCount="1">
|
||
dorsolateral ridges extend almost entire trunk length on
|
||
<specimenCode id="DBD99EE7FF9F0273FEACFBCEFE03FBF8" box="[328,477,1052,1076]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS 221484</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B173CC1FF9F0273FE03FBCEFF30FB94" ID-GBIF-Occurrence="2028521224" collectionCode="CAS" pageId="65" pageNumber="66" specimenCode="CAS 221395" specimenCount="1">
|
||
5–10% of upper tympanum surfaces are concealed by supratympanic ridges; on
|
||
<specimenCode id="DBD99EE7FF9F0273FA82FBCEFF30FB94" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS 221395</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B173CC1FF9F0273FF1EFBEDFA46FBB0" ID-GBIF-Occurrence="2028521213" collectionCode="CAS" pageId="65" pageNumber="66" specimenCode="CAS 221484" specimenCount="1">
|
||
dermal asperities moderately dense on lateral surfaces of snout, dorsal surface of eyelids and dorsally on head adjacent to supratympanic ridges, sparse on dorsal tibia, outer and ventral tarsal surfaces; on
|
||
<specimenCode id="DBD99EE7FF9F0273FAE6FBB6FA46FBB0" box="[1282,1432,1124,1148]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS 221484</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B173CC1FF9F0273FF73FB55FEF2FB24" ID-GBIF-Occurrence="2028521222" collectionCode="CAS" pageId="65" pageNumber="66" specimenCode="CAS 221484" specimenCount="1">
|
||
dermal asperities formed dense wide circummarginal band along lower jaw, sparse on tympanic regions (absent on tympanum) and on posterior lateral dorsum, absent on all remaining surfaces; V-shaped dorsal marking absent on
|
||
<specimenCode id="DBD99EE7FF9F0273FF73FB02FEF2FB24" box="[151,300,1232,1256]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="65" pageNumber="66" type="Museum">CAS 221484</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFF9F0273FF23FB26FDECFA0C" blockId="65.[151,1437,151,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B90BEA8EFF9F0273FF23FB26FDFCFAC1" bold="true" box="[199,546,1268,1293]" pageId="65" pageNumber="66">Secondary sexual characters.</emphasis>
|
||
<emphasis id="B90BEA8EFF9F0273FDCCFB26FDB2FAC1" box="[552,620,1268,1293]" italics="true" pageId="65" pageNumber="66">Males</emphasis>
|
||
: nuptial pads present, weakly raised, covered with black micro-granules, covering almost entire dorsal surface of Finger I narrowing distally where it extends to base of distal phalange on inner dorsal side; nuptial pad small to moderately large on Finger II, positioned on base of digit on inner dorsal side (extending on to base of distal phalange on some individuals); external vocal sac indistinct; internal vocal slit present on floor of mouth on each side near rear of mandible; forearms enlarged relative to upper forelimbs.
|
||
<emphasis id="B90BEA8EFF9F0273FF73FA75FF28FA0C" box="[151,246,1447,1472]" italics="true" pageId="65" pageNumber="66">Females</emphasis>
|
||
: not examined in this study.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFF9F0273FF23FA1EFD69F99C" blockId="65.[151,1437,151,2013]" pageId="65" pageNumber="66">
|
||
<emphasis id="B90BEA8EFF9F0273FF23FA1EFDC4FA29" bold="true" box="[199,538,1484,1509]" pageId="65" pageNumber="66">Morphological comparison.</emphasis>
|
||
Adult body size ranges provided for
|
||
<taxonomicName id="4C7F4D1FFF9F0273FC18FA1EFAD9FA29" authorityName="Fei, Ye and Huang" authorityYear="1990" box="[1020,1287,1484,1509]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9F0273FC18FA1EFAD9FA29" box="[1020,1287,1484,1509]" italics="true" pageId="65" pageNumber="66">Megophrys glandulosa</emphasis>
|
||
</taxonomicName>
|
||
include the measurements given in
|
||
<bibRefCitation id="EFEE4B6DFF9F0273FE4FFA3DFDB5F9C4" author="Fei, L. & Hu, S. - Q. & Ye, C. - Y. & Huang, Y. - Z." box="[427,619,1519,1544]" pageId="65" pageNumber="66" refId="ref52932" refString="Fei, L., Hu, S. - Q., Ye, C. - Y. & Huang, Y. - Z. (2009) Fauna Sinica. Amphibia. Folume 2. Anura. Chinese Academy of Science, Science, Beijing, XIII + 957 pp." type="book" year="2009">
|
||
Fei
|
||
<emphasis id="B90BEA8EFF9F0273FE3EFA23FDC8F9C4" box="[474,534,1519,1544]" italics="true" pageId="65" pageNumber="66">et al.</emphasis>
|
||
(2009)
|
||
</bibRefCitation>
|
||
and the referred specimens examined in this study (total adult males,
|
||
<emphasis id="B90BEA8EFF9F0273FF73F9C7FF74F9E0" box="[151,170,1557,1580]" italics="true" pageId="65" pageNumber="66">N</emphasis>
|
||
=13; adult females,
|
||
<emphasis id="B90BEA8EFF9F0273FE77F9C7FE78F9E0" box="[403,422,1557,1580]" italics="true" pageId="65" pageNumber="66">N</emphasis>
|
||
=3). The remaining characters were assessed based only on the referred specimens and images of the
|
||
<typeStatus id="54C4883EFF9F0273FED1F9E5FE49F99C" box="[309,407,1591,1616]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
(total: adult males,
|
||
<emphasis id="B90BEA8EFF9F0273FD92F9EBFD57F99C" box="[630,649,1593,1616]" italics="true" pageId="65" pageNumber="66">N</emphasis>
|
||
=4).
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFF9F0270FF23F98EFAE2FF34" blockId="65.[151,1437,151,2013]" lastBlockId="66.[151,1437,151,1760]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
|
||
<taxonomicName id="4C7F4D1FFF9F0273FF23F98EFE17F9B9" authorityName="Fei, Ye and Huang" authorityYear="1990" box="[199,457,1628,1653]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9F0273FF23F98EFE17F9B9" box="[199,457,1628,1653]" italics="true" pageId="65" pageNumber="66">Megophrys glandulosa</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C7F4D1FFF9F0273FDB8F98FFCDAF9B9" authorityName="Fei, Ye and Huang" authorityYear="1983" box="[604,772,1628,1653]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="medogensis">
|
||
<emphasis id="B90BEA8EFF9F0273FDB8F98FFCDAF9B9" box="[604,772,1628,1653]" italics="true" pageId="65" pageNumber="66">M. medogensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D1FFF9F0273FCF6F98FFC5CF9B9" authorityName="Ye and Fei" authorityYear="1992" box="[786,898,1628,1653]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="zhangi">
|
||
<emphasis id="B90BEA8EFF9F0273FCF6F98FFC5CF9B9" box="[786,898,1628,1653]" italics="true" pageId="65" pageNumber="66">M. zhangi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D1FFF9F0273FC74F98FFBFBF9B9" authorityName="" sensu Anderson" authorityYear="1871" box="[912,1061,1628,1653]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="monticola">
|
||
<emphasis id="B90BEA8EFF9F0273FC74F98FFBFBF9B9" box="[912,1061,1628,1653]" italics="true" pageId="65" pageNumber="66">M. monticola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D1FFF9F0273FBD7F98EFAD2F9B8" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[1075,1292,1628,1653]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFF9F0273FBD7F98EFAD2F9B8" box="[1075,1292,1628,1653]" italics="true" pageId="65" pageNumber="66">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9F0273FAF7F98EFAB3F9B9" box="[1299,1389,1628,1653]" pageId="65" pageNumber="66" rank="species">
|
||
<emphasis id="B90BEA8EFF9F0273FAF7F98EFAB3F9B9" bold="true" box="[1299,1389,1628,1653]" pageId="65" pageNumber="66">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7F4D1FFF9F0273FF73F953FECFF954" authorityName=", Rai and Anders" authorityYear="2002" box="[151,273,1663,1688]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="robusta">
|
||
<emphasis id="B90BEA8EFF9F0273FF73F953FECFF954" box="[151,273,1663,1688]" italics="true" pageId="65" pageNumber="66">M. robusta</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a distinct white upper lip stripe (vs. absent), presence of broad lateral fringes on toes (vs. absent on
|
||
<taxonomicName id="4C7F4D1FFF9F0273FEDEF977FE3CF971" authorityName="Fei, Ye and Huang" authorityYear="1983" box="[314,482,1700,1725]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="medogensis">
|
||
<emphasis id="B90BEA8EFF9F0273FEDEF977FE3CF971" box="[314,482,1700,1725]" italics="true" pageId="65" pageNumber="66">M. medogensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D1FFF9F0273FE14F977FDBFF971" authorityName="Ye and Fei" authorityYear="1992" box="[496,609,1700,1725]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="zhangi">
|
||
<emphasis id="B90BEA8EFF9F0273FE14F977FDBFF971" box="[496,609,1700,1725]" italics="true" pageId="65" pageNumber="66">M. zhangi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D1FFF9F0273FD8BF977FCDBF971" authorityName="" sensu Anderson" authorityYear="1871" box="[623,773,1700,1725]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="monticola">
|
||
<emphasis id="B90BEA8EFF9F0273FD8BF977FCDBF971" box="[623,773,1700,1725]" italics="true" pageId="65" pageNumber="66">M. monticola</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7F4D1FFF9F0273FCA7F977FC63F971" authorityName=", Rai and Anders" authorityYear="2002" box="[835,957,1700,1725]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="robusta">
|
||
<emphasis id="B90BEA8EFF9F0273FCA7F977FC63F971" box="[835,957,1700,1725]" italics="true" pageId="65" pageNumber="66">M. robusta</emphasis>
|
||
</taxonomicName>
|
||
; usually absent, though very narrow when present on
|
||
<taxonomicName id="4C7F4D1FFF9F0273FEF7F915FE33F92C" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[275,493,1735,1760]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFF9F0273FEF7F915FE33F92C" box="[275,493,1735,1760]" italics="true" pageId="65" pageNumber="66">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9F0273FE11F915FD91F92C" box="[501,591,1735,1760]" pageId="65" pageNumber="66" rank="species">
|
||
<emphasis id="B90BEA8EFF9F0273FE11F915FD91F92C" bold="true" box="[501,591,1735,1760]" pageId="65" pageNumber="66">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
), and differs further from
|
||
<taxonomicName id="4C7F4D1FFF9F0273FC9EF91BFBFAF92C" authorityName="Fei, Ye and Huang" authorityYear="1983" box="[890,1060,1735,1760]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="medogensis">
|
||
<emphasis id="B90BEA8EFF9F0273FC9EF91BFBFAF92C" box="[890,1060,1735,1760]" italics="true" pageId="65" pageNumber="66">M. medogensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D1FFF9F0273FBD6F91BFB7DF92C" authorityName="Ye and Fei" authorityYear="1992" box="[1074,1187,1735,1760]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="zhangi">
|
||
<emphasis id="B90BEA8EFF9F0273FBD6F91BFB7DF92C" box="[1074,1187,1735,1760]" italics="true" pageId="65" pageNumber="66">M. zhangi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7F4D1FFF9F0273FB06F91BFAA6F92C" authorityName="" sensu Anderson" authorityYear="1871" box="[1250,1400,1735,1760]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="monticola">
|
||
<emphasis id="B90BEA8EFF9F0273FB06F91BFAA6F92C" box="[1250,1400,1735,1760]" italics="true" pageId="65" pageNumber="66">M. monticola</emphasis>
|
||
</taxonomicName>
|
||
by its larger adult size, male SVL 76.3–81.0 mm, female SVL
|
||
<quantity id="4C879B79FF9F0273FCA1F93EFC34F8C8" box="[837,1002,1772,1797]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" metricValueMax="9.95" metricValueMin="7.65" pageId="65" pageNumber="66" unit="mm" value="88.0" valueMax="99.5" valueMin="76.5">76.5–99.5 mm</quantity>
|
||
(vs. male SVL 57.2–68.0 mm,
|
||
<emphasis id="B90BEA8EFF9F0273FAB1F93FFAB6F8C8" box="[1365,1384,1773,1796]" italics="true" pageId="65" pageNumber="66">N</emphasis>
|
||
=16; male SVL
|
||
<quantity id="4C879B79FF9F0273FEF6F8DDFE66F8EB" box="[274,440,1807,1832]" metricMagnitude="-2" metricUnit="m" metricValue="3.485" metricValueMax="3.72" metricValueMin="3.25" pageId="65" pageNumber="66" unit="mm" value="34.85" valueMax="37.2" valueMin="32.5">32.5–37.2 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFF9F0273FE21F8C3FE06F8E4" box="[453,472,1809,1832]" italics="true" pageId="65" pageNumber="66">N</emphasis>
|
||
=3; male SVL
|
||
<quantity id="4C879B79FF9F0273FD64F8DDFCF8F8EB" box="[640,806,1807,1832]" metricMagnitude="-2" metricUnit="m" metricValue="4.385" metricValueMax="4.95" metricValueMin="3.82" pageId="65" pageNumber="66" unit="mm" value="43.85" valueMax="49.5" valueMin="38.2">38.2–49.5 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFF9F0273FCD7F8C3FC98F8E4" box="[819,838,1809,1832]" italics="true" pageId="65" pageNumber="66">N</emphasis>
|
||
=17, female SVL
|
||
<quantity id="4C879B79FF9F0273FBF6F8DDFB66F8EB" box="[1042,1208,1807,1832]" metricMagnitude="-2" metricUnit="m" metricValue="4.83" metricValueMax="5.61" metricValueMin="4.05" pageId="65" pageNumber="66" unit="mm" value="48.3" valueMax="56.1" valueMin="40.5">40.5–56.1 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFF9F0273FB20F8C3FB09F8E4" box="[1220,1239,1809,1832]" italics="true" pageId="65" pageNumber="66">N</emphasis>
|
||
=6, respectively); differs from
|
||
<taxonomicName id="4C7F4D1FFF9F0273FEC2F8E6FDEFF881" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[294,561,1844,1869]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFF9F0273FEC2F8E6FDEFF881" box="[294,561,1844,1869]" italics="true" pageId="65" pageNumber="66">Megophrys himalayana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9F0273FDDDF8E6FD4DF881" box="[569,659,1844,1869]" pageId="65" pageNumber="66" rank="species">
|
||
<emphasis id="B90BEA8EFF9F0273FDDDF8E6FD4DF881" bold="true" box="[569,659,1844,1869]" pageId="65" pageNumber="66">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7F4D1FFF9F0273FD28F8E7FC40F881" authorityName="Fei & Ye" authorityYear="1990" box="[716,926,1844,1869]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="mangshanensis">
|
||
<emphasis id="B90BEA8EFF9F0273FD28F8E7FC40F881" box="[716,926,1844,1869]" italics="true" pageId="65" pageNumber="66">M. mangshanensis</emphasis>
|
||
</taxonomicName>
|
||
by its larger adult male size, SVL 76.3–81.0 mm (vs. male SVL 68.0–
|
||
<quantity id="4C879B79FF9F0273FE5CF885FDC1F8A3" box="[440,543,1879,1904]" metricMagnitude="-2" metricUnit="m" metricValue="7.35" pageId="65" pageNumber="66" unit="mm" value="73.5">73.5 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFF9F0273FDC8F88BFDE1F8BC" box="[556,575,1881,1904]" italics="true" pageId="65" pageNumber="66">N</emphasis>
|
||
=7; male SVL
|
||
<quantity id="4C879B79FF9F0273FD0DF885FC8EF8A3" box="[745,848,1879,1904]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" pageId="65" pageNumber="66" unit="mm" value="62.5">62.5 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFF9F0273FCBAF88BFCAFF8BC" box="[862,881,1881,1904]" italics="true" pageId="65" pageNumber="66">N</emphasis>
|
||
=1, respectively), and by broad lateral fringes on toes present (vs. absent); differs from
|
||
<taxonomicName id="4C7F4D1FFF9F0273FDD8F8AFFD7AF858" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[572,676,1917,1940]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="species" sensu="stricto" species="major">
|
||
<emphasis id="B90BEA8EFF9F0273FDD8F8AFFD7AF858" box="[572,676,1917,1940]" italics="true" pageId="65" pageNumber="66">M. major</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9F0273FD4EF8AFFD10F858" box="[682,718,1917,1940]" pageId="65" pageNumber="66" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7F4D1FFF9F0273FCE7F8AEFBDCF858" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[771,1026,1916,1941]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="65" pageNumber="66" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFF9F0273FCE7F8AEFBDCF858" box="[771,1026,1916,1941]" italics="true" pageId="65" pageNumber="66">Megophrys oreocrypta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9F0273FBECF8AEFBBEF859" box="[1032,1120,1916,1941]" pageId="65" pageNumber="66" rank="species">
|
||
<emphasis id="B90BEA8EFF9F0273FBECF8AEFBBEF859" bold="true" box="[1032,1120,1916,1941]" pageId="65" pageNumber="66">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by presence of broad lateral fringes on toes, and toe tips not expanded (vs. occasional presence of narrow lateral fringes on toes, and toe tips distinctly expanded relative to toe width), ventral surface of thighs densely spotted and speckled (vs. immaculate); differs from
|
||
<taxonomicName id="4C7F4D1FFF9C0270FEC7FF45FDE2FF7C" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[291,572,151,176]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFF9C0270FEC7FF45FDE2FF7C" box="[291,572,151,176]" italics="true" pageId="66" pageNumber="67">Megophrys flavipunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9C0270FDA6FF45FD42FF7C" box="[578,668,151,176]" pageId="66" pageNumber="67" rank="species">
|
||
<emphasis id="B90BEA8EFF9C0270FDA6FF45FD42FF7C" bold="true" box="[578,668,151,176]" pageId="66" pageNumber="67">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by its larger adult size, male SVL 76.3–81.0 mm, female SVL
|
||
<quantity id="4C879B79FF9C0270FAB9FF45FF22FF18" metricMagnitude="-2" metricUnit="m" metricValue="8.8" metricValueMax="9.95" metricValueMin="7.65" pageId="66" pageNumber="67" unit="mm" value="88.0" valueMax="99.5" valueMin="76.5">76.5– 99.5 mm</quantity>
|
||
(vs. male SVL
|
||
<quantity id="4C879B79FF9C0270FE4BFF6EFD8BFF18" box="[431,597,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="6.265000000000001" metricValueMax="6.840000000000001" metricValueMin="5.6899999999999995" pageId="66" pageNumber="67" unit="mm" value="62.65" valueMax="68.4" valueMin="56.9">56.9–68.4 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFF9C0270FD85FF6FFDAAFF18" box="[609,628,189,212]" italics="true" pageId="66" pageNumber="67">N</emphasis>
|
||
=4, female SVL 68.0–
|
||
<quantity id="4C879B79FF9C0270FC94FF6EFC08FF18" box="[880,982,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="7.459999999999999" pageId="66" pageNumber="67" unit="mm" value="74.6">74.6 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFF9C0270FC07FF6FFC28FF18" box="[995,1014,189,212]" italics="true" pageId="66" pageNumber="67">N</emphasis>
|
||
=3), by toe tips not expanded relative to toe width (vs. toe tips slightly expanded), ventral thighs densely spotted and speckled (vs. immaculate).
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFF9C0270FF23FED6FB7DFE39" blockId="66.[151,1437,151,1760]" pageId="66" pageNumber="67">
|
||
<emphasis id="B90BEA8EFF9C0270FF23FED6FE62FED1" bold="true" box="[199,444,260,285]" pageId="66" pageNumber="67">Systematic position.</emphasis>
|
||
The systematic position of
|
||
<taxonomicName id="4C7F4D1FFF9C0270FCF1FED6FBC0FED1" authorityName="Fei, Ye and Huang" authorityYear="1990" box="[789,1054,260,285]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FCF1FED6FBC0FED1" box="[789,1054,260,285]" italics="true" pageId="66" pageNumber="67">Megophrys glandulosa</emphasis>
|
||
</taxonomicName>
|
||
within the MMC could not be resolved in this study possibly due to incomplete sampling of nuclear markers (i.e., absence of CXCR4, TTN and SACS; Appendix I, Table 2). Analyses of the mtDNA datasets (16S and CO1 gene trees, and 16S+CO1 concatenated tree), the RHOD gene, and the concatenated mt+nuDNA dataset indicated that this species is a distinct lineage (
|
||
<figureCitation id="13442A19FF9C0270FEB2FE46FE62FE60" box="[342,444,404,429]" captionStart="FIGURE 4" captionStartId="13.[151,250,1115,1137]" captionTargetBox="[155,1430,193,1084]" captionTargetId="figure@13.[150,1436,193,1094]" captionTargetPageId="13" captionText="FIGURE 4. RAxML tree for the combined mt+nuDNA dataset expanded to include the Chen et al. (2017) sequences (Dataset J). Bootstrap support values are provided adjacent to the associated nodes. Refer to Appendix I, Table 2 for revised names for taxa based on the taxonomic review." httpUri="https://zenodo.org/record/2610210/files/figure.png" pageId="66" pageNumber="67">Figure 4</figureCitation>
|
||
; Appendix II,
|
||
<figureCitation id="13442A19FF9C0270FD82FE46FD17FE61" box="[614,713,404,429]" captionStart="FIGURE 5" captionStartId="13.[151,250,1843,1865]" captionTargetBox="[162,1426,1248,1806]" captionTargetId="figure@13.[151,1436,1248,1822]" captionTargetPageId="13" captionText="FIGURE 5. DensiTree cloudogram representations of *BEAST analyses, removing the first 10% of trees as burn-in, produced from the four nuDNA gene alignments of Dataset E: A. original unphased; B. phased. Dark blue branches represent the most frequently occurring topology, light blue branches represent the consensus trees with intensity proportional to the frequency of the topology in the dataset, and green areas represent all topologies in the dataset. Refer to Appendix I, Table 2 for revised names for taxa based on the taxonomic review." httpUri="https://zenodo.org/record/2610212/files/figure.png" pageId="66" pageNumber="67">Figure 5</figureCitation>
|
||
), however, the RAG1 gene tree (not shown) and concatenated nuDNA analyses indicated that this species may have close affinity to the species referred to here as
|
||
<taxonomicName id="4C7F4D1FFF9C0270FAB1FE6BFEF8FE38" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="maosonensis">
|
||
<emphasis id="B90BEA8EFF9C0270FAB1FE6BFAAAFE1C" box="[1365,1396,441,464]" italics="true" pageId="66" pageNumber="67">M.</emphasis>
|
||
cf.
|
||
<emphasis id="B90BEA8EFF9C0270FF73FE0FFEF8FE38" box="[151,294,477,500]" italics="true" pageId="66" pageNumber="67">maosonensis</emphasis>
|
||
</taxonomicName>
|
||
[1], though this relationship received very low support (Appendix II,
|
||
<figureCitation id="13442A19FF9C0270FBD1FE0EFB4BFE39" box="[1077,1173,476,501]" captionStart="FIGURE 6" captionStartId="18.[151,250,1977,1999]" captionTargetBox="[165,1425,193,1961]" captionTargetId="figure@18.[165,1425,193,1961]" captionTargetPageId="18" captionText="FIGURE 6. Megophrys monticola lectotype: adult female (BMNH 1947.2.25.13) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610214/files/figure.png" pageId="66" pageNumber="67">Figure 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FF9C0270FF23FE2DFC70FD49" pageId="66" pageNumber="67" type="etymology">
|
||
<paragraph id="8BC0369CFF9C0270FF23FE2DFCA5FDAC" blockId="66.[151,1437,151,1760]" pageId="66" pageNumber="67">
|
||
<emphasis id="B90BEA8EFF9C0270FF23FE2DFE90FDD4" bold="true" box="[199,334,511,536]" pageId="66" pageNumber="67">Etymology.</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="4C7F4D1FFF9C0270FDB7FE2DFD11FDD4" authorityName="Fei, Ye and Huang" authorityYear="1990" box="[595,719,511,536]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FDB7FE2DFD11FDD4" box="[595,719,511,536]" italics="true" pageId="66" pageNumber="67">glandulosa</emphasis>
|
||
</taxonomicName>
|
||
” is Latin, with
|
||
<emphasis id="B90BEA8EFF9C0270FC68FE2DFC3CFDD4" box="[908,994,511,536]" italics="true" pageId="66" pageNumber="67">glandul</emphasis>
|
||
meaning “gland”, and the suffix -
|
||
<emphasis id="B90BEA8EFF9C0270FA91FDD3FA45FDD4" box="[1397,1435,513,536]" italics="true" pageId="66" pageNumber="67">osa</emphasis>
|
||
meaning “abundant”, presumably a reference to the tubercles on the flanks of this species, a character that is not useful for diagnosing this species from other species in the MMC.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFF9C0270FF23FDBEFC70FD49" blockId="66.[151,1437,151,1760]" box="[199,942,620,645]" pageId="66" pageNumber="67">
|
||
<emphasis id="B90BEA8EFF9C0270FF23FDBEFE20FD49" bold="true" box="[199,510,620,645]" pageId="66" pageNumber="67">Suggested common name:</emphasis>
|
||
Glandular White-lipped Horned Frog.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FF9C0270FF23FD5DFCA7FCD9" pageId="66" pageNumber="67" type="distribution">
|
||
<paragraph id="8BC0369CFF9C0270FF23FD5DFCA7FCD9" blockId="66.[151,1437,151,1760]" pageId="66" pageNumber="67">
|
||
<emphasis id="B90BEA8EFF9C0270FF23FD5DFEBEFD64" bold="true" box="[199,352,655,680]" pageId="66" pageNumber="67">Distribution.</emphasis>
|
||
Besides the
|
||
<typeStatus id="54C4883EFF9C0270FE16FD42FDFCFD64" box="[498,546,656,680]" pageId="66" pageNumber="67">type</typeStatus>
|
||
locality,
|
||
<taxonomicName id="4C7F4D1FFF9C0270FD6FFD43FCF1FD64" authorityName="Fei, Ye and Huang" authorityYear="1990" box="[651,815,655,680]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FD6FFD43FCF1FD64" box="[651,815,655,680]" italics="true" pageId="66" pageNumber="67">M. glandulosa</emphasis>
|
||
</taxonomicName>
|
||
has been reported from numerous localities in Yunnan Province between
|
||
<date id="FFC1105CFF9C0270FE83FD66FDCEFD01" box="[359,528,692,717]" pageId="66" pageNumber="67" value="1900" valueMax="2500">
|
||
1900 and
|
||
<quantity id="4C879B79FF9C0270FE3CFD66FDCEFD01" box="[472,528,692,717]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="66" pageNumber="67" unit="m" value="2500.0">2500</quantity>
|
||
</date>
|
||
m asl. (e.g.,
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FD47FD66FCBCFD00" author="Jiang, J. P. & Yuan, F. R. & Xie, F. & Zheng, Z. H." box="[675,866,692,717]" pageId="66" pageNumber="67" pagination="241 - 248" refId="ref54636" refString="Jiang, J. P., Yuan, F. R., Xie, F. & Zheng, Z. H. (2003) Phylogenetic relationships of some species and genera in megophryids inferred from partial sequences of mitochondrial 12 S and 16 S rRNA genes. Zoological Research, 24 (4), 241 - 248." type="journal article" year="2003">
|
||
Jiang
|
||
<emphasis id="B90BEA8EFF9C0270FD02FD67FCC1FD01" box="[742,799,692,717]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FC8AFD67FBCEFD01" author="Fu, C. & Hua, X. & Li, J. & Chang, Z. & Pu, Z. & Chen, J." box="[878,1040,692,717]" pageId="66" pageNumber="67" pagination="919 - 927" refId="ref53295" refString="Fu, C., Hua, X., Li, J., Chang, Z., Pu, Z. & Chen, J. (2006) Elevational patterns of frog species richness and endemic richness in the Hengduan Mountains, China: geometric constraints, area and climate effects. Ecography, 29, 919 - 927 + 1 - 7 (Appendix E 4802)." type="journal article" year="2006">
|
||
Fu
|
||
<emphasis id="B90BEA8EFF9C0270FC70FD67FC13FD01" box="[916,973,692,717]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FBF8FD66FB1AFD00" author="Fei, L. & Hu, S. - Q. & Ye, C. - Y. & Huang, Y. - Z." box="[1052,1220,692,717]" pageId="66" pageNumber="67" refId="ref52932" refString="Fei, L., Hu, S. - Q., Ye, C. - Y. & Huang, Y. - Z. (2009) Fauna Sinica. Amphibia. Folume 2. Anura. Chinese Academy of Science, Science, Beijing, XIII + 957 pp." type="book" year="2009">
|
||
Fei
|
||
<emphasis id="B90BEA8EFF9C0270FBACFD67FB5FFD01" box="[1096,1153,692,717]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
), and
|
||
<collectingRegion id="49BBF87EFF9C0270FAEDFD66FA84FD01" box="[1289,1370,692,717]" country="Myanmar" name="Kachin" pageId="66" pageNumber="67">Kachin</collectingRegion>
|
||
state, northern
|
||
<collectingCountry id="F368760CFF9C0270FF18FD0AFEB7FD3C" box="[252,361,728,752]" name="Myanmar" pageId="66" pageNumber="67">Myanmar</collectingCountry>
|
||
by
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FE76FD0AFDAEFD3C" author="Wogan, G. O. U. & Vindum, J. V. & Wilkinson, J. A. & Koo, M. S. & Slowinski, J. B. & Win, H. & Thin, T. & Kyi, S. W. & Oo, S. L. & Lwin, K. S. & Shein, A. K." box="[402,624,727,752]" pageId="66" pageNumber="67" pagination="83 - 96" refId="ref59179" refString="Wogan, G. O. U., Vindum, J. V., Wilkinson, J. A., Koo, M. S., Slowinski, J. B., Win, H. Thin, T. Kyi, S. W., Oo, S. L., Lwin, K. S. & Shein, A. K. (2008) New country records and range extensions for Myanmar amphibians and reptiles. Hamadryad, 33, 83 - 96." type="journal article" year="2008">
|
||
Wogan
|
||
<emphasis id="B90BEA8EFF9C0270FE03FD0BFDC1FD3C" box="[487,543,727,752]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
(
|
||
<figureCitation id="13442A19FF9C0270FD9AFD05FD2AFD3C" box="[638,756,727,752]" captionStart="FIGURE 8" captionStartId="21.[151,250,1742,1764]" captionTargetBox="[161,1425,1085,1720]" captionTargetId="figure@21.[159,1427,1083,1721]" captionTargetPageId="21" captionText="FIGURE 8. Distribution map of the western MMSG taxa: A. All taxa (excluding M. monticola) from Northeast India and surrounding countries: diamonds represent type localities, circles represent referred localities based on molecularly sampled specimens: orange = M. zhangi; yellow = M. robusta; black = M. medogensis; dark blue = M. major; pink = Megophrys flavipunctata sp. nov.; dark green = Megophrys oreocrypta sp. nov.; light blue = Megophrys himalayana sp. nov.; light green = Megophrys periosa sp. nov.; pale blue = M. cf. periosa; purple = M. glandulosa. B. Inset rectangle in Figure A enlarged to show the distribution map for M. monticola from Darjeeling and Sikkim region, Northeast India, showing distribution of named mitochondrial haplotypes: stars represent the type localities, circles represent referred localities confirmed from sequence data: red = M. “sanu” comb. nov., light blue = M. “katabhako” comb. nov. Blue lines represent major rivers; grey lines represent international borders." httpUri="https://zenodo.org/record/2610218/files/figure.png" pageId="66" pageNumber="67">Figure 8A</figureCitation>
|
||
). Reports of this species from further west in Northeast
|
||
<collectingCountry id="F368760CFF9C0270FA86FD05FA42FD3C" box="[1378,1436,727,752]" name="India" pageId="66" pageNumber="67">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="F368760CFF9C0270FF22FD2EFEC9FCD9" box="[198,279,764,789]" name="Bhutan" pageId="66" pageNumber="67">Bhutan</collectingCountry>
|
||
are not here considered reliable (see Remarks below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FF9C0270FF23FCCDFD74F92C" pageId="66" pageNumber="67" type="discussion">
|
||
<paragraph id="8BC0369CFF9C0270FF23FCCDFAF1FC90" blockId="66.[151,1437,151,1760]" pageId="66" pageNumber="67">
|
||
<emphasis id="B90BEA8EFF9C0270FF23FCCDFEE5FCF4" bold="true" box="[199,315,799,824]" pageId="66" pageNumber="67">Remarks.</emphasis>
|
||
<taxonomicName id="4C7F4D1FFF9C0270FEA6FCCDFD9BFCF4" authorityName="Fei, Ye and Huang" authorityYear="1990" box="[322,581,799,824]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FEA6FCCDFD9BFCF4" box="[322,581,799,824]" italics="true" pageId="66" pageNumber="67">Megophrys glandulosa</emphasis>
|
||
</taxonomicName>
|
||
was described (in Chinese text) as new twice based on the same
|
||
<typeStatus id="54C4883EFF9C0270FAC1FCF2FA8BFCF4" box="[1317,1365,800,824]" pageId="66" pageNumber="67">type</typeStatus>
|
||
series (
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FF7AFC96FE9BFC90" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[158,325,836,861]" pageId="66" pageNumber="67" pagination="2" refId="ref52831" refString="Fei, L., Ye, C. - Y. & Huang, Y. - Z. (1990) Key to Chinese Amphibians. Chongqing Branch, Science and Technology Literature Publishing House, Chongqing, 364 + 2 + [iv] pp." type="journal article" year="1990">
|
||
Fei
|
||
<emphasis id="B90BEA8EFF9C0270FF2FFC97FEDDFC91" box="[203,259,836,861]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
1990
|
||
</bibRefCitation>
|
||
, 1992).
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FE7BFC96FD8DFC90" author="Fei, L. & Ye, C. - Y. & Huang, Y. - Z." box="[415,595,836,861]" pageId="66" pageNumber="67" pagination="5 - 12" refId="ref52880" refString="Fei, L., Ye, C. - Y. & Huang, Y. - Z. (1992) Two new species of the genus Megophrys, Pelobatidae (Amphibia, Anura) from China. Zoological Research, 13 (1), 5 - 12." type="journal article" year="1992">
|
||
Fei
|
||
<emphasis id="B90BEA8EFF9C0270FE2EFC97FDDCFC91" box="[458,514,836,861]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
(1992)
|
||
</bibRefCitation>
|
||
was subsequently translated into English by
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FBA8FC97FAF5FC90" author="Huang, C. - M. & Lathrop, A. & Murphy, R. W." box="[1100,1323,836,861]" pageId="66" pageNumber="67" pagination="1 - 20" refId="ref54291" refString="Huang, C. - M., Lathrop, A. & Murphy, R. W. (1998) Translations of " Two new species of amphibian in Tibet " Huang & Fei 1981 and " Description of two new species of the genus Megophrys " Fei, Ye, & Huang 1995 [1992]. Smithsonian Herpetological Information Service, 118, 1 - 20." type="journal article" year="1998">
|
||
Huang
|
||
<emphasis id="B90BEA8EFF9C0270FB7AFC97FB08FC91" box="[1182,1238,836,861]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
(1998)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFF9C0270FF23FCBAFECDFAC1" blockId="66.[151,1437,151,1760]" pageId="66" pageNumber="67">
|
||
It is highly unlikely that the
|
||
<collectingCountry id="F368760CFF9C0270FDE5FCB5FD8CFC4C" box="[513,594,871,896]" name="Bhutan" pageId="66" pageNumber="67">Bhutan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FDBBFCB5FCDAFC4C" author="Wangyal, J. T." box="[607,772,871,896]" pageId="66" pageNumber="67" pagination="4774 - 4783" refId="ref59089" refString="Wangyal, J. T. (2013) New records of reptiles and amphibians from Bhutan. Journal of Threatened Taxa, 5, 4774 - 4783. https: // doi. org / 10.11609 / JoTT. o 3539.4774 - 83" type="journal article" year="2013">Wangyal 2013</bibRefCitation>
|
||
) and Indian (
|
||
<collectingRegion id="49BBF87EFF9C0270FC73FCB5FBC6FC4C" box="[919,1048,871,896]" country="India" name="Meghalaya" pageId="66" pageNumber="67">Meghalaya</collectingRegion>
|
||
,
|
||
<collectingRegion id="49BBF87EFF9C0270FBC6FCB5FB53FC4C" box="[1058,1165,871,896]" country="India" name="Nagaland" pageId="66" pageNumber="67">Nagaland</collectingRegion>
|
||
and
|
||
<collectingRegion id="49BBF87EFF9C0270FB27FCBAFA8EFC4C" box="[1219,1360,871,896]" country="India" name="West Bengal" pageId="66" pageNumber="67">West Bengal</collectingRegion>
|
||
states:
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FF73FC5EFE93FC68" author="Ao, M. & Bordoloi, S. & Ohler, A." box="[151,333,908,933]" pageId="66" pageNumber="67" pagination="1117 - 1125" refId="ref49909" refString="Ao, M., Bordoloi, S. & Ohler, A. (2003) Amphibian fauna of Nagaland with nineteen new records from the State including five new records for India. Zoos' Print Journal, 18 (6), 1117 - 1125. https: // doi. org / 10.11609 / JoTT. ZPJ. 18.6.1117 - 25" type="journal article" year="2003">
|
||
Ao
|
||
<emphasis id="B90BEA8EFF9C0270FF22FC5FFEDAFC69" box="[198,260,908,933]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
2003
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FEBAFC5EFD40FC68" author="Deuti, K & Ayyaswamy, A. K." box="[350,670,908,933]" pageId="66" pageNumber="67" pagination="17 - 20" refId="ref52242" refString="Deuti, K & Ayyaswamy, A. K. (2008) Three new records of amphibians from West Bengal. Zoological Survey of India, 108 (4), 17 - 20." type="journal article" year="2008">Deuti & Ayyaswamy 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FD4BFC5EFC73FC68" author="Mathew, R. & Sen, N." box="[687,941,908,933]" pageId="66" pageNumber="67" refId="ref55566" refString="Mathew, R. & Sen, N. (2010) Pictorial Guide to Amphibians of North East India. Zoological Survey of India, Kolkata, 144 pp." type="book" year="2010">Mathew & Sen 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FC5BFC5EFB08FC69" author="Sangma, M. A. & Saikia, P. K." box="[959,1238,908,933]" pageId="66" pageNumber="67" pagination="2404 - 2424" refId="ref57292" refString="Sangma, M. A. & Saikia, P. K. (2015) Checklist of amphibians inventoried from Tura Peak of West Garo Hills, Meghalaya, India. Journal of Global Biosciences, 4 (5), 2404 - 2424." type="journal article" year="2015">Sangma & Saikia 2015</bibRefCitation>
|
||
) populations are conspecific with
|
||
<taxonomicName id="4C7F4D1FFF9C0270FEBCFC7DFD85FC04" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[344,603,943,968]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" sensu="stricto" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FEBCFC7DFD85FC04" box="[344,603,943,968]" italics="true" pageId="66" pageNumber="67">Megophrys glandulosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9C0270FD87FC63FD59FC04" box="[611,647,945,968]" pageId="66" pageNumber="67" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FD6BFC62FC9BFC04" author="Ao, M. & Bordoloi, S. & Ohler, A." box="[655,837,943,968]" pageId="66" pageNumber="67" pagination="1117 - 1125" refId="ref49909" refString="Ao, M., Bordoloi, S. & Ohler, A. (2003) Amphibian fauna of Nagaland with nineteen new records from the State including five new records for India. Zoos' Print Journal, 18 (6), 1117 - 1125. https: // doi. org / 10.11609 / JoTT. ZPJ. 18.6.1117 - 25" type="journal article" year="2003">
|
||
Ao
|
||
<emphasis id="B90BEA8EFF9C0270FD5DFC63FD2DFC04" box="[697,755,943,968]" italics="true" pageId="66" pageNumber="67">et al.</emphasis>
|
||
(2003)
|
||
</bibRefCitation>
|
||
reported a specimen of this species from Puliebadze (
|
||
<geoCoordinate id="EE4B505BFF9C0270FF7BFC06FEC6FC21" box="[159,280,980,1005]" direction="north" orientation="latitude" pageId="66" pageNumber="67" precision="15" value="25.616667">25°37'0"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFF9C0270FECFFC06FE4CFC21" box="[299,402,980,1005]" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="15" value="94.083336">94°5'0"E</geoCoordinate>
|
||
,
|
||
<quantity id="4C879B79FF9C0270FE40FC06FE21FC20" box="[420,511,980,1005]" metricMagnitude="3" metricUnit="m" metricValue="2.225" pageId="66" pageNumber="67" unit="m" value="2225.0">2225 m</quantity>
|
||
asl.), Kohima district,
|
||
<collectingRegion id="49BBF87EFF9C0270FCFEFC06FC5BFC21" box="[794,901,980,1005]" country="India" name="Nagaland" pageId="66" pageNumber="67">Nagaland</collectingRegion>
|
||
state, Northeast
|
||
<collectingCountry id="F368760CFF9C0270FBB2FC06FB4DFC21" box="[1110,1171,980,1005]" name="India" pageId="66" pageNumber="67">India</collectingCountry>
|
||
. Though we have not examined this specimen, we report two species of MMC from Kohima district in this study (
|
||
<taxonomicName id="4C7F4D1FFF9C0270FB27FC2BFAF0FBDC" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[1219,1326,1017,1040]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" sensu="stricto" species="major">
|
||
<emphasis id="B90BEA8EFF9C0270FB27FC2BFAF0FBDC" box="[1219,1326,1017,1040]" italics="true" pageId="66" pageNumber="67">M. major</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9C0270FAD3FC2BFA85FBDC" box="[1335,1371,1017,1040]" pageId="66" pageNumber="67" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
<quantity id="4C879B79FF9C0270FA80FC25FF2EFBF8" metricMagnitude="2" metricUnit="m" metricValue="8.525" metricValueMax="12.55" metricValueMin="4.5" pageId="66" pageNumber="67" unit="m" value="852.5" valueMax="1255.0" valueMin="450.0">450– 1255 m</quantity>
|
||
asl. and
|
||
<taxonomicName id="4C7F4D1FFF9C0270FEBAFBCEFDA5FBF9" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[350,635,1052,1077]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFF9C0270FEBAFBCEFDA5FBF9" box="[350,635,1052,1077]" italics="true" pageId="66" pageNumber="67">Megophrys flavipunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9C0270FD61FBCEFD3DFBF9" box="[645,739,1052,1077]" pageId="66" pageNumber="67" rank="species">
|
||
<emphasis id="B90BEA8EFF9C0270FD61FBCEFD3DFBF9" bold="true" box="[645,739,1052,1077]" pageId="66" pageNumber="67">
|
||
sp.
|
||
<date id="FFC1105CFF9C0270FD55FBCEFD3DFBF9" box="[689,739,1052,1077]" pageId="66" pageNumber="67" value="1575-11">nov.</date>
|
||
</emphasis>
|
||
</taxonomicNameLabel>
|
||
<quantity id="4C879B79FF9C0270FD09FBCEFC52FBF8" box="[749,908,1052,1077]" metricMagnitude="3" metricUnit="m" metricValue="1.6925" metricValueMax="1.81" metricValueMin="1.575" pageId="66" pageNumber="67" unit="m" value="1692.5" valueMax="1810.0" valueMin="1575.0">1575–1810 m</quantity>
|
||
asl.). Based on the elevation that
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FAC5FBCEFF3FFB94" author="Ao, M. & Bordoloi, S. & Ohler, A." pageId="66" pageNumber="67" pagination="1117 - 1125" refId="ref49909" refString="Ao, M., Bordoloi, S. & Ohler, A. (2003) Amphibian fauna of Nagaland with nineteen new records from the State including five new records for India. Zoos' Print Journal, 18 (6), 1117 - 1125. https: // doi. org / 10.11609 / JoTT. ZPJ. 18.6.1117 - 25" type="journal article" year="2003">
|
||
Ao
|
||
<emphasis id="B90BEA8EFF9C0270FAA9FBCFFA5DFBF9" box="[1357,1411,1052,1077]" italics="true" pageId="66" pageNumber="67">et al</emphasis>
|
||
.’s (2003)
|
||
</bibRefCitation>
|
||
specimen was collected, we suspect that it might represent
|
||
<taxonomicName id="4C7F4D1FFF9C0270FC6EFBEDFB7AFB94" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[906,1188,1087,1112]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFF9C0270FC6EFBEDFB7AFB94" box="[906,1188,1087,1112]" italics="true" pageId="66" pageNumber="67">Megophrys flavipunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9C0270FB48FBEDFAD9FB94" box="[1196,1287,1087,1112]" pageId="66" pageNumber="67" rank="species">
|
||
<emphasis id="B90BEA8EFF9C0270FB48FBEDFAD9FB94" bold="true" box="[1196,1287,1087,1112]" pageId="66" pageNumber="67">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FAEBFBEDFEC8FBB0" author="Mathew, R. & Sen, N." pageId="66" pageNumber="67" refId="ref55566" refString="Mathew, R. & Sen, N. (2010) Pictorial Guide to Amphibians of North East India. Zoological Survey of India, Kolkata, 144 pp." type="book" year="2010">Mathew and Sen (2010)</bibRefCitation>
|
||
provided an undiagnostic description of
|
||
<taxonomicName id="4C7F4D1FFF9C0270FD11FBB7FC45FBB1" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[757,923,1124,1149]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FD11FBB7FC45FBB1" box="[757,923,1124,1149]" italics="true" pageId="66" pageNumber="67">M. glandulosa</emphasis>
|
||
</taxonomicName>
|
||
for
|
||
<collectingCountry id="F368760CFF9C0270FC34FBB6FBD4FBB1" box="[976,1034,1124,1149]" name="India" pageId="66" pageNumber="67">India</collectingCountry>
|
||
with an accompanying photo of a specimen that appears to represent
|
||
<taxonomicName id="4C7F4D1FFF9C0270FDDBFB5BFD70FB6C" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[575,686,1161,1184]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" sensu="stricto" species="major">
|
||
<emphasis id="B90BEA8EFF9C0270FDDBFB5BFD70FB6C" box="[575,686,1161,1184]" italics="true" pageId="66" pageNumber="67">M. major</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9C0270FD58FB5BFD01FB6C" box="[700,735,1161,1184]" pageId="66" pageNumber="67" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
A report of two specimens of
|
||
<taxonomicName id="4C7F4D1FFF9C0270FB85FB55FB01FB6C" box="[1121,1247,1159,1184]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90BEA8EFF9C0270FB85FB55FB01FB6C" box="[1121,1247,1159,1184]" italics="true" pageId="66" pageNumber="67">Megophrys</emphasis>
|
||
</taxonomicName>
|
||
[as
|
||
<taxonomicName id="4C7F4D1FFF9C0270FAFFFB55FA4DFB6C" box="[1307,1427,1159,1184]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="12" phylum="Chordata" rank="subGenus" subGenus="Xenophrys">
|
||
<emphasis id="B90BEA8EFF9C0270FAFFFB55FA4DFB6C" box="[1307,1427,1159,1184]" italics="true" pageId="66" pageNumber="67">Xenophrys</emphasis>
|
||
</taxonomicName>
|
||
]
|
||
<taxonomicName id="4C7F4D1FFF9C0270FF73FB7EFECAFB09" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[151,276,1196,1221]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FF73FB7EFECAFB09" box="[151,276,1196,1221]" italics="true" pageId="66" pageNumber="67">glandulosa</emphasis>
|
||
</taxonomicName>
|
||
from Tura Peak, West Garo Hills, in
|
||
<collectingRegion id="49BBF87EFF9C0270FD5FFB7EFCE9FB09" box="[699,823,1196,1221]" country="India" name="Meghalaya" pageId="66" pageNumber="67">Meghalaya</collectingRegion>
|
||
state provided by
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FBEDFB7EFAF3FB09" author="Sangma, M. A. & Saikia, P. K." box="[1033,1325,1196,1221]" pageId="66" pageNumber="67" pagination="2404 - 2424" refId="ref57292" refString="Sangma, M. A. & Saikia, P. K. (2015) Checklist of amphibians inventoried from Tura Peak of West Garo Hills, Meghalaya, India. Journal of Global Biosciences, 4 (5), 2404 - 2424." type="journal article" year="2015">Sangma and Saikia (2015</bibRefCitation>
|
||
: 2407, pl. 4a) appears to represent an adult
|
||
<taxonomicName id="4C7F4D1FFF9C0270FDC7FB1DFCF8FB24" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[547,806,1231,1256]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFF9C0270FDC7FB1DFCF8FB24" box="[547,806,1231,1256]" italics="true" pageId="66" pageNumber="67">Megophrys oreocrypta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FF9C0270FCD5FB1DFC51FB24" box="[817,911,1231,1256]" pageId="66" pageNumber="67" rank="species">
|
||
<emphasis id="B90BEA8EFF9C0270FCD5FB1DFC51FB24" bold="true" box="[817,911,1231,1256]" pageId="66" pageNumber="67">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
based on the figure provided for one of the specimens.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFF9C0270FF23FACAFC96F983" blockId="66.[151,1437,151,1760]" pageId="66" pageNumber="67">
|
||
An additional report of
|
||
<taxonomicName id="4C7F4D1FFF9C0270FE07FACBFD54FAFC" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[483,650,1303,1328]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FE07FACBFD54FAFC" box="[483,650,1303,1328]" italics="true" pageId="66" pageNumber="67">M. glandulosa</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FD59FAC5FBC2FAFC" author="Deuti, K & Ayyaswamy, A. K." box="[701,1052,1303,1328]" pageId="66" pageNumber="67" pagination="17 - 20" refId="ref52242" refString="Deuti, K & Ayyaswamy, A. K. (2008) Three new records of amphibians from West Bengal. Zoological Survey of India, 108 (4), 17 - 20." type="journal article" year="2008">Deuti and Ayyaswamy (2008)</bibRefCitation>
|
||
was based on a collection from Latpanchar, Darjeeling district,
|
||
<collectingRegion id="49BBF87EFF9C0270FDF6FAEFFD79FA99" box="[530,679,1340,1365]" country="India" name="West Bengal" pageId="66" pageNumber="67">West Bengal</collectingRegion>
|
||
state, Northeast
|
||
<collectingCountry id="F368760CFF9C0270FC9FFAEEFC66FA99" box="[891,952,1340,1365]" name="India" pageId="66" pageNumber="67">India</collectingCountry>
|
||
. The figured individual displays typical characters seen on
|
||
<taxonomicName id="4C7F4D1FFF9C0270FE8FFAB3FE3AFAB4" authorityName=", Rai and Anders" authorityYear="2002" box="[363,484,1375,1400]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="robusta">
|
||
<emphasis id="B90BEA8EFF9C0270FE8FFAB3FE3AFAB4" box="[363,484,1375,1400]" italics="true" pageId="66" pageNumber="67">M. robusta</emphasis>
|
||
</taxonomicName>
|
||
, i.e., slightly enlarged posterior supratympanic ridge (vs. greatly enlarged, and beanshaped on
|
||
<taxonomicName id="4C7F4D1FFF9C0270FEF4FA57FE6AFA51" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[272,436,1412,1437]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FEF4FA57FE6AFA51" box="[272,436,1412,1437]" italics="true" pageId="66" pageNumber="67">M. glandulosa</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="13442A19FF9C0270FE26FA56FD94FA50" box="[450,586,1412,1437]" captionStart="FIGURE 24" captionStartId="64.[151,250,1907,1929]" captionTargetBox="[151,1436,553,1885]" captionTargetId="figure@64.[151,1436,553,1885]" captionTargetPageId="64" captionText="FIGURE 24. Megophrys glandulosa holotype: adult male (CIB 873112: SVL 78.4 mm [Fei et al. 1992]) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610250/files/figure.png" pageId="66" pageNumber="67">Figure 24C</figureCitation>
|
||
]); the distinctive white upper lip stripe of
|
||
<taxonomicName id="4C7F4D1FFF9C0270FBC2FA57FB14FA51" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[1062,1226,1412,1437]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FBC2FA57FB14FA51" box="[1062,1226,1412,1437]" italics="true" pageId="66" pageNumber="67">M. glandulosa</emphasis>
|
||
</taxonomicName>
|
||
is not apparent on the figured animal, and the iris colour matches that of
|
||
<taxonomicName id="4C7F4D1FFF9C0270FCE0FA7BFCA0FA0C" authorityName=", Rai and Anders" authorityYear="2002" box="[772,894,1447,1472]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="robusta">
|
||
<emphasis id="B90BEA8EFF9C0270FCE0FA7BFCA0FA0C" box="[772,894,1447,1472]" italics="true" pageId="66" pageNumber="67">M. robusta</emphasis>
|
||
</taxonomicName>
|
||
perfectly. Although some of the characters that were described, such as ventral markings, could correspond with
|
||
<taxonomicName id="4C7F4D1FFF9C0270FC98FA1FFBC1FA29" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[892,1055,1484,1509]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FC98FA1FFBC1FA29" box="[892,1055,1484,1509]" italics="true" pageId="66" pageNumber="67">M. glandulosa</emphasis>
|
||
</taxonomicName>
|
||
, other characters such as swollen digit tips, and rudimentary webbing between toes do not (see description provided above). Pending re-examination of the specimens reported by
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FE04F9C6FCEFF9E0" author="Deuti, K & Ayyaswamy, A. K." box="[480,817,1556,1581]" pageId="66" pageNumber="67" pagination="17 - 20" refId="ref52242" refString="Deuti, K & Ayyaswamy, A. K. (2008) Three new records of amphibians from West Bengal. Zoological Survey of India, 108 (4), 17 - 20." type="journal article" year="2008">Deuti and Ayyaswamy (2008)</bibRefCitation>
|
||
, we recommend that Darjeeling, and Northeast
|
||
<collectingCountry id="F368760CFF9C0270FAA2F9C6FA5EF9E1" box="[1350,1408,1556,1581]" name="India" pageId="66" pageNumber="67">India</collectingCountry>
|
||
in general, are not included in the distribution of
|
||
<taxonomicName id="4C7F4D1FFF9C0270FD7BF9EBFC9EF99C" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[671,832,1591,1616]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FD7BF9EBFC9EF99C" box="[671,832,1591,1616]" italics="true" pageId="66" pageNumber="67">M. glandulosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFF9C0270FF23F98EFD74F92C" blockId="66.[151,1437,151,1760]" pageId="66" pageNumber="67">
|
||
<bibRefCitation id="EFEE4B6DFF9C0270FF23F98EFE59F9B8" author="Wangyal, J. T." box="[199,391,1628,1653]" pageId="66" pageNumber="67" pagination="4774 - 4783" refId="ref59089" refString="Wangyal, J. T. (2013) New records of reptiles and amphibians from Bhutan. Journal of Threatened Taxa, 5, 4774 - 4783. https: // doi. org / 10.11609 / JoTT. o 3539.4774 - 83" type="journal article" year="2013">Wangyal (2013)</bibRefCitation>
|
||
reported
|
||
<taxonomicName id="4C7F4D1FFF9C0270FDEDF98FFD69F9B9" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[521,695,1628,1653]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FDEDF98FFD69F9B9" box="[521,695,1628,1653]" italics="true" pageId="66" pageNumber="67">M. glandulosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7F4D1FFF9C0270FCE0F98FFCA9F9B8" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[772,887,1629,1652]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="B90BEA8EFF9C0270FCE0F98FFCA9F9B8" box="[772,887,1629,1652]" italics="true" pageId="66" pageNumber="67">M. major</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F368760CFF9C0270FC37F98EFBF9F9B9" box="[979,1063,1628,1653]" name="Bhutan" pageId="66" pageNumber="67">Bhutan</collectingCountry>
|
||
, however, the two specimens photographed exhibit characters that allow a reasonable preliminary identification as
|
||
<taxonomicName id="4C7F4D1FFF9C0270FBB7F953FB12F954" authorityName=", Rai and Anders" authorityYear="2002" box="[1107,1228,1663,1688]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="robusta">
|
||
<emphasis id="B90BEA8EFF9C0270FBB7F953FB12F954" box="[1107,1228,1663,1688]" italics="true" pageId="66" pageNumber="67">M. robusta</emphasis>
|
||
</taxonomicName>
|
||
, i.e., both lack the light upper lip stripe seen on
|
||
<taxonomicName id="4C7F4D1FFF9C0270FE3FF977FDA3F971" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[475,637,1700,1725]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="glandulosa">
|
||
<emphasis id="B90BEA8EFF9C0270FE3FF977FDA3F971" box="[475,637,1700,1725]" italics="true" pageId="66" pageNumber="67">M. glandulosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7F4D1FFF9C0270FD56F977FCC6F970" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[690,792,1701,1724]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="B90BEA8EFF9C0270FD56F977FCC6F970" box="[690,792,1701,1724]" italics="true" pageId="66" pageNumber="67">M. major</emphasis>
|
||
</taxonomicName>
|
||
, and have pale iris colouration that is characteristic of the
|
||
<taxonomicName id="4C7F4D1FFF9C0270FF73F91BFECEF92C" authorityName=", Rai and Anders" authorityYear="2002" box="[151,272,1735,1760]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="robusta">
|
||
<emphasis id="B90BEA8EFF9C0270FF73F91BFECEF92C" box="[151,272,1735,1760]" italics="true" pageId="66" pageNumber="67">M. robusta</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7F4D1FFF9C0270FEFCF91BFE1FF92C" authorityName="Fei, Ye and Huang" authorityYear="1983" box="[280,449,1735,1760]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="medogensis">
|
||
<emphasis id="B90BEA8EFF9C0270FEFCF91BFE1FF92C" box="[280,449,1735,1760]" italics="true" pageId="66" pageNumber="67">M. medogensis</emphasis>
|
||
</taxonomicName>
|
||
clade in the MMSG.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |