treatments-xml/data/03/D0/87/03D08788E9122362FF456A0B9DE3FF28.xml

<document id="49C63A34712E7AF40772CDC107D2C4C1" ID-CLB-Dataset="45798" ID-DOI="10.11646/zootaxa.3795.3.8" ID-GBIF-Dataset="9f394f0e-b737-4ab2-9ee0-29c8ceee3790" ID-ISSN="1175-5326" ID-Zenodo-Dep="230954" ID-ZooBank="59706921-DF1D-4B7E-9615-52E8A9F2520D" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460419041812" checkinUser="plazi" docAuthor="Arias, Diomedes Quintero &amp; Esposito, Lauren A." docDate="2014" docId="03D08788E9122362FF456A0B9DE3FF28" docLanguage="en" docName="zt03795p382.pdf" docOrigin="Zootaxa 3795 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Centruroides panamensis Arias &amp; Esposito, 2014, new species" docType="treatment" docUuid="E72E2CEF-D7AD-4C17-AD15-B240E3AA9E7C" docUuidSource="ZooBank" docVersion="10" lastPageNumber="380" masterDocId="FFE9FFF0E913236AFFD26B649C04FFFD" masterDocTitle="A new species of Centruroides Marx (Scorpiones: Buthidae) from Panama and new distribution records for Centruroides bicolor (Pocock, 1898) and Centruroides granosus (Thorell, 1876)" masterLastPageNumber="382" masterPageNumber="373" pageNumber="374" updateTime="1698329475639" updateUser="ExternalLinkService">
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<mods:title id="27CC4D48D6E156C75E58CFB946C47DF7">A new species of Centruroides Marx (Scorpiones: Buthidae) from Panama and new distribution records for Centruroides bicolor (Pocock, 1898) and Centruroides granosus (Thorell, 1876)</mods:title>
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<mods:namePart id="38775B05FD17C5003963472EFB3A6215">Arias, Diomedes Quintero</mods:namePart>
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<mods:namePart id="ED4177B3A2B989E91608CFFD5F277331">Esposito, Lauren A.</mods:namePart>
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<treatment id="03D08788E9122362FF456A0B9DE3FF28" ID-DOI="http://doi.org/10.5281/zenodo.5695687" ID-GBIF-Taxon="119426402" ID-Zenodo-Dep="5695687" LSID="urn:lsid:zoobank.org:act:E72E2CEF-D7AD-4C17-AD15-B240E3AA9E7C" httpUri="http://treatment.plazi.org/id/03D08788E9122362FF456A0B9DE3FF28" lastPageId="8" lastPageNumber="380" pageId="1" pageNumber="374">
<subSubSection id="C3636515E912236BFF456A0B9D77FE51" pageId="1" pageNumber="374" type="nomenclature">
<paragraph id="8BC6369EE912236BFF456A0B9E4BFE75" blockId="1.[151,591,367,429]" box="[151,591,367,392]" pageId="1" pageNumber="374">
<heading id="D08E81F2E912236BFF456A0B9E4BFE75" bold="true" box="[151,591,367,392]" fontSize="10" level="2" pageId="1" pageNumber="374" reason="2">
<emphasis id="B90DEA8CE912236BFF456A0B9E4BFE75" bold="true" box="[151,591,367,392]" pageId="1" pageNumber="374">
<taxonomicName id="4C794D1DE912236BFF456A0B9DBAFE75" box="[151,446,367,392]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE912236BFF456A0B9DBAFE75" bold="true" box="[151,446,367,392]" italics="true" pageId="1" pageNumber="374">Centruroides panamensis</emphasis>
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<taxonomicNameLabel id="A23E57F7E912236BFE166A0B9E4BFE75" box="[452,591,367,392]" pageId="1" pageNumber="374" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8BC6369EE912236BFF456AF09D77FE51" blockId="1.[151,591,367,429]" box="[151,371,404,429]" pageId="1" pageNumber="374">
(
<figureCitation id="13422A1BE912236BFF4D6AF09D0FFE51" box="[159,267,404,429]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1928,1950]" captionStartId-1="2.[151,250,1129,1151]" captionStartId-2="4.[151,250,1948,1970]" captionStartId-3="5.[151,250,917,939]" captionTargetBox-0="[151,1436,913,1905]" captionTargetBox-1="[151,1436,193,1107]" captionTargetBox-2="[161,1425,390,1925]" captionTargetBox-3="[151,1436,445,898]" captionTargetId-0="figure@1.[151,1436,913,1906]" captionTargetId-1="figure@2.[151,1436,193,1108]" captionTargetId-2="figure@4.[160,1433,334,1982]" captionTargetId-3="figure@5.[151,1436,445,899]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Habitus of Centruroides panamensis new species: A—male paratype, dorsal aspect; B—male paratype, ventral aspect; C—female paratype, dorsal aspect; D—female paratype, ventral aspect. Scale bar represents 10 mm." captionText-1="FIGURE 2. Centruroides panamensis new species: A—male paratype dorsal carapace; B—female paratype dorsal carapace; C—female paratype pectines; D—male paratype pectines. Scale bar represents 1 mm." captionText-2="FIGURE 3. Centruroides panamensis new species, pedipalps. Female Paratype: A—dorsal aspect of femur; B—dorsal aspect of patella; C—dorsoexternal aspect of chela. Male Paratype: D—dorsal aspect of femur; E—dorsal aspect of patella; F—dorsoexternal aspect of chela. Scale bar represents 1 mm." captionText-3="FIGURE 4. Centruroides panamensis new species, metasoma. From top to bottom: lateral aspect of paratype male, ventral aspect of paratype male, lateral aspect of paratype female. Scale bar represents 2 mm." httpUri-0="https://zenodo.org/record/230955/files/figure.png" httpUri-1="https://zenodo.org/record/230956/files/figure.png" httpUri-2="https://zenodo.org/record/230957/files/figure.png" httpUri-3="https://zenodo.org/record/230959/files/figure.png" pageId="1" pageNumber="374">Figs. 1–4</figureCitation>
; 
<tableCitation id="C6FB0325E912236BFEC56AF09D6CFE51" box="[279,360,404,429]" captionStart="TABLE 1" captionStartId="7.[151,239,512,535]" captionTargetBox="[151,1414,563,1775]" captionTargetPageId="7" captionText="TABLE 1. Measurements of Centruroides panamensis new species, in mm." httpUri="http://table.plazi.org/id/DF066616E914236DFF4569649FC4FDEA" pageId="1" pageNumber="374" tableUuid="DF066616E914236DFF4569649FC4FDEA">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3636515E912236BFF456AB8997DFCC5" pageId="1" pageNumber="374" type="materials_examined">
<paragraph id="8BC6369EE912236BFF456AB8997DFCC5" blockId="1.[151,1437,476,861]" pageId="1" pageNumber="374">
<emphasis id="B90DEA8CE912236BFF456AB89E05FE08" bold="true" box="[151,513,476,501]" pageId="1" pageNumber="374">
Material examined. 
<collectingCountry id="F36E760EE912236BFE5B6AB89E05FE08" box="[393,513,476,501]" name="Panama" pageId="1" pageNumber="374">PANAMA</collectingCountry>
</emphasis>
: Chiriquí Province, Volcán Barú faldas, 
<typeStatus id="54C2883CE912236BFC1D6AB89835FE08" box="[975,1073,476,501]" pageId="1" pageNumber="374" type="holotype">holotype</typeStatus>
·♂ [
<geoCoordinate id="EE4D5059E912236BFBB56AB898BAFE09" box="[1127,1214,476,500]" direction="north" orientation="latitude" pageId="1" pageNumber="374" precision="925" value="8.8">8°48’N</geoCoordinate>
, 
<geoCoordinate id="EE4D5059E912236BFB186AB89932FE09" box="[1226,1334,476,500]" direction="west" orientation="longitude" pageId="1" pageNumber="374" precision="925" value="-82.61667">82°37’W</geoCoordinate>
], 
<date id="FFC7105EE912236BFA9B6AB89CD6FDE5" pageId="1" pageNumber="374" value="1984-07-14">14 July 1984</date>
, Arsenio Araúz (LAV-MIUP). Allotype ♀ (gravid): Finca Lukarel (see legend of Fig. 11), Volcancito [
<geoCoordinate id="EE4D5059E912236BFF4C69409CF1FDC1" box="[158,245,548,572]" direction="north" orientation="latitude" pageId="1" pageNumber="374" precision="925" value="8.783334">8°47’N</geoCoordinate>
, 82°28W], 
<date id="FFC7105EE912236BFEAB69409E2DFDC1" box="[377,553,548,573]" pageId="1" pageNumber="374" value="1984-01-02">2 January 1984</date>
, Efraín &amp; Lorena Bruña (LAV-MIUP). 
<typeStatus id="54C2883CE912236BFC396940988EFDC1" box="[1003,1162,548,572]" pageId="1" pageNumber="374" type="paratype">PARATYPES</typeStatus>
: 
<specimenCount id="9D7FFD17E912236BFB44694098C7FDC1" box="[1174,1219,548,572]" pageId="1" pageNumber="374" type="generic">3 ♂</specimenCount>
, 3 ♀, same data as 
<typeStatus id="54C2883CE912236BFF4569239CFDFD9D" box="[151,249,583,608]" pageId="1" pageNumber="374" type="holotype">holotype</typeStatus>
(AMNH-NY); 1 immature, Finca Lukarel, Volcancito, 
<date id="FFC7105EE912236BFCB9692C9833FD9D" box="[875,1079,583,608]" pageId="1" pageNumber="374" value="1984-12-09">9 December 1984</date>
, Lorena Bruña (AMNH-NY). 1 ♀, falda Volcán Barú, camino hacia Aguacate, 
<quantity id="4C819B7BE912236BFD6569089F15FD7E" box="[695,785,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="374" unit="m" value="1650.0">1650 m</quantity>
, 
<date id="FFC7105EE912236BFCCD69089FA1FD79" box="[799,933,620,644]" pageId="1" pageNumber="374" value="2012-02-04">4 Feb 2012</date>
, Arsenio Araúz (LAV-MIUP); 
<specimenCount id="9D7FFD17E912236BFAD86908995FFD79" box="[1290,1371,620,644]" pageId="1" pageNumber="374" type="male">1 male</specimenCount>
, 1 ♀, Finca Lukarel, Boquete, 
<date id="FFC7105EE912236BFE7F69F49E38FD55" box="[429,572,656,680]" pageId="1" pageNumber="374" value="1985-02-13">13 Feb 1985</date>
, Lorena Bruña (LAV-MIUP); 
<specimenCount id="9D7FFD17E912236BFC5F69F49805FD55" box="[909,1025,656,680]" pageId="1" pageNumber="374" type="subadult">1 subadult</specimenCount>
♀ under dried log, 2 ♀ under stones carrying on their dorsum 20 immatures (2–3 escaped) &amp; 22 immatures, 
<quantity id="4C819B7BE912236BFC3469D09882FD31" box="[998,1158,692,717]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="1" pageNumber="374" unit="m" value="375.0" valueMax="400.0" valueMin="350.0">350 to 400 m</quantity>
from entrance of Finca Lukarel, 
<quantity id="4C819B7BE912236BFF2C69B39D53FD12" box="[254,343,727,752]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="374" unit="m" value="1160.0">1160 m</quantity>
; Volcancito, Boquete, 
<date id="FFC7105EE912236BFD8A69BC9EE8FD0D" box="[600,748,727,752]" pageId="1" pageNumber="374" value="2012-02-22">22 Feb 2012</date>
, Arsenio Araúz (AMNH-NY); 
<specimenCount id="9D7FFD17E912236BFB8069BC98C4FD0D" box="[1106,1216,727,752]" pageId="1" pageNumber="374" type="female">2 females</specimenCount>
under stones, Paso Ancho, Bugaba, 
<date id="FFC7105EE912236BFE8E69989DADFCE9" box="[348,425,764,788]" pageId="1" pageNumber="374" value="2012-08">3 Aug</date>
, 
<date id="FFC7105EE912236BFE6B69989E47FCE9" box="[441,579,764,788]" pageId="1" pageNumber="374" value="2012-09-01">1 Sep 2012</date>
, A. Pitty, N. Santamaría (LAV-MIUP); 1 ♀ Paso Ancho, Bugaba, inside bathroom, 
<date id="FFC7105EE912236BFEC268449D90FCC5" box="[272,404,800,824]" pageId="1" pageNumber="374" value="2012-12-05">5 Dec 2012</date>
, A. Miranda (LAV-MIUP); 1 ♀, Cuesta de Piedra, 
<date id="FFC7105EE912236BFC1C68449840FCC5" box="[974,1092,800,824]" pageId="1" pageNumber="374" value="2012-07-06">6 Jul 2012</date>
, L. Espinosa (LAV-MIUP).
</paragraph>
</subSubSection>
<subSubSection id="C3636515E912236BFF1568209819FCA0" box="[199,1053,836,861]" pageId="1" pageNumber="374" type="etymology">
<paragraph id="8BC6369EE912236BFF1568209819FCA0" blockId="1.[151,1437,476,861]" box="[199,1053,836,861]" pageId="1" pageNumber="374">
<emphasis id="B90DEA8CE912236BFF1568209D4AFCA0" bold="true" box="[199,334,836,861]" pageId="1" pageNumber="374">Etymology.</emphasis>
The specific name refers to the 
<typeStatus id="54C2883CE912236BFD6468219EE2FCA0" box="[694,742,837,861]" pageId="1" pageNumber="374">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
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<paragraph id="8BC6369EE912236BFF456CEC98DDF841" blockId="1.[151,1435,1928,1980]" pageId="1" pageNumber="374">
<emphasis id="B90DEA8CE912236BFF456CEC9D11F860" bold="true" box="[151,277,1928,1950]" pageId="1" pageNumber="374">FIGURE 1.</emphasis>
Habitus of 
<taxonomicName id="4C794D1DE912236BFE406CEC9E93F863" box="[402,663,1928,1950]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="374" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE912236BFE406CEC9E93F863" box="[402,663,1928,1950]" italics="true" pageId="1" pageNumber="374">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E912236BFD726CED9F1BF863" box="[672,799,1928,1950]" pageId="1" pageNumber="374" rank="species">new species</taxonomicNameLabel>
: A—male paratype, dorsal aspect; B—male paratype, ventral aspect; C—female paratype, dorsal aspect; D—female paratype, ventral aspect. Scale bar represents 10mm.
</paragraph>
</caption>
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<paragraph id="8BC6369EE9112368FF456F0D9FF2FB63" blockId="2.[151,1436,1129,1182]" pageId="2" pageNumber="375">
<emphasis id="B90DEA8CE9112368FF456F0D9D10FB83" bold="true" box="[151,276,1129,1151]" pageId="2" pageNumber="375">FIGURE 2.</emphasis>
<taxonomicName id="4C794D1DE9112368FECE6F0D9E24FB82" box="[284,544,1129,1151]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9112368FECE6F0D9E24FB82" box="[284,544,1129,1151]" italics="true" pageId="2" pageNumber="375">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9112368FDF56F0E9EA2FB82" box="[551,678,1129,1151]" pageId="2" pageNumber="375" rank="species">new species</taxonomicNameLabel>
: A—male paratype dorsal carapace; B—female paratype dorsal carapace; C—female paratype pectines; D—male paratype pectines. Scale bar represents 1mm.
</paragraph>
</caption>
<subSubSection id="C3636515E9112369FF156FA19FAAFEE0" lastPageId="3" lastPageNumber="376" pageId="2" pageNumber="375" type="diagnosis">
<paragraph id="8BC6369EE9112368FF156FA19928FA4B" blockId="2.[151,1437,1221,2038]" pageId="2" pageNumber="375">
<emphasis id="B90DEA8CE9112368FF156FA19D45FB23" bold="true" box="[199,321,1221,1246]" pageId="2" pageNumber="375">Diagnosis.</emphasis>
Medium size scorpions, total length 
<quantity id="4C819B7BE9112368FD376FA19F7CFB20" box="[741,888,1221,1246]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="6.5" metricValueMin="5.4" pageId="2" pageNumber="375" unit="mm" value="59.5" valueMax="65.0" valueMin="54.0">54 to 65 mm</quantity>
, lightly colored (yellow) with infuscation on the pedipalp fingers, carapace, 
<taxonomicName id="4C794D1DE9112368FE1F6F8E9E42FAFC" box="[461,582,1258,1281]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">mesosoma</taxonomicName>
, and ventral surfaces of the metasoma; metasoma V and telson brown. The new species is the only Central American 
<taxonomicName id="4C794D1DE9112368FDA66E699F03FADB" box="[628,775,1293,1318]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA8CE9112368FDA66E699F03FADB" box="[628,775,1293,1318]" italics="true" pageId="2" pageNumber="375">Centruroides</emphasis>
</taxonomicName>
with eight oblique rows of granules on the dentate margin of the fixed finger of the pedipalp chela, and pectinal tooth counts of 17 to 21. Carapace and 
<taxonomicName id="4C794D1DE9112368FB1A6E56993AFAB4" box="[1224,1342,1330,1353]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">mesosoma</taxonomicName>
densely granulated with spiniform granules; pedipalp and metasoma intercarinal surfaces smooth. Pedipalp chela with five distinct carinae: dorsomedian, dorsal secondary, ventroexternal, digital, dorsointernal. Dimorphic metasoma (elongate in ♂), telson slightly elongate and non-lobate with well-developed spinate subaculear tubercle.
</paragraph>
<paragraph id="8BC6369EE9112369FF156EA59FAAFEE0" blockId="2.[151,1437,1221,2038]" lastBlockId="3.[151,1437,151,2018]" lastPageId="3" lastPageNumber="376" pageId="2" pageNumber="375">
<taxonomicName id="4C794D1DE9112368FF156EA59DE2FA24" box="[199,486,1473,1498]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9112368FF156EA59DE2FA24" box="[199,486,1473,1498]" italics="true" pageId="2" pageNumber="375">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9112368FE3C6EA59E2BFA27" box="[494,559,1473,1498]" pageId="2" pageNumber="375" rank="species">
<emphasis id="B90DEA8CE9112368FE3C6EA59E2BFA27" bold="true" box="[494,559,1473,1498]" pageId="2" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
can be distinguished from the sympatric species 
<taxonomicName id="4C794D1DE9112368FBB36EA59949FA27" box="[1121,1357,1473,1498]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE9112368FBB36EA59949FA27" box="[1121,1357,1473,1498]" italics="true" pageId="2" pageNumber="375">Centruroides bicolor</emphasis>
</taxonomicName>
by the following characters: smaller overall size (
<quantity id="4C819B7BE9112368FDA76E819F0DFA00" box="[629,777,1509,1534]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="6.5" metricValueMin="5.4" pageId="2" pageNumber="375" unit="mm" value="59.5" valueMax="65.0" valueMin="54.0">54 to 65 mm</quantity>
, 
<taxonomicName id="4C794D1DE9112368FCC66E829F82FA03" box="[788,902,1509,1534]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE9112368FCC66E829F82FA03" box="[788,902,1509,1534]" italics="true" pageId="2" pageNumber="375">C. bicolor</emphasis>
</taxonomicName>
is 
<quantity id="4C819B7BE9112368FC7A6E82985CFA03" box="[936,1112,1509,1534]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="2" pageNumber="375" unit="mm" value="130.0" valueMax="160.0" valueMin="100.0">100 to 160 mm</quantity>
), lower pectinal tooth counts (17 to 21, 
<taxonomicName id="4C794D1DE9112368FEC26D6E9D80F9DF" box="[272,388,1545,1570]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE9112368FEC26D6E9D80F9DF" box="[272,388,1545,1570]" italics="true" pageId="2" pageNumber="375">C. bicolor</emphasis>
</taxonomicName>
with 23–28), all yellow pedipalp chela manus and patella (pedipalp chela manus and distal patella are dark brown in 
<taxonomicName id="4C794D1DE9112368FE6D6D4A9E37F9BB" box="[447,563,1581,1606]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE9112368FE6D6D4A9E37F9BB" box="[447,563,1581,1606]" italics="true" pageId="2" pageNumber="375">C. bicolor</emphasis>
</taxonomicName>
), discontinuous ventrointernal carina on the pedipalp patella, pedipalp chela internodorsal carina absent (present and granular in 
<taxonomicName id="4C794D1DE9112368FD3D6D369F67F997" box="[751,867,1617,1642]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE9112368FD3D6D369F67F997" box="[751,867,1617,1642]" italics="true" pageId="2" pageNumber="375">C. bicolor</emphasis>
</taxonomicName>
), eight primary rows of granules on the pedipalp fingers in 
<taxonomicName id="4C794D1DE9112368FED96D129DB5F970" box="[267,433,1654,1677]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9112368FED96D129DB5F970" box="[267,433,1654,1677]" italics="true" pageId="2" pageNumber="375">C. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9112368FE686D119DFFF973" box="[442,507,1653,1678]" pageId="2" pageNumber="375" rank="species">
<emphasis id="B90DEA8CE9112368FE686D119DFFF973" bold="true" box="[442,507,1653,1678]" pageId="2" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
(
<taxonomicName id="4C794D1DE9112368FDDF6D129E84F973" box="[525,640,1653,1678]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE9112368FDDF6D129E84F973" box="[525,640,1653,1678]" italics="true" pageId="2" pageNumber="375">C. bicolor</emphasis>
</taxonomicName>
with nine primary rows of granules). 
<taxonomicName id="4C794D1DE9112368FBE16D119957F970" box="[1075,1363,1653,1678]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9112368FBE16D119957F970" box="[1075,1363,1653,1678]" italics="true" pageId="2" pageNumber="375">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9112368FA896D119998F973" box="[1371,1436,1653,1678]" pageId="2" pageNumber="375" rank="species">
<emphasis id="B90DEA8CE9112368FA896D119998F973" bold="true" box="[1371,1436,1653,1678]" pageId="2" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
appears to be most closely related to the allopatric species 
<taxonomicName id="4C794D1DE9112368FCE56DFE9FBEF94F" box="[823,954,1689,1714]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B90DEA8CE9112368FCE56DFE9FBEF94F" box="[823,954,1689,1714]" italics="true" pageId="2" pageNumber="375">C. limbatus</emphasis>
</taxonomicName>
but can be distinguished by the following characters: 
<taxonomicName id="4C794D1DE9112368FECB6DDA9DBBF928" box="[281,447,1726,1749]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9112368FECB6DDA9DBBF928" box="[281,447,1726,1749]" italics="true" pageId="2" pageNumber="375">C. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9112368FE1A6DD99E0DF92B" box="[456,521,1725,1750]" pageId="2" pageNumber="375" rank="species">
<emphasis id="B90DEA8CE9112368FE1A6DD99E0DF92B" bold="true" box="[456,521,1725,1750]" pageId="2" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
has strong granulation on the carapace and 
<taxonomicName id="4C794D1DE9112368FBD16DDA9878F928" box="[1027,1148,1726,1749]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="376" phylum="Arthropoda" rank="genus">mesosoma</taxonomicName>
, weak granulation on the pedipalp carinae; lack of intercarinal granulation on the internal surface of the pedipalp manus, eight primary rows of granules on the pedipalp fingers, discontinuous ventrointernal carina on the pedipalp patella (continuous in 
<taxonomicName id="4C794D1DE9112368FA506C629CF3F8BF" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B90DEA8CE9112368FA506C629CF3F8BF" italics="true" pageId="2" pageNumber="375">C. limbatus</emphasis>
</taxonomicName>
), telson smooth. 
<taxonomicName id="4C794D1DE9112368FE686C4D9EDCF8BC" box="[442,728,1833,1858]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9112368FE686C4D9EDCF8BC" box="[442,728,1833,1858]" italics="true" pageId="2" pageNumber="375">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9112368FD0D6C4D9F1AF8BF" box="[735,798,1833,1858]" pageId="2" pageNumber="375" rank="species">
<emphasis id="B90DEA8CE9112368FD0D6C4D9F1AF8BF" bold="true" box="[735,798,1833,1858]" pageId="2" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C794D1DE9112368FCFF6C4E9FB1F8BC" box="[813,949,1834,1857]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B90DEA8CE9112368FCFF6C4E9FB1F8BC" box="[813,949,1834,1857]" italics="true" pageId="2" pageNumber="375">C. granosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C794D1DE9112368FC3E6C4E9924F8BC" authority="Kraepelin, 1911" authorityName="Kraepelin" authorityYear="1911" box="[1004,1312,1833,1858]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="koesteri">
<emphasis id="B90DEA8CE9112368FC3E6C4E9862F8BF" box="[1004,1126,1833,1858]" italics="true" pageId="2" pageNumber="375">C. koesteri</emphasis>
Kraepelin, 1911
</taxonomicName>
have eight primary rows of granules on the pedipalp finger and chela fingers sparsely hirsute. The pedipalp chela manus is lighter than fingers in 
<taxonomicName id="4C794D1DE9112368FE4E6C169E47F874" box="[412,579,1906,1929]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9112368FE4E6C169E47F874" box="[412,579,1906,1929]" italics="true" pageId="2" pageNumber="375">C. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9112368FD9F6C159E8BF877" box="[589,655,1905,1930]" pageId="2" pageNumber="375" rank="species">
<emphasis id="B90DEA8CE9112368FD9F6C159E8BF877" bold="true" box="[589,655,1905,1930]" pageId="2" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
and in 
<taxonomicName id="4C794D1DE9112368FD396C169F63F877" box="[747,871,1905,1930]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="koesteri">
<emphasis id="B90DEA8CE9112368FD396C169F63F877" box="[747,871,1905,1930]" italics="true" pageId="2" pageNumber="375">C. koesteri</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C794D1DE9112368FCAA6C15989DF874" box="[888,1177,1905,1930]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9112368FCAA6C15989DF874" box="[888,1177,1905,1930]" italics="true" pageId="2" pageNumber="375">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9112368FB716C1598E1F877" box="[1187,1253,1905,1930]" pageId="2" pageNumber="375" rank="species">
<emphasis id="B90DEA8CE9112368FB716C1598E1F877" bold="true" box="[1187,1253,1905,1930]" pageId="2" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="4C794D1DE9112368FA506C169CFAF850" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B90DEA8CE9112368FA506C169CFAF850" italics="true" pageId="2" pageNumber="375">C. granosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C794D1DE9112368FEE66CF29DA9F853" box="[308,429,1941,1966]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="koesteri">
<emphasis id="B90DEA8CE9112368FEE66CF29DA9F853" box="[308,429,1941,1966]" italics="true" pageId="2" pageNumber="375">C. koesteri</emphasis>
</taxonomicName>
in having smaller size (
<quantity id="4C819B7BE9112368FD6B6CF19F33F853" box="[697,823,1941,1966]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="6.5" metricValueMin="5.4" pageId="2" pageNumber="375" unit="mm" value="59.5" valueMax="65.0" valueMin="54.0">54–65 mm</quantity>
), 
<taxonomicName id="4C794D1DE9112368FC986CF29FD6F850" box="[842,978,1942,1965]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B90DEA8CE9112368FC986CF29FD6F850" box="[842,978,1942,1965]" italics="true" pageId="2" pageNumber="375">C. granosus</emphasis>
</taxonomicName>
(males 
<quantity id="4C819B7BE9112368FBF96CF198B7F850" box="[1067,1203,1941,1966]" metricMagnitude="-1" metricUnit="m" metricValue="1.025" metricValueMax="1.1" metricValueMin="0.95" pageId="2" pageNumber="375" unit="mm" value="102.5" valueMax="110.0" valueMin="95.0">95–110 mm</quantity>
, females 
<quantity id="4C819B7BE9112368FACE6CF19993F850" box="[1308,1431,1941,1966]" metricMagnitude="-2" metricUnit="m" metricValue="8.85" metricValueMax="9.5" metricValueMin="8.2" pageId="2" pageNumber="375" unit="mm" value="88.5" valueMax="95.0" valueMin="82.0">82–95 mm</quantity>
, unpublished measurements of 
<specimenCount id="9D7FFD17E9112368FDD66CDD9EA0F82F" box="[516,676,1977,2002]" pageId="2" pageNumber="375" type="generic">86 specimens</specimenCount>
deposited LAV-MIUP), 
<taxonomicName id="4C794D1DE9112368FC1D6CDE9855F82F" box="[975,1105,1977,2002]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="koesteri">
<emphasis id="B90DEA8CE9112368FC1D6CDE9855F82F" box="[975,1105,1977,2002]" italics="true" pageId="2" pageNumber="375">C. koesteri</emphasis>
</taxonomicName>
(
<quantity id="4C819B7BE9112368FBBA6CDD98EBF82C" box="[1128,1263,1977,2002]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="2" pageNumber="375" unit="mm" value="70.0" valueMax="75.0" valueMin="65.0">65–75 mm</quantity>
). 
<taxonomicName id="4C794D1DE9112368FADB6CDD9D1FF808" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9112368FADB6CDD9D1FF808" italics="true" pageId="2" pageNumber="375">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9112368FEF06CB99D64F80B" box="[290,352,2013,2038]" pageId="2" pageNumber="375" rank="species">
<emphasis id="B90DEA8CE9112368FEF06CB99D64F80B" bold="true" box="[290,352,2013,2038]" pageId="2" pageNumber="375">n. sp.</emphasis>
</taxonomicNameLabel>
has lower pectinal tooth counts (17–21), 
<taxonomicName id="4C794D1DE9112368FCFD6CBA9FB2F808" box="[815,950,2014,2037]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="375" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B90DEA8CE9112368FCFD6CBA9FB2F808" box="[815,950,2014,2037]" italics="true" pageId="2" pageNumber="375">C. granosus</emphasis>
</taxonomicName>
(males 25–29, females 24–29, unpublished counts LAV-MIUP), 
<taxonomicName id="4C794D1DE9102369FE536BFC9DFFFF4D" box="[385,507,151,176]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="376" phylum="Arthropoda" rank="species" species="koesteri">
<emphasis id="B90DEA8CE9102369FE536BFC9DFFFF4D" box="[385,507,151,176]" italics="true" pageId="3" pageNumber="376">C. koesteri</emphasis>
</taxonomicName>
(21–26), and a central discal depression on the pectinal plate of both sexes (absent in 
<taxonomicName id="4C794D1DE9102369FF616BD99D28FF28" box="[179,300,188,213]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="376" phylum="Arthropoda" rank="species" species="koesteri">
<emphasis id="B90DEA8CE9102369FF616BD99D28FF28" box="[179,300,188,213]" italics="true" pageId="3" pageNumber="376">C. koesteri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C794D1DE9102369FEB36BD99DECFF29" box="[353,488,189,212]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="376" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B90DEA8CE9102369FEB36BD99DECFF29" box="[353,488,189,212]" italics="true" pageId="3" pageNumber="376">C. granosus</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C794D1DE9102369FE2C6BD89F1FFF29" box="[510,795,188,213]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="376" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE9102369FE2C6BD89F1FFF29" box="[510,795,188,213]" italics="true" pageId="3" pageNumber="376">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E9102369FCF06BD89F64FF28" box="[802,864,188,213]" pageId="3" pageNumber="376" rank="species">
<emphasis id="B90DEA8CE9102369FCF06BD89F64FF28" bold="true" box="[802,864,188,213]" pageId="3" pageNumber="376">n. sp.</emphasis>
</taxonomicNameLabel>
additionally differs from 
<taxonomicName id="4C794D1DE9102369FBAC6BD99901FF29" box="[1150,1285,189,212]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="376" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B90DEA8CE9102369FBAC6BD99901FF29" box="[1150,1285,189,212]" italics="true" pageId="3" pageNumber="376">C. granosus</emphasis>
</taxonomicName>
: the pedipalp chela manus not incrassate, about same width as patella, and the pectinal plate has a discal central depression; 
<taxonomicName id="4C794D1DE9102369FA506B849CFAFEE1" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="376" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B90DEA8CE9102369FA506B849CFAFEE1" italics="true" pageId="3" pageNumber="376">C. granosus</emphasis>
</taxonomicName>
has pedipalp chela manus about 1.3 times wider than patella.
</paragraph>
</subSubSection>
<subSubSection id="C3636515E910236FFF156A439FC4FE75" lastPageId="5" lastPageNumber="378" pageId="3" pageNumber="376" type="description">
<paragraph id="8BC6369EE9102369FF156A43999CFEBD" blockId="3.[151,1437,151,2018]" box="[199,1432,295,320]" pageId="3" pageNumber="376">
<emphasis id="B90DEA8CE9102369FF156A439D53FEBD" bold="true" box="[199,343,295,320]" pageId="3" pageNumber="376">Description.</emphasis>
Based on male 
<typeStatus id="54C2883CE9102369FDD56A439E6CFEBD" box="[519,616,295,320]" pageId="3" pageNumber="376" type="holotype">holotype</typeStatus>
(with differences in 
<typeStatus id="54C2883CE9102369FC9C6A4C9FA9FEBD" box="[846,941,296,320]" pageId="3" pageNumber="376" type="paratype">paratype</typeStatus>
female noted) and illustrated in figures 1–5.
</paragraph>
<paragraph id="8BC6369EE9102369FF156A289E5BFDC0" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="376">
<emphasis id="B90DEA8CE9102369FF156A289D4EFE98" box="[199,330,332,357]" italics="true" pageId="3" pageNumber="376">Coloration:</emphasis>
Chelicerae yellow with reticulated infuscation, fingers brown with non-reticulated infuscation. Carapace yellow with heavy brown infuscation, posterior margin brown. Pedipalps yellow with infuscation on the distal femur and patella, pedipalp fingers brown. Mesosomal tergites light brown and heavily infuscate, posterior margin brown. Mesosomal sternites tan with brown infuscation, distally increasing such that sternite VII is nearly entirely infuscate. Legs yellow with weak infuscation. Metasoma tan with dark brown ventral infuscation, gradually darkening increasing distally such that metasoma V is brownish-tan and nearly entirely infuscate, carinae brown. Telson brown with a reddish tint.
</paragraph>
<paragraph id="8BC6369EE9102369FF15692C98EAFCC5" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="376">
<emphasis id="B90DEA8CE9102369FF15692C9D31FDA2" box="[199,309,584,607]" italics="true" pageId="3" pageNumber="376">Carapace</emphasis>
(
<figureCitation id="13422A1BE9102369FE9A69239DAAFD9D" box="[328,430,583,608]" captionStart="FIGURE 2" captionStartId="2.[151,250,1129,1151]" captionTargetBox="[151,1436,193,1107]" captionTargetId="figure@2.[151,1436,193,1108]" captionTargetPageId="2" captionText="FIGURE 2. Centruroides panamensis new species: A—male paratype dorsal carapace; B—female paratype dorsal carapace; C—female paratype pectines; D—male paratype pectines. Scale bar represents 1 mm." httpUri="https://zenodo.org/record/230956/files/figure.png" pageId="3" pageNumber="376">Figure 2</figureCitation>
): Carapace moderately and coarsely granular with granulation medially concentrated. Anterior and posterior margins procurved with median notch, weakly granular with small granules. Lateral margins slightly widened posteriorly. Lateral ocular tubercles with three macro-ocelli each. Two pairs of fine, acuminate median ocular macrosetae. Ocular tubercle with pair of costate-granular superciliary carinae, protruding above median ocelli. Lateral ocular carinae present, weakly granular. Posterior median carinae present and costategranular. Anteromedian sulcus moderately deep, ovate; posteromedian sulcus narrow and posteriorly deep; posterolateral sulci moderately deep and slightly curved; posteromarginal sulcus shallow and wide.
</paragraph>
<paragraph id="8BC6369EE9102369FF15682198B6FCA0" blockId="3.[151,1437,151,2018]" box="[199,1202,836,861]" pageId="3" pageNumber="376">
<emphasis id="B90DEA8CE9102369FF1568219D28FCA1" box="[199,300,837,860]" italics="true" pageId="3" pageNumber="376">Sternum:</emphasis>
Subtriangular, median sulcus Y-shaped, shallow anteriorly and deep posteriorly.
</paragraph>
<paragraph id="8BC6369EE9102369FF1568039DA6FC59" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="376">
<emphasis id="B90DEA8CE9102369FF1568039DA7FC7D" box="[199,419,871,896]" italics="true" pageId="3" pageNumber="376">Genital operculum:</emphasis>
Completely divided longitudinally with marginal macrosetae (
<figureCitation id="13422A1BE9102369FBBF680398D6FC7D" box="[1133,1234,871,896]" captionStart="FIGURE 2" captionStartId="2.[151,250,1129,1151]" captionTargetBox="[151,1436,193,1107]" captionTargetId="figure@2.[151,1436,193,1108]" captionTargetPageId="2" captionText="FIGURE 2. Centruroides panamensis new species: A—male paratype dorsal carapace; B—female paratype dorsal carapace; C—female paratype pectines; D—male paratype pectines. Scale bar represents 1 mm." httpUri="https://zenodo.org/record/230956/files/figure.png" pageId="3" pageNumber="376">Figure 2</figureCitation>
). Genital papillae present (♂), absent (♀).
</paragraph>
<paragraph id="8BC6369EE9102369FF1568D59992FC11" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="376">
<emphasis id="B90DEA8CE9102369FF1568D59D28FC35" box="[199,300,945,968]" italics="true" pageId="3" pageNumber="376">Pectines:</emphasis>
Pectines elongate, small transverse basal lobe present, fulcra prominent, (
<figureCitation id="13422A1BE9102369FB5768CB98EEFC35" box="[1157,1258,943,968]" captionStart="FIGURE 2" captionStartId="2.[151,250,1129,1151]" captionTargetBox="[151,1436,193,1107]" captionTargetId="figure@2.[151,1436,193,1108]" captionTargetPageId="2" captionText="FIGURE 2. Centruroides panamensis new species: A—male paratype dorsal carapace; B—female paratype dorsal carapace; C—female paratype pectines; D—male paratype pectines. Scale bar represents 1 mm." httpUri="https://zenodo.org/record/230956/files/figure.png" pageId="3" pageNumber="376">Figure 2</figureCitation>
). Pectinal plate rectangular with a small anteromedian notch and a central discal depression. Pectinal tooth counts, 19 (♂), 17–21 (♀).
</paragraph>
<paragraph id="8BC6369EE9102369FF1568939DD8FB5B" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="376">
<emphasis id="B90DEA8CE9102369FF1568939D33FBED" box="[199,311,1015,1040]" italics="true" pageId="3" pageNumber="376">Pedipalps</emphasis>
: Orthobothriotaxic, 
<typeStatus id="54C2883CE9102369FDF5689C9E5AFBED" box="[551,606,1016,1040]" pageId="3" pageNumber="376">Type</typeStatus>
A, α configuration (femoral trichobothria 
<emphasis id="B90DEA8CE9102369FB896893986DFBED" box="[1115,1129,1015,1040]" italics="true" pageId="3" pageNumber="376">d</emphasis>
1 and 
<emphasis id="B90DEA8CE9102369FB63689398BBFBED" box="[1201,1215,1015,1040]" italics="true" pageId="3" pageNumber="376">d</emphasis>
4 situated closer to dorsoexternal carina than 
<emphasis id="B90DEA8CE9102369FE686F7A9DCCFBCA" box="[442,456,1054,1079]" italics="true" pageId="3" pageNumber="376">d</emphasis>
3) (
<figureCitation id="13422A1BE9102369FE3A6F7A9E4DFBCA" box="[488,585,1054,1079]" captionStart="FIGURE 3" captionStartId="4.[151,250,1948,1970]" captionTargetBox="[161,1425,390,1925]" captionTargetId="figure@4.[160,1433,334,1982]" captionTargetPageId="4" captionText="FIGURE 3. Centruroides panamensis new species, pedipalps. Female Paratype: A—dorsal aspect of femur; B—dorsal aspect of patella; C—dorsoexternal aspect of chela. Male Paratype: D—dorsal aspect of femur; E—dorsal aspect of patella; F—dorsoexternal aspect of chela. Scale bar represents 1 mm." httpUri="https://zenodo.org/record/230957/files/figure.png" pageId="3" pageNumber="376">Figure 3</figureCitation>
D). Femur with five carinae: externomedian carina continuous, costate with small and irregular serrate granules; dorsoexternal, dorsointernal, and ventrointernal carinae continuous with spiniform granules; internomedian carina discontinuous, comprised of isolated large spiniform granules. Intercarinal surfaces smooth.
</paragraph>
<paragraph id="8BC6369EE9102369FF156FD59DD8FACB" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="376">
Patella with seven carinae: dorsointernal, dorsomedian, dorsoexternal externomedian, ventroexternal carina continuous, costate-granular; ventrointernal and internomedian carina discontinuous, comprised of irregularly spaced spiniform granules proximally, becoming smaller distally; proximal tubercle weakly developed (
<figureCitation id="13422A1BE9102369FAC96F9D997FFAEF" box="[1307,1403,1273,1298]" captionStart="FIGURE 3" captionStartId="4.[151,250,1948,1970]" captionTargetBox="[161,1425,390,1925]" captionTargetId="figure@4.[160,1433,334,1982]" captionTargetPageId="4" captionText="FIGURE 3. Centruroides panamensis new species, pedipalps. Female Paratype: A—dorsal aspect of femur; B—dorsal aspect of patella; C—dorsoexternal aspect of chela. Male Paratype: D—dorsal aspect of femur; E—dorsal aspect of patella; F—dorsoexternal aspect of chela. Scale bar represents 1 mm." httpUri="https://zenodo.org/record/230957/files/figure.png" pageId="3" pageNumber="376">Figure 3</figureCitation>
E). Intercarinal surfaces smooth.
</paragraph>
<paragraph id="8BC6369EE9102369FF156E259950F987" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="376">
Chela manus not incrassate (
<figureCitation id="13422A1BE9102369FDCA6E259E7FFAA7" box="[536,635,1345,1370]" captionStart="FIGURE 3" captionStartId="4.[151,250,1948,1970]" captionTargetBox="[161,1425,390,1925]" captionTargetId="figure@4.[160,1433,334,1982]" captionTargetPageId="4" captionText="FIGURE 3. Centruroides panamensis new species, pedipalps. Female Paratype: A—dorsal aspect of femur; B—dorsal aspect of patella; C—dorsoexternal aspect of chela. Male Paratype: D—dorsal aspect of femur; E—dorsal aspect of patella; F—dorsoexternal aspect of chela. Scale bar represents 1 mm." httpUri="https://zenodo.org/record/230957/files/figure.png" pageId="3" pageNumber="376">Figure 3</figureCitation>
F) about same width as the patella. Chela with five distinct carinae: dorsomedian, dorsal secondary, and ventroexternal carinae continuous smooth keel (costate-granular in ♀); digital carina nearly obsolete, consisting of a few small proximal granules; dorsointernal carina nearly obsolete, consisting of a wide, smooth keel (scattered spiniform granules in ♀); other carinae absent. Intercarinal surfaces smooth. Movable finger with small lobe (eminence) proximally; movable finger length 32–36% (♂) or 61–77% (♀) greater than length along ventroexternal carina (
<tableCitation id="C6FB0325E9102369FD896E919EABF9F3" box="[603,687,1525,1550]" captionStart="TABLE 1" captionStartId="7.[151,239,512,535]" captionTargetBox="[151,1414,563,1775]" captionTargetPageId="7" captionText="TABLE 1. Measurements of Centruroides panamensis new species, in mm." httpUri="http://table.plazi.org/id/DF066616E914236DFF4569649FC4FDEA" pageId="3" pageNumber="376" tableUuid="DF066616E914236DFF4569649FC4FDEA">Table 1</tableCitation>
); dentate margins of fixed fingers with eight oblique primary rows of denticles (basal rows fused in moveable finger). In addition to the eight main rows, the movable finger has a short apical row of three-four denticles. Each denticular row terminates in a large denticle at proximal and distal ends; internal and external supernumerary denticles present; fingers each with an enlarged terminal denticle.
</paragraph>
<paragraph id="8BC6369EE9102369FF156DE29E33F91B" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="376">
<emphasis id="B90DEA8CE9102369FF156DE29D02F960" box="[199,262,1670,1693]" italics="true" pageId="3" pageNumber="376">Legs:</emphasis>
Basitarsi with paired rows of fine, acuminate macrosetae on pro- and retrolateral surfaces. Telotarsi with dense brush of acuminate macrosetae; laterodistal lobes truncated; median dorsal lobes extending to moderately long, narrow, slightly curved ungues.
</paragraph>
<paragraph id="8BC6369EE9102369FF156D969DD0F867" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="376">
<taxonomicName id="4C794D1DE9102369FF156D969DB3F8F4" authority="Tergites I" authorityName="Tergites I" box="[199,439,1777,1802]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA8CE9102369FF156D969D4CF8F4" box="[199,328,1778,1801]" italics="true" pageId="3" pageNumber="376">Mesosoma:</emphasis>
Tergites I
</taxonomicName>
–VI pretergites smooth, post-tergites coarsely granular, each with a granular dorsomedian carina. Tergite VII with costate-granular dorsomedian, dorsosubmedian, and dorsolateral carinae; intercarinal spaces finely granular. Sternites III–VI smooth, acarinate, each with pair of narrow, slit-like respiratory spiracles; V with prominent pale surface posteromedially in adult ♂. Sternite VII with pair of ventrosubmedian and ventrolateral keeled carinae.
</paragraph>
<paragraph id="8BC6369EE910236EFF156CC39979FEBD" blockId="3.[151,1437,151,2018]" lastBlockId="4.[151,1437,151,320]" lastPageId="4" lastPageNumber="377" pageId="3" pageNumber="376">
<emphasis id="B90DEA8CE9102369FF156CC39D41F843" box="[199,325,1959,1982]" italics="true" pageId="3" pageNumber="376">Metasoma:</emphasis>
Segments I–V progressively increasing in length, with ♂ having a more elongate metasoma (metasoma V length/width 2.8–3.0) than ♀ (metasoma V length/width 2.2–2.3). Intercarinal surfaces smooth (minutely granular in ♀). Segments I–IV, paired dorsosubmedian, dorsolateral, ventrolateral, ventrosubmedian, and median lateral carinae continuous, serrate to costate-granular. Segment V, dorsosubmedian carinae absent; dorsolateral carinae obsolete, granular, reduced to anterior third of segment; ventrolateral carinae continuous, weakly granular; ventrosubmedian carinae obsolete, reduced to several anterior granules; ventromedian carina continuous, weakly granular (
<figureCitation id="13422A1BE917236EFE316A439E42FEBD" box="[483,582,295,320]" captionStart="FIGURE 4" captionStartId="5.[151,250,917,939]" captionTargetBox="[151,1436,445,898]" captionTargetId="figure@5.[151,1436,445,899]" captionTargetPageId="5" captionText="FIGURE 4. Centruroides panamensis new species, metasoma. From top to bottom: lateral aspect of paratype male, ventral aspect of paratype male, lateral aspect of paratype female. Scale bar represents 2 mm." httpUri="https://zenodo.org/record/230959/files/figure.png" pageId="4" pageNumber="377">Figure 4</figureCitation>
). Number of ventrolateral macrosetae on metasomal segments 2 to 4 is 2.
</paragraph>
<caption id="DF066616E917236EFF456CF89EF9F812" httpUri="https://zenodo.org/record/230957/files/figure.png" pageId="4" pageNumber="377" targetBox="[161,1425,390,1925]" targetPageId="4">
<paragraph id="8BC6369EE917236EFF456CF89EF9F812" blockId="4.[151,1436,1948,2031]" pageId="4" pageNumber="377">
<emphasis id="B90DEA8CE917236EFF456CF89D17F84C" bold="true" box="[151,275,1948,1970]" pageId="4" pageNumber="377">FIGURE 3.</emphasis>
<taxonomicName id="4C794D1DE917236EFEC86CF89E19F84F" box="[282,541,1948,1970]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="377" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE917236EFEC86CF89E19F84F" box="[282,541,1948,1970]" italics="true" pageId="4" pageNumber="377">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E917236EFDF66CF99EA5F84F" box="[548,673,1948,1970]" pageId="4" pageNumber="377" rank="species">new species</taxonomicNameLabel>
, pedipalps. Female Paratype: A—dorsal aspect of femur; B—dorsal aspect of patella; C—dorsoexternal aspect of chela. Male Paratype: D—dorsal aspect of femur; E—dorsal aspect of patella; F—dorsoexternal aspect of chela. Scale bar represents 1mm.
</paragraph>
</caption>
<paragraph id="8BC6369EE916236FFF156BF39FD7FF05" blockId="5.[151,1436,151,392]" pageId="5" pageNumber="378">
<emphasis id="B90DEA8CE916236FFF156BF39D12FF4D" box="[199,278,151,176]" italics="true" pageId="5" pageNumber="378">Telson:</emphasis>
Slightly elongate, non-lobate, length: width ratio of 1.7–1.8, smooth (minutely granular in ♀), carinae absent (obsolete ventromedian carinae in ♀) (
<figureCitation id="13422A1BE916236FFD6B6BD89F25FF29" box="[697,801,188,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,917,939]" captionTargetBox="[151,1436,445,898]" captionTargetId="figure@5.[151,1436,445,899]" captionTargetPageId="5" captionText="FIGURE 4. Centruroides panamensis new species, metasoma. From top to bottom: lateral aspect of paratype male, ventral aspect of paratype male, lateral aspect of paratype female. Scale bar represents 2 mm." httpUri="https://zenodo.org/record/230959/files/figure.png" pageId="5" pageNumber="378">Figure 4</figureCitation>
). Subaculear tubercle well-developed, spinate, the tip oriented towards the tip of the aculeus. Aculeus strongly curved ventrally.
</paragraph>
<paragraph id="8BC6369EE916236FFF156A619FC4FE75" blockId="5.[151,1436,151,392]" pageId="5" pageNumber="378">
<emphasis id="B90DEA8CE916236FFF156A619E1AFEE0" box="[199,542,260,285]" italics="true" pageId="5" pageNumber="378">Variation. Sexual Dimorphism</emphasis>
: In addition to previously discussed characters, adult males are proportionally longer than adult females; metasomal segments sum to 61–62% of the total length of males, but to 51–58% of the total length of females. Adult males are slightly more slender than adult females: sternite VII width is 12–13% greater than its width in males and 41–51% greater in females (
<tableCitation id="C6FB0325E916236FFC896A0B9FAAFE75" box="[859,942,367,392]" captionStart="TABLE 1" captionStartId="7.[151,239,512,535]" captionTargetBox="[151,1414,563,1775]" captionTargetPageId="7" captionText="TABLE 1. Measurements of Centruroides panamensis new species, in mm." httpUri="http://table.plazi.org/id/DF066616E914236DFF4569649FC4FDEA" pageId="5" pageNumber="378" tableUuid="DF066616E914236DFF4569649FC4FDEA">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="DF066616E916236FFF4568F19FF0FC34" httpUri="https://zenodo.org/record/230959/files/figure.png" pageId="5" pageNumber="378" targetBox="[151,1436,445,898]" targetPageId="5">
<paragraph id="8BC6369EE916236FFF4568F19FF0FC34" blockId="5.[151,1435,916,969]" pageId="5" pageNumber="378">
<emphasis id="B90DEA8CE916236FFF4568F19D11FC57" bold="true" box="[151,277,917,939]" pageId="5" pageNumber="378">FIGURE 4.</emphasis>
<taxonomicName id="4C794D1DE916236FFECF68F09E27FC56" box="[285,547,916,939]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="378" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE916236FFECF68F09E27FC56" box="[285,547,916,939]" italics="true" pageId="5" pageNumber="378">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E916236FFDF968F19EAEFC57" box="[555,682,916,938]" pageId="5" pageNumber="378" rank="species">new species</taxonomicNameLabel>
, metasoma. From top to bottom: lateral aspect of paratype male, ventral aspect of paratype male, lateral aspect of paratype female. Scale bar represents 2mm.
</paragraph>
</caption>
<caption id="DF066616E916236FFF456C509992F87A" httpUri="https://zenodo.org/record/230960/files/figure.png" pageId="5" pageNumber="378" targetBox="[171,1401,1015,1824]" targetPageId="5">
<paragraph id="8BC6369EE916236FFF456C509992F87A" blockId="5.[151,1436,1844,1927]" pageId="5" pageNumber="378">
<emphasis id="B90DEA8CE916236FFF456C509D13F8B4" bold="true" box="[151,279,1844,1866]" pageId="5" pageNumber="378">FIGURE 5.</emphasis>
Panama, Chiriquí Province. Distribution of 
<taxonomicName id="4C794D1DE916236FFD266C509FFFF8B7" box="[756,1019,1844,1866]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="378" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE916236FFD266C509FFFF8B7" box="[756,1019,1844,1866]" italics="true" pageId="5" pageNumber="378">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E916236FFBD46C519883F8B7" box="[1030,1159,1844,1866]" pageId="5" pageNumber="378" rank="species">new species</taxonomicNameLabel>
. Grey circle (collected 
<taxonomicName id="4C794D1DE916236FFA566C519D0BF894" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="378" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE916236FFA566C519D0BF894" italics="true" pageId="5" pageNumber="378">C. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E916236FFEC66C379D49F895" box="[276,333,1875,1896]" pageId="5" pageNumber="378" rank="species">
<emphasis id="B90DEA8CE916236FFEC66C379D49F895" bold="true" box="[276,333,1875,1896]" pageId="5" pageNumber="378">sp. n.</emphasis>
</taxonomicNameLabel>
): dirt road to town Aguacate on SW slopes of Volcán Barú. Black circle (collected both 
<taxonomicName id="4C794D1DE916236FFB1A6C379959F894" box="[1224,1373,1875,1897]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="378" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE916236FFB1A6C379959F894" box="[1224,1373,1875,1897]" italics="true" pageId="5" pageNumber="378">C. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E916236FFAB16C379998F895" box="[1379,1436,1875,1896]" pageId="5" pageNumber="378" rank="species">
<emphasis id="B90DEA8CE916236FFAB16C379998F895" bold="true" box="[1379,1436,1875,1896]" pageId="5" pageNumber="378">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C794D1DE916236FFF136C169D2CF87A" box="[193,296,1905,1927]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="378" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE916236FFF136C169D2CF87A" box="[193,296,1905,1927]" italics="true" pageId="5" pageNumber="378">C. bicolor</emphasis>
</taxonomicName>
): Finca Lukarel, Volcancito (Finca Lukarel, is located between towns of Volcancito Arriba and Bajo Volcancito.
</paragraph>
</caption>
<subSubSection id="C3636515E9162362FF156CCA9DE3FF28" lastPageId="8" lastPageNumber="381" pageId="5" pageNumber="378" type="distribution">
<paragraph id="8BC6369EE916236CFF156CCA982AFDBD" blockId="5.[151,1436,1966,2025]" lastBlockId="6.[151,1437,151,576]" lastPageId="6" lastPageNumber="379" pageId="5" pageNumber="378">
<emphasis id="B90DEA8CE916236FFF156CCA9D64F83A" bold="true" box="[199,352,1966,1991]" pageId="5" pageNumber="378">Distribution.</emphasis>
The new species is currently known only from the Cordillera Central of Chiriquí province, 
<collectingCountry id="F36E760EE916236FFF456CB59CF7F815" box="[151,243,2001,2024]" name="Panama" pageId="5" pageNumber="378">Panama</collectingCountry>
, an uninterrupted mountainous chain above sea level until at least Miocene times (
<bibRefCitation id="EFE84B6FE916236FFB646CB49990F815" author="Montes" box="[1206,1428,2000,2025]" pageId="5" pageNumber="378" refString="Montes, C., Cardona, A., McFadden, R., Moron, S. E., Silva, C. A., Restrepo-Moreno, S., Ramirez, D. A., Hoyos, N., Wilson, J., Farris, D., Bayona, G. A., Jaramillo, C. A., Valencia, V., Bryan, J. &amp; Flores, J. A. (2012) Evidence for middle Eocene and younger land emergences in central Panama: Implications for Isthmus closure. Geological Society of America Bulletin, 124 (5 - 6), 780 - 799. http: // dx. doi. org / 10.1130 / b 30528.1" type="journal article" year="2012">
Montes 
<emphasis id="B90DEA8CE916236FFAC76CB59955F814" box="[1301,1361,2000,2025]" italics="true" pageId="5" pageNumber="378">et al.</emphasis>
2012
</bibRefCitation>
) (
<figureCitation id="13422A1BE915236CFF4C6BF39D06FF4D" box="[158,258,151,176]" captionStart="FIGURE 5" captionStartId="5.[151,250,1844,1866]" captionTargetBox="[171,1401,1015,1824]" captionTargetId="figure@5.[171,1415,1015,1828]" captionTargetPageId="5" captionText="FIGURE 5. Panama, Chiriquí Province. Distribution of Centruroides panamensis new species. Grey circle (collected C. panamensis sp. n.): dirt road to town Aguacate on SW slopes of Volcán Barú. Black circle (collected both C. panamensis sp. n. and C. bicolor): Finca Lukarel, Volcancito (Finca Lukarel, is located between towns of Volcancito Arriba and Bajo Volcancito." httpUri="https://zenodo.org/record/230960/files/figure.png" pageId="6" pageNumber="379">Figure 5</figureCitation>
). The evolutionary origin of 
<taxonomicName id="4C794D1DE915236CFD9E6BFC9EF6FF4D" box="[588,754,152,176]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B90DEA8CE915236CFD9E6BFC9EF6FF4D" box="[588,754,152,176]" italics="true" pageId="6" pageNumber="379">C. panamensis</emphasis>
</taxonomicName>
has been shaped by climatic and montane forest changes in the Cordillera de Talamanca of 
<collectingCountry id="F36E760EE915236CFE286BDD9E70FF2F" box="[506,628,185,210]" name="Costa Rica" pageId="6" pageNumber="379">Costa Rica</collectingCountry>
and 
<collectingCountry id="F36E760EE915236CFD7E6BDE9F01FF2C" box="[684,773,186,209]" name="Panama" pageId="6" pageNumber="379">Panama</collectingCountry>
brought by the last ice age that began in late Pliocene, and by past Pleistocene eruptive episodes of Volcán Barú (see 
<bibRefCitation id="EFE84B6FE915236CFCFD6BBE980BFF0E" author="Sherrod" box="[815,1039,218,243]" pageId="6" pageNumber="379" refString="Sherrod, D. R., Vallance, J. W., Tapia Espinosa, A. &amp; McGeehin, J. P. (2008) Volcan Baru: Eruptive History and Volcano- Hazards Assessment. U. S. Geological Survey, Open-File Report 2007 - 1401, 33 pp." type="book" year="2008">
Sherrod 
<emphasis id="B90DEA8CE915236CFC426BB89FCDFF0E" box="[912,969,218,243]" italics="true" pageId="6" pageNumber="379">et al.</emphasis>
2008
</bibRefCitation>
). 
<taxonomicName id="4C794D1DE915236CFBF16BBE9946FF0E" box="[1059,1346,218,243]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B90DEA8CE915236CFBF16BBE9946FF0E" box="[1059,1346,218,243]" italics="true" pageId="6" pageNumber="379">Centruroides panamensis</emphasis>
</taxonomicName>
has not shown signs of recent demographic expansion, a limitation that we interpret might have been caused by a more widespread and successful, highly polymorphic species, 
<taxonomicName id="4C794D1DE915236CFCF36A7998D5FECB" authority="Pocock 1898" authorityName="Pocock" authorityYear="1898" box="[801,1233,285,310]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B90DEA8CE915236CFCF36A79981AFECB" box="[801,1054,285,310]" italics="true" pageId="6" pageNumber="379">Centruroides limbatus</emphasis>
(Pocock 1898)
</taxonomicName>
, that ranges from 
<collectingCountry id="F36E760EE915236CFF456A5A9D0EFEAA" box="[151,266,318,343]" name="Nicaragua" pageId="6" pageNumber="379">Nicaragua</collectingCountry>
to 
<collectingCountry id="F36E760EE915236CFEEA6A5B9DBCFEAA" box="[312,440,318,343]" name="Costa Rica" pageId="6" pageNumber="379">Costa Rica</collectingCountry>
(
<bibRefCitation id="EFE84B6FE915236CFE1E6A5A9F09FEAA" author="Francke" box="[460,781,318,343]" pageId="6" pageNumber="379" refString="Francke, O. F. &amp; Stockwell, S. A. (1987) Scorpions (Arachnida) from Costa Rica. Special Publication Texas Tech University Press, 25, 1 - 64." type="journal article" year="1987">Francke &amp; Stockwell 1987</bibRefCitation>
, 
<bibRefCitation id="EFE84B6FE915236CFCCF6A5B9836FEAA" author="Armas" box="[797,1074,318,343]" pageId="6" pageNumber="379" refString="Armas, L. F. de. &amp; Trujillo, R. E. (2010) Nueva especie de Centruroides Marx, 1890 (Scorpiones: Buthidae) de Guatemala y Honduras, America Central. Boletin de la Sociedad Entomologica Aragonesa, 47, 235 - 240." type="journal article" year="2010">Armas &amp; Trujillo 2010</bibRefCitation>
) and 
<collectingCountry id="F36E760EE915236CFBA86A5B98D6FEAB" box="[1146,1234,319,342]" name="Panama" pageId="6" pageNumber="379">Panama</collectingCountry>
(Bocas del Toro Archipelago, 
<bibRefCitation id="EFE84B6FE915236CFEE26A049E4FFE85" author="Montoya" box="[304,587,352,376]" pageId="6" pageNumber="379" refString="Montoya, M. &amp; Armas, L. F. de (2002) Escorpiones (Arachnida) del Archipielago de Bocas del Toro, Panama. Revista de Biologia Tropical, 50 (1), 155 - 160." type="journal article" year="2002">Montoya &amp; Armas 2002</bibRefCitation>
). We have numerous new additional inland collections of 
<taxonomicName id="4C794D1DE915236CFB096A049959FE85" box="[1243,1373,351,376]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B90DEA8CE915236CFB096A049959FE85" box="[1243,1373,351,376]" italics="true" pageId="6" pageNumber="379">C. limbatus</emphasis>
</taxonomicName>
from Bocas del Toro (LAV-MIUP) and our data from Chiriquí Province represents a new distribution record for a second species: 
<taxonomicName id="4C794D1DE915236CFF246AC69E96FE46" authority="Pocock, 1898" authorityName="Pocock" authorityYear="1898" box="[246,658,418,443]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE915236CFF246AC69DE5FE46" box="[246,481,418,443]" italics="true" pageId="6" pageNumber="379">Centruroides bicolor</emphasis>
(Pocock, 1898)
</taxonomicName>
(collecting data listed apart, Appendix 1). We have found individuals of the dark morph of 
<taxonomicName id="4C794D1DE915236CFE4D6AA19E12FE20" box="[415,534,452,477]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE915236CFE4D6AA19E12FE20" box="[415,534,452,477]" italics="true" pageId="6" pageNumber="379">C. bicolor</emphasis>
</taxonomicName>
present in Volcancito and Boquete, Chiriquí Province, in close vicinity and sympatric with 
<taxonomicName id="4C794D1DE915236CFE9D6A829DFCFE00" box="[335,504,486,509]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B90DEA8CE915236CFE9D6A829DFCFE00" box="[335,504,486,509]" italics="true" pageId="6" pageNumber="379">C. panamensis</emphasis>
</taxonomicName>
in Finca Lukarel, Volcancito. 
<taxonomicName id="4C794D1DE915236CFCB66A829FDFFE03" box="[868,987,485,510]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B90DEA8CE915236CFCB66A829FDFFE03" box="[868,987,485,510]" italics="true" pageId="6" pageNumber="379">C. bicolor</emphasis>
</taxonomicName>
is the only species found in sporadic collections of scorpions carried out by Arsenio Araúz from the years 2008 until 
<quantity id="4C819B7BE915236CFBC569639868FDE2" box="[1047,1132,518,543]" metricMagnitude="1" metricUnit="m" metricValue="5.11048" pageId="6" pageNumber="379" unit="in" value="2012.0">2012 in</quantity>
the fields surroundings the town of Balita de Dolega, Dolega District, southeast from Volcán Barú (
<figureCitation id="13422A1BE915236CFC6C69439824FDBD" box="[958,1056,551,576]" captionStart="FIGURE 5" captionStartId="5.[151,250,1844,1866]" captionTargetBox="[171,1401,1015,1824]" captionTargetId="figure@5.[171,1415,1015,1828]" captionTargetPageId="5" captionText="FIGURE 5. Panama, Chiriquí Province. Distribution of Centruroides panamensis new species. Grey circle (collected C. panamensis sp. n.): dirt road to town Aguacate on SW slopes of Volcán Barú. Black circle (collected both C. panamensis sp. n. and C. bicolor): Finca Lukarel, Volcancito (Finca Lukarel, is located between towns of Volcancito Arriba and Bajo Volcancito." httpUri="https://zenodo.org/record/230960/files/figure.png" pageId="6" pageNumber="379">Figure 5</figureCitation>
).
</paragraph>
<caption id="DF066616E915236CFF456C199FEFF812" httpUri="https://zenodo.org/record/230961/files/figure.png" pageId="6" pageNumber="379" targetBox="[151,1436,627,1895]" targetPageId="6">
<paragraph id="8BC6369EE915236CFF456C199FEFF812" blockId="6.[151,1437,1917,2031]" pageId="6" pageNumber="379">
<emphasis id="B90DEA8CE915236CFF456C199D17F86F" bold="true" box="[151,275,1917,1939]" pageId="6" pageNumber="379">FIGURE 6.</emphasis>
Habitat of 
<taxonomicName id="4C794D1DE915236CFE5A6C1A9E1AF86E" box="[392,542,1918,1939]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="379" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE915236CFE5A6C1A9E1AF86E" box="[392,542,1918,1939]" italics="true" pageId="6" pageNumber="379">C. panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E915236CFDF76C1A9EA7F86E" box="[549,675,1917,1939]" pageId="6" pageNumber="379" rank="species">new species</taxonomicNameLabel>
: A—Collection site of male holotype and 7 paratypes; the southwest flank of Volcán Barú looms in the background. B—Dirt road between towns of Volcancito Arriba and Bajo Volcancito; Coffee plantation across the road with row of 
<taxonomicName id="4C794D1DE915236CFDE16CDF9EC7F82D" box="[563,707,1978,2000]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="379" phylum="Tracheophyta" rank="species" species="caribae">
<emphasis id="B90DEA8CE915236CFDE16CDF9EC7F82D" box="[563,707,1978,2000]" italics="true" pageId="6" pageNumber="379">Pinus caribae</emphasis>
</taxonomicName>
trees. C—Under these dried logs one subadult female was collected. D—Bank road where one female carrying 20 immatures was collected (under rock).
</paragraph>
</caption>
<paragraph id="8BC6369EE914236DFF156BF39E03FE2D" blockId="7.[151,1436,151,464]" pageId="7" pageNumber="380">
<emphasis id="B90DEA8CE914236DFF156BF39D2FFF4D" bold="true" box="[199,299,151,176]" pageId="7" pageNumber="380">Ecology.</emphasis>
The 
<typeStatus id="54C2883CE914236DFEB46BF39DCCFF4D" box="[358,456,151,176]" pageId="7" pageNumber="380" type="holotype">holotype</typeStatus>
male and 7 
<typeStatus id="54C2883CE914236DFD816BFC9EBAFF4D" box="[595,702,152,176]" pageId="7" pageNumber="380" type="paratype">paratypes</typeStatus>
of the new species were found in the foothills of Volcán Barú, at 
<quantity id="4C819B7BE914236DFF456BD89D31FF29" box="[151,309,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.675" metricValueMax="1.7" metricValueMin="1.65" pageId="7" pageNumber="380" unit="m" value="1675.0" valueMax="1700.0" valueMin="1650.0">1650–1700 m</quantity>
, in an open, treeless habitat (
<figureCitation id="13422A1BE914236DFD516BD89EE2FF28" box="[643,742,188,213]" captionStart="FIGURE 6" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[151,1436,627,1895]" captionTargetId="figure@6.[151,1436,626,1895]" captionTargetPageId="6" captionText="FIGURE 6. Habitat of C. panamensis new species: A—Collection site of male holotype and 7 paratypes; the southwest flank of Volcán Barú looms in the background. B—Dirt road between towns of Volcancito Arriba and Bajo Volcancito; Coffee plantation across the road with row of Pinus caribae trees. C—Under these dried logs one subadult female was collected. D—Bank road where one female carrying 20 immatures was collected (under rock)." httpUri="https://zenodo.org/record/230961/files/figure.png" pageId="7" pageNumber="380">Figure 6</figureCitation>
), under rocks, and inside shallow earth ground fissures, near the border of dirt paths. Two 
<typeStatus id="54C2883CE914236DFDD66B849E60FF05" box="[516,612,224,248]" pageId="7" pageNumber="380" type="paratype">paratype</typeStatus>
females carrying immatures (20 collected, 2–3 escaped; 22 younger immatures) were found under stones (one was close to the dirt road margin, the other, about 
<quantity id="4C819B7BE914236DFB136A6098EAFEE1" box="[1217,1262,260,284]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="380" unit="m" value="2.0">2 m</quantity>
away from the road), at 
<quantity id="4C819B7BE914236DFED26A439D9FFEC2" box="[256,411,295,320]" metricMagnitude="2" metricUnit="m" metricValue="3.7" metricValueMax="3.9" metricValueMin="3.5" pageId="7" pageNumber="380" unit="m" value="370.0" valueMax="390.0" valueMin="350.0">350 to 390 m</quantity>
from the entrance to Finca Lukarel, Boquete, at an elevation of 
<quantity id="4C819B7BE914236DFB546A4398DAFEC2" box="[1158,1246,295,320]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="7" pageNumber="380" unit="m" value="1160.0">1160 m</quantity>
, on 
<date id="FFC7105EE914236DFAC06A4C9CD7FE99" pageId="7" pageNumber="380" value="2012-02-22">22 February 2012</date>
. On the same date, one subadult female was found under a dried log, 
<quantity id="4C819B7BE914236DFBC36A28985FFE99" box="[1041,1115,332,356]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="380" unit="m" value="400.0">400 m</quantity>
from the entrance to Finca Lukarel. This area has numerous coffee plantations and rows of 
<taxonomicName id="4C794D1DE914236DFCB76A159814FE75" box="[869,1040,367,392]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="380" phylum="Tracheophyta" rank="species" species="caribaea">
<emphasis id="B90DEA8CE914236DFCB76A159814FE75" box="[869,1040,367,392]" italics="true" pageId="7" pageNumber="380">Pinus caribaea</emphasis>
</taxonomicName>
trees are planted on one side of the dirt road. The ground was covered with dried grasses, pine needles, and under the stones diplopods, centipedes, roaches and crickets were found.
</paragraph>
<caption id="DF066616E914236DFF4569649FC4FDEA" ID-Table-UUID="DF066616E914236DFF4569649FC4FDEA" box="[151,960,511,535]" httpUri="http://table.plazi.org/id/DF066616E914236DFF4569649FC4FDEA" pageId="7" pageNumber="380" targetBox="[151,1414,563,1775]" targetIsTable="true" targetPageId="7">
<paragraph id="8BC6369EE914236DFF4569649FC4FDEA" blockId="7.[151,960,511,535]" box="[151,960,511,535]" pageId="7" pageNumber="380">
<emphasis id="B90DEA8CE914236DFF4569649D0EFDEA" bold="true" box="[151,266,512,535]" pageId="7" pageNumber="380">TABLE 1.</emphasis>
Measurements of 
<taxonomicName id="4C794D1DE914236DFE1D6A9B9EE4FDEA" box="[463,736,511,535]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="380" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B90DEA8CE914236DFE1D6A9B9EE4FDEA" box="[463,736,511,535]" italics="true" pageId="7" pageNumber="380">Centruroides panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F7E914236DFD3469649F6DFDEA" box="[742,873,511,535]" pageId="7" pageNumber="380" rank="species">new species</taxonomicNameLabel>
, in mm.
</paragraph>
</caption>
<paragraph id="8BC6369EE914236DFE0669579D1FF912" pageId="7" pageNumber="380">
<table id="F979C43EE914DC95FF4569579982F912" box="[151,1414,563,1775]" gridcols="9" gridrows="25" pageId="7" pageNumber="380">
<tr id="354934DCE914DC95FF4569579982FD9A" box="[151,1414,563,615]" gridrow="0" pageId="7" pageNumber="380" rowspan-0="1">
<th id="76985DA0E914DC95FE0669579E36FD9A" box="[468,562,563,615]" gridcol="1" gridrow="0" pageId="7" pageNumber="380">♂ Holotype</th>
<th id="76985DA0E914DC95FD9C69579EA3FD9A" box="[590,679,563,615]" gridcol="2" gridrow="0" pageId="7" pageNumber="380">♀ Paratype</th>
<th id="76985DA0E914DC95FD1569579F1BFD9A" box="[711,799,563,615]" gridcol="3" gridrow="0" pageId="7" pageNumber="380">♂ Paratype</th>
<th id="76985DA0E914DC95FC9469579FADFD9A" box="[838,937,563,615]" gridcol="4" gridrow="0" pageId="7" pageNumber="380">♀ Paratype</th>
<th id="76985DA0E914DC95FC126957981DFD9A" box="[960,1049,563,615]" gridcol="5" gridrow="0" pageId="7" pageNumber="380">♀ Paratype</th>
<th id="76985DA0E914DC95FBEB69579897FD9A" box="[1081,1171,563,615]" gridcol="6" gridrow="0" pageId="7" pageNumber="380">♂ Paratype</th>
<th id="76985DA0E914DC95FB6169579908FD9A" box="[1203,1292,563,615]" gridcol="7" gridrow="0" pageId="7" pageNumber="380">♀ Paratype</th>
<th id="76985DA0E914DC95FAFF69579982FD9A" box="[1325,1414,563,615]" gridcol="8" gridrow="0" pageId="7" pageNumber="380">♀ Paratype</th>
</tr>
<tr id="354934DCE914DC95FF45691E9982FD6D" box="[151,1414,634,656]" gridrow="1" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF45691E9DBFFD6D" box="[151,443,634,656]" gridcol="0" gridrow="1" pageId="7" pageNumber="380">Carapace length</th>
<td id="76985DA0E914DC95FE06691E9E36FD6D" box="[468,562,634,656]" gridcol="1" gridrow="1" pageId="7" pageNumber="380">5.3</td>
<td id="76985DA0E914DC95FD9C691E9EA3FD6D" box="[590,679,634,656]" gridcol="2" gridrow="1" pageId="7" pageNumber="380">5.8</td>
<td id="76985DA0E914DC95FD15691E9F1BFD6D" box="[711,799,634,656]" gridcol="3" gridrow="1" pageId="7" pageNumber="380">5.3</td>
<td id="76985DA0E914DC95FC94691E9FADFD6D" box="[838,937,634,656]" gridcol="4" gridrow="1" pageId="7" pageNumber="380">6.1</td>
<td id="76985DA0E914DC95FC12691E981DFD6D" box="[960,1049,634,656]" gridcol="5" gridrow="1" pageId="7" pageNumber="380">5.2</td>
<td id="76985DA0E914DC95FBEB691E9897FD6D" box="[1081,1171,634,656]" gridcol="6" gridrow="1" pageId="7" pageNumber="380">5.5</td>
<td id="76985DA0E914DC95FB61691E9908FD6D" box="[1203,1292,634,656]" gridcol="7" gridrow="1" pageId="7" pageNumber="380">6.2</td>
<td id="76985DA0E914DC95FAFF691E9982FD6D" box="[1325,1414,634,656]" gridcol="8" gridrow="1" pageId="7" pageNumber="380">6.8</td>
</tr>
<tr id="354934DCE914DC95FF4569C09982FD47" box="[151,1414,676,698]" gridrow="2" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF4569C09DBFFD47" box="[151,443,676,698]" gridcol="0" gridrow="2" pageId="7" pageNumber="380">Anterior width</th>
<td id="76985DA0E914DC95FE0669C09E36FD47" box="[468,562,676,698]" gridcol="1" gridrow="2" pageId="7" pageNumber="380">2.8</td>
<td id="76985DA0E914DC95FD9C69C09EA3FD47" box="[590,679,676,698]" gridcol="2" gridrow="2" pageId="7" pageNumber="380">3.2</td>
<td id="76985DA0E914DC95FD1569C09F1BFD47" box="[711,799,676,698]" gridcol="3" gridrow="2" pageId="7" pageNumber="380">2</td>
<td id="76985DA0E914DC95FC9469C09FADFD47" box="[838,937,676,698]" gridcol="4" gridrow="2" pageId="7" pageNumber="380">2.5</td>
<td id="76985DA0E914DC95FC1269C0981DFD47" box="[960,1049,676,698]" gridcol="5" gridrow="2" pageId="7" pageNumber="380">2.3</td>
<td id="76985DA0E914DC95FBEB69C09897FD47" box="[1081,1171,676,698]" gridcol="6" gridrow="2" pageId="7" pageNumber="380">2.1</td>
<td id="76985DA0E914DC95FB6169C09908FD47" box="[1203,1292,676,698]" gridcol="7" gridrow="2" pageId="7" pageNumber="380">2.3</td>
<td id="76985DA0E914DC95FAFF69C09982FD47" box="[1325,1414,676,698]" gridcol="8" gridrow="2" pageId="7" pageNumber="380">2.7</td>
</tr>
<tr id="354934DCE914DC95FF4569A99982FD1E" box="[151,1414,717,739]" gridrow="3" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF4569A99DBFFD1E" box="[151,443,717,739]" gridcol="0" gridrow="3" pageId="7" pageNumber="380">Posterior width</th>
<td id="76985DA0E914DC95FE0669A99E36FD1E" box="[468,562,717,739]" gridcol="1" gridrow="3" pageId="7" pageNumber="380">5.3</td>
<td id="76985DA0E914DC95FD9C69A99EA3FD1E" box="[590,679,717,739]" gridcol="2" gridrow="3" pageId="7" pageNumber="380">5.8</td>
<td id="76985DA0E914DC95FD1569A99F1BFD1E" box="[711,799,717,739]" gridcol="3" gridrow="3" pageId="7" pageNumber="380">4.98</td>
<td id="76985DA0E914DC95FC9469A99FADFD1E" box="[838,937,717,739]" gridcol="4" gridrow="3" pageId="7" pageNumber="380">6.2</td>
<td id="76985DA0E914DC95FC1269A9981DFD1E" box="[960,1049,717,739]" gridcol="5" gridrow="3" pageId="7" pageNumber="380">5.4</td>
<td id="76985DA0E914DC95FBEB69A99897FD1E" box="[1081,1171,717,739]" gridcol="6" gridrow="3" pageId="7" pageNumber="380">5.2</td>
<td id="76985DA0E914DC95FB6169A99908FD1E" box="[1203,1292,717,739]" gridcol="7" gridrow="3" pageId="7" pageNumber="380">6.0</td>
<td id="76985DA0E914DC95FAFF69A99982FD1E" box="[1325,1414,717,739]" gridcol="8" gridrow="3" pageId="7" pageNumber="380">6.7</td>
</tr>
<tr id="354934DCE914DC95FF4569929982FCF1" box="[151,1414,758,780]" gridrow="4" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF4569929DBFFCF1" box="[151,443,758,780]" gridcol="0" gridrow="4" pageId="7" pageNumber="380">Metasoma length</th>
<td id="76985DA0E914DC95FE0669929E36FCF1" box="[468,562,758,780]" gridcol="1" gridrow="4" pageId="7" pageNumber="380">34.5</td>
<td id="76985DA0E914DC95FD9C69929EA3FCF1" box="[590,679,758,780]" gridcol="2" gridrow="4" pageId="7" pageNumber="380">31.0</td>
<td id="76985DA0E914DC95FD1569929F1BFCF1" box="[711,799,758,780]" gridcol="3" gridrow="4" pageId="7" pageNumber="380">34.6</td>
<td id="76985DA0E914DC95FC9469929FADFCF1" box="[838,937,758,780]" gridcol="4" gridrow="4" pageId="7" pageNumber="380">26.9</td>
<td id="76985DA0E914DC95FC126992981DFCF1" box="[960,1049,758,780]" gridcol="5" gridrow="4" pageId="7" pageNumber="380">27.3</td>
<td id="76985DA0E914DC95FBEB69929897FCF1" box="[1081,1171,758,780]" gridcol="6" gridrow="4" pageId="7" pageNumber="380">35.7</td>
<td id="76985DA0E914DC95FB6169929908FCF1" box="[1203,1292,758,780]" gridcol="7" gridrow="4" pageId="7" pageNumber="380">27.5</td>
<td id="76985DA0E914DC95FAFF69929982FCF1" box="[1325,1414,758,780]" gridcol="8" gridrow="4" pageId="7" pageNumber="380">31.8</td>
</tr>
<tr id="354934DCE914DC95FF4568449982FCCB" box="[151,1414,800,822]" gridrow="5" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF4568449DBFFCCB" box="[151,443,800,822]" gridcol="0" gridrow="5" pageId="7" pageNumber="380">Segment I length (width)</th>
<td id="76985DA0E914DC95FE0668449E36FCCB" box="[468,562,800,822]" gridcol="1" gridrow="5" pageId="7" pageNumber="380">5.2 (2.8)</td>
<td id="76985DA0E914DC95FD9C68449EA3FCCB" box="[590,679,800,822]" gridcol="2" gridrow="5" pageId="7" pageNumber="380">3.9 (3.1)</td>
<td id="76985DA0E914DC95FD1568449F1BFCCB" box="[711,799,800,822]" gridcol="3" gridrow="5" pageId="7" pageNumber="380">5.1 (2.6)</td>
<td id="76985DA0E914DC95FC9468449FADFCCB" box="[838,937,800,822]" gridcol="4" gridrow="5" pageId="7" pageNumber="380">4.2 (3.3)</td>
<td id="76985DA0E914DC95FC126844981DFCCB" box="[960,1049,800,822]" gridcol="5" gridrow="5" pageId="7" pageNumber="380">4.3 (3.2)</td>
<td id="76985DA0E914DC95FBEB68449897FCCB" box="[1081,1171,800,822]" gridcol="6" gridrow="5" pageId="7" pageNumber="380">5.3 (2.7)</td>
<td id="76985DA0E914DC95FB6168449908FCCB" box="[1203,1292,800,822]" gridcol="7" gridrow="5" pageId="7" pageNumber="380">4.4 (3.0)</td>
<td id="76985DA0E914DC95FAFF68449982FCCB" box="[1325,1414,800,822]" gridcol="8" gridrow="5" pageId="7" pageNumber="380">5.2 (3.6)</td>
</tr>
<tr id="354934DCE914DC95FF45682D9982FCA2" box="[151,1414,841,863]" gridrow="6" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF45682D9DBFFCA2" box="[151,443,841,863]" gridcol="0" gridrow="6" pageId="7" pageNumber="380">Segment II length (width)</th>
<td id="76985DA0E914DC95FE06682D9E36FCA2" box="[468,562,841,863]" gridcol="1" gridrow="6" pageId="7" pageNumber="380">6.1 (2.5)</td>
<td id="76985DA0E914DC95FD9C682D9EA3FCA2" box="[590,679,841,863]" gridcol="2" gridrow="6" pageId="7" pageNumber="380">4.7 (3.1)</td>
<td id="76985DA0E914DC95FD15682D9F1BFCA2" box="[711,799,841,863]" gridcol="3" gridrow="6" pageId="7" pageNumber="380">6.6 (2.5)</td>
<td id="76985DA0E914DC95FC94682D9FADFCA2" box="[838,937,841,863]" gridcol="4" gridrow="6" pageId="7" pageNumber="380">5.0 (3.1)</td>
<td id="76985DA0E914DC95FC12682D981DFCA2" box="[960,1049,841,863]" gridcol="5" gridrow="6" pageId="7" pageNumber="380">5.1 (3.0)</td>
<td id="76985DA0E914DC95FBEB682D9897FCA2" box="[1081,1171,841,863]" gridcol="6" gridrow="6" pageId="7" pageNumber="380">6.6 (2.5)</td>
<td id="76985DA0E914DC95FB61682D9908FCA2" box="[1203,1292,841,863]" gridcol="7" gridrow="6" pageId="7" pageNumber="380">5.3 (2.9)</td>
<td id="76985DA0E914DC95FAFF682D9982FCA2" box="[1325,1414,841,863]" gridcol="8" gridrow="6" pageId="7" pageNumber="380">6.1 (3.5)</td>
</tr>
<tr id="354934DCE914DC95FF4568169982FC75" box="[151,1414,882,904]" gridrow="7" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF4568169DBFFC75" box="[151,443,882,904]" gridcol="0" gridrow="7" pageId="7" pageNumber="380">Segment III length (width)</th>
<td id="76985DA0E914DC95FE0668169E36FC75" box="[468,562,882,904]" gridcol="1" gridrow="7" pageId="7" pageNumber="380">6.8 (2.6)</td>
<td id="76985DA0E914DC95FD9C68169EA3FC75" box="[590,679,882,904]" gridcol="2" gridrow="7" pageId="7" pageNumber="380">5.5 (2.7)</td>
<td id="76985DA0E914DC95FD1568169F1BFC75" box="[711,799,882,904]" gridcol="3" gridrow="7" pageId="7" pageNumber="380">7.5 (2.4)</td>
<td id="76985DA0E914DC95FC9468169FADFC75" box="[838,937,882,904]" gridcol="4" gridrow="7" pageId="7" pageNumber="380">5.4 (3.02)</td>
<td id="76985DA0E914DC95FC126816981DFC75" box="[960,1049,882,904]" gridcol="5" gridrow="7" pageId="7" pageNumber="380">5.5 (2.9)</td>
<td id="76985DA0E914DC95FBEB68169897FC75" box="[1081,1171,882,904]" gridcol="6" gridrow="7" pageId="7" pageNumber="380">7.7 (2.5)</td>
<td id="76985DA0E914DC95FB6168169908FC75" box="[1203,1292,882,904]" gridcol="7" gridrow="7" pageId="7" pageNumber="380">5.4 (2.9)</td>
<td id="76985DA0E914DC95FAFF68169982FC75" box="[1325,1414,882,904]" gridcol="8" gridrow="7" pageId="7" pageNumber="380">6.4 (3.3)</td>
</tr>
<tr id="354934DCE914DC95FF4568F89982FC4F" box="[151,1414,924,946]" gridrow="8" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF4568F89DBFFC4F" box="[151,443,924,946]" gridcol="0" gridrow="8" pageId="7" pageNumber="380">Segment IV length (width)</th>
<td id="76985DA0E914DC95FE0668F89E36FC4F" box="[468,562,924,946]" gridcol="1" gridrow="8" pageId="7" pageNumber="380">7.3 (2.6)</td>
<td id="76985DA0E914DC95FD9C68F89EA3FC4F" box="[590,679,924,946]" gridcol="2" gridrow="8" pageId="7" pageNumber="380">5.6 (3.1)</td>
<td id="76985DA0E914DC95FD1568F89F1BFC4F" box="[711,799,924,946]" gridcol="3" gridrow="8" pageId="7" pageNumber="380">7.6 (2.3)</td>
<td id="76985DA0E914DC95FC9468F89FADFC4F" box="[838,937,924,946]" gridcol="4" gridrow="8" pageId="7" pageNumber="380">5.7 (3.1)</td>
<td id="76985DA0E914DC95FC1268F8981DFC4F" box="[960,1049,924,946]" gridcol="5" gridrow="8" pageId="7" pageNumber="380">6.0 (2.9)</td>
<td id="76985DA0E914DC95FBEB68F89897FC4F" box="[1081,1171,924,946]" gridcol="6" gridrow="8" pageId="7" pageNumber="380">7.8 (2.5)</td>
<td id="76985DA0E914DC95FB6168F89908FC4F" box="[1203,1292,924,946]" gridcol="7" gridrow="8" pageId="7" pageNumber="380">5.8 (2.9)</td>
<td id="76985DA0E914DC95FAFF68F89982FC4F" box="[1325,1414,924,946]" gridcol="8" gridrow="8" pageId="7" pageNumber="380">6.6 (3.5)</td>
</tr>
<tr id="354934DCE914DC95FF4568A19982FC26" box="[151,1414,965,987]" gridrow="9" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF4568A19DBFFC26" box="[151,443,965,987]" gridcol="0" gridrow="9" pageId="7" pageNumber="380">Segment V length (width)</th>
<td id="76985DA0E914DC95FE0668A19E36FC26" box="[468,562,965,987]" gridcol="1" gridrow="9" pageId="7" pageNumber="380">7.9 (2.6)</td>
<td id="76985DA0E914DC95FD9C68A19EA3FC26" box="[590,679,965,987]" gridcol="2" gridrow="9" pageId="7" pageNumber="380">6.2 (3.1)</td>
<td id="76985DA0E914DC95FD1568A19F1BFC26" box="[711,799,965,987]" gridcol="3" gridrow="9" pageId="7" pageNumber="380">7.8 (2.7)</td>
<td id="76985DA0E914DC95FC9468A19FADFC26" box="[838,937,965,987]" gridcol="4" gridrow="9" pageId="7" pageNumber="380">6.6 (3.0)</td>
<td id="76985DA0E914DC95FC1268A1981DFC26" box="[960,1049,965,987]" gridcol="5" gridrow="9" pageId="7" pageNumber="380">6.5 (2.8)</td>
<td id="76985DA0E914DC95FBEB68A19897FC26" box="[1081,1171,965,987]" gridcol="6" gridrow="9" pageId="7" pageNumber="380">8.2 (3.0)</td>
<td id="76985DA0E914DC95FB6168A19908FC26" box="[1203,1292,965,987]" gridcol="7" gridrow="9" pageId="7" pageNumber="380">6.6 (2.9)</td>
<td id="76985DA0E914DC95FAFF68A19982FC26" box="[1325,1414,965,987]" gridcol="8" gridrow="9" pageId="7" pageNumber="380">7.5 (3.4)</td>
</tr>
<tr id="354934DCE914DC95FF45688A9982FBF9" box="[151,1414,1006,1028]" gridrow="10" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF45688A9DBFFBF9" box="[151,443,1006,1028]" gridcol="0" gridrow="10" pageId="7" pageNumber="380">Telson vesicle length</th>
<td id="76985DA0E914DC95FE06688A9E36FBF9" box="[468,562,1006,1028]" gridcol="1" gridrow="10" pageId="7" pageNumber="380">3</td>
<td id="76985DA0E914DC95FD9C688A9EA3FBF9" box="[590,679,1006,1028]" gridcol="2" gridrow="10" pageId="7" pageNumber="380">3.1</td>
<td id="76985DA0E914DC95FD15688A9F1BFBF9" box="[711,799,1006,1028]" gridcol="3" gridrow="10" pageId="7" pageNumber="380">3.8</td>
<td id="76985DA0E914DC95FC94688A9FADFBF9" box="[838,937,1006,1028]" gridcol="4" gridrow="10" pageId="7" pageNumber="380">3.8</td>
<td id="76985DA0E914DC95FC12688A981DFBF9" box="[960,1049,1006,1028]" gridcol="5" gridrow="10" pageId="7" pageNumber="380">3.3</td>
<td id="76985DA0E914DC95FBEB688A9897FBF9" box="[1081,1171,1006,1028]" gridcol="6" gridrow="10" pageId="7" pageNumber="380">4.5</td>
<td id="76985DA0E914DC95FB61688A9908FBF9" box="[1203,1292,1006,1028]" gridcol="7" gridrow="10" pageId="7" pageNumber="380">3.8</td>
<td id="76985DA0E914DC95FAFF688A9982FBF9" box="[1325,1414,1006,1028]" gridcol="8" gridrow="10" pageId="7" pageNumber="380">4.6</td>
</tr>
<tr id="354934DCE914DC95FF456F7C9982FBD3" box="[151,1414,1048,1070]" gridrow="11" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456F7C9DBFFBD3" box="[151,443,1048,1070]" gridcol="0" gridrow="11" pageId="7" pageNumber="380">Aculeus length</th>
<td id="76985DA0E914DC95FE066F7C9E36FBD3" box="[468,562,1048,1070]" gridcol="1" gridrow="11" pageId="7" pageNumber="380">2.7</td>
<td id="76985DA0E914DC95FD9C6F7C9EA3FBD3" box="[590,679,1048,1070]" gridcol="2" gridrow="11" pageId="7" pageNumber="380">1.9</td>
<td id="76985DA0E914DC95FD156F7C9F1BFBD3" box="[711,799,1048,1070]" gridcol="3" gridrow="11" pageId="7" pageNumber="380">2.6</td>
<td id="76985DA0E914DC95FC946F7C9FADFBD3" box="[838,937,1048,1070]" gridcol="4" gridrow="11" pageId="7" pageNumber="380">2.7</td>
<td id="76985DA0E914DC95FC126F7C981DFBD3" box="[960,1049,1048,1070]" gridcol="5" gridrow="11" pageId="7" pageNumber="380">2.5</td>
<td id="76985DA0E914DC95FBEB6F7C9897FBD3" box="[1081,1171,1048,1070]" gridcol="6" gridrow="11" pageId="7" pageNumber="380">2.4</td>
<td id="76985DA0E914DC95FB616F7C9908FBD3" box="[1203,1292,1048,1070]" gridcol="7" gridrow="11" pageId="7" pageNumber="380">2.5</td>
<td id="76985DA0E914DC95FAFF6F7C9982FBD3" box="[1325,1414,1048,1070]" gridcol="8" gridrow="11" pageId="7" pageNumber="380">2.6</td>
</tr>
<tr id="354934DCE914DC95FF456F259982FBAA" box="[151,1414,1089,1111]" gridrow="12" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456F259DBFFBAA" box="[151,443,1089,1111]" gridcol="0" gridrow="12" pageId="7" pageNumber="380">Vesicle width (depth)</th>
<td id="76985DA0E914DC95FE066F259E36FBAA" box="[468,562,1089,1111]" gridcol="1" gridrow="12" pageId="7" pageNumber="380">2.3 (1.7)</td>
<td id="76985DA0E914DC95FD9C6F259EA3FBAA" box="[590,679,1089,1111]" gridcol="2" gridrow="12" pageId="7" pageNumber="380">2.4 (1.9)</td>
<td id="76985DA0E914DC95FD156F259F1BFBAA" box="[711,799,1089,1111]" gridcol="3" gridrow="12" pageId="7" pageNumber="380">2.4 (1.9)</td>
<td id="76985DA0E914DC95FC946F259FADFBAA" box="[838,937,1089,1111]" gridcol="4" gridrow="12" pageId="7" pageNumber="380">2.3 (1.7)</td>
<td id="76985DA0E914DC95FC126F25981DFBAA" box="[960,1049,1089,1111]" gridcol="5" gridrow="12" pageId="7" pageNumber="380">2.1 (1.5)</td>
<td id="76985DA0E914DC95FBEB6F259897FBAA" box="[1081,1171,1089,1111]" gridcol="6" gridrow="12" pageId="7" pageNumber="380">2.5 (1.9)</td>
<td id="76985DA0E914DC95FB616F259908FBAA" box="[1203,1292,1089,1111]" gridcol="7" gridrow="12" pageId="7" pageNumber="380">2.1 (1.5)</td>
<td id="76985DA0E914DC95FAFF6F259982FBAA" box="[1325,1414,1089,1111]" gridcol="8" gridrow="12" pageId="7" pageNumber="380">2.5 (2.0)</td>
</tr>
<tr id="354934DCE914DC95FF456F0E9982FB7D" box="[151,1414,1130,1152]" gridrow="13" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456F0E9DBFFB7D" box="[151,443,1130,1152]" gridcol="0" gridrow="13" pageId="7" pageNumber="380">Pedipalp length</th>
<td id="76985DA0E914DC95FE066F0E9E36FB7D" box="[468,562,1130,1152]" gridcol="1" gridrow="13" pageId="7" pageNumber="380">21.2</td>
<td id="76985DA0E914DC95FD9C6F0E9EA3FB7D" box="[590,679,1130,1152]" gridcol="2" gridrow="13" pageId="7" pageNumber="380">23</td>
<td id="76985DA0E914DC95FD156F0E9F1BFB7D" box="[711,799,1130,1152]" gridcol="3" gridrow="13" pageId="7" pageNumber="380">21.3</td>
<td id="76985DA0E914DC95FC946F0E9FADFB7D" box="[838,937,1130,1152]" gridcol="4" gridrow="13" pageId="7" pageNumber="380">21.1</td>
<td id="76985DA0E914DC95FC126F0E981DFB7D" box="[960,1049,1130,1152]" gridcol="5" gridrow="13" pageId="7" pageNumber="380">19.4</td>
<td id="76985DA0E914DC95FBEB6F0E9897FB7D" box="[1081,1171,1130,1152]" gridcol="6" gridrow="13" pageId="7" pageNumber="380">22.4</td>
<td id="76985DA0E914DC95FB616F0E9908FB7D" box="[1203,1292,1130,1152]" gridcol="7" gridrow="13" pageId="7" pageNumber="380">21.5</td>
<td id="76985DA0E914DC95FAFF6F0E9982FB7D" box="[1325,1414,1130,1152]" gridcol="8" gridrow="13" pageId="7" pageNumber="380">24.2</td>
</tr>
<tr id="354934DCE914DC95FF456FF09982FB57" box="[151,1414,1172,1194]" gridrow="14" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456FF09DBFFB57" box="[151,443,1172,1194]" gridcol="0" gridrow="14" pageId="7" pageNumber="380">Femur length (width)</th>
<td id="76985DA0E914DC95FE066FF09E36FB57" box="[468,562,1172,1194]" gridcol="1" gridrow="14" pageId="7" pageNumber="380">5.4 (1.2)</td>
<td id="76985DA0E914DC95FD9C6FF09EA3FB57" box="[590,679,1172,1194]" gridcol="2" gridrow="14" pageId="7" pageNumber="380">4.7 (1.3)</td>
<td id="76985DA0E914DC95FD156FF09F1BFB57" box="[711,799,1172,1194]" gridcol="3" gridrow="14" pageId="7" pageNumber="380">5.4 (1.4)</td>
<td id="76985DA0E914DC95FC946FF09FADFB57" box="[838,937,1172,1194]" gridcol="4" gridrow="14" pageId="7" pageNumber="380">5.2 (1.6)</td>
<td id="76985DA0E914DC95FC126FF0981DFB57" box="[960,1049,1172,1194]" gridcol="5" gridrow="14" pageId="7" pageNumber="380">4.9 (1.4)</td>
<td id="76985DA0E914DC95FBEB6FF09897FB57" box="[1081,1171,1172,1194]" gridcol="6" gridrow="14" pageId="7" pageNumber="380">5.7 (1.4)</td>
<td id="76985DA0E914DC95FB616FF09908FB57" box="[1203,1292,1172,1194]" gridcol="7" gridrow="14" pageId="7" pageNumber="380">5.3 (1.7)</td>
<td id="76985DA0E914DC95FAFF6FF09982FB57" box="[1325,1414,1172,1194]" gridcol="8" gridrow="14" pageId="7" pageNumber="380">6.0 (1.9)</td>
</tr>
<tr id="354934DCE914DC95FF456FD99982FB2E" box="[151,1414,1213,1235]" gridrow="15" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456FD99DBFFB2E" box="[151,443,1213,1235]" gridcol="0" gridrow="15" pageId="7" pageNumber="380">Patella length (width)</th>
<td id="76985DA0E914DC95FE066FD99E36FB2E" box="[468,562,1213,1235]" gridcol="1" gridrow="15" pageId="7" pageNumber="380">6.2 (1.6)</td>
<td id="76985DA0E914DC95FD9C6FD99EA3FB2E" box="[590,679,1213,1235]" gridcol="2" gridrow="15" pageId="7" pageNumber="380">6.1 (1.6)</td>
<td id="76985DA0E914DC95FD156FD99F1BFB2E" box="[711,799,1213,1235]" gridcol="3" gridrow="15" pageId="7" pageNumber="380">6.2 (1.9)</td>
<td id="76985DA0E914DC95FC946FD99FADFB2E" box="[838,937,1213,1235]" gridcol="4" gridrow="15" pageId="7" pageNumber="380">6.1 (2.1)</td>
<td id="76985DA0E914DC95FC126FD9981DFB2E" box="[960,1049,1213,1235]" gridcol="5" gridrow="15" pageId="7" pageNumber="380">5.7 (1.9)</td>
<td id="76985DA0E914DC95FBEB6FD99897FB2E" box="[1081,1171,1213,1235]" gridcol="6" gridrow="15" pageId="7" pageNumber="380">6.2 (1.9)</td>
<td id="76985DA0E914DC95FB616FD99908FB2E" box="[1203,1292,1213,1235]" gridcol="7" gridrow="15" pageId="7" pageNumber="380">6.1 (2.0)</td>
<td id="76985DA0E914DC95FAFF6FD99982FB2E" box="[1325,1414,1213,1235]" gridcol="8" gridrow="15" pageId="7" pageNumber="380">6.8 (2.4)</td>
</tr>
<tr id="354934DCE914DC95FF456F829982FAE6" box="[151,1414,1254,1307]" gridrow="16" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456F829DBFFAE6" box="[151,443,1254,1307]" gridcol="0" gridrow="16" pageId="7" pageNumber="380">Chela external carina length (width)</th>
<td id="76985DA0E914DC95FE066F829E36FAE6" box="[468,562,1254,1307]" gridcol="1" gridrow="16" pageId="7" pageNumber="380">4.2 (2.3)</td>
<td id="76985DA0E914DC95FD9C6F829EA3FAE6" box="[590,679,1254,1307]" gridcol="2" gridrow="16" pageId="7" pageNumber="380">3.8 (2.3)</td>
<td id="76985DA0E914DC95FD156F829F1BFAE6" box="[711,799,1254,1307]" gridcol="3" gridrow="16" pageId="7" pageNumber="380">4.2 (2.4)</td>
<td id="76985DA0E914DC95FC946F829FADFAE6" box="[838,937,1254,1307]" gridcol="4" gridrow="16" pageId="7" pageNumber="380">3.8 (2.4)</td>
<td id="76985DA0E914DC95FC126F82981DFAE6" box="[960,1049,1254,1307]" gridcol="5" gridrow="16" pageId="7" pageNumber="380">3.4 (2.0)</td>
<td id="76985DA0E914DC95FBEB6F829897FAE6" box="[1081,1171,1254,1307]" gridcol="6" gridrow="16" pageId="7" pageNumber="380">4.4 (2.5)</td>
<td id="76985DA0E914DC95FB616F829908FAE6" box="[1203,1292,1254,1307]" gridcol="7" gridrow="16" pageId="7" pageNumber="380">3.6 (2.3)</td>
<td id="76985DA0E914DC95FAFF6F829982FAE6" box="[1325,1414,1254,1307]" gridcol="8" gridrow="16" pageId="7" pageNumber="380">4.2 (2.3)</td>
</tr>
<tr id="354934DCE914DC95FF456E4A9982FAB9" box="[151,1414,1326,1348]" gridrow="17" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456E4A9DBFFAB9" box="[151,443,1326,1348]" gridcol="0" gridrow="17" pageId="7" pageNumber="380">Movable finger length</th>
<td id="76985DA0E914DC95FE066E4A9E36FAB9" box="[468,562,1326,1348]" gridcol="1" gridrow="17" pageId="7" pageNumber="380">6.1</td>
<td id="76985DA0E914DC95FD9C6E4A9EA3FAB9" box="[590,679,1326,1348]" gridcol="2" gridrow="17" pageId="7" pageNumber="380">6.2</td>
<td id="76985DA0E914DC95FD156E4A9F1BFAB9" box="[711,799,1326,1348]" gridcol="3" gridrow="17" pageId="7" pageNumber="380">5.6</td>
<td id="76985DA0E914DC95FC946E4A9FADFAB9" box="[838,937,1326,1348]" gridcol="4" gridrow="17" pageId="7" pageNumber="380">6.0</td>
<td id="76985DA0E914DC95FC126E4A981DFAB9" box="[960,1049,1326,1348]" gridcol="5" gridrow="17" pageId="7" pageNumber="380">5.5</td>
<td id="76985DA0E914DC95FBEB6E4A9897FAB9" box="[1081,1171,1326,1348]" gridcol="6" gridrow="17" pageId="7" pageNumber="380">6.0</td>
<td id="76985DA0E914DC95FB616E4A9908FAB9" box="[1203,1292,1326,1348]" gridcol="7" gridrow="17" pageId="7" pageNumber="380">6.5</td>
<td id="76985DA0E914DC95FAFF6E4A9982FAB9" box="[1325,1414,1326,1348]" gridcol="8" gridrow="17" pageId="7" pageNumber="380">7.2</td>
</tr>
<tr id="354934DCE914DC95FF456E3C9982FA71" box="[151,1414,1368,1420]" gridrow="18" pageId="7" pageNumber="380" rowspan-1="1" rowspan-2="1">
<th id="76985DA0E914DC95FF456E3C9DBFFA71" box="[151,443,1368,1420]" gridcol="0" gridrow="18" pageId="7" pageNumber="380">Pectines (length along denticle margin)</th>
<td id="76985DA0E914DC95FD156E3C9F1BFA71" box="[711,799,1368,1420]" gridcol="3" gridrow="18" pageId="7" pageNumber="380">3.9</td>
<td id="76985DA0E914DC95FC946E3C9FADFA71" box="[838,937,1368,1420]" gridcol="4" gridrow="18" pageId="7" pageNumber="380">3.4</td>
<td id="76985DA0E914DC95FC126E3C981DFA71" box="[960,1049,1368,1420]" gridcol="5" gridrow="18" pageId="7" pageNumber="380">3.8</td>
<td id="76985DA0E914DC95FBEB6E3C9897FA71" box="[1081,1171,1368,1420]" gridcol="6" gridrow="18" pageId="7" pageNumber="380">4.4</td>
<td id="76985DA0E914DC95FB616E3C9908FA71" box="[1203,1292,1368,1420]" gridcol="7" gridrow="18" pageId="7" pageNumber="380">4.5</td>
<td id="76985DA0E914DC95FAFF6E3C9982FA71" box="[1325,1414,1368,1420]" gridcol="8" gridrow="18" pageId="7" pageNumber="380">4.4</td>
</tr>
<tr id="354934DCE914DC95FF456EC49982FA4B" box="[151,1414,1440,1462]" gridrow="19" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456EC49DBFFA4B" box="[151,443,1440,1462]" gridcol="0" gridrow="19" pageId="7" pageNumber="380">Pectinal teeth L/R</th>
<td id="76985DA0E914DC95FE066EC49E36FA4B" box="[468,562,1440,1462]" gridcol="1" gridrow="19" pageId="7" pageNumber="380">17/17</td>
<td id="76985DA0E914DC95FD9C6EC49EA3FA4B" box="[590,679,1440,1462]" gridcol="2" gridrow="19" pageId="7" pageNumber="380">18/18</td>
<td id="76985DA0E914DC95FD156EC49F1BFA4B" box="[711,799,1440,1462]" gridcol="3" gridrow="19" pageId="7" pageNumber="380">17/17</td>
<td id="76985DA0E914DC95FC946EC49FADFA4B" box="[838,937,1440,1462]" gridcol="4" gridrow="19" pageId="7" pageNumber="380">17/17</td>
<td id="76985DA0E914DC95FC126EC4981DFA4B" box="[960,1049,1440,1462]" gridcol="5" gridrow="19" pageId="7" pageNumber="380">19/19</td>
<td id="76985DA0E914DC95FBEB6EC49897FA4B" box="[1081,1171,1440,1462]" gridcol="6" gridrow="19" pageId="7" pageNumber="380">19/19</td>
<td id="76985DA0E914DC95FB616EC49908FA4B" box="[1203,1292,1440,1462]" gridcol="7" gridrow="19" pageId="7" pageNumber="380">20/20</td>
<td id="76985DA0E914DC95FAFF6EC49982FA4B" box="[1325,1414,1440,1462]" gridcol="8" gridrow="19" pageId="7" pageNumber="380">20/19</td>
</tr>
<tr id="354934DCE914DC95FF456EAD9982FA22" box="[151,1414,1481,1503]" gridrow="20" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456EAD9DBFFA22" box="[151,443,1481,1503]" gridcol="0" gridrow="20" pageId="7" pageNumber="380">
<taxonomicName id="4C794D1DE914236DFF4D6EAE9D09FA22" box="[159,269,1482,1503]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="376" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
I-VI length
</th>
<td id="76985DA0E914DC95FE066EAD9E36FA22" box="[468,562,1481,1503]" gridcol="1" gridrow="20" pageId="7" pageNumber="380">12.1</td>
<td id="76985DA0E914DC95FD9C6EAD9EA3FA22" box="[590,679,1481,1503]" gridcol="2" gridrow="20" pageId="7" pageNumber="380">12.9</td>
<td id="76985DA0E914DC95FD156EAD9F1BFA22" box="[711,799,1481,1503]" gridcol="3" gridrow="20" pageId="7" pageNumber="380">12.2</td>
<td id="76985DA0E914DC95FC946EAD9FADFA22" box="[838,937,1481,1503]" gridcol="4" gridrow="20" pageId="7" pageNumber="380">15.6</td>
<td id="76985DA0E914DC95FC126EAD981DFA22" box="[960,1049,1481,1503]" gridcol="5" gridrow="20" pageId="7" pageNumber="380">11.4</td>
<td id="76985DA0E914DC95FBEB6EAD9897FA22" box="[1081,1171,1481,1503]" gridcol="6" gridrow="20" pageId="7" pageNumber="380">12.4</td>
<td id="76985DA0E914DC95FB616EAD9908FA22" box="[1203,1292,1481,1503]" gridcol="7" gridrow="20" pageId="7" pageNumber="380">11.8</td>
<td id="76985DA0E914DC95FAFF6EAD9982FA22" box="[1325,1414,1481,1503]" gridcol="8" gridrow="20" pageId="7" pageNumber="380">13.2</td>
</tr>
<tr id="354934DCE914DC95FF456E969982F9F5" box="[151,1414,1522,1544]" gridrow="21" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456E969DBFF9F5" box="[151,443,1522,1544]" gridcol="0" gridrow="21" pageId="7" pageNumber="380">
<taxonomicName id="4C794D1DE914236DFF4D6E979D09F9F5" box="[159,269,1523,1544]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="376" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
VII length
</th>
<td id="76985DA0E914DC95FE066E969E36F9F5" box="[468,562,1522,1544]" gridcol="1" gridrow="21" pageId="7" pageNumber="380">4.2</td>
<td id="76985DA0E914DC95FD9C6E969EA3F9F5" box="[590,679,1522,1544]" gridcol="2" gridrow="21" pageId="7" pageNumber="380">4.0</td>
<td id="76985DA0E914DC95FD156E969F1BF9F5" box="[711,799,1522,1544]" gridcol="3" gridrow="21" pageId="7" pageNumber="380">4.2</td>
<td id="76985DA0E914DC95FC946E969FADF9F5" box="[838,937,1522,1544]" gridcol="4" gridrow="21" pageId="7" pageNumber="380">4.1</td>
<td id="76985DA0E914DC95FC126E96981DF9F5" box="[960,1049,1522,1544]" gridcol="5" gridrow="21" pageId="7" pageNumber="380">4.1</td>
<td id="76985DA0E914DC95FBEB6E969897F9F5" box="[1081,1171,1522,1544]" gridcol="6" gridrow="21" pageId="7" pageNumber="380">4.0</td>
<td id="76985DA0E914DC95FB616E969908F9F5" box="[1203,1292,1522,1544]" gridcol="7" gridrow="21" pageId="7" pageNumber="380">4.3</td>
<td id="76985DA0E914DC95FAFF6E969982F9F5" box="[1325,1414,1522,1544]" gridcol="8" gridrow="21" pageId="7" pageNumber="380">5.2</td>
</tr>
<tr id="354934DCE914DC95FF456D789982F9AD" box="[151,1414,1564,1616]" gridrow="22" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456D789DBFF9AD" box="[151,443,1564,1616]" gridcol="0" gridrow="22" pageId="7" pageNumber="380">
<taxonomicName id="4C794D1DE914236DFF4D6D799D09F9CF" box="[159,269,1565,1586]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="376" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
VII anterior width
</th>
<td id="76985DA0E914DC95FE066D789E36F9AD" box="[468,562,1564,1616]" gridcol="1" gridrow="22" pageId="7" pageNumber="380">4.8</td>
<td id="76985DA0E914DC95FD9C6D789EA3F9AD" box="[590,679,1564,1616]" gridcol="2" gridrow="22" pageId="7" pageNumber="380">6.0</td>
<td id="76985DA0E914DC95FD156D789F1BF9AD" box="[711,799,1564,1616]" gridcol="3" gridrow="22" pageId="7" pageNumber="380">4.8</td>
<td id="76985DA0E914DC95FC946D789FADF9AD" box="[838,937,1564,1616]" gridcol="4" gridrow="22" pageId="7" pageNumber="380">6.2</td>
<td id="76985DA0E914DC95FC126D78981DF9AD" box="[960,1049,1564,1616]" gridcol="5" gridrow="22" pageId="7" pageNumber="380">5.7</td>
<td id="76985DA0E914DC95FBEB6D789897F9AD" box="[1081,1171,1564,1616]" gridcol="6" gridrow="22" pageId="7" pageNumber="380">4.4</td>
<td id="76985DA0E914DC95FB616D789908F9AD" box="[1203,1292,1564,1616]" gridcol="7" gridrow="22" pageId="7" pageNumber="380">6.2</td>
<td id="76985DA0E914DC95FAFF6D789982F9AD" box="[1325,1414,1564,1616]" gridcol="8" gridrow="22" pageId="7" pageNumber="380">7.4</td>
</tr>
<tr id="354934DCE914DC95FF456D009982F987" box="[151,1414,1636,1658]" gridrow="23" pageId="7" pageNumber="380">
<th id="76985DA0E914DC95FF456D009DBFF987" box="[151,443,1636,1658]" gridcol="0" gridrow="23" pageId="7" pageNumber="380">Total length</th>
<td id="76985DA0E914DC95FE066D009E36F987" box="[468,562,1636,1658]" gridcol="1" gridrow="23" pageId="7" pageNumber="380">65.4</td>
<td id="76985DA0E914DC95FD9C6D009EA3F987" box="[590,679,1636,1658]" gridcol="2" gridrow="23" pageId="7" pageNumber="380">64.1</td>
<td id="76985DA0E914DC95FD156D009F1BF987" box="[711,799,1636,1658]" gridcol="3" gridrow="23" pageId="7" pageNumber="380">60.6</td>
<td id="76985DA0E914DC95FC946D009FADF987" box="[838,937,1636,1658]" gridcol="4" gridrow="23" pageId="7" pageNumber="380">58.6</td>
<td id="76985DA0E914DC95FC126D00981DF987" box="[960,1049,1636,1658]" gridcol="5" gridrow="23" pageId="7" pageNumber="380">52.9</td>
<td id="76985DA0E914DC95FBEB6D009897F987" box="[1081,1171,1636,1658]" gridcol="6" gridrow="23" pageId="7" pageNumber="380">62.5</td>
<td id="76985DA0E914DC95FB616D009908F987" box="[1203,1292,1636,1658]" gridcol="7" gridrow="23" pageId="7" pageNumber="380">55.5</td>
<td id="76985DA0E914DC95FAFF6D009982F987" box="[1325,1414,1636,1658]" gridcol="8" gridrow="23" pageId="7" pageNumber="380">63.7</td>
</tr>
<tr id="354934DCE914DC95FF456DB19982F912" box="[151,1414,1749,1775]" gridrow="24" pageId="7" pageNumber="380" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76985DA0E914DC95FF456DB19DBFF912" box="[151,443,1749,1775]" gridcol="0" gridrow="24" pageId="7" pageNumber="380">Discussion</th>
</tr>
</table>
</paragraph>
<paragraph id="8BC6369EE9142362FF456C789DE3FF28" blockId="7.[151,1436,1820,2025]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="381" pageId="7" pageNumber="380">
Separation of species in 
<taxonomicName id="4C794D1DE914236DFE7B6C789E38F8C8" box="[425,572,1820,1845]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA8CE914236DFE7B6C789E38F8C8" box="[425,572,1820,1845]" italics="true" pageId="7" pageNumber="380">Centruroides</emphasis>
</taxonomicName>
based heavily on coloration patterns and morphometric characters, known to have high intraspecific variation, has led to a proliferation of names and misidentifications. Using a combination of molecular and morphological data, the genus 
<taxonomicName id="4C794D1DE914236DFD776C009F3CF880" box="[677,824,1892,1917]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA8CE914236DFD776C009F3CF880" box="[677,824,1892,1917]" italics="true" pageId="7" pageNumber="380">Centruroides</emphasis>
</taxonomicName>
is currently being revised by coauthor LAE. 
<collectingCountry id="F36E760EE914236DFA946C00999FF880" box="[1350,1435,1892,1917]" name="Mexico" pageId="7" pageNumber="380">Mexico</collectingCountry>
and the southwestern 
<collectingCountry id="F36E760EE914236DFE586CEC9DC6F85D" box="[394,450,1928,1952]" name="United States of America" pageId="7" pageNumber="380">USA</collectingCountry>
have some 
<taxonomicName id="4C794D1DE914236DFD956CE39EDEF85D" box="[583,730,1927,1952]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA8CE914236DFD956CE39EDEF85D" box="[583,730,1927,1952]" italics="true" pageId="7" pageNumber="380">Centruroides</emphasis>
</taxonomicName>
species with venoms highly toxic to humans, however none of the presently known species of 
<taxonomicName id="4C794D1DE914236DFE2C6CC89E95F838" box="[510,657,1964,1989]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="380" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA8CE914236DFE2C6CC89E95F838" box="[510,657,1964,1989]" italics="true" pageId="7" pageNumber="380">Centruroides</emphasis>
</taxonomicName>
from 
<collectingCountry id="F36E760EE914236DFD0A6CC99F35F839" box="[728,817,1965,1988]" name="Panama" pageId="7" pageNumber="380">Panama</collectingCountry>
cause medically significant envenomations in humans (
<bibRefCitation id="EFE84B6FE914236DFF4C6CB49D7BF815" author="Borges" box="[158,383,2000,2025]" pageId="7" pageNumber="380" refString="Borges, A., Miranda, R. J. &amp; Pascale, J. M. (2012) Scorpionism in Central America, with special reference to the case of Panama. Journal of Venomous Animals and Toxins including Tropical Diseases, 18, 130 - 143. http: // dx. doi. org / 10.1590 / s 1678 - 91992012000200002" type="journal article" year="2012">
Borges 
<emphasis id="B90DEA8CE914236DFF286CB59D33F814" box="[250,311,2000,2025]" italics="true" pageId="7" pageNumber="380">et al.</emphasis>
2012
</bibRefCitation>
). This new species, with a distribution limited only to the uppermost limit of the highest mountain range in 
<collectingCountry id="F36E760EE91B2362FEA76BFC9DD5FF52" box="[373,465,152,175]" name="Panama" pageId="8" pageNumber="381">Panama</collectingCountry>
, provides evidence for arachnid endemism in this unique region and has important implications for conservation.
</paragraph>
</subSubSection>
</treatment>
</document>