treatments-xml/data/03/A9/87/03A987F75412FFADC4B22BFDACD0FB0F.xml
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<document id="4649799F20C34269F8C5CA206EF4FF7D" ID-CLB-Dataset="297560" ID-DOI="10.5252/zoosystema2024v46a13" ID-GBIF-Dataset="9d97ce13-f0b1-4c8e-a281-034d4df4fe41" ID-ISSN="1638-9387" ID-Zenodo-Dep="11391415" ID-ZooBank="7547F492-D2B2-4881-B3D9-12F730B4DA65" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1716971933471" checkinUser="plazi" docAuthor="Langlois, Dominique &amp; Daugeron, Christophe" docDate="2024" docId="03A987F75412FFADC4B22BFDACD0FB0F" docLanguage="en" docName="zoosystema.2024.46.13.pdf" docOrigin="Zoosystema 46 (13)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a13.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Empis (Empis) cartusiana Langlois &amp; Daugeron 2024, n. sp." docType="treatment" docUuid="98CB22EC-C56D-4172-9186-D550D4CAE76A" docUuidSource="ZooBank" docVersion="3" lastPageNumber="322" masterDocId="FF90FF8F5411FFA8C773296BAD09FFA1" masterDocTitle="Two new species of the genus Empis Linnaeus, 1758 (Diptera, Empididae, Empidinae) from French nature reserves" masterLastPageNumber="325" masterPageNumber="319" pageNumber="320" updateTime="1717016358231" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="57FB3BC3845B13EDF47053BDF855266E">Two new species of the genus Empis Linnaeus, 1758 (Diptera, Empididae, Empidinae) from French nature reserves</mods:title>
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<mods:namePart id="039AB6BA968ADA0D021FD1AC69FB3021">Langlois, Dominique</mods:namePart>
<mods:affiliation id="BC026D76BB52224EF9396B31BFF35FC6">Foucherans, Bonnevaux-le-Prieuré, F- 25620 Ornans (France) dl. entomo @ gmx. com</mods:affiliation>
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<mods:namePart id="AD6438C14425175A90883DED9BE3798E">Daugeron, Christophe</mods:namePart>
<mods:affiliation id="3256FA19AD1CC6B01F8F5DC43648B7FD">Muséum national dHistoire naturelle, Centre national de la Recherche scientifique, Mécanismes adaptatifs et évolution, UMR 7179 MNHN-CNRS, case postale 50, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) christophe. daugeron @ mnhn. fr (corresponding author)</mods:affiliation>
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<mods:date id="FB52475FC13CA57BE8D5FE260AB9BFEE">2024</mods:date>
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<treatment id="03A987F75412FFADC4B22BFDACD0FB0F" ID-DOI="http://doi.org/10.5281/zenodo.11391425" ID-Zenodo-Dep="11391425" LSID="urn:lsid:zoobank.org:act:98CB22EC-C56D-4172-9186-D550D4CAE76A" httpUri="http://treatment.plazi.org/id/03A987F75412FFADC4B22BFDACD0FB0F" lastPageId="5" lastPageNumber="322" pageId="3" pageNumber="320">
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<taxonomicName id="4C004D625412FFABC4B22BFDA9D0FD11" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" box="[961,1241,662,688]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="species" species="cartusiana" status="sp. nov." subGenus="Empis">
<emphasis id="B974EAF35412FFABC4B22BFDA90DFD11" bold="true" box="[961,1028,662,688]" italics="true" pageId="3" pageNumber="320">Empis</emphasis>
(
<emphasis id="B974EAF35412FFABC3672BFDA95EFD11" bold="true" box="[1044,1111,662,688]" italics="true" pageId="3" pageNumber="320">Empis</emphasis>
)
<emphasis id="B974EAF35412FFABC3142BFDA9D0FD11" bold="true" box="[1127,1241,662,688]" italics="true" pageId="3" pageNumber="320">cartusiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757885412FFABC3932BFDA812FD11" box="[1248,1307,662,688]" pageId="3" pageNumber="320" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="133B2A645412FFABC34F2BDCA979FD6E" box="[1084,1136,695,719]" captionStart="FIG" captionStartId="4.[133,143,1307,1325]" captionTargetBox="[132,1455,215,1264]" captionTargetId="figure-383@4.[132,820,215,788]" captionTargetPageId="4" captionText="FIG. 1. — Empis (Empis) cartusiana n. sp.: A, male habitus; B, male hypopygium; C, female habitus. Abbreviations: see Material and methods. Scale bars: A, C, 1 mm; B, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11391417" httpUri="https://zenodo.org/record/11391417/files/figure.png" pageId="3" pageNumber="320">Fig. 1</figureCitation>
A-C)
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
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<paragraph id="8BBF36E15412FFABC45E2A4EA832FC07" blockId="3.[811,1457,805,1199]" pageId="3" pageNumber="320">
<materialsCitation id="3B683CBC5412FFABC45E2A4EA832FC07" collectingDate="2019-07-22" collectionCode="TYPE, MATERIAL" collectorName="J. Claude" country="France" county="Dent de Crolles" elevation="1762" latitude="45.31889" location="Saint-Pierre de Chartreuse" longLatPrecision="19" longitude="5.8569446" municipality="Hauts de Chartreuse" pageId="3" pageNumber="320" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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. —
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<typeStatus id="54BB88435412FFABC4812A4EA958FC9C" box="[1010,1105,805,829]" pageId="3" pageNumber="320" type="holotype">Holotype</typeStatus>
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.
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<collectingCountry id="F31776715412FFABC32D2A4EA9A9FC9C" box="[1118,1184,805,829]" name="France" pageId="3" pageNumber="320">France</collectingCountry>
</emphasis>
<specimenCount id="9D06FD685412FFABC3CA2A43A9C4FC9D" box="[1209,1229,808,828]" count="1" pageId="3" pageNumber="320" type="male"></specimenCount>
;
<location id="8EDF603A5412FFABC3AE2A4EAE6FFCF6" LSID="urn:lsid:plazi:treatment:03A987F75412FFADC4B22BFDACD0FB0F:8EDF603A5412FFABC3AE2A4EAE6FFCF6" country="France" county="Dent de Crolles" latitude="45.31889" longLatPrecision="19" longitude="5.8569446" municipality="Hauts de Chartreuse" name="Saint-Pierre de Chartreuse" pageId="3" pageNumber="320">
Saint-Pierre
<collectorName id="26F553375412FFABC2242A4EAE6FFCF6" pageId="3" pageNumber="320">de Chartreuse</collectorName>
</location>
(38),
<location id="8EDF603A5412FFABC4D62A54AEE6FCF6" LSID="urn:lsid:plazi:treatment:03A987F75412FFADC4B22BFDACD0FB0F:8EDF603A5412FFABC4D62A54AEE6FCF6" box="[933,1007,831,855]" country="France" county="Dent de Crolles" latitude="45.31889" longLatPrecision="19" longitude="5.8569446" municipality="Hauts de Chartreuse" name="Reserve" pageId="3" pageNumber="320">Réserve</location>
naturelle nationale des
<collectingMunicipality id="6BDBAC9B5412FFABC3AF2A2BA8A2FCF6" box="[1244,1451,831,855]" pageId="3" pageNumber="320">Hauts de Chartreuse</collectingMunicipality>
,
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Dent
<collectorName id="26F553375412FFABC4172A31AECCFCD3" box="[868,965,858,882]" pageId="3" pageNumber="320">de Crolles</collectorName>
</collectingCounty>
;
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,
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;
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<quantity id="4CF89B045412FFABC3DC2A31A9F0FCD0" box="[1199,1273,858,882]" metricMagnitude="3" metricUnit="m" metricValue="1.7619999999999998" pageId="3" pageNumber="320" unit="m" value="1762.0">1762 m</quantity>
a.s.l.
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;
<date id="FFBE10215412FFABC2412A31A8A5FCD3" box="[1330,1452,858,882]" pageId="3" pageNumber="320" value="2019-07-22">
<collectingDate id="EFFAE9C95412FFABC2412A31A8A5FCD3" box="[1330,1452,858,882]" pageId="3" pageNumber="320" value="2019-07-22">22.VII.2019</collectingDate>
</date>
;
<collectorName id="26F553375412FFABC4582A1EAE8CFC2D" box="[811,901,884,908]" pageId="3" pageNumber="320">J. Claude</collectorName>
leg.;
<collectingMethod id="52414EF65412FFABC4C52A1FA925FC2D" box="[950,1068,884,908]" pageId="3" pageNumber="320">Malaise trap</collectingMethod>
no. 118, subalpine meadows and moorland, nestled in a pine forest; MNHN-ED-ED10932.
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</paragraph>
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<materialsCitation id="3B683CBC5412FFABC45E2AC2A87DFC7A" collectingDate="2019-07-22" collectionCode="TYPE, MATERIAL" collectorName="J. Claude" country="France" county="Dent de Crolles" elevation="1762" latitude="45.31889" location="Saint-Pierre de Chartreuse" longLatPrecision="19" longitude="5.8569446" municipality="Hauts de Chartreuse" pageId="3" pageNumber="320" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
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<typeStatus id="54BB88435412FFABC45E2AC2AE84FC60" box="[813,909,937,961]" pageId="3" pageNumber="320" type="paratype">Paratypes</typeStatus>
</emphasis>
.
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<collectingCountry id="F31776715412FFABC4EA2AC2AED2FC60" box="[921,987,937,961]" name="France" pageId="3" pageNumber="320">France</collectingCountry>
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<specimenCount id="9D06FD685412FFABC4802AC2A91FFC61" box="[1011,1046,937,961]" count="3" pageId="3" pageNumber="320" type="male">3♂</specimenCount>
; same data as for holotype; MNHN-EDED10933, MNHN-ED-ED10934, MNHN-ED-ED10935
</materialsCitation>
<materialsCitation id="3B683CBC5412FFABC2FB2AAFA87BFBB0" collectingDate="2019-07-22" collectionCode="TYPE, MATERIAL" collectorName="J. Claude" country="France" county="Dent de Crolles" elevation="1762" latitude="45.31889" location="Saint-Pierre de Chartreuse" longLatPrecision="19" longitude="5.8569446" municipality="Hauts de Chartreuse" pageId="3" pageNumber="320" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<specimenCount id="9D06FD685412FFABC2FB2AAFA8A1FC7A" box="[1416,1448,964,988]" count="4" pageId="3" pageNumber="320" type="female">4♀</specimenCount>
; same data as for holotype; MNHN-ED-ED11399, MNHN-EDED11400, MNHN-ED-ED11401, MNHN-ED-ED11402
</materialsCitation>
<materialsCitation id="3B683CBC5412FFABC2F62A93A910FB8B" collectingDate="2019-07-22" collectionCode="TYPE, MATERIAL" collectorName="Coll. DL" country="France" county="Dent de Crolles" elevation="1762" latitude="45.31889" location="Saint-Pierre de Chartreuse" longLatPrecision="19" longitude="5.8569446" municipality="Hauts de Chartreuse" pageId="3" pageNumber="320" specimenCount="11" specimenCount-female="5" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="9D06FD685412FFABC2F62A93A8A0FBB1" box="[1413,1449,1016,1040]" count="6" pageId="3" pageNumber="320" type="male">6 ♂</specimenCount>
,
<specimenCount id="9D06FD685412FFABC45E2D78AE46FB8B" box="[813,847,1043,1067]" count="5" pageId="3" pageNumber="320" type="female">5 ♀</specimenCount>
; same data;
<collectorName id="26F553375412FFABC4B12D78A910FB8B" box="[962,1049,1043,1067]" pageId="3" pageNumber="320">Coll. DL</collectorName>
</materialsCitation>
<materialsCitation id="3B683CBC5412FFABC35F2D78A864FBE4" collectingDate="2020-06-17" collectionCode="VI" collectorName="Coll. DL" country="France" county="Dent de Crolles" elevation="1755" latitude="45.32" location="Saint-Pierre de Chartreuse" longLatPrecision="19" longitude="5.8569446" municipality="Hauts de Chartreuse" pageId="3" pageNumber="320" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D06FD685412FFABC35F2D78A946FB8B" box="[1068,1103,1043,1067]" count="1" pageId="3" pageNumber="320" type="male">1♂</specimenCount>
; same data;
<geoCoordinate id="EE3450265412FFABC3B12D78A832FB8A" box="[1218,1339,1043,1067]" degrees="45" direction="north" minutes="19" orientation="latitude" pageId="3" pageNumber="320" precision="15" seconds="12" value="45.32">45°1912”N</geoCoordinate>
,
<geoCoordinate id="EE3450265412FFABC2372D78A8A0FB8A" box="[1348,1449,1043,1067]" degrees="5" direction="east" minutes="51" orientation="longitude" pageId="3" pageNumber="320" precision="15" seconds="25" value="5.8569446">5°5125”E</geoCoordinate>
;
<elevation id="002DD1D25412FFABC45E2D46AEA7FBE4" box="[813,942,1069,1093]" metricMagnitude="3" metricUnit="m" metricValue="1.755" pageId="3" pageNumber="320" unit="m" value="1755.0">
<quantity id="4CF89B045412FFABC45E2D46AE73FBE4" box="[813,890,1069,1093]" metricMagnitude="3" metricUnit="m" metricValue="1.755" pageId="3" pageNumber="320" unit="m" value="1755.0">1755 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBE10215412FFABC4C92D45A926FBE7" box="[954,1071,1070,1094]" pageId="3" pageNumber="320" value="2020-06-17">
<collectingDate id="EFFAE9C95412FFABC4C92D45A926FBE7" box="[954,1071,1070,1094]" pageId="3" pageNumber="320" value="2020-06-17">
17.
<collectionCode id="ED11AE245412FFABC4A52D45AEFAFBE7" box="[982,1011,1070,1094]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="320">VI</collectionCode>
.2020
</collectingDate>
</date>
;
<collectingMethod id="52414EF65412FFABC3492D46A9BBFBE4" box="[1082,1202,1069,1093]" pageId="3" pageNumber="320">Malaise trap</collectingMethod>
no. 127;
<collectorName id="26F553375412FFABC2602D46A864FBE4" box="[1299,1389,1069,1093]" pageId="3" pageNumber="320">Coll. DL</collectorName>
</materialsCitation>
<materialsCitation id="3B683CBC5412FFABC2F62D45A951FBDB" collectingDate="2020-06-17" collectionCode="VI" collectorName="Coll. DL" country="France" county="Dent de Crolles" elevation="1776" latitude="45.32" location="Saint-Pierre de Chartreuse" longLatPrecision="19" longitude="5.855" municipality="Hauts de Chartreuse" pageId="3" pageNumber="320" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D06FD685412FFABC2F62D45A8A0FBE4" box="[1413,1449,1070,1094]" count="1" pageId="3" pageNumber="320" type="male">1 ♂</specimenCount>
; same data;
<geoCoordinate id="EE3450265412FFABC4EC2D23A912FBC1" box="[927,1051,1096,1120]" degrees="45" direction="north" minutes="19" orientation="latitude" pageId="3" pageNumber="320" precision="15" seconds="12" value="45.32">45°1912”N</geoCoordinate>
,
<geoCoordinate id="EE3450265412FFABC35B2D23A998FBC1" box="[1064,1169,1096,1120]" degrees="5" direction="east" minutes="51" orientation="longitude" pageId="3" pageNumber="320" precision="15" seconds="18" value="5.855">5°5118”E</geoCoordinate>
;
<elevation id="002DD1D25412FFABC3EC2D23A82FFBC1" box="[1183,1318,1096,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.7759999999999998" pageId="3" pageNumber="320" unit="m" value="1776.0">
<quantity id="4CF89B045412FFABC3EC2D23A9E6FBFE" box="[1183,1263,1096,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.7759999999999998" pageId="3" pageNumber="320" unit="m" value="1776.0">1776 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBE10215412FFABC2402D23A8A3FBC1" box="[1331,1450,1096,1120]" pageId="3" pageNumber="320" value="2020-06-17">
<collectingDate id="EFFAE9C95412FFABC2402D23A8A3FBC1" box="[1331,1450,1096,1120]" pageId="3" pageNumber="320" value="2020-06-17">
17.
<collectionCode id="ED11AE245412FFABC23C2D23A864FBC1" box="[1359,1389,1096,1120]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="320">VI</collectionCode>
.2020
</collectingDate>
</date>
;
<collectingMethod id="52414EF65412FFABC45D2D09AEACFBDB" box="[814,933,1122,1146]" pageId="3" pageNumber="320">Malaise trap</collectingMethod>
no. 128;
<collectorName id="26F553375412FFABC3732D09A951FBDB" box="[1024,1112,1122,1146]" pageId="3" pageNumber="320">Coll. DL</collectorName>
</materialsCitation>
<materialsCitation id="3B683CBC5412FFABC3182D09AEEAFB0E" collectingDate="2020-07-01" collectorName="Coll. DL" country="France" elevation="1832" latitude="45.45" location="Chapareillan" longLatPrecision="19" longitude="5.925833" municipality="Mont Granier" pageId="3" pageNumber="320" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D06FD685412FFABC3182D09A984FBDB" box="[1131,1165,1122,1146]" count="1" pageId="3" pageNumber="320" type="male">1♂</specimenCount>
;
<location id="8EDF603A5412FFABC3EA2D09A810FBDB" LSID="urn:lsid:plazi:treatment:03A987F75412FFADC4B22BFDACD0FB0F:8EDF603A5412FFABC3EA2D09A810FBDB" box="[1177,1305,1122,1146]" country="France" latitude="45.45" longLatPrecision="19" longitude="5.925833" municipality="Mont Granier" name="Chapareillan" pageId="3" pageNumber="320">Chapareillan</location>
,
<collectingMunicipality id="6BDBAC9B5412FFABC2522D08A8A5FBDB" box="[1313,1452,1122,1146]" pageId="3" pageNumber="320">Mont Granier</collectingMunicipality>
;
<geoCoordinate id="EE3450265412FFABC4582D16AE92FB34" box="[811,923,1149,1173]" degrees="45" direction="north" minutes="27" orientation="latitude" pageId="3" pageNumber="320" precision="15" seconds="0" value="45.45">45°270”N</geoCoordinate>
,
<geoCoordinate id="EE3450265412FFABC4D42D16A906FB34" box="[935,1039,1149,1173]" degrees="5" direction="east" minutes="55" orientation="longitude" pageId="3" pageNumber="320" precision="15" seconds="33" value="5.925833">5°5533”E</geoCoordinate>
;
<elevation id="002DD1D25412FFABC36D2D16A9ACFB34" box="[1054,1189,1149,1173]" metricMagnitude="3" metricUnit="m" metricValue="1.832" pageId="3" pageNumber="320" unit="m" value="1832.0">
<quantity id="4CF89B045412FFABC36D2D16A967FB35" box="[1054,1134,1149,1173]" metricMagnitude="3" metricUnit="m" metricValue="1.832" pageId="3" pageNumber="320" unit="m" value="1832.0">1832 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBE10215412FFABC3C12D16A82CFB34" box="[1202,1317,1149,1173]" pageId="3" pageNumber="320" value="2020-07-01">
<collectingDate id="EFFAE9C95412FFABC3C12D16A82CFB34" box="[1202,1317,1149,1173]" pageId="3" pageNumber="320" value="2020-07-01">1.VII.2020</collectingDate>
</date>
;
<collectingMethod id="52414EF65412FFABC2412D16A8A7FB35" box="[1330,1454,1149,1173]" pageId="3" pageNumber="320">Malaise trap</collectingMethod>
no. 125;
<collectorName id="26F553375412FFABC4F42DFCAEEAFB0E" box="[903,995,1175,1199]" pageId="3" pageNumber="320">Coll. DL</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A656A5412FFABC45E2DACAEADFAE8" pageId="3" pageNumber="320" type="etymology">
<paragraph id="8BBF36E15412FFABC45E2DACAEADFAE8" blockId="3.[813,1457,1223,1353]" pageId="3" pageNumber="320">
<smallCapsWord id="8D59A03D5412FFABC45E2DACAEACFB7C" baselines="1240,1241" box="[813,933,1223,1246]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="3" pageNumber="320">ETYMOLOGY</smallCapsWord>
. — From the Latin word
<taxonomicName id="4C004D625412FFABC3EB2DACA9E1FB7F" authorityName="Mai, J.R.Johansen &amp; Pietrasiak" authorityYear="2018" box="[1176,1256,1223,1246]" class="Cyanobacteriia" family="Oculatellaceae" genus="Cartusia" higherTaxonomySource="GBIF" kingdom="Bacteria" order="Synechococcales" pageId="3" pageNumber="320" phylum="Cyanobacteria" rank="genus">
<emphasis id="B974EAF35412FFABC3EB2DACA9E1FB7F" box="[1176,1256,1223,1246]" italics="true" pageId="3" pageNumber="320">Cartusia</emphasis>
</taxonomicName>
meaning
<emphasis id="B974EAF35412FFABC2362DACA8A1FB7E" box="[1349,1448,1223,1247]" italics="true" pageId="3" pageNumber="320">Chartreuse</emphasis>
, in reference to the Chartreuse mountain range, an Alpine massif located in the French departments of Isère and Savoie, in which the specimens were collected (between the cities of Grenoble and Chambéry).
</paragraph>
</subSubSection>
<subSubSection id="C31A656A5412FFABC45E2C0AA894FA43" pageId="3" pageNumber="320" type="diagnosis">
<paragraph id="8BBF36E15412FFABC45E2C0AA894FA43" blockId="3.[811,1457,1376,1506]" pageId="3" pageNumber="320">
<smallCapsWord id="8D59A03D5412FFABC45E2C0AAE91FAD6" baselines="1394,1395" box="[813,920,1377,1400]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="3" pageNumber="320">DIAGNOSIS</smallCapsWord>
. — Rather small greyish dusted species of the
<emphasis id="B974EAF35412FFABC2062C0AAE88FA33" italics="true" pageId="3" pageNumber="320">
<taxonomicName id="4C004D625412FFABC2062C0AA8A6FAD9" authorityName="Linnaeus" authorityYear="1758" box="[1397,1455,1377,1400]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="genus">Empis</taxonomicName>
pennipes-
</emphasis>
group of the subgenus
<taxonomicName id="4C004D625412FFABC33C2C10A84BFA32" authority="(sensu Chvala 2011)" baseAuthorityName="Chvala" baseAuthorityYear="2011" box="[1103,1346,1403,1427]" pageId="3" pageNumber="320" rank="subGenus" subGenus="Empis">
<emphasis id="B974EAF35412FFABC33C2C10A98FFA33" box="[1103,1158,1403,1426]" italics="true" pageId="3" pageNumber="320">Empis</emphasis>
(
<emphasis id="B974EAF35412FFABC3E12C17A9B6FA32" box="[1170,1215,1404,1427]" italics="true" pageId="3" pageNumber="320">sensu</emphasis>
<bibRefCitation id="EF914B105412FFABC3B72C10A835FA32" author="CHVALA M." box="[1220,1340,1403,1427]" pageId="3" pageNumber="320" pagination="173 - 193" refId="ref3813" refString="CHVALA M. 2011. - Revision of European species of the Empis (s. str.) pennipes group of species (Diptera: Empididae), with descriptions of five new species from the Mediterranean and the Caucasus. Studia Dipterologica 18: 173 - 193." type="journal article" year="2011">Chvála 2011</bibRefCitation>
)
</taxonomicName>
, with white abdominal pubescence, dark halteres, an anterodorsal tuft of black hairs at the base of male cercus; phallus well visible, long, thin, curved; wing clear in male, dark in female; legs of female with pennation.
</paragraph>
</subSubSection>
<subSubSection id="C31A656A5412FFABC45E2C91A9BDF9B3" box="[813,1204,1530,1554]" pageId="3" pageNumber="320" type="distribution">
<paragraph id="8BBF36E15412FFABC45E2C91A9BDF9B3" blockId="3.[813,1204,1530,1554]" box="[813,1204,1530,1554]" pageId="3" pageNumber="320">
<smallCapsWord id="8D59A03D5412FFABC45E2C91AEB3F9B1" baselines="1547,1548" box="[813,954,1530,1553]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="3" pageNumber="320">DISTRIBUTION</smallCapsWord>
. —
<collectingCountry id="F31776715412FFABC4952C91A923F9B0" box="[998,1066,1530,1553]" name="France" pageId="3" pageNumber="320">France</collectingCountry>
: Alps (
<figureCitation id="133B2A645412FFABC31F2C91A9AFF9B3" box="[1132,1190,1530,1554]" captionStart="FIG" captionStartId="7.[133,144,948,966]" captionTargetBox="[132,1456,94,878]" captionTargetId="figure-528@7.[130,1453,-323,1260]" captionTargetPageId="7" captionText="FIG. 3. — Map of France showing the distribution of Empis (Empis) cartusiana n. sp. (●) and Empis (Xanthempis) montsegurensis n. sp. (Ω). Background map: Sémhur, Wikimedia Commons." figureDoi="http://doi.org/10.5281/zenodo.11391423" httpUri="https://zenodo.org/record/11391423/files/figure.png" pageId="3" pageNumber="320">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A656A5412FFADC45E2F58AC7DFC8E" lastPageId="5" lastPageNumber="322" pageId="3" pageNumber="320" type="description">
<paragraph id="8BBF36E15412FFABC45E2F58AEB4F9EA" blockId="3.[811,1457,1587,1868]" box="[813,957,1587,1613]" pageId="3" pageNumber="320">
<smallCapsWord id="8D59A03D5412FFABC45E2F58AEB4F9EA" baselines="1606,1606" box="[813,957,1587,1613]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="3" pageNumber="320">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BBF36E15412FFABC45E2F39AEE7F9CC" blockId="3.[811,1457,1587,1868]" box="[813,1006,1618,1645]" pageId="3" pageNumber="320">
<emphasis id="B974EAF35412FFABC45E2F39AEE7F9CC" box="[813,1006,1618,1645]" italics="true" pageId="3" pageNumber="320">
Male (
<figureCitation id="133B2A645412FFABC4022F38AEEBF9CC" box="[881,994,1618,1645]" captionStart="FIG" captionStartId="4.[133,143,1307,1325]" captionTargetBox="[132,1455,215,1264]" captionTargetId="figure-383@4.[132,820,215,788]" captionTargetPageId="4" captionText="FIG. 1. — Empis (Empis) cartusiana n. sp.: A, male habitus; B, male hypopygium; C, female habitus. Abbreviations: see Material and methods. Scale bars: A, C, 1 mm; B, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11391417" httpUri="https://zenodo.org/record/11391417/files/figure.png" pageId="3" pageNumber="320">Fig. 1A, B</figureCitation>
).
</emphasis>
</paragraph>
<paragraph id="8BBF36E15412FFABC45E2F19A93AF8EA" blockId="3.[811,1457,1587,1868]" pageId="3" pageNumber="320">
<emphasis id="B974EAF35412FFABC45E2F19AE79F92D" bold="true" box="[813,880,1650,1676]" pageId="3" pageNumber="320">Head.</emphasis>
Occiput dull black to dusted grey, covered with many black occipital setae, a distinct row of postoculars, postgena with yellowish fine setae. Frons reduced to small triangle above antennae. Face dull black. Antennae blackish, scape as long as pedicel, postpedicel three times scape length. Proboscis blackish, labrum about twice head height, palpus black with a few black bristly-hairs.
</paragraph>
<paragraph id="8BBF36E15412FFACC45E2E1AAC1BF90D" blockId="3.[813,1456,1905,2028]" lastBlockId="4.[131,776,1427,1709]" lastPageId="4" lastPageNumber="321" pageId="3" pageNumber="320">
<emphasis id="B974EAF35412FFABC45E2E1AAE77F82A" bold="true" box="[813,894,1905,1931]" pageId="3" pageNumber="320">Thorax</emphasis>
. Dull black with grey pruinosity. Antepronotum with row of short black setae. Proepisternum and prosternum with a few rather pale yellow setae. Postpronotum with one strong, long apical seta and a few fine, short anteriors. Acrostichals biserial, rather short, absent in prescutellar depression; dorsocentrals irregularly biserial, a little longer than acrostichals, ending in two strong, long setae in prescutellar depression. Laterotergite with fan of mix black and yellowish setae. Scutellum with two strong, long apical setae, two thinner, shorter subapicals. Other strong, long setae as follows: three notopleurals, two pre- and one postsutural supraalars, one postalar, two distinct intraalars in the front of scutum on each side.
</paragraph>
<caption id="DF7F66695415FFACC7F62C70AC40FAE5" ID-DOI="http://doi.org/10.5281/zenodo.11391417" ID-Zenodo-Dep="11391417" httpUri="https://zenodo.org/record/11391417/files/figure.png" pageId="4" pageNumber="321" startId="4.[133,143,1307,1325]" targetBox="[132,1455,215,1264]" targetPageId="4" targetType="figure">
<paragraph id="8BBF36E15415FFACC7F62C70AC40FAE5" blockId="4.[132,1456,1307,1348]" pageId="4" pageNumber="321">
<smallCapsWord id="8D59A03D5415FFACC7F62C70AD94FA8D" baselines="1320,1321" box="[133,157,1307,1325]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="4" pageNumber="321">FIG</smallCapsWord>
. 1. —
<taxonomicName id="4C004D625415FFACC7A42C70ACA4FA8C" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" box="[215,429,1307,1325]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="321" phylum="Arthropoda" rank="species" species="cartusiana" status="sp. nov." subGenus="Empis">
<emphasis id="B974EAF35415FFACC7A42C70AC02FA8C" box="[215,267,1307,1325]" italics="true" pageId="4" pageNumber="321">Empis</emphasis>
(
<emphasis id="B974EAF35415FFACC6642C70AC42FA8C" box="[279,331,1307,1325]" italics="true" pageId="4" pageNumber="321">Empis</emphasis>
)
<emphasis id="B974EAF35415FFACC6242C70ACA4FA8C" box="[343,429,1307,1325]" italics="true" pageId="4" pageNumber="321">cartusiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757885415FFACC6C72C70ACE1FA8C" box="[436,488,1307,1325]" pageId="4" pageNumber="321" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B974EAF35415FFACC6822C70ACF7FA8C" bold="true" box="[497,510,1307,1325]" pageId="4" pageNumber="321">A</emphasis>
, male habitus;
<emphasis id="B974EAF35415FFACC5F02C70AF99FA8C" bold="true" box="[643,656,1307,1325]" pageId="4" pageNumber="321">B</emphasis>
, male hypopygium;
<emphasis id="B974EAF35415FFACC4332C70AE47FA8C" bold="true" box="[832,846,1307,1325]" pageId="4" pageNumber="321">C</emphasis>
, female habitus. Abbreviations: see Material and methods. Scale bars: A, C, 1 mm; B, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BBF36E15415FFACC7F72FB9A89DF92C" blockId="4.[130,777,1746,2028]" lastBlockId="4.[812,1455,1427,1677]" pageId="4" pageNumber="321">
<emphasis id="B974EAF35415FFACC7F72FB9ADBCF94D" bold="true" box="[132,181,1746,1772]" pageId="4" pageNumber="321">Legs</emphasis>
. Dark brown blackish. Fore and mid coxae with mainly pale yellow setae, hind coxa with rather strong black setae mixed with a few pale yellow setae. Fore femur with short dorsal setae, longer fine ventrals; fore tibia with dorsal rows of distinct setae especially one posterodorsal row of rather strong, longer than tibia depth setae; first four fore tarsomeres with rather strong apical setae. Mid femur with dorsal setae shorter than femur depth, one antero- and one posteroventral rows of strong, long setae except apically; mid tibia with one anteroand one posteroventral rows of rather strong, short setae, one strong, long dorsal seta at middle; first four mid tarsomeres with rather strong apical setae. Hind femur with many dorsal and ventral setae shorter or as long as femur depth; hind tibia with many ventral setae, shorter than tibia depth, dorsal row of strong, long setae mixed with many thinner, shorter other setae; first hind tarsomere somewhat swollen with distinct apical setae, remaining tarsomeres with rather short setae.
</paragraph>
<paragraph id="8BBF36E15415FFACC45E2FD9A9CDF94D" blockId="4.[813,1455,1714,1772]" pageId="4" pageNumber="321">
<emphasis id="B974EAF35415FFACC45E2FD9AE62F96D" bold="true" box="[813,875,1714,1740]" pageId="4" pageNumber="321">Wing</emphasis>
. Length:
<quantity id="4CF89B045415FFACC4A22FD9A923F96C" box="[977,1066,1714,1741]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="321" unit="mm" value="4.4">4.4 mm</quantity>
. Clear, distinct brown stigma. Sc abbreviated, A1 complete. Haltere dark.
</paragraph>
<paragraph id="8BBF36E15415FFACC45E2E7AA88DF8CA" blockId="4.[813,1456,1809,1900]" pageId="4" pageNumber="321">
<emphasis id="B974EAF35415FFACC45E2E7AAE90F88A" bold="true" box="[813,921,1809,1835]" pageId="4" pageNumber="321">Abdomen</emphasis>
. Dull to dusted greyish, with whitish setae at base. First five tergites with many rather long yellowish marginal and lateral setae; sternites with yellowish setae ventrally.
</paragraph>
<paragraph id="8BBF36E15415FFADC45E2EFAAC08FE90" blockId="4.[813,1455,1937,2028]" lastBlockId="5.[132,775,215,305]" lastPageId="5" lastPageNumber="322" pageId="4" pageNumber="321">
<emphasis id="B974EAF35415FFACC45E2EFAAECCF80A" bold="true" box="[813,965,1937,1963]" pageId="4" pageNumber="321">Hypopygium</emphasis>
. Dull black with black setae, cercus with anterodorsal fan of dense thin setae; epandrial lamella about 2.5 times higher than wide with strong, long setae on the posterior margin; hypandrium reduced to simple ventral plate; phallus yellowish brown, thin, forming a long posterior loop.
</paragraph>
<paragraph id="8BBF36E15414FFADC7F7283CAC3CFED0" blockId="5.[130,776,343,816]" box="[132,309,343,369]" pageId="5" pageNumber="322">
<emphasis id="B974EAF35414FFADC7F7283CAC3CFED0" box="[132,309,343,369]" italics="true" pageId="5" pageNumber="322">
Female (
<figureCitation id="133B2A645414FFADC7AF283CAC25FED0" box="[220,300,343,369]" captionStart="FIG" captionStartId="4.[133,143,1307,1325]" captionTargetBox="[132,1455,215,1264]" captionTargetId="figure-383@4.[132,820,215,788]" captionTargetPageId="4" captionText="FIG. 1. — Empis (Empis) cartusiana n. sp.: A, male habitus; B, male hypopygium; C, female habitus. Abbreviations: see Material and methods. Scale bars: A, C, 1 mm; B, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11391417" httpUri="https://zenodo.org/record/11391417/files/figure.png" pageId="5" pageNumber="322">Fig. 1C</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BBF36E15414FFADC7F7281CAC7DFC8E" blockId="5.[130,776,343,816]" pageId="5" pageNumber="322">Similar to male except for the usual sexual differences and the following characters: setae shorter; eyes dichoptic, all ommatidia of equal size; frons as wide as face with a few distinct setae along margin of eyes; wing darkened; legs with many pennate setae: fore femur with a few anterodorsal subpennate setae basally, fore tibia with short posterodorsal pennate setae; mid femur with long dorsal and ventral pennation, mid tibia with short dorsal pennation except at tips, short ventral pennation at base, first mid tarsomere with a few dorsal subpennate setae; hind femur with long dorsal and ventral pennation, hind tibia with rather long dorsal pennation and ventral pennation at basal half, first hind tarsomere slightly longer, less swollen with dorsal and ventral subpennate setae, more distinct dorsally.</paragraph>
</subSubSection>
<subSubSection id="C31A656A5414FFADC7F72A3DACD0FB0F" pageId="5" pageNumber="322" type="discussion">
<paragraph id="8BBF36E15414FFADC7F72A3DADE1FCCF" blockId="5.[132,777,854,1198]" box="[132,232,854,880]" pageId="5" pageNumber="322">
<smallCapsWord id="8D59A03D5414FFADC7F72A3DADE1FCCF" baselines="873,873" box="[132,232,854,880]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="5" pageNumber="322">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BBF36E15414FFADC7F72A1EACD0FB0F" blockId="5.[132,777,854,1198]" pageId="5" pageNumber="322">
In Chválas key to the
<taxonomicName id="4C004D625414FFADC6152A1DACDEFC2E" authorityName="Linnaeus" authorityYear="1758" box="[358,471,886,911]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="pennipes">
<emphasis id="B974EAF35414FFADC6152A1DACDEFC2E" box="[358,471,886,911]" italics="true" pageId="5" pageNumber="322">E. pennipes</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EF914B105414FFADC55A2A1EAFB9FC2E" author="CHVALA M." box="[553,688,885,911]" pageId="5" pageNumber="322" pagination="173 - 193" refId="ref3813" refString="CHVALA M. 2011. - Revision of European species of the Empis (s. str.) pennipes group of species (Diptera: Empididae), with descriptions of five new species from the Mediterranean and the Caucasus. Studia Dipterologica 18: 173 - 193." type="journal article" year="2011">Chvála 2011</bibRefCitation>
),
<taxonomicName id="4C004D625414FFADC5B72A1DADD8FC0E" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="cartusiana" status="n. sp.">
<emphasis id="B974EAF35414FFADC5B72A1DADD8FC0E" italics="true" pageId="5" pageNumber="322">E. cartusiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757885414FFADC7A92AFEAC1EFC0E" box="[218,279,917,943]" pageId="5" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
is near the pair
<taxonomicName id="4C004D625414FFADC6BC2AFDAFF4FC0E" authority="Meigen, 1838" authorityName="Meigen" authorityYear="1838" box="[463,765,917,943]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="rufiventris">
<emphasis id="B974EAF35414FFADC6BC2AFDAF53FC0E" box="[463,602,917,943]" italics="true" pageId="5" pageNumber="322">E. rufiventris</emphasis>
Meigen, 1838
</taxonomicName>
/
<taxonomicName id="4C004D625414FFADC7F52ADEACA4FC6E" authority="Chvala, 2011" authorityName="Chvala" authorityYear="2011" box="[134,429,949,975]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="provencalis">
<emphasis id="B974EAF35414FFADC7F52ADEAC1DFC6E" box="[134,276,949,975]" italics="true" pageId="5" pageNumber="322">E. provencalis</emphasis>
<bibRefCitation id="EF914B105414FFADC66A2ADEACA4FC6E" author="CHVALA M." box="[281,429,949,975]" pageId="5" pageNumber="322" pagination="173 - 193" refId="ref3813" refString="CHVALA M. 2011. - Revision of European species of the Empis (s. str.) pennipes group of species (Diptera: Empididae), with descriptions of five new species from the Mediterranean and the Caucasus. Studia Dipterologica 18: 173 - 193." type="journal article" year="2011">Chvála, 2011</bibRefCitation>
</taxonomicName>
, but it must be compared to the latter as both species have a mix of yellowish and black setae on the laterotergite; however the labrum of
<taxonomicName id="4C004D625414FFADC5042A9EAE0FFBAF" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" box="[631,774,1013,1038]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="cartusiana" status="n. sp.">
<emphasis id="B974EAF35414FFADC5042A9EAE0FFBAF" box="[631,774,1013,1038]" italics="true" pageId="5" pageNumber="322">E. cartusiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757885414FFADC7F72D7EADC8FB8E" box="[132,193,1045,1071]" pageId="5" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
is distinctly shorter, the females have well-developed pennation on legs and the male genitalia are different. On the other hand
<taxonomicName id="4C004D625414FFADC6412D3EACB7FBCF" authority="Langlois &amp; Daugeron, 2024" authorityName="Langlois &amp; Daugeron" authorityYear="2024" box="[306,446,1109,1134]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="cartusiana" status="n. sp.">
<emphasis id="B974EAF35414FFADC6412D3EACB7FBCF" box="[306,446,1109,1134]" italics="true" pageId="5" pageNumber="322">E. cartusiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757885414FFADC6B52D3EAF0AFBCE" box="[454,515,1109,1135]" pageId="5" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
does not correspond to any Caucasian species of the
<taxonomicName id="4C004D625414FFADC6B52D1EAF34FB2F" authorityName="Linnaeus" authorityYear="1758" box="[454,573,1141,1166]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="pennipes">
<emphasis id="B974EAF35414FFADC6B52D1EAF34FB2F" box="[454,573,1141,1166]" italics="true" pageId="5" pageNumber="322">E. pennipes</emphasis>
</taxonomicName>
group as reviewed by
<bibRefCitation id="EF914B105414FFADC7D62DFFACDAFB0F" author="KUSTOV S. Y. &amp; SHAMSHEV I. V." box="[165,467,1172,1198]" pageId="5" pageNumber="322" pagination="170 - 184" refId="ref4106" refString="KUSTOV S. Y. &amp; SHAMSHEV I. V. 2014. - A review of dance-flies of the Empis (s. str.) pennipes group of species (Diptera: Empididae) from the Caucasus, with descriptions of five new species. Caucasian Entomological Bulletin 10 (1): 170 - 184. https: // doi. org / 10.23885 / 1814 - 3326 - 2014 - 10 - 1 - 170 - 184" type="journal article" year="2014">Kustov &amp; Shamshev (2014)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>