treatments-xml/data/03/A9/87/03A987E2A973350CFF3FFEBEFBF4FEEB.xml

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<mods:title id="C01ED7E164E798675CB208109B4D36CE">Leiocassis rudicula (Teleostei: Siluriformes: Bagridae), a new catfish from eastern Borneo</mods:title>
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<mods:namePart id="5BAD680AD622AAD31F06BED1AC9DC518">Ng, Heok Hee</mods:namePart>
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<taxonomicName id="4C004D77A973350BFF3FFEBEFE5CFEC0" authority="Ng &amp; Hadiaty, 2019" authorityName="Ng &amp; Hadiaty" authorityYear="2019" box="[151,385,333,360]" class="Actinopterygii" family="Bagridae" genus="Leiocassis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="192" phylum="Chordata" rank="species" species="rudicula" status="sp. nov.">
<emphasis id="B974EAE6A973350BFF3FFEBEFE5CFEC0" bold="true" box="[151,385,333,360]" italics="true" pageId="1" pageNumber="192">Leiocassis rudicula</emphasis>
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, 
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(
<figureCitation id="133B2A71A973350BFF37FE82FF30FE23" box="[159,237,369,395]" captionStart="FIGURE 1" captionStartId="2.[151,250,1601,1626]" captionTargetBox="[151,1436,189,1567]" captionTargetId="figure@2.[151,1436,189,1567]" captionTargetPageId="2" captionText="FIGURE 1. Leiocassis rudicula, MZB 16529, holotype, 102.8 mm SL; Borneo: Kalimantan Timur, Sungai Petung Kanan." httpUri="https://zenodo.org/record/3358942/files/figure.png" pageId="1" pageNumber="192">Figs. 1</figureCitation>
, 
<figureCitation id="133B2A71A973350BFF53FE82FECBFE23" box="[251,278,369,395]" captionStart="FIGURE 2" captionStartId="3.[151,250,1960,1985]" captionTargetBox="[178,1409,166,1940]" captionTargetId="figure@3.[178,1409,166,1940]" captionTargetPageId="3" captionText="FIGURE 2. Lateral views of heads of: a. Leiocassis rudicula, holotype, MZB 16529, 102.8 mm SL; b. other congeners (L. micropogon, ZRC 41846, 143.5 mm SL illustrated). Arrow indicates concavity above orbit. Images not to scale." httpUri="https://zenodo.org/record/3358944/files/figure.png" pageId="1" pageNumber="192">2a</figureCitation>
)
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, 102.8 mm SL; 
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: 
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, 
<collectingMunicipality id="6BDBAC8EA973350BFC3CFE4AFB3AFE7B" box="[916,1255,441,467]" pageId="1" pageNumber="192">Kabupaten Kutai Kartanegara</collectingMunicipality>
, 
<location id="8EDF602FA973350BFB5AFE4AFF0DFE5F" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFB5AFE4AFF0DFE5F" county="Borneo" latitude="0.535" longLatPrecision="21" longitude="116.18166" municipality="Kabupaten Kutai Kartanegara" name="Kecamatan Tabang" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Kecamatan Tabang</location>
, 
<location id="8EDF602FA973350BFF74FE2EFE0CFE5F" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFF74FE2EFE0CFE5F" box="[220,465,477,503]" county="Borneo" latitude="0.535" longLatPrecision="21" longitude="116.18166" municipality="Kabupaten Kutai Kartanegara" name="Sungai Petung Kanan" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Sungai Petung Kanan</location>
, 
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<geoCoordinate id="EE345033A973350BFDE5FE2FFD3DFE5F" box="[589,736,476,503]" degrees="116" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="192" precision="15" seconds="54" value="116.18166">116°10′54′′E</geoCoordinate>
; 
<collectorName id="26F55322A973350BFD43FE2EFC5CFE5F" box="[747,897,477,503]" pageId="1" pageNumber="192">R.K. Hadiaty</collectorName>
, 
<collectorName id="26F55322A973350BFC23FE2EFBDBFE5F" box="[907,1030,476,503]" pageId="1" pageNumber="192">A. Mun′im</collectorName>
&amp; 
<collectorName id="26F55322A973350BFB81FE2EFB42FE5F" box="[1065,1183,477,503]" pageId="1" pageNumber="192">D. Wowor</collectorName>
, 
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<date id="FFBE1034A973350BFB02FE2EFAE8FE5F" box="[1194,1333,477,503]" pageId="1" pageNumber="192" value="2008-06-06">6 June 2008</date>
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.
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<typeStatus id="54BB8856A973350BFF6FFDF2FE9CFDB3" box="[199,321,513,539]" pageId="1" pageNumber="192" type="paratype">Paratypes</typeStatus>
.
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<collectionCode id="ED11AE31A973350BFEE7FDF2FE51FDB3" box="[335,396,513,539]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
16476 (1), 52.0 mm SL; 
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: 
<collectingRegion id="49C4F816A973350BFCB0FDF2FC37FDB3" box="[792,1002,513,539]" country="Indonesia" name="Kalimantan Timur" pageId="1" pageNumber="192">Kalimantan Timur</collectingRegion>
, 
<collectingMunicipality id="6BDBAC8EA973350BFC5FFDF2FA90FDB3" box="[1015,1357,513,539]" pageId="1" pageNumber="192">Kabupaten Kutai Kartanegara</collectingMunicipality>
, 
<location id="8EDF602FA973350BFAF1FDF2FEE7FD97" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFAF1FDF2FEE7FD97" county="Borneo" latitude="0.5341667" longLatPrecision="21" longitude="116.20611" municipality="Kabupaten Kutai Kartanegara" name="Kecamatan Tabang" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Kecamatan Tabang</location>
, 
<location id="8EDF602FA973350BFEE0FDD6FDDDFD97" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFEE0FDD6FDDDFD97" box="[328,512,549,575]" county="Borneo" latitude="0.5341667" longLatPrecision="21" longitude="116.20611" municipality="Kabupaten Kutai Kartanegara" name="Mahakam River" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Mahakam River</location>
drainage, 
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, a tributary of 
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, itself a tributary of 
<location id="8EDF602FA973350BFF3FFDBAFE93FDCB" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFF3FFDBAFE93FDCB" box="[151,334,585,611]" county="Borneo" latitude="0.5341667" longLatPrecision="21" longitude="116.20611" municipality="Kabupaten Kutai Kartanegara" name="Sungai Belayan" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Sungai Belayan</location>
, 
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; 
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, 
<collectorName id="26F55322A973350BFCBCFDBAFC4FFDCB" box="[788,914,584,611]" pageId="1" pageNumber="192">A. Mun′im</collectorName>
&amp; 
<collectorName id="26F55322A973350BFC13FDBAFBE9FDCB" box="[955,1076,585,611]" pageId="1" pageNumber="192">D. Wowor</collectorName>
, 
<collectingDate id="EFFAE9DCA973350BFBE9FDBAFB0FFDCB" box="[1089,1234,585,611]" pageId="1" pageNumber="192" value="2008-06-06">
<date id="FFBE1034A973350BFBE9FDBAFB0FFDCB" box="[1089,1234,585,611]" pageId="1" pageNumber="192" value="2008-06-06">6 June 2008</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="3B683CA9A973350BFB76FDBAFADBFD2F" ID-GBIF-Occurrence="2302318126" collectingDate="2009-04-21" collectingDateMax="2009-04-22" collectingDateMin="2009-04-21" collectionCode="MZB" collectorName="R. K. Hadiaty &amp; S. Sauri &amp; Wowor" county="Borneo" latitude="0.5341667" location="Kecamatan Tabang" longLatPrecision="21" longitude="116.20611" municipality="Kabupaten Kutai Kartanegara" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Kalimantan Timur" typeStatus="paratype">
<collectionCode id="ED11AE31A973350BFB76FDBAFAC6FDCB" box="[1246,1307,585,611]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
17480 (2), 43.8–51.0 mm SL; data as for MZB 16476; 
<collectorName id="26F55322A973350BFD39FD9EFCF5FD2F" box="[657,808,621,647]" pageId="1" pageNumber="192">R.K. Hadiaty</collectorName>
, 
<collectorName id="26F55322A973350BFC9CFD9EFC53FD2F" box="[820,910,621,647]" pageId="1" pageNumber="192">S. Sauri</collectorName>
&amp; D. 
<collectorName id="26F55322A973350BFC71FD9EFBF0FD2F" box="[985,1069,621,647]" pageId="1" pageNumber="192">Wowor</collectorName>
, 
<collectingDate id="EFFAE9DCA973350BFB91FD9EFADBFD2F" box="[1081,1286,621,647]" pageId="1" pageNumber="192" value="2009-04-21" valueMax="2009-04-22" valueMin="2009-04-21">
<date id="FFBE1034A973350BFB91FD9EFADBFD2F" box="[1081,1286,621,647]" pageId="1" pageNumber="192" value="2009-04-21" valueMax="2009-04-22" valueMin="2009-04-21">21–22 April 2009</date>
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. 
<materialsCitation id="3B683CA9A973350BFAB9FD9EFB92FD67" ID-GBIF-Occurrence="2302318128" collectingDate="2009-04-22" collectionCode="MZB" collectorName="R. K. Hadiaty &amp; S. Sauri &amp; Wowor" county="Borneo" latitude="0.5188889" location="Kecamatan Tabang" longLatPrecision="21" longitude="116.22639" municipality="Kabupaten Kutai Kartanegara" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Kalimantan Timur" typeStatus="paratype">
<collectionCode id="ED11AE31A973350BFAB9FD9EFA93FD2F" box="[1297,1358,621,647]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
17500 (1), 
<quantity id="4CF89B11A973350BFF60FD62FEF2FD03" box="[200,303,657,683]" metricMagnitude="-2" metricUnit="m" metricValue="8.37" pageId="1" pageNumber="192" unit="mm" value="83.7">83.7 mm</quantity>
SL; 
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: 
<collectingRegion id="49C4F816A973350BFE65FD62FD42FD03" box="[461,671,657,683]" country="Indonesia" name="Kalimantan Timur" pageId="1" pageNumber="192">Kalimantan Timur</collectingRegion>
, 
<collectingMunicipality id="6BDBAC8EA973350BFD04FD62FBDEFD03" box="[684,1027,657,683]" pageId="1" pageNumber="192">Kabupaten Kutai Kartanegara</collectingMunicipality>
, 
<location id="8EDF602FA973350BFBB8FD62FB31FD03" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFBB8FD62FB31FD03" box="[1040,1260,657,683]" county="Borneo" latitude="0.5188889" longLatPrecision="21" longitude="116.22639" municipality="Kabupaten Kutai Kartanegara" name="Kecamatan Tabang" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Kecamatan Tabang</location>
, 
<location id="8EDF602FA973350BFB51FD62FF3EFD67" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFB51FD62FF3EFD67" county="Borneo" latitude="0.5188889" longLatPrecision="21" longitude="116.22639" municipality="Kabupaten Kutai Kartanegara" name="Sungai Senget Kanan" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Sungai Senget Kanan</location>
, 
<geoCoordinate id="EE345033A973350BFF58FD47FE87FD67" box="[240,346,692,719]" degrees="0" direction="north" minutes="31" orientation="latitude" pageId="1" pageNumber="192" precision="15" seconds="8" value="0.5188889">0°31′8′′N</geoCoordinate>
<geoCoordinate id="EE345033A973350BFECAFD47FE2BFD67" box="[354,502,692,719]" degrees="116" direction="east" minutes="13" orientation="longitude" pageId="1" pageNumber="192" precision="15" seconds="35" value="116.22639">116°13′35′′E</geoCoordinate>
; 
<collectorName id="26F55322A973350BFDABFD46FD47FD67" box="[515,666,693,719]" pageId="1" pageNumber="192">R.K. Hadiaty</collectorName>
, 
<collectorName id="26F55322A973350BFD0FFD46FCDCFD67" box="[679,769,693,719]" pageId="1" pageNumber="192">S. Sauri</collectorName>
&amp; D. 
<collectorName id="26F55322A973350BFCE4FD46FC7DFD67" box="[844,928,693,719]" pageId="1" pageNumber="192">Wowor</collectorName>
, 
<collectingDate id="EFFAE9DCA973350BFC04FD46FB92FD67" box="[940,1103,693,719]" pageId="1" pageNumber="192" value="2009-04-22">
<date id="FFBE1034A973350BFC04FD46FB92FD67" box="[940,1103,693,719]" pageId="1" pageNumber="192" value="2009-04-22">22 April 2009</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="3B683CA9A973350BFBF3FD46FBF6FCBF" ID-GBIF-Occurrence="2302318116" collectingDate="2010-07-18" collectionCode="MZB" collectorName="R. K. Hadiaty &amp; Mulyadi &amp; D. Wowor" county="Borneo" latitude="0.58055556" location="Kecamatan Tabang" longLatPrecision="21" longitude="16.12111" municipality="Kabupaten Kutai Kartanegara" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Kalimantan Timur" typeStatus="paratype">
<collectionCode id="ED11AE31A973350BFBF3FD46FB45FD67" box="[1115,1176,693,719]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
19210 (3), 62.6–118.0 mm SL; 
<collectingCounty id="62DE4E78A973350BFF5FFD2AFE93FD5B" box="[247,334,729,755]" pageId="1" pageNumber="192">Borneo</collectingCounty>
: 
<collectingRegion id="49C4F816A973350BFEFFFD2AFDFBFD5B" box="[343,550,729,755]" country="Indonesia" name="Kalimantan Timur" pageId="1" pageNumber="192">Kalimantan Timur</collectingRegion>
, 
<collectingMunicipality id="6BDBAC8EA973350BFD98FD2AFCA2FD5B" box="[560,895,729,755]" pageId="1" pageNumber="192">Kabupaten Kutai Kartanegara</collectingMunicipality>
, 
<location id="8EDF602FA973350BFC21FD2AFBBCFD5B" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFC21FD2AFBBCFD5B" box="[905,1121,729,755]" county="Borneo" latitude="0.58055556" longLatPrecision="21" longitude="16.12111" municipality="Kabupaten Kutai Kartanegara" name="Kecamatan Tabang" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Kecamatan Tabang</location>
, 
<location id="8EDF602FA973350BFBC3FD2AFAD5FD5B" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFBC3FD2AFAD5FD5B" box="[1131,1288,729,755]" county="Borneo" latitude="0.58055556" longLatPrecision="21" longitude="16.12111" municipality="Kabupaten Kutai Kartanegara" name="Sungai Mejuk" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Sungai Mejuk</location>
at 
<location id="8EDF602FA973350BFA80FD2AFF05FCBF" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFA80FD2AFF05FCBF" county="Borneo" latitude="0.58055556" longLatPrecision="21" longitude="16.12111" municipality="Kabupaten Kutai Kartanegara" name="Desa Uma Diam" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Desa Uma Diam</location>
, 
<geoCoordinate id="EE345033A973350BFF4DFD0FFE81FCBF" box="[229,348,764,791]" degrees="0" direction="north" minutes="34" orientation="latitude" pageId="1" pageNumber="192" precision="15" seconds="50" value="0.58055556">0°34′50′′N</geoCoordinate>
<geoCoordinate id="EE345033A973350BFECDFD0FFE01FCBF" box="[357,476,764,791]" degrees="16" direction="east" minutes="7" orientation="longitude" pageId="1" pageNumber="192" precision="15" seconds="16" value="16.12111">16°7′16′′E</geoCoordinate>
; 
<collectorName id="26F55322A973350BFE41FD0EFD5DFCBF" box="[489,640,765,791]" pageId="1" pageNumber="192">R.K. Hadiaty</collectorName>
, 
<collectorName id="26F55322A973350BFD24FD0EFD36FCBF" box="[652,747,765,791]" pageId="1" pageNumber="192">Mulyadi</collectorName>
&amp; 
<collectorName id="26F55322A973350BFCB9FD0EFC55FCBF" box="[785,904,765,791]" pageId="1" pageNumber="192">D. Wowor</collectorName>
, 
<collectingDate id="EFFAE9DCA973350BFC3CFD0EFBF6FCBF" box="[916,1067,765,791]" pageId="1" pageNumber="192" value="2010-07-18">
<date id="FFBE1034A973350BFC3CFD0EFBF6FCBF" box="[916,1067,765,791]" pageId="1" pageNumber="192" value="2010-07-18">18 July 2010</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="3B683CA9A973350BFB9EFD0EFEF6FC2B" ID-GBIF-Occurrence="2302318129" collectingDate="2010-07-20" collectionCode="MZB" collectorName="R. K. Hadiaty &amp; Mulyadi &amp; D. Wowor" county="Borneo" latitude="0.41194445" location="Kecamatan Tabang" longLatPrecision="21" longitude="116.06556" municipality="Kabupaten Kutai Kartanegara" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Kalimantan Timur" typeStatus="paratype">
<collectionCode id="ED11AE31A973350BFB9EFD0EFBAEFCBF" box="[1078,1139,765,791]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
19264 (1), 
<quantity id="4CF89B11A973350BFB51FD0EFAB1FCBF" box="[1273,1388,765,791]" metricMagnitude="-1" metricUnit="m" metricValue="1.061" pageId="1" pageNumber="192" unit="mm" value="106.1">106.1 mm</quantity>
SL; 
<collectingCounty id="62DE4E78A973350BFF3FFCD2FF33FC93" box="[151,238,801,827]" pageId="1" pageNumber="192">Borneo</collectingCounty>
: 
<collectingRegion id="49C4F816A973350BFF50FCD2FE1AFC93" box="[248,455,801,827]" country="Indonesia" name="Kalimantan Timur" pageId="1" pageNumber="192">Kalimantan Timur</collectingRegion>
, 
<collectingMunicipality id="6BDBAC8EA973350BFE79FCD2FCFCFC93" box="[465,801,801,827]" pageId="1" pageNumber="192">Kabupaten Kutai Kartanegara</collectingMunicipality>
, 
<location id="8EDF602FA973350BFC82FCD2FBDEFC93" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFC82FCD2FBDEFC93" box="[810,1027,801,827]" county="Borneo" latitude="0.41194445" longLatPrecision="21" longitude="116.06556" municipality="Kabupaten Kutai Kartanegara" name="Kecamatan Tabang" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Kecamatan Tabang</location>
, second right tributary of a branch of 
<location id="8EDF602FA973350BFF3FFCB6FEEDFCF7" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFF3FFCB6FEEDFCF7" box="[151,304,837,863]" county="Borneo" latitude="0.41194445" longLatPrecision="21" longitude="116.06556" municipality="Kabupaten Kutai Kartanegara" name="Sungai Tahap" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Sungai Tahap</location>
at 
<location id="8EDF602FA973350BFEFDFCB6FDCFFCF7" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFEFDFCB6FDCFFCF7" box="[341,530,837,863]" county="Borneo" latitude="0.41194445" longLatPrecision="21" longitude="116.06556" municipality="Kabupaten Kutai Kartanegara" name="Desa Ritan Baru" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Desa Ritan Baru</location>
, in 
<location id="8EDF602FA973350BFD94FCB6FD00FCF7" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFD94FCB6FD00FCF7" box="[572,733,837,863]" county="Borneo" latitude="0.41194445" longLatPrecision="21" longitude="116.06556" municipality="Kabupaten Kutai Kartanegara" name="Batu Pek area" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur">Batu Pek area</location>
; 
<geoCoordinate id="EE345033A973350BFD41FCB7FCBCFCF7" box="[745,865,836,863]" degrees="0" direction="north" minutes="24" orientation="latitude" pageId="1" pageNumber="192" precision="15" seconds="43" value="0.41194445">0°24′43′′N</geoCoordinate>
<geoCoordinate id="EE345033A973350BFCC1FCB7FC33FCF7" box="[873,1006,836,863]" degrees="116" direction="east" minutes="3" orientation="longitude" pageId="1" pageNumber="192" precision="15" seconds="56" value="116.06556">116°3′56′′E</geoCoordinate>
; 
<collectorName id="26F55322A973350BFC53FCB6FB4CFCF7" box="[1019,1169,837,863]" pageId="1" pageNumber="192">R.K. Hadiaty</collectorName>
, 
<collectorName id="26F55322A973350BFB35FCB6FB21FCF7" box="[1181,1276,837,863]" pageId="1" pageNumber="192">Mulyadi</collectorName>
&amp; 
<collectorName id="26F55322A973350BFA8AFCB6FA44FCF7" box="[1314,1433,837,863]" pageId="1" pageNumber="192">D. Wowor</collectorName>
, 
<collectingDate id="EFFAE9DCA973350BFF3FFC9AFEF6FC2B" box="[151,299,873,899]" pageId="1" pageNumber="192" value="2010-07-20">
<date id="FFBE1034A973350BFF3FFC9AFEF6FC2B" box="[151,299,873,899]" pageId="1" pageNumber="192" value="2010-07-20">20 July 2010</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="3B683CA9A973350BFE9DFC9AFE02FC63" ID-GBIF-Occurrence="2302318127" collectingDate="2011-08-06" collectionCode="MZB" collectorName="R. K. Hadiaty &amp; S. Sauri &amp; D. Wowor" county="Borneo" latitude="0.50416666" location="Sungai Senyiur" longLatPrecision="21" longitude="116.388054" municipality="Kecamatan Muara Ancalong" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Kalimantan Timur, Kabupaten Kutai Timur" typeStatus="paratype">
<collectionCode id="ED11AE31A973350BFE9DFC9AFEAFFC2B" box="[309,370,873,899]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
20147 (2), 
<quantity id="4CF89B11A973350BFE5CFC9BFD4AFC2B" box="[500,663,872,899]" metricMagnitude="-2" metricUnit="m" metricValue="6.975" metricValueMax="8.58" metricValueMin="5.37" pageId="1" pageNumber="192" unit="mm" value="69.75" valueMax="85.8" valueMin="53.7">53.7–85.8 mm</quantity>
SL; 
<collectingCounty id="62DE4E78A973350BFD64FC9AFCFEFC2B" box="[716,803,873,899]" pageId="1" pageNumber="192">Borneo</collectingCounty>
: 
<collectingRegion id="49C4F816A973350BFC85FC9AFAC9FC2B" box="[813,1300,873,899]" isEnumeration="true" pageId="1" pageNumber="192">Kalimantan Timur, Kabupaten Kutai Timur</collectingRegion>
, 
<collectingMunicipality id="6BDBAC8EA973350BFAB6FC9AFE8AFC0F" pageId="1" pageNumber="192">Kecamatan Muara Ancalong</collectingMunicipality>
, 
<location id="8EDF602FA973350BFECAFC7EFDD2FC0F" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFECAFC7EFDD2FC0F" box="[354,527,909,935]" county="Borneo" latitude="0.50416666" longLatPrecision="21" longitude="116.388054" municipality="Kecamatan Muara Ancalong" name="Sungai Senyiur" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur, Kabupaten Kutai Timur">Sungai Senyiur</location>
in the vicinity of 
<location id="8EDF602FA973350BFD74FC7EFCA8FC0F" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFD74FC7EFCA8FC0F" box="[732,885,909,935]" county="Borneo" latitude="0.50416666" longLatPrecision="21" longitude="116.388054" municipality="Kecamatan Muara Ancalong" name="Desa Senyiur" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur, Kabupaten Kutai Timur">Desa Senyiur</location>
, 
<geoCoordinate id="EE345033A973350BFC29FC7FFC24FC0F" box="[897,1017,908,935]" degrees="0" direction="north" minutes="30" orientation="latitude" pageId="1" pageNumber="192" precision="15" seconds="15" value="0.50416666">0°30′15′′N</geoCoordinate>
<geoCoordinate id="EE345033A973350BFBA9FC7FFB49FC0F" box="[1025,1172,908,935]" degrees="116" direction="east" minutes="23" orientation="longitude" pageId="1" pageNumber="192" precision="15" seconds="17" value="116.388054">116°23′17′′E</geoCoordinate>
; 
<collectorName id="26F55322A973350BFB08FC7EFAEAFC0F" box="[1184,1335,909,935]" pageId="1" pageNumber="192">R.K. Hadiaty</collectorName>
, 
<collectorName id="26F55322A973350BFAEBFC7EFA41FC0F" box="[1347,1436,909,935]" pageId="1" pageNumber="192">S. Sauri</collectorName>
&amp; 
<collectorName id="26F55322A973350BFF1DFC42FEF1FC63" box="[181,300,945,971]" pageId="1" pageNumber="192">D. Wowor</collectorName>
, 
<collectingDate id="EFFAE9DCA973350BFE9FFC42FE02FC63" box="[311,479,945,971]" pageId="1" pageNumber="192" value="2011-08-06">
<date id="FFBE1034A973350BFE9FFC42FE02FC63" box="[311,479,945,971]" pageId="1" pageNumber="192" value="2011-08-06">6 August 2011</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="3B683CA9A973350BFE43FC42FD9FFBBB" ID-GBIF-Occurrence="2302318122" collectingDate="2011-08-07" collectionCode="MZB" collectorName="R. K. Hadiaty &amp; S. Sauri &amp; D. Wowor" county="Borneo" latitude="0.53194445" location="Sungai Senyiur" longLatPrecision="21" longitude="116.36889" municipality="Kecamatan Muara Ancalong" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Kalimantan Timur, Kabupaten Kutai Timur" typeStatus="paratype">
<collectionCode id="ED11AE31A973350BFE43FC42FDF5FC63" box="[491,552,945,971]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
20168 (1), 
<quantity id="4CF89B11A973350BFD06FC42FCCEFC63" box="[686,787,945,971]" metricMagnitude="-2" metricUnit="m" metricValue="8.23" pageId="1" pageNumber="192" unit="mm" value="82.3">82.3 mm</quantity>
SL; 
<collectingCounty id="62DE4E78A973350BFCE3FC42FC7FFC63" box="[843,930,945,971]" pageId="1" pageNumber="192">Borneo</collectingCounty>
: 
<collectingRegion id="49C4F816A973350BFC05FC42FA45FC63" box="[941,1432,945,971]" isEnumeration="true" pageId="1" pageNumber="192">Kalimantan Timur, Kabupaten Kutai Timur</collectingRegion>
, 
<collectingMunicipality id="6BDBAC8EA973350BFF3FFC26FE04FC47" box="[151,473,981,1007]" pageId="1" pageNumber="192">Kecamatan Muara Ancalong</collectingMunicipality>
, 
<location id="8EDF602FA973350BFE4CFC26FD52FC47" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFE4CFC26FD52FC47" box="[484,655,981,1007]" county="Borneo" latitude="0.53194445" longLatPrecision="21" longitude="116.36889" municipality="Kecamatan Muara Ancalong" name="Sungai Senyiur" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur, Kabupaten Kutai Timur">Sungai Senyiur</location>
in the vicinity of 
<location id="8EDF602FA973350BFCFBFC26FC36FC47" LSID="urn:lsid:plazi:treatment:03A987E2A973350CFF3FFEBEFBF4FEEB:8EDF602FA973350BFCFBFC26FC36FC47" box="[851,1003,981,1007]" county="Borneo" latitude="0.53194445" longLatPrecision="21" longitude="116.36889" municipality="Kecamatan Muara Ancalong" name="Desa Senyiur" pageId="1" pageNumber="192" stateProvince="Kalimantan Timur, Kabupaten Kutai Timur">Desa Senyiur</location>
, 
<geoCoordinate id="EE345033A973350BFC5DFC27FBB0FC47" box="[1013,1133,980,1007]" degrees="0" direction="north" minutes="31" orientation="latitude" pageId="1" pageNumber="192" precision="15" seconds="55" value="0.53194445">0°31′55′′N</geoCoordinate>
<geoCoordinate id="EE345033A973350BFBDBFC27FB25FC47" box="[1139,1272,980,1007]" degrees="116" direction="east" minutes="22" orientation="longitude" pageId="1" pageNumber="192" precision="15" seconds="8" value="116.36889">116°22′8′′E</geoCoordinate>
; 
<collectorName id="26F55322A973350BFAABFC26FA45FC47" box="[1283,1432,981,1007]" pageId="1" pageNumber="192">R.K. Hadiaty</collectorName>
, 
<collectorName id="26F55322A973350BFF3FFC0AFF2CFBBB" box="[151,241,1017,1043]" pageId="1" pageNumber="192">S. Sauri</collectorName>
&amp; 
<collectorName id="26F55322A973350BFEBFFC0AFE53FBBB" box="[279,398,1017,1043]" pageId="1" pageNumber="192">D. Wowor</collectorName>
, 
<collectingDate id="EFFAE9DCA973350BFE32FC0AFD9FFBBB" box="[410,578,1017,1043]" pageId="1" pageNumber="192" value="2011-08-07">
<date id="FFBE1034A973350BFE32FC0AFD9FFBBB" box="[410,578,1017,1043]" pageId="1" pageNumber="192" value="2011-08-07">7 August 2011</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="3B683CA9A973350BFDE6FC0AFC7FFBBB" ID-GBIF-Occurrence="2302318119" box="[590,930,1017,1043]" collectionCode="MZB" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Kalimantan Timur, Kabupaten Kutai Timur" typeStatus="paratype">
<collectionCode id="ED11AE31A973350BFDE6FC0AFD56FBBB" box="[590,651,1017,1043]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
20233 (1), 
<quantity id="4CF89B11A973350BFCB9FC0AFCABFBBB" box="[785,886,1017,1043]" metricMagnitude="-2" metricUnit="m" metricValue="4.87" pageId="1" pageNumber="192" unit="mm" value="48.7">48.7 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B683CA9A973350BFC07FC0AFB8FFBF3" ID-GBIF-Occurrence="2302318123" collectingDate="2011-08-09" collectionCode="MZB" collectorName="R. K. Hadiaty &amp; S. Sauri &amp; D. Wowor" county="Borneo" latitude="0.3025" location="Sungai Loa Katong" longLatPrecision="21" longitude="116.61083" municipality="Kecamatan Muara Ancalong" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Kalimantan Timur, Kabupaten Kutai Timur" typeStatus="paratype">
<collectionCode id="ED11AE31A973350BFC07FC0AFC31FBBB" box="[943,1004,1017,1043]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
20278 (1), 
<quantity id="4CF89B11A973350BFBDAFC0AFB0AFBBB" box="[1138,1239,1017,1043]" metricMagnitude="-2" metricUnit="m" metricValue="9.809999999999999" pageId="1" pageNumber="192" unit="mm" value="98.1">98.1 mm</quantity>
SL; 
<collectingCounty id="62DE4E78A973350BFAB8FC0AFABAFBBB" box="[1296,1383,1017,1043]" pageId="1" pageNumber="192">Borneo</collectingCounty>
: 
<collectingRegion id="49C4F816A973350BFADAFC0AFDBCFB9F" isEnumeration="true" pageId="1" pageNumber="192">Kalimantan Timur, Kabupaten Kutai Timur</collectingRegion>
, 
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in the vicinity of 
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, 
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; 
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, 
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&amp; 
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, 
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<date id="FFBE1034A973350BFC04FBB2FB8FFBF3" box="[940,1106,1089,1115]" pageId="1" pageNumber="192" value="2011-08-09">9 August 2011</date>
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. 
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<collectionCode id="ED11AE31A973350BFBF6FBB2FB46FBF3" box="[1118,1179,1089,1115]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
20296 (3), 
<quantity id="4CF89B11A973350BFAB6FBB2FF1EFBD7" metricMagnitude="-2" metricUnit="m" metricValue="7.17" metricValueMax="10.42" metricValueMin="3.92" pageId="1" pageNumber="192" unit="mm" value="71.7" valueMax="104.2" valueMin="39.2">39.2–104.2 mm</quantity>
SL; 
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: 
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in the vicinity of 
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; 
<collectorName id="26F55322A973350BFD57FB7AFC48FB0B" box="[767,917,1161,1187]" pageId="1" pageNumber="192">R.K. Hadiaty</collectorName>
, 
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&amp; 
<collectorName id="26F55322A973350BFBB6FB7AFB48FB0B" box="[1054,1173,1161,1187]" pageId="1" pageNumber="192">D. Wowor</collectorName>
, 
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<date id="FFBE1034A973350BFB08FB7AFA89FB0B" box="[1184,1364,1161,1187]" pageId="1" pageNumber="192" value="2011-08-10">10 August 2011</date>
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<collectionCode id="ED11AE31A973350BFAF7FB7AFA41FB0B" box="[1375,1436,1161,1187]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
21004 (3), 
<quantity id="4CF89B11A973350BFEBEFB5EFE66FB6F" box="[278,443,1197,1223]" metricMagnitude="-2" metricUnit="m" metricValue="8.169999999999998" metricValueMax="9.86" metricValueMin="6.4799999999999995" pageId="1" pageNumber="192" unit="mm" value="81.69999999999999" valueMax="98.6" valueMin="64.8">64.8–98.6 mm</quantity>
SL; 
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<collectingMunicipality id="6BDBAC8EA973350BFBF9FB5EFA45FB6F" box="[1105,1432,1197,1223]" pageId="1" pageNumber="192">Kecamatan Muara Ancalong</collectingMunicipality>
, 
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in the vicinity of 
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, 
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; 
<collectorName id="26F55322A973350BFC64FB22FBB8FB43" box="[972,1125,1233,1259]" pageId="1" pageNumber="192">R.K. Hadiaty</collectorName>
, 
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&amp; 
<collectorName id="26F55322A973350BFB51FB22FAAFFB43" box="[1273,1394,1233,1259]" pageId="1" pageNumber="192">D. Wowor</collectorName>
, 
<collectingDate id="EFFAE9DCA973350BFA28FB22FEF4FAA7" pageId="1" pageNumber="192" value="2011-08-10">
<date id="FFBE1034A973350BFA28FB22FEF4FAA7" pageId="1" pageNumber="192" value="2011-08-10">10 August 2011</date>
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. 
<materialsCitation id="3B683CA9A973350BFE9CFB06FD7DFAFF" ID-GBIF-Occurrence="2302318124" collectingDate="2012-08-06" collectionCode="MZB" collectorName="K. Hadiaty &amp; S. Sauri &amp; D. Wowor" county="Borneo" latitude="0.4725" location="Sungai Senyiur" longLatPrecision="21" longitude="116.39083" municipality="Kecamatan Muara Ancalong" pageId="1" pageNumber="192" specimenCount="1" stateProvince="Kalimantan Timur, Kabupaten Kutai Timur" typeStatus="paratype">
<collectionCode id="ED11AE31A973350BFE9CFB06FEACFAA7" box="[308,369,1269,1295]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="192" type="Museum">MZB</collectionCode>
21099 (2), 
<quantity id="4CF89B11A973350BFE5BFB07FD4BFAA7" box="[499,662,1268,1295]" metricMagnitude="-2" metricUnit="m" metricValue="6.775" metricValueMax="7.92" metricValueMin="5.63" pageId="1" pageNumber="192" unit="mm" value="67.75" valueMax="79.2" valueMin="56.3">56.3–79.2 mm</quantity>
SL; 
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: 
<collectingRegion id="49C4F816A973350BFC85FB06FAC9FAA7" box="[813,1300,1269,1295]" isEnumeration="true" pageId="1" pageNumber="192">Kalimantan Timur, Kabupaten Kutai Timur</collectingRegion>
, 
<collectingMunicipality id="6BDBAC8EA973350BFAB6FB06FE84FA9B" pageId="1" pageNumber="192">Kecamatan Muara Ancalong</collectingMunicipality>
, lower stretch of 
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in the vicinity of 
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, 
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;. 
<collectorName id="26F55322A973350BFAD6FAEAFF2CFAFF" pageId="1" pageNumber="192">K. Hadiaty</collectorName>
, 
<collectorName id="26F55322A973350BFF54FACEFE88FAFF" box="[252,341,1341,1367]" pageId="1" pageNumber="192">S. Sauri</collectorName>
&amp; 
<collectorName id="26F55322A973350BFED0FACEFE33FAFF" box="[376,494,1341,1367]" pageId="1" pageNumber="192">D. Wowor</collectorName>
, 
<collectingDate id="EFFAE9DCA973350BFE51FACEFD7DFAFF" box="[505,672,1341,1367]" pageId="1" pageNumber="192" value="2012-08-06">
<date id="FFBE1034A973350BFE51FACEFD7DFAFF" box="[505,672,1341,1367]" pageId="1" pageNumber="192" value="2012-08-06">6 August 2012</date>
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.
</paragraph>
<paragraph id="8BBF36F4A973350BFF6FFA93FB33F9DF" blockId="1.[151,1437,440,2016]" pageId="1" pageNumber="192">
<emphasis id="B974EAE6A973350BFF6FFA93FE9FFAD3" bold="true" box="[199,322,1376,1403]" pageId="1" pageNumber="192">Diagnosis.</emphasis>
<taxonomicName id="4C004D77A973350BFEE2FA91FDFBFAD3" authorityName="Ng &amp; Hadiaty" authorityYear="2019" box="[330,550,1377,1403]" class="Actinopterygii" family="Bagridae" genus="Leiocassis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="192" phylum="Chordata" rank="species" species="rudicula">
<emphasis id="B974EAE6A973350BFEE2FA91FDFBFAD3" box="[330,550,1377,1403]" italics="true" pageId="1" pageNumber="192">Leiocassis rudicula</emphasis>
</taxonomicName>
can be distinguished from congeners by the following unique combination of characters: distinct concavity at level of orbit in dorsolateral profile of head (
<figureCitation id="133B2A71A973350BFBBBFA76FB87FA37" box="[1043,1114,1413,1439]" captionStart="FIGURE 2" captionStartId="3.[151,250,1960,1985]" captionTargetBox="[178,1409,166,1940]" captionTargetId="figure@3.[178,1409,166,1940]" captionTargetPageId="3" captionText="FIGURE 2. Lateral views of heads of: a. Leiocassis rudicula, holotype, MZB 16529, 102.8 mm SL; b. other congeners (L. micropogon, ZRC 41846, 143.5 mm SL illustrated). Arrow indicates concavity above orbit. Images not to scale." httpUri="https://zenodo.org/record/3358944/files/figure.png" pageId="1" pageNumber="192">Fig. 2</figureCitation>
); tip of supraoccipital spine not reaching anterior nuchal plate element; smooth posterior margin of dorsal spine; presence of an irregular pale band on sides of body between dorsal and adipose fins; anal fin with narrow brown band across middle third of fin; narrow caudal-fin lobes with nearly straight posterior margins; snout length 36–40% HL; eye diameter 14–20% HL; head depth 17.6–20.0% SL; length of dorsal-fin spine 14.1–18.5% SL; length of pectoral-fin spine 15.1–17.7 %SL; dorsal-to-adipose distance 10.9–14.3% SL; body depth at anus 16.4–19.5% SL; length of adipose-fin base 18.9–24.1% SL; length of anal-fin base 18.8–22.6% SL; and caudal peduncle depth 9.1–10.9% SL.
</paragraph>
<paragraph id="8BBF36F4A973350BFF6FF973FEE7F8E7" blockId="1.[151,1437,440,2016]" pageId="1" pageNumber="192">
<emphasis id="B974EAE6A973350BFF6FF973FE84F933" bold="true" box="[199,345,1664,1691]" pageId="1" pageNumber="192">Description.</emphasis>
Morphometric data as in 
<tableCitation id="C682034FA973350BFDD3F972FD12F933" box="[635,719,1665,1691]" captionStart="TABLE 1" captionStartId="5.[151,239,154,178]" captionTargetBox="[166,1296,191,1380]" captionText="TABLE 1. Morphometric data for the type series of Leiocassis rudicula (n=9)." httpUri="http://table.plazi.org/id/DF7F667CA977350FFF3FFF69FC3DFF1A" pageId="1" pageNumber="192" tableUuid="DF7F667CA977350FFF3FFF69FC3DFF1A">Table 1</tableCitation>
. Head and body somewhat compressed, body moderately deep; dorsal profile sloping almost evenly, with distinct concavity above orbit; ventral profile almost straight. Dorsal profile rising evenly and steeply from tip of snout to origin of dorsal fin and sloping gently ventrally from origin of dorsal fin to posterior insertion of adipose fin. Ventral profile slightly convex to anal-fin base, then sloping slightly dorsally to end of caudal peduncle. Skin smooth. Lateral line complete and midlateral in position. Vertebrae 19+21=40 (1).
</paragraph>
<paragraph id="8BBF36F4A973350BFF6FF8AAFAA3F877" blockId="1.[151,1437,440,2016]" pageId="1" pageNumber="192">Bony elements of dorsal surface of head covered with thick skin. Midline of cranium with elongate fossa extending from behind snout to base of supraoccipital spine. Supraoccipital spine very short and narrow, reaching well short of nuchal plate. Eye ovoid, horizontal axis longest, with free margin; located entirely in dorsal half of head. Gill openings wide, extending from posttemporal to isthmus. First gill arch with 12 (6), 13* (2) or 15 (1) rakers.</paragraph>
<caption id="DF7F667CA9703508FF3FF9B2FAAEF9F1" box="[151,1395,1601,1626]" httpUri="https://zenodo.org/record/3358942/files/figure.png" pageId="2" pageNumber="193" startId="2.[151,250,1601,1626]" targetBox="[151,1436,189,1567]" targetPageId="2">
<paragraph id="8BBF36F4A9703508FF3FF9B2FAAEF9F1" blockId="2.[151,1395,1601,1626]" box="[151,1395,1601,1626]" pageId="2" pageNumber="193">
<emphasis id="B974EAE6A9703508FF3FF9B2FECEF9F2" bold="true" box="[151,275,1601,1626]" pageId="2" pageNumber="193">FIGURE 1.</emphasis>
<taxonomicName id="4C004D77A9703508FEB1F9B0FE02F9F2" authorityName="Ng &amp; Hadiaty" authorityYear="2019" box="[281,479,1602,1626]" class="Actinopterygii" family="Bagridae" genus="Leiocassis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="193" phylum="Chordata" rank="species" species="rudicula">
<emphasis id="B974EAE6A9703508FEB1F9B0FE02F9F2" box="[281,479,1602,1626]" italics="true" pageId="2" pageNumber="193">Leiocassis rudicula</emphasis>
</taxonomicName>
, MZB 16529, holotype, 102.8 mm SL; Borneo: Kalimantan Timur, Sungai Petung Kanan.
</paragraph>
</caption>
<paragraph id="8BBF36F4A9703508FF6FF97BFB62F89A" blockId="2.[151,1437,1672,2022]" pageId="2" pageNumber="193">Mouth inferior, snout strongly projecting. Oral teeth small and viliform, in irregular rows on all tooth-bearing surfaces. Premaxillary tooth band rounded, of equal width throughout. Dentary tooth band much narrower than premaxillary tooth band at symphysis, tapering laterally. Vomerine tooth band unpaired, continuous across midline; smoothly arched along anterior margin, tapering laterally to point extending posteriorly well past level of premaxillary band; band width broader than premaxillary band at midline, and tapering posterolaterally.</paragraph>
<paragraph id="8BBF36F4A9703508FF6FF8CFFEE3F84E" blockId="2.[151,1437,1672,2022]" pageId="2" pageNumber="193">Barbels slender, short and in four pairs. Nasal barbel extending just beyond posterior orbital margin. Maxillary barbel extending to approximately two-fifths of distance between barbel base and base of pectoral-fin spine. Inner mandibular-barbel origin closer to midline, extending to one-third of distance between barbel base and isthmus. Outer mandibular barbel originating posterolateral of inner mandibular barbel, extending to vertical through anterior orbital margin.</paragraph>
<caption id="DF7F667CA9713509FF3FF85BFAD8F84D" httpUri="https://zenodo.org/record/3358944/files/figure.png" pageId="3" pageNumber="194" startId="3.[151,250,1960,1985]" targetBox="[178,1409,166,1940]" targetPageId="3">
<paragraph id="8BBF36F4A9713509FF3FF85BFAD8F84D" blockId="3.[151,1437,1960,2021]" pageId="3" pageNumber="194">
<emphasis id="B974EAE6A9713509FF3FF85BFEC8F869" bold="true" box="[151,277,1960,1985]" pageId="3" pageNumber="194">FIGURE 2.</emphasis>
Lateral views of heads of: a. 
<taxonomicName id="4C004D77A9713509FDE7F85AFCCAF868" authorityName="Ng &amp; Hadiaty" authorityYear="2019" box="[591,791,1960,1984]" class="Actinopterygii" family="Bagridae" genus="Leiocassis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="rudicula">
<emphasis id="B974EAE6A9713509FDE7F85AFCCAF868" box="[591,791,1960,1984]" italics="true" pageId="3" pageNumber="194">Leiocassis rudicula</emphasis>
</taxonomicName>
, holotype, MZB 16529, 102.8 mm SL; b. other congeners (
<taxonomicName id="4C004D77A9713509FA20F85AFECDF84C" baseAuthorityName="Bleeker" baseAuthorityYear="1852" class="Actinopterygii" family="Bagridae" genus="Leiocassis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="micropogon">
<emphasis id="B974EAE6A9713509FA20F85AFECDF84C" italics="true" pageId="3" pageNumber="194">L. micropogon</emphasis>
</taxonomicName>
, ZRC 41846, 143.5 mm SL illustrated). Arrow indicates concavity above orbit. Images not to scale.
</paragraph>
</caption>
<caption id="DF7F667CA976350EFF3FF9FCFA4BF980" box="[151,1430,1551,1576]" httpUri="https://zenodo.org/record/3358947/files/figure.png" pageId="4" pageNumber="195" startId="4.[151,250,1551,1576]" targetBox="[152,1375,181,1526]" targetPageId="4">
<paragraph id="8BBF36F4A976350EFF3FF9FCFA4BF980" blockId="4.[151,1430,1551,1576]" box="[151,1430,1551,1576]" pageId="4" pageNumber="195">
<emphasis id="B974EAE6A976350EFF3FF9FCFECEF980" bold="true" box="[151,275,1551,1576]" pageId="4" pageNumber="195">FIGURE 3.</emphasis>
Map showing collection localities of 
<taxonomicName id="4C004D77A976350EFD3AF9E2FC85F980" authorityName="Ng &amp; Hadiaty" authorityYear="2019" box="[658,856,1552,1576]" class="Actinopterygii" family="Bagridae" genus="Leiocassis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="195" phylum="Chordata" rank="species" species="rudicula">
<emphasis id="B974EAE6A976350EFD3AF9E2FC85F980" box="[658,856,1552,1576]" italics="true" pageId="4" pageNumber="195">Leiocassis rudicula</emphasis>
</taxonomicName>
. Each symbol may represent more than a single locality.
</paragraph>
</caption>
<paragraph id="8BBF36F4A976350EFF6FF9A5FC0CF910" blockId="4.[151,1436,1622,2044]" pageId="4" pageNumber="195">Dorsal fin with spinelet, spine, and 7* (7) or 8 (2) rays. Dorsal fin margin convex, usually with anterior branch of fin rays longer than other branches. Dorsal fin spine moderately long, straight and slender, posterior edge smooth, without serrations. Nuchal plate narrow, tapering to pointed tip anteriorly.</paragraph>
<paragraph id="8BBF36F4A976350EFF6FF931FD24F88C" blockId="4.[151,1436,1622,2044]" pageId="4" pageNumber="195">Pectoral fin with stout spine, sharply pointed at tip, and 7* (4), 8 (4) or 9 (1) rays. Anterior spine margin smooth; posterior spine margin with 14–16 large serrations along entire length (serrations fewer in smaller specimens). Pectoral fin margin straight anteriorly, convex posteriorly.</paragraph>
<paragraph id="8BBF36F4A976350EFF6FF8DDFB6FF839" blockId="4.[151,1436,1622,2044]" pageId="4" pageNumber="195">Pelvic-fin origin at vertical through posterior end of dorsal-fin base, with i,5 (9) rays and slightly convex margin; tip of adpressed fin reaching anal-fin origin. Anus and urogenital openings located at vertical through middle of pelvic fin. Males with elongate genital papilla reaching to base of second or third anal-fin ray.</paragraph>
<paragraph id="8BBF36F4A976350EFF6FF869FE76F854" blockId="4.[151,1436,1622,2044]" pageId="4" pageNumber="195">Adipose fin with convex margin for entire length and with deeply-incised posterior portion; length of base approximately equal to length of anal-fin base. Anal-fin base roughly in line with adipose fin, with iv,12 (9) rays and curved posterior margin.</paragraph>
<caption id="DF7F667CA977350FFF3FFF69FC3DFF1A" ID-Table-UUID="DF7F667CA977350FFF3FFF69FC3DFF1A" box="[151,992,154,178]" httpUri="http://table.plazi.org/id/DF7F667CA977350FFF3FFF69FC3DFF1A" pageId="5" pageNumber="196" targetBox="[166,1296,191,1380]" targetIsTable="true" targetPageId="5">
<paragraph id="8BBF36F4A977350FFF3FFF69FC3DFF1A" blockId="5.[151,992,154,178]" box="[151,992,154,178]" pageId="5" pageNumber="196">
<emphasis id="B974EAE6A977350FFF3FFF69FED7FF1A" bold="true" box="[151,266,154,178]" pageId="5" pageNumber="196">TABLE 1.</emphasis>
Morphometric data for the type series of 
<taxonomicName id="4C004D77A977350FFD6FFF69FC4BFF1A" authorityName="Ng &amp; Hadiaty" authorityYear="2019" box="[711,918,154,178]" class="Actinopterygii" family="Bagridae" genus="Leiocassis" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="rudicula">
<emphasis id="B974EAE6A977350FFD6FFF69FC4BFF1A" box="[711,918,154,178]" italics="true" pageId="5" pageNumber="196">Leiocassis rudicula</emphasis>
</taxonomicName>
(n=9).
</paragraph>
</caption>
<paragraph id="8BBF36F4A977350FFDCCFF4CFB06FACC" pageId="5" pageNumber="196">
<table id="F900C454A977CAF5FF0EFF4CFACDFACC" box="[166,1296,191,1380]" gridcols="4" gridrows="32" pageId="5" pageNumber="196">
<tr id="353034B6A977CAF5FF0EFF4CFACDFF70" box="[166,1296,191,216]" gridrow="0" pageId="5" pageNumber="196" rowspan-0="1">
<th id="76E15DCAA977CAF5FDCCFF4CFD3DFF70" box="[612,736,191,216]" gridcol="1" gridrow="0" pageId="5" pageNumber="196">
<typeStatus id="54BB8856A977350FFDCCFF4CFD1FFF70" box="[612,706,191,216]" pageId="5" pageNumber="196" type="holotype">Holotype</typeStatus>
</th>
<th id="76E15DCAA977CAF5FCD4FF4CFC25FF70" box="[892,1016,191,216]" gridcol="2" gridrow="0" pageId="5" pageNumber="196">Range</th>
<th id="76E15DCAA977CAF5FB3CFF4CFACDFF70" box="[1172,1296,191,216]" gridcol="3" gridrow="0" pageId="5" pageNumber="196">Mean±SD</th>
</tr>
<tr id="353034B6A977CAF5FF0EFF10FACDFF54" box="[166,1296,227,252]" gridrow="1" pageId="5" pageNumber="196" rowspan-0="1">
<td id="76E15DCAA977CAF5FDCCFF10FD3DFF54" box="[612,736,227,252]" gridcol="1" gridrow="1" pageId="5" pageNumber="196">
<collectionCode id="ED11AE31A977350FFDCCFF17FD46FF53" box="[612,667,228,251]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="196" type="Museum">MZB</collectionCode>
16529
</td>
<td id="76E15DCAA977CAF5FCD4FF10FC25FF54" box="[892,1016,227,252]" gridcol="2" gridrow="1" pageId="5" pageNumber="196">
(
<typeStatus id="54BB8856A977350FFC2BFF10FC6DFF53" box="[899,944,227,251]" pageId="5" pageNumber="196">type</typeStatus>
series)
</td>
<td id="76E15DCAA977CAF5FB3CFF10FACDFF54" box="[1172,1296,227,252]" gridcol="3" gridrow="1" pageId="5" pageNumber="196">
<materialsCitation id="3B683CA9A977350FFB3CFF10FACDFF54" ID-GBIF-Occurrence="2413160532" box="[1172,1296,227,252]" collectionCode="MZB" pageId="5" pageNumber="196" specimenCount="2" typeStatus="holotype">
(
<typeStatus id="54BB8856A977350FFB33FF10FB15FF53" box="[1179,1224,227,251]" pageId="5" pageNumber="196">type</typeStatus>
series)
</materialsCitation>
</td>
</tr>
<tr id="353034B6A977CAF5FF0EFEF0FACDFEB4" box="[166,1296,259,284]" gridrow="2" pageId="5" pageNumber="196" rowspan-3="1">
<th id="76E15DCAA977CAF5FF0EFEF0FE39FEB4" box="[166,484,259,284]" gridcol="0" gridrow="2" pageId="5" pageNumber="196">Standard length (mm)</th>
<td id="76E15DCAA977CAF5FDCCFEF0FD3DFEB4" box="[612,736,259,284]" gridcol="1" gridrow="2" pageId="5" pageNumber="196">102.8</td>
<td id="76E15DCAA977CAF5FCD4FEF0FC25FEB4" box="[892,1016,259,284]" gridcol="2" gridrow="2" pageId="5" pageNumber="196">43.8–118.0</td>
</tr>
<tr id="353034B6A977CAF5FF0EFEDAFACDFEEA" box="[166,1296,297,322]" gridrow="3" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFEDAFACDFEEA" box="[166,1296,297,322]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="5" pageNumber="196">Percent of standard length</th>
</tr>
<tr id="353034B6A977CAF5FF0EFEBCFACDFEC0" box="[166,1296,335,360]" gridrow="4" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFEBCFE39FEC0" box="[166,484,335,360]" gridcol="0" gridrow="4" pageId="5" pageNumber="196">Predorsal length</th>
<td id="76E15DCAA977CAF5FDCCFEBCFD3DFEC0" box="[612,736,335,360]" gridcol="1" gridrow="4" pageId="5" pageNumber="196">45.3</td>
<td id="76E15DCAA977CAF5FCD4FEBCFC25FEC0" box="[892,1016,335,360]" gridcol="2" gridrow="4" pageId="5" pageNumber="196">43.1–46.8</td>
<td id="76E15DCAA977CAF5FB3CFEBCFACDFEC0" box="[1172,1296,335,360]" gridcol="3" gridrow="4" pageId="5" pageNumber="196">44.8±1.11</td>
</tr>
<tr id="353034B6A977CAF5FF0EFE86FACDFE26" box="[166,1296,373,398]" gridrow="5" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFE86FE39FE26" box="[166,484,373,398]" gridcol="0" gridrow="5" pageId="5" pageNumber="196">Preanal length</th>
<td id="76E15DCAA977CAF5FDCCFE86FD3DFE26" box="[612,736,373,398]" gridcol="1" gridrow="5" pageId="5" pageNumber="196">68.8</td>
<td id="76E15DCAA977CAF5FCD4FE86FC25FE26" box="[892,1016,373,398]" gridcol="2" gridrow="5" pageId="5" pageNumber="196">67.1–69.7</td>
<td id="76E15DCAA977CAF5FB3CFE86FACDFE26" box="[1172,1296,373,398]" gridcol="3" gridrow="5" pageId="5" pageNumber="196">68.2±0.98</td>
</tr>
<tr id="353034B6A977CAF5FF0EFE68FACDFE1C" box="[166,1296,411,436]" gridrow="6" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFE68FE39FE1C" box="[166,484,411,436]" gridcol="0" gridrow="6" pageId="5" pageNumber="196">Prepelvic length</th>
<td id="76E15DCAA977CAF5FDCCFE68FD3DFE1C" box="[612,736,411,436]" gridcol="1" gridrow="6" pageId="5" pageNumber="196">55.3</td>
<td id="76E15DCAA977CAF5FCD4FE68FC25FE1C" box="[892,1016,411,436]" gridcol="2" gridrow="6" pageId="5" pageNumber="196">53.0–58.7</td>
<td id="76E15DCAA977CAF5FB3CFE68FACDFE1C" box="[1172,1296,411,436]" gridcol="3" gridrow="6" pageId="5" pageNumber="196">55.1±1.69</td>
</tr>
<tr id="353034B6A977CAF5FF0EFE33FACDFE71" box="[166,1296,448,473]" gridrow="7" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFE33FE39FE71" box="[166,484,448,473]" gridcol="0" gridrow="7" pageId="5" pageNumber="196">Prepectoral length</th>
<td id="76E15DCAA977CAF5FDCCFE33FD3DFE71" box="[612,736,448,473]" gridcol="1" gridrow="7" pageId="5" pageNumber="196">28.1</td>
<td id="76E15DCAA977CAF5FCD4FE33FC25FE71" box="[892,1016,448,473]" gridcol="2" gridrow="7" pageId="5" pageNumber="196">27.9–32.2</td>
<td id="76E15DCAA977CAF5FB3CFE33FACDFE71" box="[1172,1296,448,473]" gridcol="3" gridrow="7" pageId="5" pageNumber="196">29.5±1.58</td>
</tr>
<tr id="353034B6A977CAF5FF0EFE15FACDFE57" box="[166,1296,486,511]" gridrow="8" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFE15FE39FE57" box="[166,484,486,511]" gridcol="0" gridrow="8" pageId="5" pageNumber="196">Length of dorsal-fin base</th>
<td id="76E15DCAA977CAF5FDCCFE15FD3DFE57" box="[612,736,486,511]" gridcol="1" gridrow="8" pageId="5" pageNumber="196">12.4</td>
<td id="76E15DCAA977CAF5FCD4FE15FC25FE57" box="[892,1016,486,511]" gridcol="2" gridrow="8" pageId="5" pageNumber="196">11.5–15.3</td>
<td id="76E15DCAA977CAF5FB3CFE15FACDFE57" box="[1172,1296,486,511]" gridcol="3" gridrow="8" pageId="5" pageNumber="196">13.0±1.26</td>
</tr>
<tr id="353034B6A977CAF5FF0EFDFFFACDFD8D" box="[166,1296,524,549]" gridrow="9" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFDFFFE39FD8D" box="[166,484,524,549]" gridcol="0" gridrow="9" pageId="5" pageNumber="196">Dorsal-spine length</th>
<td id="76E15DCAA977CAF5FDCCFDFFFD3DFD8D" box="[612,736,524,549]" gridcol="1" gridrow="9" pageId="5" pageNumber="196">16.8</td>
<td id="76E15DCAA977CAF5FCD4FDFFFC25FD8D" box="[892,1016,524,549]" gridcol="2" gridrow="9" pageId="5" pageNumber="196">14.1–18.5</td>
<td id="76E15DCAA977CAF5FB3CFDFFFACDFD8D" box="[1172,1296,524,549]" gridcol="3" gridrow="9" pageId="5" pageNumber="196">16.7±1.41</td>
</tr>
<tr id="353034B6A977CAF5FF0EFDC1FACDFDE3" box="[166,1296,562,587]" gridrow="10" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFDC1FE39FDE3" box="[166,484,562,587]" gridcol="0" gridrow="10" pageId="5" pageNumber="196">Anal-fin length</th>
<td id="76E15DCAA977CAF5FDCCFDC1FD3DFDE3" box="[612,736,562,587]" gridcol="1" gridrow="10" pageId="5" pageNumber="196">19.1</td>
<td id="76E15DCAA977CAF5FCD4FDC1FC25FDE3" box="[892,1016,562,587]" gridcol="2" gridrow="10" pageId="5" pageNumber="196">18.8–22.6</td>
<td id="76E15DCAA977CAF5FB3CFDC1FACDFDE3" box="[1172,1296,562,587]" gridcol="3" gridrow="10" pageId="5" pageNumber="196">20.0±1.24</td>
</tr>
<tr id="353034B6A977CAF5FF0EFDABFACDFDD9" box="[166,1296,600,625]" gridrow="11" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFDABFE39FDD9" box="[166,484,600,625]" gridcol="0" gridrow="11" pageId="5" pageNumber="196">Pelvic-fin length</th>
<td id="76E15DCAA977CAF5FDCCFDABFD3DFDD9" box="[612,736,600,625]" gridcol="1" gridrow="11" pageId="5" pageNumber="196">13.5</td>
<td id="76E15DCAA977CAF5FCD4FDABFC25FDD9" box="[892,1016,600,625]" gridcol="2" gridrow="11" pageId="5" pageNumber="196">12.8–14.8</td>
<td id="76E15DCAA977CAF5FB3CFDABFACDFDD9" box="[1172,1296,600,625]" gridcol="3" gridrow="11" pageId="5" pageNumber="196">13.8±0.59</td>
</tr>
<tr id="353034B6A977CAF5FF0EFD8EFACDFD3E" box="[166,1296,637,662]" gridrow="12" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFD8EFE39FD3E" box="[166,484,637,662]" gridcol="0" gridrow="12" pageId="5" pageNumber="196">Pectoral-fin length</th>
<td id="76E15DCAA977CAF5FDCCFD8EFD3DFD3E" box="[612,736,637,662]" gridcol="1" gridrow="12" pageId="5" pageNumber="196">20.1</td>
<td id="76E15DCAA977CAF5FCD4FD8EFC25FD3E" box="[892,1016,637,662]" gridcol="2" gridrow="12" pageId="5" pageNumber="196">18.2–20.5</td>
<td id="76E15DCAA977CAF5FB3CFD8EFACDFD3E" box="[1172,1296,637,662]" gridcol="3" gridrow="12" pageId="5" pageNumber="196">19.6±0.70</td>
</tr>
<tr id="353034B6A977CAF5FF0EFD50FACDFD14" box="[166,1296,675,700]" gridrow="13" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFD50FE39FD14" box="[166,484,675,700]" gridcol="0" gridrow="13" pageId="5" pageNumber="196">Pectoral-spine length</th>
<td id="76E15DCAA977CAF5FDCCFD50FD3DFD14" box="[612,736,675,700]" gridcol="1" gridrow="13" pageId="5" pageNumber="196">17.7</td>
<td id="76E15DCAA977CAF5FCD4FD50FC25FD14" box="[892,1016,675,700]" gridcol="2" gridrow="13" pageId="5" pageNumber="196">15.1–17.7</td>
<td id="76E15DCAA977CAF5FB3CFD50FACDFD14" box="[1172,1296,675,700]" gridcol="3" gridrow="13" pageId="5" pageNumber="196">16.1±0.83</td>
</tr>
<tr id="353034B6A977CAF5FF0EFD3AFACDFD4A" box="[166,1296,713,738]" gridrow="14" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFD3AFE39FD4A" box="[166,484,713,738]" gridcol="0" gridrow="14" pageId="5" pageNumber="196">Caudal-fin length</th>
<td id="76E15DCAA977CAF5FDCCFD3AFD3DFD4A" box="[612,736,713,738]" gridcol="1" gridrow="14" pageId="5" pageNumber="196">25.8</td>
<td id="76E15DCAA977CAF5FCD4FD3AFC25FD4A" box="[892,1016,713,738]" gridcol="2" gridrow="14" pageId="5" pageNumber="196">23.8–27.0</td>
<td id="76E15DCAA977CAF5FB3CFD3AFACDFD4A" box="[1172,1296,713,738]" gridcol="3" gridrow="14" pageId="5" pageNumber="196">25.6±1.08</td>
</tr>
<tr id="353034B6A977CAF5FF0EFD1CFACDFCA0" box="[166,1296,751,776]" gridrow="15" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFD1CFE39FCA0" box="[166,484,751,776]" gridcol="0" gridrow="15" pageId="5" pageNumber="196">Length of adipose-fin base</th>
<td id="76E15DCAA977CAF5FDCCFD1CFD3DFCA0" box="[612,736,751,776]" gridcol="1" gridrow="15" pageId="5" pageNumber="196">22.2</td>
<td id="76E15DCAA977CAF5FCD4FD1CFC25FCA0" box="[892,1016,751,776]" gridcol="2" gridrow="15" pageId="5" pageNumber="196">18.9–24.1</td>
<td id="76E15DCAA977CAF5FB3CFD1CFACDFCA0" box="[1172,1296,751,776]" gridcol="3" gridrow="15" pageId="5" pageNumber="196">21.9±1.86</td>
</tr>
<tr id="353034B6A977CAF5FF0EFCE6FACDFC86" box="[166,1296,789,814]" gridrow="16" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFCE6FE39FC86" box="[166,484,789,814]" gridcol="0" gridrow="16" pageId="5" pageNumber="196">Dorsal to adipose distance</th>
<td id="76E15DCAA977CAF5FDCCFCE6FD3DFC86" box="[612,736,789,814]" gridcol="1" gridrow="16" pageId="5" pageNumber="196">14.3</td>
<td id="76E15DCAA977CAF5FCD4FCE6FC25FC86" box="[892,1016,789,814]" gridcol="2" gridrow="16" pageId="5" pageNumber="196">10.9–14.3</td>
<td id="76E15DCAA977CAF5FB3CFCE6FACDFC86" box="[1172,1296,789,814]" gridcol="3" gridrow="16" pageId="5" pageNumber="196">13.0±1.33</td>
</tr>
<tr id="353034B6A977CAF5FF0EFCC9FACDFCFB" box="[166,1296,826,851]" gridrow="17" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFCC9FE39FCFB" box="[166,484,826,851]" gridcol="0" gridrow="17" pageId="5" pageNumber="196">Post-adipose distance</th>
<td id="76E15DCAA977CAF5FDCCFCC9FD3DFCFB" box="[612,736,826,851]" gridcol="1" gridrow="17" pageId="5" pageNumber="196">15.7</td>
<td id="76E15DCAA977CAF5FCD4FCC9FC25FCFB" box="[892,1016,826,851]" gridcol="2" gridrow="17" pageId="5" pageNumber="196">12.3–16.9</td>
<td id="76E15DCAA977CAF5FB3CFCC9FACDFCFB" box="[1172,1296,826,851]" gridcol="3" gridrow="17" pageId="5" pageNumber="196">15.2±1.56</td>
</tr>
<tr id="353034B6A977CAF5FF0EFC93FACDFCD1" box="[166,1296,864,889]" gridrow="18" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFC93FE39FCD1" box="[166,484,864,889]" gridcol="0" gridrow="18" pageId="5" pageNumber="196">Caudal peduncle length</th>
<td id="76E15DCAA977CAF5FDCCFC93FD3DFCD1" box="[612,736,864,889]" gridcol="1" gridrow="18" pageId="5" pageNumber="196">16.8</td>
<td id="76E15DCAA977CAF5FCD4FC93FC25FCD1" box="[892,1016,864,889]" gridcol="2" gridrow="18" pageId="5" pageNumber="196">15.4–18.6</td>
<td id="76E15DCAA977CAF5FB3CFC93FACDFCD1" box="[1172,1296,864,889]" gridcol="3" gridrow="18" pageId="5" pageNumber="196">17.1±1.09</td>
</tr>
<tr id="353034B6A977CAF5FF0EFC75FACDFC37" box="[166,1296,902,927]" gridrow="19" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFC75FE39FC37" box="[166,484,902,927]" gridcol="0" gridrow="19" pageId="5" pageNumber="196">Caudal peduncle depth</th>
<td id="76E15DCAA977CAF5FDCCFC75FD3DFC37" box="[612,736,902,927]" gridcol="1" gridrow="19" pageId="5" pageNumber="196">9.6</td>
<td id="76E15DCAA977CAF5FCD4FC75FC25FC37" box="[892,1016,902,927]" gridcol="2" gridrow="19" pageId="5" pageNumber="196">9.1–10.9</td>
<td id="76E15DCAA977CAF5FB3CFC75FACDFC37" box="[1172,1296,902,927]" gridcol="3" gridrow="19" pageId="5" pageNumber="196">9.5±0.56</td>
</tr>
<tr id="353034B6A977CAF5FF0EFC5FFACDFC6D" box="[166,1296,940,965]" gridrow="20" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFC5FFE39FC6D" box="[166,484,940,965]" gridcol="0" gridrow="20" pageId="5" pageNumber="196">Body depth at anus</th>
<td id="76E15DCAA977CAF5FDCCFC5FFD3DFC6D" box="[612,736,940,965]" gridcol="1" gridrow="20" pageId="5" pageNumber="196">18.7</td>
<td id="76E15DCAA977CAF5FCD4FC5FFC25FC6D" box="[892,1016,940,965]" gridcol="2" gridrow="20" pageId="5" pageNumber="196">16.4–19.5</td>
<td id="76E15DCAA977CAF5FB3CFC5FFACDFC6D" box="[1172,1296,940,965]" gridcol="3" gridrow="20" pageId="5" pageNumber="196">18.3±1.24</td>
</tr>
<tr id="353034B6A977CAF5FF0EFC21FACDFC43" box="[166,1296,978,1003]" gridrow="21" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFC21FE39FC43" box="[166,484,978,1003]" gridcol="0" gridrow="21" pageId="5" pageNumber="196">Head length</th>
<td id="76E15DCAA977CAF5FDCCFC21FD3DFC43" box="[612,736,978,1003]" gridcol="1" gridrow="21" pageId="5" pageNumber="196">30.9</td>
<td id="76E15DCAA977CAF5FCD4FC21FC25FC43" box="[892,1016,978,1003]" gridcol="2" gridrow="21" pageId="5" pageNumber="196">29.3–33.2</td>
<td id="76E15DCAA977CAF5FB3CFC21FACDFC43" box="[1172,1296,978,1003]" gridcol="3" gridrow="21" pageId="5" pageNumber="196">30.7±1.23</td>
</tr>
<tr id="353034B6A977CAF5FF0EFC04FACDFBB8" box="[166,1296,1015,1040]" gridrow="22" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFC04FE39FBB8" box="[166,484,1015,1040]" gridcol="0" gridrow="22" pageId="5" pageNumber="196">Head width</th>
<td id="76E15DCAA977CAF5FDCCFC04FD3DFBB8" box="[612,736,1015,1040]" gridcol="1" gridrow="22" pageId="5" pageNumber="196">18.0</td>
<td id="76E15DCAA977CAF5FCD4FC04FC25FBB8" box="[892,1016,1015,1040]" gridcol="2" gridrow="22" pageId="5" pageNumber="196">17.6–20.0</td>
<td id="76E15DCAA977CAF5FB3CFC04FACDFBB8" box="[1172,1296,1015,1040]" gridcol="3" gridrow="22" pageId="5" pageNumber="196">18.5±0.87</td>
</tr>
<tr id="353034B6A977CAF5FF0EFBEEFACDFB9E" box="[166,1296,1053,1078]" gridrow="23" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFBEEFE39FB9E" box="[166,484,1053,1078]" gridcol="0" gridrow="23" pageId="5" pageNumber="196">Head depth</th>
<td id="76E15DCAA977CAF5FDCCFBEEFD3DFB9E" box="[612,736,1053,1078]" gridcol="1" gridrow="23" pageId="5" pageNumber="196">17.2</td>
<td id="76E15DCAA977CAF5FCD4FBEEFC25FB9E" box="[892,1016,1053,1078]" gridcol="2" gridrow="23" pageId="5" pageNumber="196">17.2–20.5</td>
<td id="76E15DCAA977CAF5FB3CFBEEFACDFB9E" box="[1172,1296,1053,1078]" gridcol="3" gridrow="23" pageId="5" pageNumber="196">18.5±1.05</td>
</tr>
<tr id="353034B6A977CAF5FF0EFBB0FACDFBF4" box="[166,1296,1091,1116]" gridrow="24" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFBB0FACDFBF4" box="[166,1296,1091,1116]" colspan="4" colspanRight="3" gridcol="0" gridrow="24" pageId="5" pageNumber="196">Percent of head length</th>
</tr>
<tr id="353034B6A977CAF5FF0EFB9AFACDFB2A" box="[166,1296,1129,1154]" gridrow="25" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFB9AFE39FB2A" box="[166,484,1129,1154]" gridcol="0" gridrow="25" pageId="5" pageNumber="196">Snout length</th>
<td id="76E15DCAA977CAF5FDCCFB9AFD3DFB2A" box="[612,736,1129,1154]" gridcol="1" gridrow="25" pageId="5" pageNumber="196">38</td>
<td id="76E15DCAA977CAF5FCD4FB9AFC25FB2A" box="[892,1016,1129,1154]" gridcol="2" gridrow="25" pageId="5" pageNumber="196">36–40</td>
<td id="76E15DCAA977CAF5FB3CFB9AFACDFB2A" box="[1172,1296,1129,1154]" gridcol="3" gridrow="25" pageId="5" pageNumber="196">38±1.6</td>
</tr>
<tr id="353034B6A977CAF5FF0EFB7DFACDFB0F" box="[166,1296,1166,1191]" gridrow="26" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFB7DFE39FB0F" box="[166,484,1166,1191]" gridcol="0" gridrow="26" pageId="5" pageNumber="196">Interorbital distance</th>
<td id="76E15DCAA977CAF5FDCCFB7DFD3DFB0F" box="[612,736,1166,1191]" gridcol="1" gridrow="26" pageId="5" pageNumber="196">22</td>
<td id="76E15DCAA977CAF5FCD4FB7DFC25FB0F" box="[892,1016,1166,1191]" gridcol="2" gridrow="26" pageId="5" pageNumber="196">21–28</td>
<td id="76E15DCAA977CAF5FB3CFB7DFACDFB0F" box="[1172,1296,1166,1191]" gridcol="3" gridrow="26" pageId="5" pageNumber="196">24±2.2</td>
</tr>
<tr id="353034B6A977CAF5FF0EFB47FACDFB65" box="[166,1296,1204,1229]" gridrow="27" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFB47FE39FB65" box="[166,484,1204,1229]" gridcol="0" gridrow="27" pageId="5" pageNumber="196">Eye diameter</th>
<td id="76E15DCAA977CAF5FDCCFB47FD3DFB65" box="[612,736,1204,1229]" gridcol="1" gridrow="27" pageId="5" pageNumber="196">15</td>
<td id="76E15DCAA977CAF5FCD4FB47FC25FB65" box="[892,1016,1204,1229]" gridcol="2" gridrow="27" pageId="5" pageNumber="196">14–20</td>
<td id="76E15DCAA977CAF5FB3CFB47FACDFB65" box="[1172,1296,1204,1229]" gridcol="3" gridrow="27" pageId="5" pageNumber="196">16±2.1</td>
</tr>
<tr id="353034B6A977CAF5FF0EFB29FACDFB5B" box="[166,1296,1242,1267]" gridrow="28" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFB29FE39FB5B" box="[166,484,1242,1267]" gridcol="0" gridrow="28" pageId="5" pageNumber="196">Nasal barbel length</th>
<td id="76E15DCAA977CAF5FDCCFB29FD3DFB5B" box="[612,736,1242,1267]" gridcol="1" gridrow="28" pageId="5" pageNumber="196">26</td>
<td id="76E15DCAA977CAF5FCD4FB29FC25FB5B" box="[892,1016,1242,1267]" gridcol="2" gridrow="28" pageId="5" pageNumber="196">13–26</td>
<td id="76E15DCAA977CAF5FB3CFB29FACDFB5B" box="[1172,1296,1242,1267]" gridcol="3" gridrow="28" pageId="5" pageNumber="196">17±4.5</td>
</tr>
<tr id="353034B6A977CAF5FF0EFAF3FACDFAB1" box="[166,1296,1280,1305]" gridrow="29" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFAF3FE39FAB1" box="[166,484,1280,1305]" gridcol="0" gridrow="29" pageId="5" pageNumber="196">Maxillary barbel length</th>
<td id="76E15DCAA977CAF5FDCCFAF3FD3DFAB1" box="[612,736,1280,1305]" gridcol="1" gridrow="29" pageId="5" pageNumber="196">42</td>
<td id="76E15DCAA977CAF5FCD4FAF3FC25FAB1" box="[892,1016,1280,1305]" gridcol="2" gridrow="29" pageId="5" pageNumber="196">23–42</td>
<td id="76E15DCAA977CAF5FB3CFAF3FACDFAB1" box="[1172,1296,1280,1305]" gridcol="3" gridrow="29" pageId="5" pageNumber="196">32±6.8</td>
</tr>
<tr id="353034B6A977CAF5FF0EFAD5FACDFA97" box="[166,1296,1318,1343]" gridrow="30" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFAD5FE39FA97" box="[166,484,1318,1343]" gridcol="0" gridrow="30" pageId="5" pageNumber="196">Inner mandibular barbel length</th>
<td id="76E15DCAA977CAF5FDCCFAD5FD3DFA97" box="[612,736,1318,1343]" gridcol="1" gridrow="30" pageId="5" pageNumber="196">15</td>
<td id="76E15DCAA977CAF5FCD4FAD5FC25FA97" box="[892,1016,1318,1343]" gridcol="2" gridrow="30" pageId="5" pageNumber="196">14–20</td>
<td id="76E15DCAA977CAF5FB3CFAD5FACDFA97" box="[1172,1296,1318,1343]" gridcol="3" gridrow="30" pageId="5" pageNumber="196">17±2.0</td>
</tr>
<tr id="353034B6A977CAF5FF0EFAB8FACDFACC" box="[166,1296,1355,1380]" gridrow="31" pageId="5" pageNumber="196">
<th id="76E15DCAA977CAF5FF0EFAB8FE39FACC" box="[166,484,1355,1380]" gridcol="0" gridrow="31" pageId="5" pageNumber="196">Outer mandibular barbel length</th>
<td id="76E15DCAA977CAF5FDCCFAB8FD3DFACC" box="[612,736,1355,1380]" gridcol="1" gridrow="31" pageId="5" pageNumber="196">24</td>
<td id="76E15DCAA977CAF5FCD4FAB8FC25FACC" box="[892,1016,1355,1380]" gridcol="2" gridrow="31" pageId="5" pageNumber="196">20–30</td>
<td id="76E15DCAA977CAF5FB3CFAB8FACDFACC" box="[1172,1296,1355,1380]" gridcol="3" gridrow="31" pageId="5" pageNumber="196">24±2.8</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBF36F4A977350FFF6FFA65FA92FA7C" blockId="5.[151,1437,1430,2033]" pageId="5" pageNumber="196">Caudal peduncle moderately slender. Caudal fin deeply forked, with i,7,8,i (9) principal rays; lobes slender and pointed, with upper lobe slightly longer than lower. Procurrent rays extend only slightly anterior to fin base.</paragraph>
<paragraph id="8BBF36F4A977350FFF6FFA2DFB88F858" blockId="5.[151,1437,1430,2033]" pageId="5" pageNumber="196">
<emphasis id="B974EAE6A977350FFF6FFA2DFE92FA51" bold="true" box="[199,335,1502,1529]" pageId="5" pageNumber="196">Coloration.</emphasis>
In 70% ethanol: dorsal and lateral surfaces of head and body chocolate brown, with thin, diffuse irregular yellow band encircling dorsal half of body at nape. Ventral surfaces of head and body yellow, fading to lighter cream ventrally. Lateral line visible as thin pale midaxial stripe running along sides of body. Diffuse ovoid yellow patch present on lateral surfaces of body between nape and base of dorsal-fin spine. Lateral surfaces of body with irregular yellow band at region between dorsal and adipose fins and a pair of horizontally elongate cream patches located dorsal and ventral to midaxial stripe at caudal peduncle. Nasal and maxillary barbels chocolate brown dorsally, cream ventrally; mandibular barbels cream. Dorsal spine mostly chocolate brown in some individuals and mostly yellow in others; dorsal fin hyaline, with chocolate brown band on distal third to distal half of fin. Pectoral spine chocolate brown; pectoral fin with thin irregular chocolate brown band on proximal third of fin, and thicker chocolate brown band subdistally. Adipose fin chocolate brown, with three horizontally elongate ovoid yellow patches along distal margin: first at dorsally-sloping region of fin, second along apex of fin and third at incised posterior insertion of fin. Pelvic and anal fins with diffuse brown base and thin irregular chocolate brown band along middle third. Horizontally elongate ovoid yellow patch on dorsal and ventral procurrent caudal-fin rays. Caudal fin hyaline, with irregular chocolate brown markings on middle of each lobe and thin, subdistal W-shaped chocolate brown band along posterior edge. Color in life similar, but with strong yellowish hue.
</paragraph>
<paragraph id="8BBF36F4A974350CFF6FFF6BFBAFFF1B" blockId="6.[151,1436,152,323]" box="[199,1138,152,179]" pageId="6" pageNumber="197">
<emphasis id="B974EAE6A974350CFF6FFF6BFEBCFF1B" bold="true" box="[199,353,152,179]" pageId="6" pageNumber="197">Distribution.</emphasis>
Known from the Mahakam River drainage in eastern Borneo (
<figureCitation id="133B2A71A974350CFB88FF6AFBB9FF1B" box="[1056,1124,153,179]" captionStart="FIGURE 3" captionStartId="4.[151,250,1551,1576]" captionTargetBox="[152,1375,181,1526]" captionTargetId="figure@4.[151,1436,181,1532]" captionTargetPageId="4" captionText="FIGURE 3. Map showing collection localities of Leiocassis rudicula. Each symbol may represent more than a single locality." httpUri="https://zenodo.org/record/3358947/files/figure.png" pageId="6" pageNumber="197">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BBF36F4A974350CFF6FFF4FFAEDFF53" blockId="6.[151,1436,152,323]" pageId="6" pageNumber="197">
<emphasis id="B974EAE6A974350CFF6FFF4FFEF7FF7F" bold="true" box="[199,298,188,215]" pageId="6" pageNumber="197">Habitat.</emphasis>
<taxonomicName id="4C004D77A974350CFE99FF4DFDD1FF7F" authorityName="Ng &amp; Hadiaty" authorityYear="2019" box="[305,524,189,215]" class="Actinopterygii" family="Bagridae" genus="Leiocassis" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="rudicula">
<emphasis id="B974EAE6A974350CFE99FF4DFDD1FF7F" box="[305,524,189,215]" italics="true" pageId="6" pageNumber="197">Leiocassis rudicula</emphasis>
</taxonomicName>
is known from streams with a moderate or fast current with clear, and sometimes tannin-stained, water. The substrate in habitats it has been collected in typically consist of gravel or sand.
</paragraph>
<paragraph id="8BBF36F4A974350CFF6FFEF6FBF4FEEB" blockId="6.[151,1436,152,323]" pageId="6" pageNumber="197">
<emphasis id="B974EAE6A974350CFF6FFEF6FE93FEB7" bold="true" box="[199,334,261,287]" pageId="6" pageNumber="197">Etymology.</emphasis>
The specific epithet is the Latin diminutive for a wooden spoon. The name alludes to the distinct concavity on the dorsolateral profile of the head, which is reminiscent of a spoon.
</paragraph>
</subSubSection>
</treatment>
</document>