247 lines
28 KiB
XML
247 lines
28 KiB
XML
<document id="EA9805B49A2D453A09A64B7F5F0BD5BE" ID-DOI="10.11646/zootaxa.4012.1.1" ID-GBIF-Dataset="0b7d3317-e0a6-4491-a068-bc0378359441" ID-ISSN="1175-5326" ID-Zenodo-Dep="289354" ID-ZooBank="4DE5E609-AC90-4AA5-84D1-AA0D86B5C4DB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461224212127" checkinUser="plazi" docAuthor="Braun, Holger" docDate="2015" docId="03A987CF0556A90FFF4528AAFDB02563" docLanguage="en" docName="zt04012p032.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typophyllum vignoni Braun, 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="21" masterDocId="FF90FFB70544A91BFFD22C5EFFA42676" masterDocTitle="Little walking leaves from southeast Ecuador: biology and taxonomy of Typophyllum species (Orthoptera, Tettigoniidae, Pterochrozinae)" masterLastPageNumber="32" masterPageNumber="1" pageNumber="19" updateTime="1698604814266" updateUser="plazi">
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<mods:titleInfo id="57CA6E14E5CFCF75CB8FA70C643FB3EA">
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<mods:title id="6871FD8D847AD2DE4093754E03949302">Little walking leaves from southeast Ecuador: biology and taxonomy of Typophyllum species (Orthoptera, Tettigoniidae, Pterochrozinae)</mods:title>
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<mods:name id="6541EC9A6D8FA7E8FDEA51B2B6D76B4E" type="personal">
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<mods:namePart id="DE9AD5E3342326479E1B923209FA5022">Braun, Holger</mods:namePart>
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<mods:title id="4DA6758E9398E5EBDABCDF06E72492C1">Zootaxa</mods:title>
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<mods:part id="BAFA0CBABC4C90E88C0466A63BC49C04">
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<mods:date id="B431028B3C37A9B0E1C5A5AC069C6AB4">2015</mods:date>
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<mods:number id="51D9BD1C4E3E897A49F83873DA4B4539">4012</mods:number>
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<mods:classification id="DD4553B6EA1053E3B4335DDF7CAAE752">journal article</mods:classification>
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<mods:identifier id="5912E33F98212B0D25A2DE7AB7F72418" type="DOI">10.11646/zootaxa.4012.1.1</mods:identifier>
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<mods:identifier id="71B221323A8EDBEF42F1B28F4D6C6FC6" type="GBIF-Dataset">0b7d3317-e0a6-4491-a068-bc0378359441</mods:identifier>
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<mods:identifier id="1FD8AFBE16D11687F99DDCE1E6095125" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A987CF0556A90FFF4528AAFDB02563" ID-DOI="http://doi.org/10.5281/zenodo.6103130" ID-GBIF-Taxon="127692952" ID-Zenodo-Dep="6103130" LSID="urn:lsid:plazi:treatment:03A987CF0556A90FFF4528AAFDB02563" httpUri="http://treatment.plazi.org/id/03A987CF0556A90FFF4528AAFDB02563" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
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<subSubSection id="C31A65520556A909FF4528AAFC6C2302" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8BBF36D90556A909FF4528AAFE592378" blockId="18.[151,509,1268,1327]" box="[151,509,1268,1294]" pageId="18" pageNumber="19">
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<heading id="D0F781B50556A909FF4528AAFE592378" bold="true" box="[151,509,1268,1294]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<emphasis id="B974EACB0556A909FF4528AAFE592378" bold="true" box="[151,509,1268,1294]" pageId="18" pageNumber="19">
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<taxonomicName id="4C004D5A0556A909FF4528AAFE3C2378" box="[151,408,1268,1294]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vignoni" status="sp. nov.">
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<emphasis id="B974EACB0556A909FF4528AAFE3C2378" bold="true" box="[151,408,1268,1294]" italics="true" pageId="18" pageNumber="19">Typophyllum vignoni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24757B00556A909FE4D28AAFE592378" box="[415,509,1268,1294]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBF36D90556A909FF452948FF5D2359" blockId="18.[151,509,1268,1327]" box="[151,249,1302,1327]" pageId="18" pageNumber="19">
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(
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<figureCitation id="133B2A5C0556A909FF4D2948FF552359" box="[159,241,1302,1327]" captionStart="FIGURE 16" captionStartId="19.[151,250,1836,1858]" captionTargetBox="[160,1419,201,1799]" captionTargetId="figure@19.[151,1436,193,1814]" captionTargetPageId="19" captionText="FIGURE 16. Typophyllum vignoni sp. nov.: male holotype (tegmen length 26 mm)." httpUri="https://zenodo.org/record/289370/files/figure.png" pageId="18" pageNumber="19">Fig. 16</figureCitation>
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)
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</paragraph>
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<paragraph id="8BBF36D90556A909FF452900FC6C2302" blockId="18.[151,968,1374,1396]" box="[151,968,1374,1396]" pageId="18" pageNumber="19">
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http://lsid.speciesfile.org/urn:lsid:
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<taxonomicName id="4C004D5A0556A909FE372900FDFD2302" box="[485,601,1374,1396]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
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.speciesfile.org:TaxonName:470367
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A65520556A909FF4529FFFA9A21C4" pageId="18" pageNumber="19" type="etymology">
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<paragraph id="8BBF36D90556A909FF4529FFFF7F2074" blockId="18.[151,1437,1441,1970]" pageId="18" pageNumber="19">
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<emphasis id="B974EACB0556A909FF4529FFFEBA23CC" bold="true" box="[151,286,1441,1466]" pageId="18" pageNumber="19">Etymology.</emphasis>
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Dedicated to the French biologist Paul Vignon (
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<date id="FFBE10190556A909FC8029FFFC7223CC" box="[850,982,1441,1466]" pageId="18" pageNumber="19" value="1865" valueMax="1943">1865–1943</date>
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), specialist of “sauterelles-feuilles” and author of 50 species and varieties of “Ptérochrozées”. He is also renowned for his investigations on the Shroud of Turin.
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</paragraph>
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<paragraph id="8BBF36D90556A909FF152A53FAC0203F" blockId="18.[151,1437,1441,1970]" pageId="18" pageNumber="19">
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<emphasis id="B974EACB0556A909FF152A53FE652050" bold="true" box="[199,449,1549,1574]" pageId="18" pageNumber="19">Examined specimen.</emphasis>
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Male
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<typeStatus id="54BB887B0556A909FDC22A53FDD62050" box="[528,626,1549,1574]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
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cbt007s02, Alto Nangaritza,
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<quantity id="4CF89B3C0556A909FC192A53FBC82053" box="[971,1132,1549,1574]" metricMagnitude="3" metricUnit="m" metricValue="1.325" metricValueMax="1.35" metricValueMin="1.3" pageId="18" pageNumber="19" unit="m" value="1325.0" valueMax="1350.0" valueMin="1300.0">1300–1350 m</quantity>
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,
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<date id="FFBE10190556A909FBAE2A50FAB22050" box="[1148,1302,1549,1574]" pageId="18" pageNumber="19" value="2009-04-07">7 April 2009</date>
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, leg. Elicio Tapia, deposited in the Museo de La Plata,
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<collectingCountry id="F31776490556A909FDA82A6FFD4E203C" box="[634,746,1585,1610]" name="Argentina" pageId="18" pageNumber="19">Argentina</collectingCountry>
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[erroneous species code cbt007 refers to
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<taxonomicName id="4C004D5A0556A909FB682A6CFAF7203C" box="[1210,1363,1585,1610]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="erosifolium">
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<emphasis id="B974EACB0556A909FB682A6CFAF7203C" box="[1210,1363,1585,1610]" italics="true" pageId="18" pageNumber="19">T. erosifolium</emphasis>
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</taxonomicName>
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].
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</paragraph>
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<paragraph id="8BBF36D90556A909FF152A0BFA9A21C4" blockId="18.[151,1437,1441,1970]" pageId="18" pageNumber="19">
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<emphasis id="B974EACB0556A909FF152A0BFD842018" bold="true" box="[199,544,1621,1646]" pageId="18" pageNumber="19">Notes on two similar species:</emphasis>
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The male specimen resembles to a certain degree the unique male
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<typeStatus id="54BB887B0556A909FAC32A0BFADA2018" box="[1297,1406,1621,1646]" pageId="18" pageNumber="19" type="holotype">holotypes</typeStatus>
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of the two remaining species in the third group (aside from the preceding
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<taxonomicName id="4C004D5A0556A909FC642A24FBF420E4" box="[950,1104,1657,1682]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="erosifolium">
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<emphasis id="B974EACB0556A909FC642A24FBF420E4" box="[950,1104,1657,1682]" italics="true" pageId="18" pageNumber="19">T. erosifolium</emphasis>
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</taxonomicName>
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),
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<taxonomicName id="4C004D5A0556A909FBBA2A24FAC820E4" authority="Vignon 1926" authorityName="Vignon" authorityYear="1926" box="[1128,1388,1657,1682]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="curtum">
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<emphasis id="B974EACB0556A909FBBA2A24FB7520E7" box="[1128,1233,1658,1681]" italics="true" pageId="18" pageNumber="19">T. curtum</emphasis>
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Vignon 1926
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</taxonomicName>
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and
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<taxonomicName id="4C004D5A0556A909FF452AC0FE6920C0" authority="Walker 1870" authorityName="Walker" authorityYear="1870" box="[151,461,1693,1718]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="mutilatum">
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<emphasis id="B974EACB0556A909FF452AC0FE8D20C0" box="[151,297,1693,1718]" italics="true" pageId="18" pageNumber="19">T. mutilatum</emphasis>
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<bibRefCitation id="EF914B280556A909FEE72AC3FE6920C0" author="Walker" box="[309,461,1693,1718]" pageId="18" pageNumber="19" refString="Walker, F. (1870) Catalogue of the Specimens of Dermaptera Saltatoria in the Collection of the British Museum, 3, 425 - 604." type="journal article" year="1870">Walker 1870</bibRefCitation>
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</taxonomicName>
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, which are mutually very similar (photos in OSF). In the
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<taxonomicName id="4C004D5A0556A909FB552AC0FB5120C3" box="[1159,1269,1694,1717]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="curtum">
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<emphasis id="B974EACB0556A909FB552AC0FB5120C3" box="[1159,1269,1694,1717]" italics="true" pageId="18" pageNumber="19">T. curtum</emphasis>
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</taxonomicName>
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specimen the emargination of the costal margin of the tegmina is fairly narrow, U-shaped, with the contour distally almost parallel. In the
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<taxonomicName id="4C004D5A0556A909FE802AB8FE422088" box="[338,486,1765,1790]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="mutilatum">
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<emphasis id="B974EACB0556A909FE802AB8FE422088" box="[338,486,1765,1790]" italics="true" pageId="18" pageNumber="19">T. mutilatum</emphasis>
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</taxonomicName>
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specimen this diagnostic emargination is much broader, with distally diverging contour. In addition, the costal margin of the tegmina is almost parallel to the main veins (Subcosta+Radius), whereas in the
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<taxonomicName id="4C004D5A0556A909FE922B70FE0C2133" box="[320,424,1838,1861]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="curtum">
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<emphasis id="B974EACB0556A909FE922B70FE0C2133" box="[320,424,1838,1861]" italics="true" pageId="18" pageNumber="19">T. curtum</emphasis>
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</taxonomicName>
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male costal margin and radial vein are diverging (this is true despite that the costal area is basally slightly bent in the expanded left tegmen). So the two males described as
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<taxonomicName id="4C004D5A0556A909FB8F2B0CFB6E211F" box="[1117,1226,1874,1897]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="curtum">
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<emphasis id="B974EACB0556A909FB8F2B0CFB6E211F" box="[1117,1226,1874,1897]" italics="true" pageId="18" pageNumber="19">T. curtum</emphasis>
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</taxonomicName>
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from “Cumbasi”, probably Distrito San Roque de Cumbaza in northern
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<collectingCountry id="F31776490556A909FCD22B28FC9221FB" box="[768,822,1910,1933]" name="Peru" pageId="18" pageNumber="19">Peru</collectingCountry>
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, and
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<taxonomicName id="4C004D5A0556A909FCA62B28FBA621F8" box="[884,1026,1909,1934]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="mutilatum">
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<emphasis id="B974EACB0556A909FCA62B28FBA621F8" box="[884,1026,1909,1934]" italics="true" pageId="18" pageNumber="19">T. mutilatum</emphasis>
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</taxonomicName>
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from Tefé (in description under old name Ega), some
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<quantity id="4CF89B3C0556A909FE8D2BC4FE6621C4" box="[351,450,1945,1970]" metricMagnitude="6" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="19" unit="km" value="1300.0">1300 km</quantity>
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to the east in northwestern
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<collectingCountry id="F31776490556A909FD2A2BC7FC9B21C4" box="[760,831,1945,1970]" name="Brazil" pageId="18" pageNumber="19">Brazil</collectingCountry>
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, could in fact belong to two different species.
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</paragraph>
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</subSubSection>
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<caption id="DF7F66510557A908FF452B72FC4F2134" box="[151,1003,1836,1858]" httpUri="https://zenodo.org/record/289370/files/figure.png" pageId="19" pageNumber="20" targetBox="[160,1419,201,1799]" targetPageId="19">
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<paragraph id="8BBF36D90557A908FF452B72FC4F2134" blockId="19.[151,1003,1836,1858]" box="[151,1003,1836,1858]" pageId="19" pageNumber="20">
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<emphasis id="B974EACB0557A908FF452B72FEBB2137" bold="true" box="[151,287,1836,1858]" pageId="19" pageNumber="20">FIGURE 16.</emphasis>
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<taxonomicName id="4C004D5A0557A908FEF72B72FE5E2134" box="[293,506,1836,1858]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="vignoni" status="sp. nov.">
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<emphasis id="B974EACB0557A908FEF72B72FE5E2134" box="[293,506,1836,1858]" italics="true" pageId="19" pageNumber="20">Typophyllum vignoni</emphasis>
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</taxonomicName>
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<emphasis id="B974EACB0557A908FDD22B72FDF52137" bold="true" box="[512,593,1836,1857]" pageId="19" pageNumber="20">
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<taxonomicNameLabel id="A24757B00557A908FDD22B72FDF52137" box="[512,593,1836,1857]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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: male holotype (tegmen length 26 mm).
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</paragraph>
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</caption>
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<subSubSection id="C31A65520557A90FFF152B30FC4724BA" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
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<paragraph id="8BBF36D90557A90FFF152B30FDE62429" blockId="19.[151,1436,1902,2035]" lastBlockId="20.[151,1437,151,789]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
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<emphasis id="B974EACB0557A908FF152B30FEFD21F1" bold="true" box="[199,345,1902,1927]" pageId="19" pageNumber="20">Description.</emphasis>
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The internal tympanal chamber walls are conspicuously expanded (
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<figureCitation id="133B2A5C0557A908FB882B30FB0921F1" box="[1114,1197,1902,1927]" captionStart="FIGURE 16" captionStartId="19.[151,250,1836,1858]" captionTargetBox="[160,1419,201,1799]" captionTargetId="figure@19.[151,1436,193,1814]" captionTargetPageId="19" captionText="FIGURE 16. Typophyllum vignoni sp. nov.: male holotype (tegmen length 26 mm)." httpUri="https://zenodo.org/record/289370/files/figure.png" pageId="19" pageNumber="20">Fig. 16</figureCitation>
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B,C), as is diagnostic for the third group according to
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<bibRefCitation id="EF914B280557A908FDDB2BCCFD1921DD" author="Vignon" box="[521,701,1938,1963]" pageId="19" pageNumber="20" refString="Vignon, P. (1925 a) Essai de classification du genre Typophyllum Serville (Orth. Phasgon.). A propos des quatre especes nouvelles dont les types sont au Musee de Madrid. EOS, Revista Espanola de Entomologia, 1, 249 - 281." type="journal article" year="1925" yearSuffix="a">Vignon (1925a)</bibRefCitation>
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, a character shared with the usually somewhat bigger species of
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<taxonomicName id="4C004D5A0557A908FF452BE9FEA521B8" box="[151,257,1975,1998]" class="Insecta" family="Phaneropteridae" genus="Mimetica" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
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<emphasis id="B974EACB0557A908FF452BE9FEA521B8" box="[151,257,1975,1998]" italics="true" pageId="19" pageNumber="20">Mimetica</emphasis>
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</taxonomicName>
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. In tegmina of
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<taxonomicName id="4C004D5A0557A908FE612BE8FDE721B9" box="[435,579,1974,1999]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
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<emphasis id="B974EACB0557A908FE612BE8FDE721B9" box="[435,579,1974,1999]" italics="true" pageId="19" pageNumber="20">Typophyllum</emphasis>
|
||
</taxonomicName>
|
||
the first big cubital cell at the base of the anal area borders on the big cell located at the Rs origin, while in
|
||
<emphasis id="B974EACB0557A908FDDB2B85FDC12184" box="[521,613,2011,2034]" italics="true" pageId="19" pageNumber="20">Mimetic</emphasis>
|
||
a these cells are separated (
|
||
<bibRefCitation id="EF914B280557A908FC412B84FB8C2185" author="Vignon" box="[915,1064,2010,2035]" pageId="19" pageNumber="20" refString="Vignon, P. (1931) Recherches sur les Sauterelles-Feuilles de l'Amerique tropicale. Archives du Museum d'Histoire Naturelle, Paris, 5, 57 - 212." type="journal article" year="1931">Vignon 1931</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF914B280557A908FBE62B84FB172185" author="Beier" box="[1076,1203,2010,2035]" pageId="19" pageNumber="20" refString="Beier, M. (1960) Orthoptera, Tettigoniidae, Pseudophyllinae II. Das Tierreich, 74, 1 - 396." type="journal article" year="1960">Beier 1960</bibRefCitation>
|
||
), or touch in a single point in males. In the male found in the Alto Nangaritza valley these cells share a common border along
|
||
<quantity id="4CF89B3C0550A90FFAFA2CC9FA2526D9" box="[1320,1409,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="mm" value="2.5">2.5 mm</quantity>
|
||
. It is considerably bigger than the
|
||
<taxonomicName id="4C004D5A0550A90FFDD52CE3FDD026A2" box="[519,628,189,212]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="curtum">
|
||
<emphasis id="B974EACB0550A90FFDD52CE3FDD026A2" box="[519,628,189,212]" italics="true" pageId="20" pageNumber="21">T. curtum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C004D5A0550A90FFD662CE3FCE126A3" box="[692,837,188,213]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mutilatum">
|
||
<emphasis id="B974EACB0550A90FFD662CE3FCE126A3" box="[692,837,188,213]" italics="true" pageId="20" pageNumber="21">T. mutilatum</emphasis>
|
||
</taxonomicName>
|
||
specimens: tegmen length
|
||
<quantity id="4CF89B3C0550A90FFB562CE2FB7226A2" box="[1156,1238,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="20" pageNumber="21" unit="mm" value="26.0">26 mm</quantity>
|
||
vs.
|
||
<quantity id="4CF89B3C0550A90FFAD92CE2FA3926A2" box="[1291,1437,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.575" metricValueMax="1.7" metricValueMin="1.45" pageId="20" pageNumber="21" unit="mm" value="15.75" valueMax="17.0" valueMin="14.5">14.5–17 mm</quantity>
|
||
(
|
||
<bibRefCitation id="EF914B280550A90FFF4C2C81FEE6268E" author="Vignon" box="[158,322,223,248]" pageId="20" pageNumber="21" refString="Vignon, P. (1925 a) Essai de classification du genre Typophyllum Serville (Orth. Phasgon.). A propos des quatre especes nouvelles dont les types sont au Musee de Madrid. EOS, Revista Espanola de Entomologia, 1, 249 - 281." type="journal article" year="1925" yearSuffix="a">Vignon 1925a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF914B280550A90FFE822C81FE69268E" author="Beier" box="[336,461,223,248]" pageId="20" pageNumber="21" refString="Beier, M. (1960) Orthoptera, Tettigoniidae, Pseudophyllinae II. Das Tierreich, 74, 1 - 396." type="journal article" year="1960">Beier 1960</bibRefCitation>
|
||
—the “17 lines” in the original description of
|
||
<taxonomicName id="4C004D5A0550A90FFC332C81FBF7268E" box="[993,1107,223,248]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mutilatum">
|
||
<emphasis id="B974EACB0550A90FFC332C81FBF7268E" box="[993,1107,223,248]" italics="true" pageId="20" pageNumber="21">mutilatum</emphasis>
|
||
</taxonomicName>
|
||
must be wrong according to detailed measurements by Vignon, who apparently inspected this specimen). The most obvious difference to these two species and all other
|
||
<taxonomicName id="4C004D5A0550A90FFE652D79FDE32736" box="[439,583,295,320]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B974EACB0550A90FFE652D79FDE32736" box="[439,583,295,320]" italics="true" pageId="20" pageNumber="21">Typophyllum</emphasis>
|
||
</taxonomicName>
|
||
males are the two emarginations in the distal half of the anal margin of the tegmina (
|
||
<figureCitation id="133B2A5C0550A90FFED12D12FEFC2713" box="[259,344,332,357]" captionStart="FIGURE 16" captionStartId="19.[151,250,1836,1858]" captionTargetBox="[160,1419,201,1799]" captionTargetId="figure@19.[151,1436,193,1814]" captionTargetPageId="19" captionText="FIGURE 16. Typophyllum vignoni sp. nov.: male holotype (tegmen length 26 mm)." httpUri="https://zenodo.org/record/289370/files/figure.png" pageId="20" pageNumber="21">Fig. 16</figureCitation>
|
||
C). Only the male of the otherwise very different
|
||
<taxonomicName id="4C004D5A0550A90FFC4A2D13FBAB2713" box="[920,1039,332,357]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bolivari">
|
||
<emphasis id="B974EACB0550A90FFC4A2D13FBAB2713" box="[920,1039,332,357]" italics="true" pageId="20" pageNumber="21">T. bolivarí</emphasis>
|
||
</taxonomicName>
|
||
has there also deep emarginations (photos of
|
||
<typeStatus id="54BB887B0550A90FFECB2D31FEDF27FE" box="[281,379,367,392]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
|
||
and additional males in OSF). But the contour of this part of the tegmina as well as the development of its tip (pointed in contrast to
|
||
<taxonomicName id="4C004D5A0550A90FFD5E2DCBFD7F27DA" box="[652,731,405,428]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="curtum">
|
||
<emphasis id="B974EACB0550A90FFD5E2DCBFD7F27DA" box="[652,731,405,428]" italics="true" pageId="20" pageNumber="21">curtum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C004D5A0550A90FFCC32DCAFC2727DB" box="[785,899,404,429]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mutilatum">
|
||
<emphasis id="B974EACB0550A90FFCC32DCAFC2727DB" box="[785,899,404,429]" italics="true" pageId="20" pageNumber="21">mutilatum</emphasis>
|
||
</taxonomicName>
|
||
) could be variable, as is shown here for females of
|
||
<taxonomicName id="4C004D5A0550A90FFF672DE7FE9D27A6" box="[181,313,441,464]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="egregium">
|
||
<emphasis id="B974EACB0550A90FFF672DE7FE9D27A6" box="[181,313,441,464]" italics="true" pageId="20" pageNumber="21">T. egregium</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C004D5A0550A90FFE952DE7FE1027A6" box="[327,436,441,464]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="morrisi" status="sp. nov.">
|
||
<emphasis id="B974EACB0550A90FFE952DE7FE1027A6" box="[327,436,441,464]" italics="true" pageId="20" pageNumber="21">T. morrisi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B974EACB0550A90FFE692DE9FDB127A6" bold="true" box="[443,533,439,464]" pageId="20" pageNumber="21">
|
||
<taxonomicNameLabel id="A24757B00550A90FFE692DE9FDB127A6" box="[443,533,439,464]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C004D5A0550A90FFD992DE7FD4027A6" box="[587,740,439,464]" class="Insecta" family="Phaneropteridae" genus="Typophyllum" kingdom="Animalia" order="Orthoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="erosifolium">
|
||
<emphasis id="B974EACB0550A90FFD992DE7FD4027A6" box="[587,740,439,464]" italics="true" pageId="20" pageNumber="21">T. erosifolium</emphasis>
|
||
</taxonomicName>
|
||
. However, no such variation was found in males. The legs are considerably longer and more slender as in other species, and the spines on fore and middle femora are much more separated. The speculum on the base of the right tegmen is fully transparent. Spines on prosternum are very small, much shorter than the width of the tip of the maxillary palp, meso- and metasternum bear spines that are slightly longer than this comparative measure.
|
||
</paragraph>
|
||
<paragraph id="8BBF36D90550A90FFF152E32FA2E24DE" blockId="20.[151,1437,151,789]" pageId="20" pageNumber="21">
|
||
<emphasis id="B974EACB0550A90FFF152E32FEEB24F3" bold="true" box="[199,335,620,645]" pageId="20" pageNumber="21">Coloration.</emphasis>
|
||
The only specimen is dark brown with light brown legs and antennae, a light spot behind each eye continues into a thin light anterio-ventral margin of the lateral lobe of the pronotum. The palps are light greenish.
|
||
</paragraph>
|
||
<paragraph id="8BBF36D90550A90FFF152EEAFC4724BA" blockId="20.[151,1437,151,789]" box="[199,995,692,717]" pageId="20" pageNumber="21">
|
||
<emphasis id="B974EACB0550A90FFF152EEAFED924BB" bold="true" box="[199,381,692,717]" pageId="20" pageNumber="21">Measurements.</emphasis>
|
||
Tegmen length in male
|
||
<quantity id="4CF89B3C0550A90FFD5E2EEAFD7924BA" box="[652,733,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="20" pageNumber="21" unit="mm" value="26.0">26 mm</quantity>
|
||
, hind femora
|
||
<quantity id="4CF89B3C0550A90FFCAA2EEAFC7B24BA" box="[888,991,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="20" pageNumber="21" unit="mm" value="18.5">18.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A65520550A90FFF152E89FDB02563" pageId="20" pageNumber="21" type="distribution">
|
||
<paragraph id="8BBF36D90550A90FFF152E89FDB02563" blockId="20.[151,1437,151,789]" pageId="20" pageNumber="21">
|
||
<emphasis id="B974EACB0550A90FFF152E89FEC42486" bold="true" box="[199,352,727,752]" pageId="20" pageNumber="21">Distribution.</emphasis>
|
||
Only known from
|
||
<typeStatus id="54BB887B0550A90FFD902E86FDD62486" box="[578,626,728,752]" pageId="20" pageNumber="21">type</typeStatus>
|
||
locality on the Cordillera del Condor near the border with
|
||
<collectingCountry id="F31776490550A90FFAF42E86FAF82499" box="[1318,1372,728,751]" name="Peru" pageId="20" pageNumber="21">Peru</collectingCountry>
|
||
, Alto Nangaritza, above Las Orquídeas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |