142 lines
17 KiB
XML
142 lines
17 KiB
XML
<document id="06DFD06FCB032C607F446E9EDE5C572D" ID-DOI="10.1649/1176.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4924356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623349980529" checkinUser="carolina" docAuthor="Liebherr, James K." docDate="2009" docId="03A987B8FFFBEF2F628FFACFFE3DFD8A" docLanguage="en" docName="TheColeopBull.63.4.383-411.pdf.imf" docOrigin="The Coleopterists Bulletin 63 (4)" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Gnathaphanus picipes" docType="treatment" docVersion="4" lastPageNumber="404" masterDocId="FF90FFC0FFEFEF3A632AFFC5FFEAFFE5" masterDocTitle="Native And Alien Carabidae (Coleoptera) Share Lanai, An Ecologically Devastated Island" masterLastPageNumber="411" masterPageNumber="383" pageNumber="403" updateTime="1698962403354" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="401A2E22CE1116D27D7703E64508B840" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B2C1FE83D53A4C3C07101C36145ABA89">
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<mods:title id="DA526D0EE9F1D8F026EF270F24FF3A69">Native And Alien Carabidae (Coleoptera) Share Lanai, An Ecologically Devastated Island</mods:title>
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</mods:titleInfo>
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<mods:name id="0B6191F3B559F20EEBAD58CA109D5FC6" type="personal">
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<mods:role id="430147166570BAD3BD7947C0A700A3FB">
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<mods:roleTerm id="47F194281F63CEF4E3B1F335DF2644B4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9C6429DDE3DDDED99BB7EA3E5CB32BB2">Liebherr, James K.</mods:namePart>
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<mods:affiliation id="0E6BA271CB64B2D025A02AF177799F9B">Department of Entomology, John H. and Anna B. Comstock Hall Cornell University Ithaca, NY 14853 - 2601, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="AC45040B6A30BA8256AF3C87F01AA658" type="email">JKL5@cornell.edu</mods:nameIdentifier>
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<mods:typeOfResource id="812B4FC1FC07E63C94B8F95F21F2DDA6">text</mods:typeOfResource>
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<mods:titleInfo id="233DA90E1D0A21BAFBB283FB1E2F4ED1">
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<mods:title id="2F9E06E83A7478F25637240E2AE2402B">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="A857CD4E167AFB33BA74B86DF78BE85B">
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<mods:date id="B05E82EC554FC5535C6F5C55892DF6AD">2009</mods:date>
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<mods:detail id="0595CA2431C9E2AD8BC3537D34B5C6B9" type="pubDate">
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<mods:number id="48D9B28EAB51EC43874378A18D634CF6">2009-12-29</mods:number>
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</mods:detail>
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<mods:detail id="57DB2E24ADB5075047D8D3F24F525814" type="volume">
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<mods:number id="3E851D20F6A783071B0DBA8A2D975212">63</mods:number>
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<mods:detail id="74F701C55E09BDA9C8F31A5020B65841" type="issue">
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<mods:number id="F665939EEF6729320C42DFE5EF61759E">4</mods:number>
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<mods:extent id="3C6B17F1F429403356E4B67B30B70747" unit="page">
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<mods:start id="518B9B6BA76AFBB53855BBC1F495D6E8">383</mods:start>
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<mods:end id="FCF7E73DCD7BE3233E80243D93894362">411</mods:end>
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<mods:classification id="80DD523DD27C1FA9D50736BBE591E419">journal article</mods:classification>
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<mods:identifier id="83CF5CBF649D14797A1377B243B76F90" type="DOI">10.1649/1176.1</mods:identifier>
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<mods:identifier id="85035C29B7DEDDE59B2E57BE461D5260" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="AA855991C272C1528FBA3A97252B4F9A" type="Zenodo-Dep">4924356</mods:identifier>
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<treatment id="03A987B8FFFBEF2F628FFACFFE3DFD8A" ID-DOI="http://doi.org/10.5281/zenodo.4924678" ID-Zenodo-Dep="4924678" LSID="urn:lsid:plazi:treatment:03A987B8FFFBEF2F628FFACFFE3DFD8A" httpUri="http://treatment.plazi.org/id/03A987B8FFFBEF2F628FFACFFE3DFD8A" lastPageId="21" lastPageNumber="404" pageId="20" pageNumber="403">
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<subSubSection id="C31A6525FFFBEF2E628FFACFFD1DFAC5" box="[421,759,1290,1312]" pageId="20" pageNumber="403" type="nomenclature">
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<paragraph id="8BBF36AEFFFBEF2E628FFACFFD1DFAC5" blockId="20.[421,759,1290,1340]" box="[421,759,1290,1312]" pageId="20" pageNumber="403">
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<heading id="D0F781C2FFFBEF2E628FFACFFD1DFAC5" box="[421,759,1290,1312]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="403" reason="2">
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<taxonomicName id="4C004D2DFFFBEF2E628FFACFFD1DFAC5" ID-CoL="3GL8Y" authority="(MacLeay)" baseAuthorityName="MacLeay" box="[421,759,1290,1312]" class="Insecta" family="Carabidae" genus="Gnathaphanus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="403" phylum="Arthropoda" rank="species" species="picipes">
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<emphasis id="B974EABCFFFBEF2E628FFACFFD1DFAC5" bold="true" box="[421,759,1290,1312]" pageId="20" pageNumber="403">
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<emphasis id="B974EABCFFFBEF2E628FFACFFD94FAC5" bold="true" box="[421,638,1290,1312]" italics="true" pageId="20" pageNumber="403">Gnathaphanus picipes</emphasis>
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(MacLeay)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A6525FFFBEF2E6120FAE3FD7BFAD9" box="[522,657,1318,1340]" pageId="20" pageNumber="403" type="description">
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<paragraph id="8BBF36AEFFFBEF2E6120FAE3FD7BFAD9" blockId="20.[421,759,1290,1340]" box="[522,657,1318,1340]" pageId="20" pageNumber="403">
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(
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<figureCitation id="133B2A2BFFFBEF2E6138FAE3FD8BFAD9" box="[530,609,1318,1340]" captionStart="Fig" captionStartId="17.[167,198,1308,1327]" captionTargetBox="[172,949,194,1286]" captionTargetId="figure-66@17.[171,949,194,1286]" captionTargetPageId="17" captionText="Fig. 6. Known distributions of carabid beetle species on Lanai. A. Blackburnia lanaihalensis. B. B. depressa. C. B. filipes. D. Metacolpodes buchanani. E. Gnathaphanus picipes (small circles), G. picipes plus G. multipunctatus (larger bullseye). F. Perigona nigriceps." figureDoi="http://doi.org/10.5281/zenodo.4924370" httpUri="https://zenodo.org/record/4924370/files/figure.png" pageId="20" pageNumber="403">Fig. 6E</figureCitation>
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,
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<figureCitation id="133B2A2BFFFBEF2E6147FAE3FD60FAD9" box="[621,650,1318,1340]" captionStart="Fig" captionStartId="19.[167,198,1396,1415]" captionTargetBox="[172,948,195,1371]" captionTargetId="figure-1@19.[171,950,194,1374]" captionTargetPageId="19" captionText="Fig. 7. Non-native carabid beetle species known from Lanai (vignettes represent actual body size). A. Metacolpodes buchanani. B. Gnathaphanus picipes. C. G. multipunctatus. D. Perigona nigriceps." figureDoi="http://doi.org/10.5281/zenodo.4924374" httpUri="https://zenodo.org/record/4924374/files/figure.png" pageId="20" pageNumber="403">7B</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A6525FFFBEF2F63EDFA94FE3DFD8A" lastPageId="21" lastPageNumber="404" pageId="20" pageNumber="403" type="diagnosis">
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<paragraph id="8BBF36AEFFFBEF2E63EDFA94FD17FA09" blockId="20.[175,1005,1361,1570]" pageId="20" pageNumber="403">
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<emphasis id="B974EABCFFFBEF2E63EDFA94FEC5FA82" bold="true" box="[199,303,1361,1383]" pageId="20" pageNumber="403">Diagnosis.</emphasis>
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Easily distinguished among Hawaiian
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<taxonomicName id="4C004D2DFFFBEF2E61EFFA94FCC2FA82" box="[709,808,1361,1383]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="403" phylum="Arthropoda" rank="tribe" tribe="Harpalini">Harpalini</taxonomicName>
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by the presence of five to eight dorsal elytral setae in the third elytral interval, two to three of the setae situated very near the elytral apex, in combination with a glabrous fifth elytral stria. As in
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<taxonomicName id="4C004D2DFFFBEF2E6245FA64FDCBFA53" box="[367,545,1441,1462]" class="Insecta" family="Carabidae" genus="Gnathaphanus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="403" phylum="Arthropoda" rank="species" species="multipunctatus">
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<emphasis id="B974EABCFFFBEF2E6245FA64FDCBFA53" box="[367,545,1441,1462]" italics="true" pageId="20" pageNumber="403">G. multipunctatus</emphasis>
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</taxonomicName>
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, the tibiae and tarsi are flavous, however, in contrast to that species the femora are darker, brunneous, and concolorous with the body venter. Standardized body length
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<quantity id="4CF89B4BFFFBEF2E615BFA13FD13FA09" box="[625,761,1494,1516]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" metricValueMax="1.1199999999999999" metricValueMin="0.94" pageId="20" pageNumber="403" unit="mm" value="10.3" valueMax="11.2" valueMin="9.4">9.4–11.2 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BBF36AEFFFBEF2F63EDFA37FDB9FEBA" blockId="20.[175,1005,1361,1570]" lastBlockId="21.[146,975,194,623]" lastPageId="21" lastPageNumber="404" pageId="20" pageNumber="403">
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<emphasis id="B974EABCFFFBEF2E63EDFA37FE67F9ED" bold="true" box="[199,397,1522,1544]" pageId="20" pageNumber="403">Lanai Distribution.</emphasis>
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This Australian and New Guinean species (
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<bibRefCitation id="EF914B5FFFFBEF2E6050FA37FF0BF9C7" author="Darlington" pageId="20" pageNumber="403" pagination="1 - 253" refId="ref13828" refString="Darlington, P. J., Jr. 1968. The carabid beetles of New Guinea, Part III. Harpalinae (continued): Perigonini to Pseudomorphini. Bulletin of the Museum of Comparative Zoology 137: 1 - 253." type="journal article" year="1968">
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<collectingRegion id="49C4F84CFFFBEF2E6050FA37FC06F9ED" box="[890,1004,1522,1544]" country="United Kingdom" name="Darlington" pageId="20" pageNumber="403">Darlington</collectingRegion>
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1968
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</bibRefCitation>
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;
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<bibRefCitation id="EF914B5FFFFBEF2E63C7F9C9FE4DF9C7" author="Moore & Weir & Pyke & Coleoptera & Adephaga & Rhysodidae and Carabidae & In & Zoological Catalogue of Australia" box="[237,423,1548,1570]" pageId="20" pageNumber="403" pagination="17 - 320" refId="ref15604" refString="Moore, B. P., T. A. Weir, and J. E. Pyke. 1987. Coleoptera: Adephaga: Rhysodidae and Carabidae [pp. 17 - 320]. In: Zoological Catalogue of Australia 4 (D. W. Walton, editor). Australian Government Printing Service, Canberra, Australia. 444 pp." type="book chapter" year="1987">
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Moore
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<emphasis id="B974EABCFFFBEF2E6211F9C8FE85F9C7" box="[315,367,1549,1570]" italics="true" pageId="20" pageNumber="403">et al.</emphasis>
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1987
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</bibRefCitation>
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) was first collected on Lanai in Maunalei Gulch,
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<quantity id="4CF89B4BFFFBEF2E6087F9C9FC07F9C7" box="[941,1005,1548,1570]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="20" pageNumber="403" unit="m" value="490.0">490 m</quantity>
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elevation,
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<date id="FFBE106EFFFAEF2F63D6FF07FE9DFF3D" box="[252,375,194,216]" pageId="21" pageNumber="404" value="1993-05-03">3 May 1993</date>
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(Liebherr, CUIC). The specimen comprises a pterothorax and abdomen collected under a spider web suspended from the water pipe coming out of the Gulch headwall.
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<bibRefCitation id="EF914B5FFFFAEF2F6282FF3DFD7FFEEB" author="Samuelson" box="[424,661,248,270]" pageId="21" pageNumber="404" pagination="22 - 23" refId="ref16114" refString="Samuelson, G. A., J. K. Liebherr, and K. W. Will. 1997. Gnathaphanus picipes, an established adventive in Hawaii (Coleoptera: Carabidae). Bishop Museum Occasional Papers 49: 22 - 23." type="journal article" year="1997">
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Samuelson
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<emphasis id="B974EABCFFFAEF2F6137FF3DFDBAFEE8" box="[541,592,248,269]" italics="true" pageId="21" pageNumber="404">et al.</emphasis>
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(1997)
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</bibRefCitation>
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report a 1995 Maunalei Gulch specimen as the first Lanai record. Subsequently, the species has been collected across a broad distribution on Lanai (
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<figureCitation id="133B2A2BFFFAEF2F610BFEEBFD84FEA1" box="[545,622,302,324]" captionStart="Fig" captionStartId="17.[167,198,1308,1327]" captionTargetBox="[172,949,194,1286]" captionTargetId="figure-66@17.[171,949,194,1286]" captionTargetPageId="17" captionText="Fig. 6. Known distributions of carabid beetle species on Lanai. A. Blackburnia lanaihalensis. B. B. depressa. C. B. filipes. D. Metacolpodes buchanani. E. Gnathaphanus picipes (small circles), G. picipes plus G. multipunctatus (larger bullseye). F. Perigona nigriceps." figureDoi="http://doi.org/10.5281/zenodo.4924370" httpUri="https://zenodo.org/record/4924370/files/figure.png" pageId="21" pageNumber="404">Fig. 6E</figureCitation>
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) including the cloud forest, mesic forest, and dry forest communities (
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<figureCitation id="133B2A2BFFFAEF2F612DFE8CFDA9FEBA" box="[519,579,329,351]" captionStart="Fig" captionStartId="3.[167,198,915,934]" captionTargetBox="[181,941,194,893]" captionTargetId="figure-200@3.[181,941,194,893]" captionTargetPageId="3" captionText="Fig. 1. Pre-Polynesian vegetation communities on the island of Lanai (after Hobdy 1993): 1) cloud forest; 2) mesic forest; 3) dry forest; 4) arid grassland and shrubland; 5) coast and strand." figureDoi="http://doi.org/10.5281/zenodo.4924358" httpUri="https://zenodo.org/record/4924358/files/figure.png" pageId="21" pageNumber="404">Fig. 1</figureCitation>
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).
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</paragraph>
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<paragraph id="8BBF36AEFFFAEF2F6380FEA0FE3DFD8A" blockId="21.[146,975,194,623]" pageId="21" pageNumber="404">
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<emphasis id="B974EABCFFFAEF2F6380FEA0FF14FE9E" bold="true" box="[170,254,357,379]" pageId="21" pageNumber="404">Habitat.</emphasis>
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This species has been collected at the following anthropogenically disturbed sites (CUIC): 1) the seasonally dry Maunalei Gulch bottom; 2) running on a sidewalk at the Lanai Airport; 3) in the company of
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<taxonomicName id="4C004D2DFFFAEF2F6033FE5EFC25FE55" box="[793,975,411,432]" class="Insecta" family="Carabidae" genus="Gnathaphanus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="404" phylum="Arthropoda" rank="species" species="multipunctatus">
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<emphasis id="B974EABCFFFAEF2F6033FE5EFC25FE55" box="[793,975,411,432]" italics="true" pageId="21" pageNumber="404">G. multipunctatus</emphasis>
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</taxonomicName>
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running on a clay road after dark at the Kanepuu Dry Forest; 4) under a rock along the road at Puu Aalii on the summit ridge; 5) dead on the road adjacent to a
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<taxonomicName id="4C004D2DFFFAEF2F63B8FE28FF23FDE7" box="[146,201,493,514]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="21" pageNumber="404" phylum="Tracheophyta" rank="genus">
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<emphasis id="B974EABCFFFAEF2F63B8FE28FF23FDE7" box="[146,201,493,514]" italics="true" pageId="21" pageNumber="404">Pinus</emphasis>
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</taxonomicName>
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plantation at Haalelepaakai; and 6) in company with
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<taxonomicName id="4C004D2DFFFAEF2F602BFE28FC66FDE7" baseAuthorityName="Hope" baseAuthorityYear="1831" box="[769,908,493,514]" class="Insecta" family="Carabidae" genus="Metacolpodes" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="404" phylum="Arthropoda" rank="species" species="buchanani">
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<emphasis id="B974EABCFFFAEF2F602BFE28FC66FDE7" box="[769,908,493,514]" italics="true" pageId="21" pageNumber="404">M. buchanani</emphasis>
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</taxonomicName>
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under bark of downed
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<taxonomicName id="4C004D2DFFFAEF2F6276FDCDFE20FDF8" box="[348,458,520,541]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="21" pageNumber="404" phylum="Tracheophyta" rank="genus">
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<emphasis id="B974EABCFFFAEF2F6276FDCDFE20FDF8" box="[348,458,520,541]" italics="true" pageId="21" pageNumber="404">Eucalyptus</emphasis>
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</taxonomicName>
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trunks near Lehua,
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<quantity id="4CF89B4BFFFAEF2F61EFFDCDFCEFFDFB" box="[709,773,520,542]" metricMagnitude="2" metricUnit="m" metricValue="6.83" pageId="21" pageNumber="404" unit="m" value="683.0">683 m</quantity>
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elevation on the southeastern end of the summit ridge. On one occasion this species was collected with native species—
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<taxonomicName id="4C004D2DFFFAEF2F625CFDFBFE04FDB6" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[374,494,574,595]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="404" phylum="Arthropoda" rank="species" species="depressa">
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<emphasis id="B974EABCFFFAEF2F625CFDFBFE04FDB6" box="[374,494,574,595]" italics="true" pageId="21" pageNumber="404">B. depressa</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C004D2DFFFAEF2F6129FDFBFD88FDB6" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[515,610,574,595]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="404" phylum="Arthropoda" rank="species" species="filipes">
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<emphasis id="B974EABCFFFAEF2F6129FDFBFD88FDB6" box="[515,610,574,595]" italics="true" pageId="21" pageNumber="404">B. filipes</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C004D2DFFFAEF2F6180FDFBFCFBFDB6" baseAuthorityName="Sharp" baseAuthorityYear="1903" box="[682,785,574,595]" class="Insecta" family="Carabidae" genus="Blackburnia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="404" phylum="Arthropoda" rank="species" species="filipes">
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<emphasis id="B974EABCFFFAEF2F6180FDFBFCFBFDB6" box="[682,785,574,595]" italics="true" pageId="21" pageNumber="404">M. filipes</emphasis>
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</taxonomicName>
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—by beating soft understory fern in cloud forest.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |