176 lines
21 KiB
XML
176 lines
21 KiB
XML
<document id="0C60BD1890C1DE9D0F315517B96A6476" ID-CLB-Dataset="290635" ID-DOI="10.11646/zootaxa.5419.2.1" ID-GBIF-Dataset="d4a40af2-a5bf-43ed-95e3-98c1fd6c2942" ID-ISSN="1175-5326" ID-Zenodo-Dep="10792055" ID-ZooBank="88353CBA-6C4D-40E3-8475-B1FCA2C48637" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1709795503569" checkinUser="plazi" docAuthor="Holzmann, Maria, Barrenechea-Angeles, Inés, Lim, Swee-Cheng & Pawlowski, Jan" docDate="2024" docId="03A987A17B59AF7866C4456EFAAFD889" docLanguage="en" docName="zootaxa.5419.2.1.pdf" docOrigin="Zootaxa 5419 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5419.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psammina limbata Kamenskaya, Gooday & Tendal 2015" docType="treatment" docVersion="7" lastPageNumber="178" masterDocId="FF90FFD97B43AF636653407FFF99D97C" masterDocTitle="New xenophyophores (Foraminifera, Monothalamea) from the eastern Clarion-Clipperton Zone (equatorial Pacific)" masterLastPageNumber="188" masterPageNumber="151" pageNumber="177" updateTime="1716474086662" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EE4935C628E0831ACA9E5D3258538A64">New xenophyophores (Foraminifera, Monothalamea) from the eastern Clarion-Clipperton Zone (equatorial Pacific)</mods:title>
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<mods:namePart id="9BF0024D4B8B4CFF9ECAB49088F2095D">Holzmann, Maria</mods:namePart>
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<mods:nameIdentifier id="E027D8A08F69047707536661B889E307" type="ORCID">0000-0003-2460-6210</mods:nameIdentifier>
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<mods:affiliation id="1D0346FB1046229ABA41BCCFCAA0F503">Department of Genetics and Evolution, University of Geneva, Quai Ernest Ansermet 30, 1211 Geneva 4, Geneva, Switzerland.</mods:affiliation>
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<mods:nameIdentifier id="009D7D3DA2F4BE328DF720219C358A04" type="email">olzmann@unige.ch</mods:nameIdentifier>
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<mods:namePart id="E4258C6A03C12EF51BC3FA5F9AE0EFEF">Barrenechea-Angeles, Inés</mods:namePart>
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<mods:affiliation id="A3BF58A5E84519CF0EF5F30254275AD8">Department of Genetics and Evolution, University of Geneva, Quai Ernest Ansermet 30, 1211 Geneva 4, Geneva, Switzerland. & Department of Geosciences, The Arctic University of Norway, Dramsvegen 201, 9037 TromsØ, Norway.</mods:affiliation>
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<mods:namePart id="D8CABB5299FAE1E8CE2B32770E83D17A">Lim, Swee-Cheng</mods:namePart>
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<mods:affiliation id="73E44D8230AF67641D96B85316B41FA6">Tropical Marine Science Institute, National University of Singapore, 18 Kent Ridge Road, Singapore 119227, Singapore. & Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377</mods:affiliation>
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<mods:namePart id="F51D53DAE7EBEC762FEBB13C24AEEAAF">Pawlowski, Jan</mods:namePart>
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<mods:affiliation id="B7915F037378B77827760606A1C9D6F7">Institute of Oceanology, Polish Academy of Sciences, 81 - 712 Sopot, Poland. & ID-Gene Ecodiagnostics, Chemin du Pont-du-Centenaire 109, 1228 Plan-les-Ouates, Switzerland.</mods:affiliation>
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<mods:title id="3E0B4381FA425D8A6FD248AE264903D0">Zootaxa</mods:title>
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<mods:date id="076A7D052DD86DB2318367FBA8530811">2024</mods:date>
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<treatment id="03A987A17B59AF7866C4456EFAAFD889" ID-DOI="http://doi.org/10.5281/zenodo.11262843" ID-GBIF-Taxon="222869756" ID-Zenodo-Dep="11262843" LSID="urn:lsid:plazi:treatment:03A987A17B59AF7866C4456EFAAFD889" httpUri="http://treatment.plazi.org/id/03A987A17B59AF7866C4456EFAAFD889" lastPageId="27" lastPageNumber="178" pageId="26" pageNumber="177">
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<subSubSection id="C31A653C7B59AF7966C4456EFCDEDC57" box="[151,839,1297,1324]" pageId="26" pageNumber="177" type="nomenclature">
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<paragraph id="8BBF36B77B59AF7966C4456EFCDEDC57" blockId="26.[151,839,1297,1359]" box="[151,839,1297,1324]" pageId="26" pageNumber="177">
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<heading id="D0F781DB7B59AF7966C4456EFCDEDC57" bold="true" box="[151,839,1297,1324]" fontSize="11" level="1" pageId="26" pageNumber="177" reason="1">
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<emphasis id="B974EAA57B59AF7966C4456EFCDEDC57" bold="true" box="[151,839,1297,1324]" pageId="26" pageNumber="177">
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<taxonomicName id="4C004D347B59AF7966C4456EFE32DC50" ID-CoL="4NBCD" authority="Kamenskaya, Gooday & Tendal, 2015" authorityName="Kamenskaya, Gooday & Tendal" authorityYear="2015" box="[151,427,1297,1324]" class="Monothalamea" family="Psamminidae" genus="Psammina" isUncertain="true" kingdom="Chromista" pageId="26" pageNumber="177" phylum="Foraminifera" rank="species" species="limbata">
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<emphasis id="B974EAA57B59AF7966C4456EFE81DC57" bold="true" box="[151,280,1297,1323]" italics="true" pageId="26" pageNumber="177">Psammina</emphasis>
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aff.
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<emphasis id="B974EAA57B59AF796703456EFE32DC50" bold="true" box="[336,427,1297,1324]" italics="true" pageId="26" pageNumber="177">limbata</emphasis>
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</taxonomicName>
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form 2 sensu Gooday
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<emphasis id="B974EAA57B59AF796490456DFD67DC50" bold="true" box="[707,766,1297,1324]" italics="true" pageId="26" pageNumber="177">et al.</emphasis>
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, 2018
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A653C7B59AF7966C4454AFBDEDCEE" pageId="26" pageNumber="177" type="description">
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<paragraph id="8BBF36B77B59AF7966C4454AFE01DC33" blockId="26.[151,839,1297,1359]" box="[151,408,1333,1359]" pageId="26" pageNumber="177">
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Supplementary
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<figureCitation id="133B2A327B59AF79671B454AFE01DC33" box="[328,408,1333,1359]" captionStart="FIGURE 7" captionStartId="14.[151,250,1153,1177]" captionTargetBox="[151,1436,181,1129]" captionTargetId="figure-20@14.[151,1436,181,1129]" captionTargetPageId="14" captionText="FIGURE 7. Abyssalia aff. foliformis, laboratory photographs; Specimen RC0615 (BC011). A, Fragment. B, Detail of edge. C–D, Broken edge showing granellare embedded in dak stercomare masses." figureDoi="http://doi.org/10.5281/zenodo.10792073" httpUri="https://zenodo.org/record/10792073/files/figure.png" pageId="26" pageNumber="177">Fig. S7</figureCitation>
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</paragraph>
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<paragraph id="8BBF36B77B59AF7966C44505FBDEDCEE" blockId="26.[151,1095,1402,1426]" box="[151,1095,1402,1426]" pageId="26" pageNumber="177">
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<taxonomicName id="4C004D347B59AF7966C44505FEE7DCEE" authorityName="Kamenskaya, Gooday & Tendal" authorityYear="2015" box="[151,382,1402,1426]" class="Monothalamea" family="Psamminidae" genus="Psammina" isUncertain="true" kingdom="Chromista" pageId="26" pageNumber="177" phylum="Foraminifera" rank="species" species="limbata">
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<emphasis id="B974EAA57B59AF7966C44505FE9BDCED" box="[151,258,1402,1425]" italics="true" pageId="26" pageNumber="177">Psammina</emphasis>
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aff.
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<emphasis id="B974EAA57B59AF7967624505FEE7DCEE" box="[305,382,1402,1426]" italics="true" pageId="26" pageNumber="177">limbata</emphasis>
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</taxonomicName>
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form 2. Gooday
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<emphasis id="B974EAA57B59AF79647F4504FDC0DCEE" box="[556,601,1402,1426]" italics="true" pageId="26" pageNumber="177">et al</emphasis>
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., 2018, 934–935,
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||
<figureCitation id="133B2A327B59AF7965434505FCD5DCEE" box="[784,844,1402,1426]" captionStart="FIGURE 6" captionStartId="13.[151,250,1983,2007]" captionTargetBox="[151,1436,341,1960]" captionTargetId="figure-100@13.[151,1436,341,1960]" captionTargetPageId="13" captionText="FIGURE 6. Abyssalia aff. foliformis, shipboard photographs; RC0615 (BC011).A, B, Two different side views. C, Test viewed from above. D–F, Details of test; the yellow tube in E is probably part of a Saccorhiza ramosa test." figureDoi="http://doi.org/10.5281/zenodo.10792071" httpUri="https://zenodo.org/record/10792071/files/figure.png" pageId="26" pageNumber="177">Fig. 6</figureCitation>
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, Supplementary
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<figureCitation id="133B2A327B59AF7965AB4505FBDDDCEE" box="[1016,1092,1402,1426]" captionStart="FIGURE 2" captionStartId="7.[151,250,1598,1622]" captionTargetBox="[151,1436,181,1574]" captionTargetId="figure-18@7.[151,1436,181,1574]" captionTargetPageId="7" captionText="FIGURE 2. Aschemonella monilis Gooday & Holzmann 2017. A–D, Sequenced specimens. A, Dark specimen attached to nodule; RC1056 (BC026). B, Interior of test fragment with red granellare (after removal of most pieces for sequencing) and dark stercomare; RC0056 (BC001). C, Fragment with two segments; RC0490B (BC010).D, E, RC1900.1 (BC045). D, Fragment with broken segment containing granellare. E, Detail of dome and tubular apertural structure. F, G, Specimens without cytoplasm; RC1900.2 (BC045). F, Complete fragment. G, Detail of apertural structures. No scale is available for B." figureDoi="http://doi.org/10.5281/zenodo.10792059" httpUri="https://zenodo.org/record/10792059/files/figure.png" pageId="26" pageNumber="177">Fig. S2</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31A653C7B59AF7966C445BFFBBBDC82" pageId="26" pageNumber="177" type="materials_examined">
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<paragraph id="8BBF36B77B59AF7966C445BFFBBBDC82" blockId="26.[151,1437,1472,2039]" pageId="26" pageNumber="177">
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<emphasis id="B974EAA57B59AF7966C445BFFEE7DCA6" bold="true" box="[151,382,1472,1498]" pageId="26" pageNumber="177">Material examined</emphasis>
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. Box core 034. Specimen RC1492 (morphology only). Box core 040. Specimen RC1337 (morphology only). Box core 042. Specimen RC1743 (morphology only). Dried.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A653C7B59AF7866944677FCCAD819" lastPageId="27" lastPageNumber="178" pageId="26" pageNumber="177" type="description">
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<paragraph id="8BBF36B77B59AF7966944677FEE5DE63" blockId="26.[151,1437,1472,2039]" pageId="26" pageNumber="177">
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<emphasis id="B974EAA57B59AF7966944677FEC8DF5E" bold="true" box="[199,337,1544,1570]" pageId="26" pageNumber="177">Description</emphasis>
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. RC1337 (Supplementary
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<figureCitation id="133B2A327B59AF79642C4677FC90DF5E" box="[639,777,1544,1570]" captionStart="FIGURE 7" captionStartId="14.[151,250,1153,1177]" captionTargetBox="[151,1436,181,1129]" captionTargetId="figure-20@14.[151,1436,181,1129]" captionTargetPageId="14" captionText="FIGURE 7. Abyssalia aff. foliformis, laboratory photographs; Specimen RC0615 (BC011). A, Fragment. B, Detail of edge. C–D, Broken edge showing granellare embedded in dak stercomare masses." figureDoi="http://doi.org/10.5281/zenodo.10792073" httpUri="https://zenodo.org/record/10792073/files/figure.png" pageId="26" pageNumber="177">Fig. S7A–C</figureCitation>
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). The test comprises a semicircular, fan-shaped upper part, almost
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<quantity id="4CF89B527B59AF7966B94653FEA2DF3B" box="[234,315,1580,1607]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="26" pageNumber="177" unit="mm" value="33.0">33 mm</quantity>
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wide, merging into a tapered stalk attached to a nodule. The test (stalk plus fan) is around
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<quantity id="4CF89B527B59AF79631F4653FA04DF3B" box="[1356,1437,1580,1607]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="26" pageNumber="177" unit="mm" value="26.0">26 mm</quantity>
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high and almost intact apart from some damage to the margin and several holes within the fan. The upper part is distinctly asymmetrical, being better developed on one side than the other. As a result, the stalk is also asymmetrical and longer on the side where the fan is less well developed. The base of the stalk extends into at least three branched root-like structures that spread across the nodule surface. The fan displays clearly developed ‘growth lines’ that delimit concentric zones. The upper part appears to be empty, the lower part is filled with dark material, probably decayed stercomare.
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</paragraph>
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<paragraph id="8BBF36B77B59AF7966944757FC84DE8A" blockId="26.[151,1437,1472,2039]" pageId="26" pageNumber="177">
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RC1492 (Supplementary
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<figureCitation id="133B2A327B59AF7967B94757FDCDDE3E" box="[490,596,1832,1858]" captionStart="FIGURE 7" captionStartId="14.[151,250,1153,1177]" captionTargetBox="[151,1436,181,1129]" captionTargetId="figure-20@14.[151,1436,181,1129]" captionTargetPageId="14" captionText="FIGURE 7. Abyssalia aff. foliformis, laboratory photographs; Specimen RC0615 (BC011). A, Fragment. B, Detail of edge. C–D, Broken edge showing granellare embedded in dak stercomare masses." figureDoi="http://doi.org/10.5281/zenodo.10792073" httpUri="https://zenodo.org/record/10792073/files/figure.png" pageId="26" pageNumber="177">Fig. S7D</figureCitation>
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). The upper part of the test is damaged with little or none of the original margin surviving. What remains is
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<quantity id="4CF89B527B59AF79647D4733FDE6DE1B" box="[558,639,1868,1895]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="177" unit="mm" value="18.0">18 mm</quantity>
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wide and displays a well-developed concentric zonation. The overall height is also
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<quantity id="4CF89B527B59AF796761470FFE1CDEF7" box="[306,389,1904,1931]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="177" unit="mm" value="18.0">18 mm</quantity>
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. However, the lower part is intact and tapers to a very short (
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<quantity id="4CF89B527B59AF79627C470FFB11DEF6" box="[1071,1160,1904,1930]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="177" unit="mm" value="1.2">1.2 mm</quantity>
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) but relatively wide (3.0 mm) stalk. In addition to the main stalk there is a wide secondary support developed from the base of the fan, and two long, straight, bar-like processes (6.5 and
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<quantity id="4CF89B527B59AF7964CC47C6FD6CDEAF" box="[671,757,1977,2003]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="26" pageNumber="177" unit="mm" value="7.7">7.7 mm</quantity>
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long and
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<quantity id="4CF89B527B59AF79653647C7FC75DEAF" box="[869,1004,1976,2003]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="26" pageNumber="177" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
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wide), arise from the intact margin on the side of the test and are directed obliquely downwards.
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</paragraph>
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<paragraph id="8BBF36B77B58AF78669440E8FCCAD819" blockId="27.[151,1436,151,501]" pageId="27" pageNumber="178">
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RC1743 (dried). The specimen is badly cracked, but basically forms a semicircular fan with a fairly straight lower margin and the base of a stem. It measures
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<quantity id="4CF89B527B58AF78648F40C4FCDAD9A9" box="[732,835,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="27" pageNumber="178" unit="mm" value="29.3">29.3 mm</quantity>
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(width) by
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<quantity id="4CF89B527B58AF78659D40C5FBADD9A9" box="[974,1076,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="27" pageNumber="178" unit="mm" value="25.8">25.8 mm</quantity>
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(height). The wall has a rough surface with concentric growth zones. It comprises a mesh of sponge spicules, radiolarians and subordinate numbers of agglutinated foraminiferal tests and fragments, mainly yellow or orange in colour. Parts of the test interior are visible through gaps in the cracked wall, revealing parts of several very thin partitions corresponding to the external furrows that define the zones. Otherwise, the interior is empty.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A653C7B58AF7866944110FAAFD889" pageId="27" pageNumber="178" type="discussion">
|
||
<paragraph id="8BBF36B77B58AF7866944110FAAFD889" blockId="27.[151,1436,151,501]" pageId="27" pageNumber="178">
|
||
<emphasis id="B974EAA57B58AF7866944110FEA2D8F5" bold="true" box="[199,315,367,393]" pageId="27" pageNumber="178">Remarks.</emphasis>
|
||
These
|
||
<specimenCount id="9D06FD3E7B58AF7867D94110FDA4D8F5" box="[394,573,367,393]" count="3" pageId="27" pageNumber="178" type="generic">three specimens</specimenCount>
|
||
are very similar to
|
||
<taxonomicName id="4C004D347B58AF7865404110FB88D8F5" authorityName="Kamenskaya, Gooday & Tendal" authorityYear="2015" box="[787,1041,367,393]" class="Monothalamea" family="Psamminidae" genus="Psammina" isUncertain="true" kingdom="Chromista" pageId="27" pageNumber="178" phylum="Foraminifera" rank="species" species="limbata">
|
||
<emphasis id="B974EAA57B58AF7865404110FC10D8F4" box="[787,905,367,392]" italics="true" pageId="27" pageNumber="178">Psammina</emphasis>
|
||
aff.
|
||
<emphasis id="B974EAA57B58AF7865E84110FB88D8F5" box="[955,1041,367,393]" italics="true" pageId="27" pageNumber="178">limbata</emphasis>
|
||
</taxonomicName>
|
||
form 2 from the UK-1 license area, as illustrated in
|
||
<figureCitation id="133B2A327B58AF78671941ECFE5BD8D1" box="[330,450,403,429]" captionStart="FIGURE 6" captionStartId="13.[151,250,1983,2007]" captionTargetBox="[151,1436,341,1960]" captionTargetId="figure-100@13.[151,1436,341,1960]" captionTargetPageId="13" captionText="FIGURE 6. Abyssalia aff. foliformis, shipboard photographs; RC0615 (BC011).A, B, Two different side views. C, Test viewed from above. D–F, Details of test; the yellow tube in E is probably part of a Saccorhiza ramosa test." figureDoi="http://doi.org/10.5281/zenodo.10792071" httpUri="https://zenodo.org/record/10792071/files/figure.png" pageId="27" pageNumber="178">Figs 6E, G</figureCitation>
|
||
of Gooday
|
||
<emphasis id="B974EAA57B58AF78641B41ECFDE0D8D1" box="[584,633,403,429]" italics="true" pageId="27" pageNumber="178">et al</emphasis>
|
||
. (2018). The concentric zonation with corresponding internal partitions is a typical feature and distinguishes this form from other stalked
|
||
<taxonomicName id="4C004D347B58AF7865DE41C8FBC7D8AD" box="[909,1118,439,465]" pageId="27" pageNumber="178">
|
||
<emphasis id="B974EAA57B58AF7865DE41C8FB9AD8AC" box="[909,1027,439,464]" italics="true" pageId="27" pageNumber="178">Psammina</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
described by Gooday
|
||
<emphasis id="B974EAA57B58AF78633141C8FA0CD8AD" box="[1378,1429,439,465]" italics="true" pageId="27" pageNumber="178">et al</emphasis>
|
||
. (2018). The straight, bar-like processes (Supplementary
|
||
<figureCitation id="133B2A327B58AF78655D41A4FCEDD889" box="[782,884,475,501]" captionStart="FIGURE 7" captionStartId="14.[151,250,1153,1177]" captionTargetBox="[151,1436,181,1129]" captionTargetId="figure-20@14.[151,1436,181,1129]" captionTargetPageId="14" captionText="FIGURE 7. Abyssalia aff. foliformis, laboratory photographs; Specimen RC0615 (BC011). A, Fragment. B, Detail of edge. C–D, Broken edge showing granellare embedded in dak stercomare masses." figureDoi="http://doi.org/10.5281/zenodo.10792073" httpUri="https://zenodo.org/record/10792073/files/figure.png" pageId="27" pageNumber="178">Fig. S7D</figureCitation>
|
||
) are also present in the UK-
|
||
<specimenCount id="9D06FD3E7B58AF7862E641A4FAABD889" box="[1205,1330,475,501]" count="1" pageId="27" pageNumber="178" type="generic">1 specimen</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |