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<document id="E22B9AF741670339CE3D0DC387B2B826" ID="10.11646/zootaxa.4225.1.1" ID-CLB-Dataset="37263" ID-DOI="10.5281/zenodo.259087" ID-GBIF-Dataset="05be601b-f9f1-4952-ac09-7012ef31aab0" ID-ISSN="1175-5326" ID-Zenodo-Dep="259087" ID-ZooBank="4676EE59-569B-4197-B108-9B32FD88E68E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1485243787505" checkinUser="plazi" docAuthor="Skartveit, John &amp; Nel, André" docDate="2017" docId="03A9879CFFB9F47545D334D97CFC3911" docLanguage="en" docName="zootaxa.4225.1.1.pdf" docOrigin="Zootaxa 4225 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Bibio spectabilis Theobald 1937" docType="treatment" docVersion="8" lastPageNumber="74" masterDocId="FF90FFE4FFFEF43C4544344E7D193E27" masterDocTitle="Revision of fossil Bibionidae (Insecta: Diptera) from French Oligocene deposits" masterLastPageNumber="83" masterPageNumber="1" pageNumber="72" updateTime="1698436270113" updateUser="ExternalLinkService">
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<mods:title id="9D88B4AAC7FEA29D7A5D4954AA815A9F">Revision of fossil Bibionidae (Insecta: Diptera) from French Oligocene deposits</mods:title>
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<mods:namePart id="2AB102C60A4D3B19158500F99DEE30E7">Skartveit, John</mods:namePart>
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<mods:namePart id="AC9AEBC32934B70D63B7B8A3D57F3772">Nel, André</mods:namePart>
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<mods:date id="F0A084FD47FE9D789BA44A22FE850F1C">2017</mods:date>
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<treatment id="03A9879CFFB9F47545D334D97CFC3911" ID-DOI="http://doi.org/10.5281/zenodo.6039685" ID-GBIF-Taxon="127624375" ID-Zenodo-Dep="6039685" LSID="urn:lsid:plazi:treatment:03A9879CFFB9F47545D334D97CFC3911" httpUri="http://treatment.plazi.org/id/03A9879CFFB9F47545D334D97CFC3911" lastPageId="73" lastPageNumber="74" pageId="71" pageNumber="72">
<subSubSection id="C31A6501FFB9F47B45D334D97C743EF4" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="8BBF368AFFB9F47B45D334D97F3D3E95" blockId="71.[151,548,151,211]" box="[151,548,151,178]" pageId="71" pageNumber="72">
<heading id="D0F781E6FFB9F47B45D334D97F3D3E95" bold="true" box="[151,548,151,178]" fontSize="11" level="1" pageId="71" pageNumber="72" reason="1">
<taxonomicName id="4C004D09FFB9F47B45D334D97F3D3E95" ID-CoL="8B76F" authority="Theobald, 1937" authorityName="Theobald" authorityYear="1937" box="[151,548,151,178]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B974EA98FFB9F47B45D334D97F3D3E95" bold="true" box="[151,548,151,178]" pageId="71" pageNumber="72">
<emphasis id="B974EA98FFB9F47B45D334D97C453E96" bold="true" box="[151,348,151,177]" italics="true" pageId="71" pageNumber="72">Bibio spectabilis</emphasis>
<bibRefCitation id="EF914B7BFFB9F47B442034D67F3D3E95" author="Theobald" box="[356,548,152,178]" pageId="71" pageNumber="72" refString="Theobald, N. (1937) Les insectes fossiles des terrains oligocenes de France. Bulletin mensuel (Memoires) de la Societe des Sciences de Nancy 1, 1 - 473." type="journal article" year="1937">Théobald, 1937</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BBF368AFFB9F47B45D334F47C743EF4" blockId="71.[151,548,151,211]" box="[151,365,186,211]" pageId="71" pageNumber="72">
(figs. 68, 198
<emphasis id="B974EA98FFB9F47B446834F57C233EF5" box="[300,314,187,210]" italics="true" pageId="71" pageNumber="72"></emphasis>
206)
</paragraph>
</subSubSection>
<subSubSection id="C31A6501FFB9F47B45D3354C7F063F3F" box="[151,543,258,280]" pageId="71" pageNumber="72" type="reference_group">
<paragraph id="8BBF368AFFB9F47B45D3354C7F063F3F" blockId="71.[151,543,258,280]" box="[151,543,258,280]" pageId="71" pageNumber="72">
<treatmentCitationGroup id="AB1011A4FFB9F47B45D3354C7F063F3F" box="[151,543,258,280]" pageId="71" pageNumber="72">
<taxonomicName id="4C004D09FFB9F47B45D3354C7F053F3F" ID-CoL="8B76F" authority="Theobald, 1937: 333" authorityName="Theobald" authorityPageNumber="333" authorityYear="1937" box="[151,540,258,280]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B974EA98FFB9F47B45D3354C7C583F3F" box="[151,321,258,280]" italics="true" pageId="71" pageNumber="72">Bibio spectabilis</emphasis>
<treatmentCitation id="0AA1109BFFB9F47B4402354C7F053F3F" author="Theobald" box="[326,540,258,280]" page="333" pageId="71" pageNumber="72" year="1937">
<bibRefCitation id="EF914B7BFFB9F47B4402354C7CF03F3F" author="Theobald" box="[326,489,258,280]" pageId="71" pageNumber="72" refString="Theobald, N. (1937) Les insectes fossiles des terrains oligocenes de France. Bulletin mensuel (Memoires) de la Societe des Sciences de Nancy 1, 1 - 473." type="journal article" year="1937">Théobald, 1937</bibRefCitation>
: 333
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31A6501FFB9F47545D3350B7F1E3EDF" lastPageId="73" lastPageNumber="74" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="8BBF368AFFB9F47B45D3350B7C333FA6" blockId="71.[151,1436,325,385]" pageId="71" pageNumber="72">
<emphasis id="B974EA98FFB9F47B45D3350B7C9B3F79" bold="true" box="[151,386,325,350]" pageId="71" pageNumber="72">Material examined.</emphasis>
Holotype (female) of
<taxonomicName id="4C004D09FFB9F47B47C2350B7E5D3F7A" box="[646,836,325,349]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B974EA98FFB9F47B47C2350B7E5D3F7A" box="[646,836,325,349]" italics="true" pageId="71" pageNumber="72">Bibio spectabilis</emphasis>
</taxonomicName>
FSL 391911. Paratype (female) of
<taxonomicName id="4C004D09FFB9F47B419A350B78853F7A" box="[1246,1436,325,349]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B974EA98FFB9F47B419A350B78853F7A" box="[1246,1436,325,349]" italics="true" pageId="71" pageNumber="72">Bibio spectabilis</emphasis>
</taxonomicName>
FSL 391912.
</paragraph>
<caption id="DF7F6602FFB9F47B45D333147F0B39B7" pageId="71" pageNumber="72">
<paragraph id="8BBF368AFFB9F47B45D333147F0B39B7" blockId="71.[151,705,1882,2028]" pageId="71" pageNumber="72">
<emphasis id="B974EA98FFB9F47B45D333147C753957" bold="true" box="[151,364,1882,1905]" pageId="71" pageNumber="72">FIGURES 198206.</emphasis>
<taxonomicName id="4C004D09FFB9F47B443733157FD83956" authority="Theobald, 1937" authorityName="Theobald" authorityYear="1937" box="[371,705,1883,1905]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B974EA98FFB9F47B443733157F043956" box="[371,541,1883,1905]" italics="true" pageId="71" pageNumber="72">Bibio spectabilis</emphasis>
Théobald, 1937
</taxonomicName>
Fig. 198. Female, wing. FSL 391911.
</paragraph>
</caption>
<paragraph id="8BBF368AFFB9F47B45D333D67F283989" blockId="71.[151,705,1882,2028]" box="[151,561,1944,1966]" pageId="71" pageNumber="72">
<figureCitation id="133B2A0FFFB9F47B45D333D67DF73989" box="[151,238,1944,1966]" captionStart="FIGURES 198 206" captionStartId="71.[151,264,1882,1905]" captionText="FIGURES 198 206. Bibio spectabilis Théobald, 1937 Fig. 198. Female, wing. FSL 391911." pageId="71" pageNumber="72">Fig. 199</figureCitation>
. Male, wing. MNHN Aix 1436.
</paragraph>
<paragraph id="8BBF368AFFB9F47B45D333F97F2239EA" blockId="71.[151,705,1882,2028]" box="[151,571,1975,1997]" pageId="71" pageNumber="72">
<figureCitation id="133B2A0FFFB9F47B45D333F97DF739EA" box="[151,238,1975,1997]" captionStart="FIGURES 198 206" captionStartId="71.[151,264,1882,1905]" captionText="FIGURES 198 206. Bibio spectabilis Théobald, 1937 Fig. 198. Female, wing. FSL 391911." pageId="71" pageNumber="72">Fig. 200</figureCitation>
. Male, habitus. MNHN Aix 313.
</paragraph>
<paragraph id="8BBF368AFFB9F47B45D333987FB939CB" blockId="71.[151,705,1882,2028]" box="[151,672,2006,2028]" pageId="71" pageNumber="72">
<figureCitation id="133B2A0FFFB9F47B45D333987DF739CB" box="[151,238,2006,2028]" captionStart="FIGURES 198 206" captionStartId="71.[151,264,1882,1905]" captionText="FIGURES 198 206. Bibio spectabilis Théobald, 1937 Fig. 198. Female, wing. FSL 391911." pageId="71" pageNumber="72">Fig. 201</figureCitation>
. Male, head and thorax. MNHN Aix 1436.
</paragraph>
<paragraph id="8BBF368AFFB6F47445D333017F483942" blockId="72.[151,695,1871,2016]" box="[151,593,1871,1893]" pageId="72" pageNumber="73">
<figureCitation id="133B2A0FFFB6F47445D333017DF73942" box="[151,238,1871,1893]" captionStart="FIGURES 198 206" captionStartId="71.[151,264,1882,1905]" captionText="FIGURES 198 206. Bibio spectabilis Théobald, 1937 Fig. 198. Female, wing. FSL 391911." pageId="72" pageNumber="73">Fig. 202</figureCitation>
. Male, hind leg. MNHN Aix 1377.
</paragraph>
<paragraph id="8BBF368AFFB6F47445D333207F7C39A3" blockId="72.[151,695,1871,2016]" box="[151,613,1902,1924]" pageId="72" pageNumber="73">
<figureCitation id="133B2A0FFFB6F47445D333207DF739A3" box="[151,238,1902,1924]" captionStart="FIGURES 198 206" captionStartId="71.[151,264,1882,1905]" captionText="FIGURES 198 206. Bibio spectabilis Théobald, 1937 Fig. 198. Female, wing. FSL 391911." pageId="72" pageNumber="73">Fig. 203</figureCitation>
. Male, terminalia. MNHN Aix 1436.
</paragraph>
<paragraph id="8BBF368AFFB6F47445D333C27F4239E6" blockId="72.[151,695,1871,2016]" pageId="72" pageNumber="73">
<figureCitation id="133B2A0FFFB6F47445D333C27DF73985" box="[151,238,1932,1954]" captionStart="FIGURES 198 206" captionStartId="71.[151,264,1882,1905]" captionText="FIGURES 198 206. Bibio spectabilis Théobald, 1937 Fig. 198. Female, wing. FSL 391911." pageId="72" pageNumber="73">Fig. 204</figureCitation>
. Female, head and thorax. MNHN Aix 1299.
<figureCitation id="133B2A0FFFB6F47445D333E57DF739E6" box="[151,238,1963,1985]" captionStart="FIGURES 198 206" captionStartId="71.[151,264,1882,1905]" captionText="FIGURES 198 206. Bibio spectabilis Théobald, 1937 Fig. 198. Female, wing. FSL 391911." pageId="72" pageNumber="73">Fig. 205</figureCitation>
. Female, hind leg. MNHN Aix 845.
</paragraph>
<paragraph id="8BBF368AFFB6F47445D333847F7639C7" blockId="72.[151,695,1871,2016]" box="[151,623,1994,2016]" pageId="72" pageNumber="73">
<figureCitation id="133B2A0FFFB6F47445D333847DF739C7" box="[151,238,1994,2016]" captionStart="FIGURES 198 206" captionStartId="71.[151,264,1882,1905]" captionText="FIGURES 198 206. Bibio spectabilis Théobald, 1937 Fig. 198. Female, wing. FSL 391911." pageId="72" pageNumber="73">Fig. 206</figureCitation>
. Female, terminalia. MNHN Aix 845.
</paragraph>
<paragraph id="8BBF368AFFB7F475458334D979D33EF2" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">Additional material: Male (tentatively associated) EM 70215 (1.2.2) (stored in FSL). Males (MNHN Aix 52, 1377). Females (MNHN Aix 6, 10, 11, 107, 161, 309, 845, 1090, 1299, 1307, 1446, 1447, P25).</paragraph>
<paragraph id="8BBF368AFFB7F475458334917F1E3EDF" blockId="73.[151,1437,151,1846]" box="[199,519,223,248]" pageId="73" pageNumber="74">
<emphasis id="B974EA98FFB7F475458334917C583EDF" bold="true" box="[199,321,223,248]" pageId="73" pageNumber="74">Localities.</emphasis>
Aix-en-Provence
</paragraph>
</subSubSection>
<subSubSection id="C31A6501FFB7F4754583354A7CC03FAD" pageId="73" pageNumber="74" type="diagnosis">
<paragraph id="8BBF368AFFB7F4754583354A7CC03FAD" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">
<emphasis id="B974EA98FFB7F4754583354A7C583F3A" bold="true" box="[199,321,260,285]" pageId="73" pageNumber="74">Diagnosis.</emphasis>
A medium-sized to large (
<quantity id="4CF89B6FFFB7F475472D354A7FF43F3B" box="[617,749,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.2" metricValueMin="0.85" pageId="73" pageNumber="74" unit="mm" value="10.25" valueMax="12.0" valueMin="8.5">
8.5
<emphasis id="B974EA98FFB7F47547C9354B7F823F3B" box="[653,667,261,284]" italics="true" pageId="73" pageNumber="74"></emphasis>
12 mm
</quantity>
), robust
<taxonomicName id="4C004D09FFB7F4754608354A7E903F3B" box="[844,905,260,284]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B974EA98FFB7F4754608354A7E903F3B" box="[844,905,260,284]" italics="true" pageId="73" pageNumber="74">Bibio</emphasis>
</taxonomicName>
species. The wing has a short but distinctive R- M crossvein, about one fifth as long as basal part of R s, and CuA1 and CuA2 are apically straight or nearly so. Antenna robust with 8-segmented flagellum. Male epandrium not bilobate, apically straight with slightly protruding posterior corners.
</paragraph>
</subSubSection>
<subSubSection id="C31A6501FFB7F475458335D87CFC3911" pageId="73" pageNumber="74" type="description">
<paragraph id="8BBF368AFFB7F475458335D879BE3F88" blockId="73.[151,1437,151,1846]" box="[199,1191,406,431]" pageId="73" pageNumber="74">
<emphasis id="B974EA98FFB7F475458335D87C6D3F88" bold="true" box="[199,372,406,431]" pageId="73" pageNumber="74">Redescription.</emphasis>
Male (tentatively associated, fig. 200): Total length 9.0
<emphasis id="B974EA98FFB7F47546A735D67EE83F88" box="[995,1009,408,431]" italics="true" pageId="73" pageNumber="74"></emphasis>
10.0 mm (N=3).
</paragraph>
<paragraph id="8BBF368AFFB7F475458335F478063FD0" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">
Head (fig. 201): Length
<quantity id="4CF89B6FFFB7F47544AD35F57F6A3FF5" box="[489,627,442,467]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="73" pageNumber="74" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">
1.3
<emphasis id="B974EA98FFB7F475474835F57F033FF5" box="[524,538,443,466]" italics="true" pageId="73" pageNumber="74"></emphasis>
1.6 mm
</quantity>
(N=2), width
<quantity id="4CF89B6FFFB7F475466435F47E633FF5" box="[800,890,442,467]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="73" pageNumber="74" unit="mm" value="1.5">1.5 mm</quantity>
(N=1). Dark brown, presumably black in life. Antennal flagellum 8-segmented. Palp relatively short and robust. Ocellar triangle large and prominent.
</paragraph>
<paragraph id="8BBF368AFFB7F4754583364C788C3C3C" blockId="73.[151,1437,151,1846]" box="[199,1429,514,539]" pageId="73" pageNumber="74">
Thorax: Length
<quantity id="4CF89B6FFFB7F4754439364D7F1D3C3D" box="[381,516,515,539]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="73" pageNumber="74" unit="mm" value="2.95" valueMax="3.1" valueMin="2.8">
2.8
<emphasis id="B974EA98FFB7F47544E4364D7CB73C3D" box="[416,430,515,538]" italics="true" pageId="73" pageNumber="74"></emphasis>
3.1 mm
</quantity>
(N=2), width
<quantity id="4CF89B6FFFB7F47547E0364D7FE33C3D" box="[676,762,515,539]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="73" pageNumber="74" unit="mm" value="2.3">2.3 mm</quantity>
(N=1). Dark brown, presumably black in life. Haltere pale.
</paragraph>
<paragraph id="8BBF368AFFB7F475458336687FBD3C89" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">
Wing (fig. 199): Length
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5.8
<emphasis id="B974EA98FFB7F475475736667F383C18" box="[531,545,552,575]" italics="true" pageId="73" pageNumber="74"></emphasis>
6.9 mm
</quantity>
(N=3), width
<quantity id="4CF89B6FFFB7F475466936687EA03C18" box="[813,953,550,575]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="73" pageNumber="74" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">
2.5
<emphasis id="B974EA98FFB7F475461436667E473C18" box="[848,862,552,575]" italics="true" pageId="73" pageNumber="74"></emphasis>
2.8 mm
</quantity>
(N=3), length/width = 2.31
<emphasis id="B974EA98FFB7F47540413666780A3C18" box="[1285,1299,552,575]" italics="true" pageId="73" pageNumber="74"></emphasis>
2.73 (N=3). Hyaline? Membrane pigmentation not well preserved, veins distinctive throughout. R4+5 arched forwards, running more or less parallel to
<collectingRegion id="49C4F868FFB7F47544EF363C7CF23CAD" box="[427,491,626,650]" country="New Zealand" name="West Coast" pageId="73" pageNumber="74">Costa</collectingRegion>
in distal half. Crossvein R-M short but distinctive. CuA-veins apically straight or nearly so. Wing vein measurements in
<tableCitation id="C6820331FFB7F475470E36DB7FB93C89" box="[586,672,661,686]" captionStart="TABLE 4" captionStartId="58.[164,225,1207,1223]" captionTargetBox="[164,1421,1249,1825]" captionTargetPageId="58" captionText="TABLE 4. Bibio spp., wing measurements" httpUri="http://table.plazi.org/id/DF7F6602FFC4F40645E030F97CC73AE0" pageId="73" pageNumber="74" tableUuid="DF7F6602FFC4F40645E030F97CC73AE0">Table 4</tableCitation>
.
</paragraph>
<paragraph id="8BBF368AFFB7F475458336F47E203D19" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">
Legs (fig. 202): Fore tibia with spur a little less than half as long as anterior spine. Hind femur stout, clavate. Hind tibia rather short and robust with long, curved, apically blunt spur. Hind tarsus robust, first tarsomere approximately three times as long as wide. Hind femur length
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2.5
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2.7 mm
</quantity>
(N=3), width
<quantity id="4CF89B6FFFB7F4754134374F780B3D3E" box="[1136,1298,769,794]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" metricValueMax="6.0" metricValueMin="5.2" pageId="73" pageNumber="74" unit="mm" value="0.56" valueMax="0.6" valueMin="0.52">
0.52
<emphasis id="B974EA98FFB7F47541E5374C79B63D3E" box="[1185,1199,770,793]" italics="true" pageId="73" pageNumber="74"></emphasis>
0.60 mm
</quantity>
(N=3), hind tibia length
<quantity id="4CF89B6FFFB7F475445937687CBD3D1A" box="[285,420,806,830]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.9" metricValueMin="2.3" pageId="73" pageNumber="74" unit="mm" value="2.5999999999999996" valueMax="2.9" valueMin="2.3">
2.3
<emphasis id="B974EA98FFB7F475447B37687C543D1A" box="[319,333,806,829]" italics="true" pageId="73" pageNumber="74"></emphasis>
2.9 mm
</quantity>
(N=3), width
<quantity id="4CF89B6FFFB7F475470037687FFE3D1A" box="[580,743,806,830]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="73" pageNumber="74" unit="mm" value="0.42" valueMax="0.44" valueMin="0.4">
0.40
<emphasis id="B974EA98FFB7F475473037687F9B3D1A" box="[628,642,806,829]" italics="true" pageId="73" pageNumber="74"></emphasis>
0.44 mm
</quantity>
(N=3).
</paragraph>
<paragraph id="8BBF368AFFB7F4754583370778623D45" blockId="73.[151,1437,151,1846]" box="[199,1403,841,866]" pageId="73" pageNumber="74">
Abdomen: Length
<quantity id="4CF89B6FFFB7F47544DE37077F383D46" box="[410,545,841,866]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.2" metricValueMin="5.8" pageId="73" pageNumber="74" unit="mm" value="6.0" valueMax="6.2" valueMin="5.8">
5.8
<emphasis id="B974EA98FFB7F47544F937047CD23D46" box="[445,459,842,865]" italics="true" pageId="73" pageNumber="74"></emphasis>
6.2 mm
</quantity>
(N=3), width
<quantity id="4CF89B6FFFB7F475478537047E0E3D46" box="[705,791,842,866]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="73" pageNumber="74" unit="mm" value="1.3">1.3 mm</quantity>
(N=1). Dark brown, presumably black in life, conical.
</paragraph>
<paragraph id="8BBF368AFFB7F4754583372379DE3D8D" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">
Terminalia (fig. 203): Hypopygium width
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0.92
<emphasis id="B974EA98FFB7F47547A137207FEA3DA2" box="[741,755,878,901]" italics="true" pageId="73" pageNumber="74"></emphasis>
1.10 mm
</quantity>
(N=2). Epandrium not bilobate, posterior margin nearly straight with rounded, slightly protruding posterior corners. Gonostylus large and robust.
</paragraph>
<paragraph id="8BBF368AFFB7F475458337FB7FBA3DE9" blockId="73.[151,1437,151,1846]" box="[199,675,949,974]" pageId="73" pageNumber="74">
Female: Total length 9.3
<emphasis id="B974EA98FFB7F475449C37F87CFF3DEA" box="[472,486,950,973]" italics="true" pageId="73" pageNumber="74"></emphasis>
12.0 mm (N=13)
</paragraph>
<paragraph id="8BBF368AFFB7F475458337977F233A1D" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">
Head (fig. 204): Length
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1.19
<emphasis id="B974EA98FFB7F475474B37947F043DD6" box="[527,541,986,1009]" italics="true" pageId="73" pageNumber="74"></emphasis>
1.39 mm
</quantity>
(N=6), width
<quantity id="4CF89B6FFFB7F475466C37947ED53DD6" box="[808,972,986,1010]" metricMagnitude="-3" metricUnit="m" metricValue="1.125" metricValueMax="1.32" metricValueMin="0.93" pageId="73" pageNumber="74" unit="mm" value="1.125" valueMax="1.32" valueMin="0.93">
0.93
<emphasis id="B974EA98FFB7F475461D37947E7E3DD6" box="[857,871,986,1009]" italics="true" pageId="73" pageNumber="74"></emphasis>
1.32 mm
</quantity>
(N=6). Black. Antennal flagellum
<quantity id="4CF89B6FFFB7F475401937977DE43A32" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.2" metricValueMin="5.5" pageId="73" pageNumber="74" unit="mm" value="0.585" valueMax="0.62" valueMin="0.55">
0.55
<emphasis id="B974EA98FFB7F47540CA379478853DD6" box="[1422,1436,986,1009]" italics="true" pageId="73" pageNumber="74"></emphasis>
0.62 mm
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long (N=2), 8-segmented. Ocellar tubercle small and low. Palp short and robust. Compound eye rather small, about one third length of head.
</paragraph>
<paragraph id="8BBF368AFFB7F4754583300B796A3A79" blockId="73.[151,1437,151,1846]" box="[199,1139,1093,1118]" pageId="73" pageNumber="74">
Thorax: Length
<quantity id="4CF89B6FFFB7F475443930087F1D3A7A" box="[381,516,1094,1118]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.8" metricValueMin="2.8" pageId="73" pageNumber="74" unit="mm" value="3.3" valueMax="3.8" valueMin="2.8">
2.8
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3.8 mm
</quantity>
(N=13), width
<quantity id="4CF89B6FFFB7F47547F630087E203A7A" box="[690,825,1094,1118]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="3.1" metricValueMin="2.1" pageId="73" pageNumber="74" unit="mm" value="2.6" valueMax="3.1" valueMin="2.1">
2.1
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3.1 mm
</quantity>
(N=5). Haltere dark brown.
</paragraph>
<paragraph id="8BBF368AFFB7F4754583302778333AD6" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">
Wing (fig. 198): Length
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6.7
<emphasis id="B974EA98FFB7F475474730247F083AA6" box="[515,529,1130,1153]" italics="true" pageId="73" pageNumber="74"></emphasis>
10.6 mm
</quantity>
(N=14), width 3.0
<emphasis id="B974EA98FFB7F475460830247E433AA6" box="[844,858,1130,1153]" italics="true" pageId="73" pageNumber="74"></emphasis>
<quantity id="4CF89B6FFFB7F475461E30247EA83AA6" box="[858,945,1130,1154]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="73" pageNumber="74" unit="mm" value="4.3">4.3 mm</quantity>
(N=14), length/width = 2.08
<emphasis id="B974EA98FFB7F47541BF302478103AA6" box="[1275,1289,1130,1153]" italics="true" pageId="73" pageNumber="74"></emphasis>
2.62 (N=14). Grayish hyaline, pterostigma oval, dark brown, wing mesally with a chocolate-brown band stretching from the origin of R s distad to pterostigma, posteriorly to CuA1.Crossvein R-M short but distinctive, about one fifth as long as basal section of R s. The CuA-veins apically straight or nearly so. Wing vein measurements in
<tableCitation id="C6820331FFB7F47541943096783F3AD6" box="[1232,1318,1240,1265]" captionStart="TABLE 4" captionStartId="58.[164,225,1207,1223]" captionTargetBox="[164,1421,1249,1825]" captionTargetPageId="58" captionText="TABLE 4. Bibio spp., wing measurements" httpUri="http://table.plazi.org/id/DF7F6602FFC4F40645E030F97CC73AE0" pageId="73" pageNumber="74" tableUuid="DF7F6602FFC4F40645E030F97CC73AE0">Table 4</tableCitation>
.
</paragraph>
<paragraph id="8BBF368AFFB7F4754583314E7EDC3BEB" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">
Legs (fig. 205): femorae and tibiae brownish yellow, tibiae apically darker. Fore femur relatively slender, about three times as long as wide in lateral view. Fore tibia with spur about one third the length of anterior spine. All tarsi slender, tarsomeres brownish yellow, apically darkened. Hind femur slightly clavate, hind tibia slender. Metatibial spur rather long, blunt. Fore femur length
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1.8
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2.4 mm
</quantity>
(N=9), width
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0.69
<emphasis id="B974EA98FFB7F475468031227ECB3BA4" box="[964,978,1388,1411]" italics="true" pageId="73" pageNumber="74"></emphasis>
0.86 mm
</quantity>
(N=7), fore tibia length
<quantity id="4CF89B6FFFB7F475400C31257DDA3B80" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" pageId="73" pageNumber="74" unit="mm" value="2.05" valueMax="2.4" valueMin="1.7">
1.7
<emphasis id="B974EA98FFB7F475402F312278603BA4" box="[1387,1401,1388,1411]" italics="true" pageId="73" pageNumber="74"></emphasis>
2.4 mm
</quantity>
(N=11), width
<quantity id="4CF89B6FFFB7F475442B31C17F0B3B80" box="[367,530,1423,1448]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.699999999999999" metricValueMin="4.6000000000000005" pageId="73" pageNumber="74" unit="mm" value="0.46499999999999997" valueMax="0.47" valueMin="0.46">
0.46
<emphasis id="B974EA98FFB7F47544E431DE7CB73B80" box="[416,430,1424,1447]" italics="true" pageId="73" pageNumber="74"></emphasis>
0.47 mm
</quantity>
(N=2), hind femur length
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2.6
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3.6 mm
</quantity>
(N=11), width
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0.54
<emphasis id="B974EA98FFB7F47541DB31DE79B43B80" box="[1183,1197,1424,1447]" italics="true" pageId="73" pageNumber="74"></emphasis>
0.76 mm
</quantity>
(N=9), hind tibia length
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2.7
<emphasis id="B974EA98FFB7F475447B31FA7C543BEC" box="[319,333,1460,1483]" italics="true" pageId="73" pageNumber="74"></emphasis>
3.9 mm
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(N=11), hind first tarsomere 1.0
<emphasis id="B974EA98FFB7F475464B31FA7E043BEC" box="[783,797,1460,1483]" italics="true" pageId="73" pageNumber="74"></emphasis>
<quantity id="4CF89B6FFFB7F475465931FD7E6A3BEC" box="[797,883,1459,1483]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="73" pageNumber="74" unit="mm" value="1.2">1.2 mm</quantity>
(N=5).
</paragraph>
<paragraph id="8BBF368AFFB7F4754583319979713BD7" blockId="73.[151,1437,151,1846]" box="[199,1128,1495,1520]" pageId="73" pageNumber="74">
Abdomen: Length
<quantity id="4CF89B6FFFB7F47544DE31997F383BC8" box="[410,545,1495,1520]" metricMagnitude="-3" metricUnit="m" metricValue="7.6499999999999995" metricValueMax="9.2" metricValueMin="6.1" pageId="73" pageNumber="74" unit="mm" value="7.6499999999999995" valueMax="9.2" valueMin="6.1">
6.1
<emphasis id="B974EA98FFB7F47544F931967CD23BC8" box="[445,459,1496,1519]" italics="true" pageId="73" pageNumber="74"></emphasis>
9.2 mm
</quantity>
(N=15), width
<quantity id="4CF89B6FFFB7F475478B31997E4F3BC8" box="[719,854,1495,1520]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="73" pageNumber="74" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">
2.5
<emphasis id="B974EA98FFB7F47547B631967E193BC8" box="[754,768,1496,1519]" italics="true" pageId="73" pageNumber="74"></emphasis>
2.8 mm
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(N=6). Blackish brown.
</paragraph>
<paragraph id="8BBF368AFFB7F475458331B57DEB381F" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">Terminalia (fig. 206): Cercus slender, rounded. It may appear to have two segments but this is probably an artefact.</paragraph>
<paragraph id="8BBF368AFFB7F4754583320D7CFC3911" blockId="73.[151,1437,151,1846]" pageId="73" pageNumber="74">
There is also a specimen in FSL (FSL 391912) labeled as a
<typeStatus id="54BB8828FFB7F475461A320A7EA7387B" box="[862,958,1604,1628]" pageId="73" pageNumber="74" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C004D09FFB7F47546A6320A7960387C" box="[994,1145,1603,1627]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B974EA98FFB7F47546A6320A7960387C" box="[994,1145,1603,1627]" italics="true" pageId="73" pageNumber="74">B. spectabilis</emphasis>
</taxonomicName>
. This specimen, however, has a slender fore tibia and a distinctive vein R2+3 and belongs in the genus
<emphasis id="B974EA98FFB7F475414A322979443858" box="[1038,1117,1639,1663]" italics="true" pageId="73" pageNumber="74">
<taxonomicName id="4C004D09FFB7F475414A322979403858" box="[1038,1113,1639,1663]" class="Insecta" family="Bibionidae" genus="Plecia" kingdom="Animalia" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">Plecia</taxonomicName>
.
</emphasis>
In ETHZ there is a female specimen labeled “
<taxonomicName id="4C004D09FFB7F475443532C07F683881" authority="Heer" authorityName="Heer" box="[369,625,1678,1702]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B974EA98FFB7F475443532C07F363881" box="[369,559,1678,1702]" italics="true" pageId="73" pageNumber="74">Bibio spectabilis</emphasis>
Heer
</taxonomicName>
” (ETHZ Pl. I 855) from the Miocene site of Öhningen. The specimen actually belongs to
<taxonomicName id="4C004D09FFB7F475443D32FC7E2038ED" authority="Skartveit &amp; Pika 2014" authorityName="Skartveit &amp; Pika" authorityYear="2014" box="[377,825,1714,1739]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="moestus">
<emphasis id="B974EA98FFB7F475443D32FC7F0038ED" box="[377,537,1714,1738]" italics="true" pageId="73" pageNumber="74">Bibio moestus</emphasis>
(
<bibRefCitation id="EF914B7BFFB7F475476E32FC7E2838ED" author="Skartveit" box="[554,817,1714,1739]" pageId="73" pageNumber="74" refString="Skartveit, J. &amp; Pika, M. (2014) Revision of Bibionidae (Diptera) named by Oswald Heer from the Miocene of Ohningen, Southern Germany. Mitteilungen der schweizerischen entomologischen Gesellschaft, 87, 103 - 134." type="journal article" year="2014">Skartveit &amp; Pika 2014</bibRefCitation>
)
</taxonomicName>
, and the name
<taxonomicName id="4C004D09FFB7F47546AE32FC79FE38ED" authority="Heer" authorityName="Heer" box="[1002,1255,1714,1738]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="spectabilis">
<emphasis id="B974EA98FFB7F47546AE32FC79B138ED" box="[1002,1192,1714,1738]" italics="true" pageId="73" pageNumber="74">Bibio spectabilis</emphasis>
Heer
</taxonomicName>
was apparently never formally published and so does not affect the nomenclature of this species. Using the dark-banded wings as an identification character is problematic since the vast majority of the specimens from Aix-en-Provence have no preserved wing pigmentation.
</paragraph>
</subSubSection>
</treatment>
</document>