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<document ID-DOI="10.17109/AZH.68.3.231.2022" ID-ISSN="2064-2474" ID-Zenodo-Dep="7160685" approvalRequired="3" approvalRequired_for_textStreams="3" checkinTime="1665198218098" checkinUser="felipe" docAuthor="Ermilov, Sergey G., Subías, Luis S., Shtanchaeva, Umukusum Ya. &amp; Friedrich, Stefan" docDate="2022" docId="03A9878DA2371E49FE5259D726AD3472" docLanguage="en" docName="ActaZoolAcadSciH.68.3.231-238.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 68 (3)" docSource="http://dx.doi.org/10.17109/azh.68.3.231.2022" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.1:ActaZoolAcadSciH.2015-.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-.journal_article" docStyleVersion="1" docTitle="Ceratobates pachiteaensis Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich 2022, sp. n." docType="treatment" docVersion="2" lastPageNumber="236" masterDocId="FF90FFF5A2351E4CFFC3580E253F3135" masterDocTitle="A New Species Of Ceratobates (Acari, Oribatida) From Peru And A Key To Known Species Of The Genus" masterLastPageNumber="238" masterPageNumber="231" pageNumber="233" updateTime="1665199574211" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:titleInfo>
<mods:title>A New Species Of Ceratobates (Acari, Oribatida) From Peru And A Key To Known Species Of The Genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ermilov, Sergey G.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Subías, Luis S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shtanchaeva, Umukusum Ya.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Friedrich, Stefan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>68</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
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<mods:start>231</mods:start>
<mods:end>238</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.17109/azh.68.3.231.2022</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.17109/AZH.68.3.231.2022</mods:identifier>
<mods:identifier type="ISSN">2064-2474</mods:identifier>
<mods:identifier type="Zenodo-Dep">7160685</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472" httpUri="http://treatment.plazi.org/id/03A9878DA2371E49FE5259D726AD3472" lastPageId="5" lastPageNumber="236" pageId="2" pageNumber="233">
<subSubSection box="[401,824,473,500]" pageId="2" pageNumber="233" type="nomenclature">
<paragraph blockId="2.[401,824,473,500]" box="[401,824,473,500]" pageId="2" pageNumber="233">
<heading box="[401,824,473,500]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="233" reason="2">
<taxonomicName authority="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich, 2022" authorityName="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich" authorityYear="2022" box="[401,747,473,499]" class="Arachnida" family="Chamobatidae" genus="Ceratobates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="pachiteaensis" status="sp. n.">
<emphasis bold="true" box="[401,747,473,499]" pageId="2" pageNumber="233">Ceratobates pachiteaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[754,824,474,500]" pageId="2" pageNumber="233" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="236" pageId="2" pageNumber="233" type="description">
<paragraph blockId="2.[550,675,508,535]" box="[550,675,508,535]" pageId="2" pageNumber="233">
<heading box="[550,675,508,535]" centered="true" fontSize="11" level="3" pageId="2" pageNumber="233" reason="3">
(
<figureCitation box="[559,636,508,535]" captionStart="Fig" captionStartId="3.[174,210,1523,1545]" captionTargetBox="[176,1114,395,1493]" captionTargetId="figure-108@3.[174,1116,393,1497]" captionTargetPageId="3" captionText="Fig. 1. Ceratobates pachiteaensis sp. n., adult, legs not shown: A = dorsal view; B = ventral view; C = right lateral view; D = posterior view, right half. Scale bar 50 μm" figureDoi="http://doi.org/10.5281/zenodo.7160687" httpUri="https://zenodo.org/record/7160687/files/figure.png" pageId="2" pageNumber="233">Figs 1</figureCitation>
,
<figureCitation box="[650,666,509,535]" captionStart="Fig" captionStartId="4.[136,172,1467,1489]" captionTargetBox="[139,1085,356,1440]" captionTargetId="figure-90@4.[136,1088,353,1443]" captionTargetPageId="4" captionText="Fig. 2. Ceratobates pachiteaensis sp. n., adult: A = subcapitulum, ventral view; B = palp, right, antiaxial view; C = chelicera, left, paraxial view; D = leg I, right, antiaxial view; E = leg II, without tarsus, right, antiaxial view; F = leg III, without tarsus, left, antiaxial view; G = leg IV, left, antiaxial view. Scale bar 20 μm" figureDoi="http://doi.org/10.5281/zenodo.7160691" httpUri="https://zenodo.org/record/7160691/files/figure.png" pageId="2" pageNumber="233">2</figureCitation>
)
</heading>
</paragraph>
<paragraph blockId="2.[136,1089,565,794]" pageId="2" pageNumber="233">
Diagnosis. Body length: 365381. Rostrum bidentate. Lamellar cusp with long lateral tooth. Rostral seta setiform, barbed; lamellar and interlamellar setae thick, barbed;
<emphasis box="[358,381,634,659]" italics="true" pageId="2" pageNumber="233">ro</emphasis>
shortest,
<emphasis box="[502,525,634,659]" italics="true" pageId="2" pageNumber="233">in</emphasis>
longest. Bothridial seta long, clavate, slightly barbed; head elongate oval. All notogastral setae setiform, slightly barbed. Epimeres III, IV with two and three pairs of setae, respectively;
<emphasis box="[920,945,700,726]" italics="true" pageId="2" pageNumber="233">1c</emphasis>
of medium length, thickened, barbed; other setae and anogenital setae short, setiform, slightly barbed. Leg tarsi with one (in typical case) or three claws.
</paragraph>
<paragraph blockId="2.[136,1089,828,1570]" pageId="2" pageNumber="233">
Description of adult.
<emphasis box="[435,578,829,850]" italics="true" pageId="2" pageNumber="233">Measurements</emphasis>
. Body length: 381 (
<typeStatus box="[798,895,828,850]" pageId="2" pageNumber="233">holotype</typeStatus>
, female), 365381 (
<specimenCount box="[143,304,857,879]" pageId="2" pageNumber="233" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
,
<specimenCount box="[317,428,857,879]" pageId="2" pageNumber="233" type="male">two males</specimenCount>
and
<specimenCount box="[485,618,857,879]" pageId="2" pageNumber="233" type="female">two females</specimenCount>
); notogaster width: 265 (
<typeStatus box="[887,985,857,879]" pageId="2" pageNumber="233">holotype</typeStatus>
), 249265 (
<specimenCount box="[143,304,886,908]" pageId="2" pageNumber="233" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
). No difference between males and females in size (in studied population).
</paragraph>
<paragraph blockId="2.[136,1089,828,1570]" pageId="2" pageNumber="233">
<emphasis box="[197,310,915,936]" italics="true" pageId="2" pageNumber="233">Integument</emphasis>
. Natural body color brown. Surface of body microsculpturing microgran- ulate; epimeral region, genital plate and subcapitular mentum hardly striate; antiaxial side of leg femora I-IV and trochanters III, IV with rounded end elongate tubercles. Podosomal part of body partially with granulate cerotegument.
</paragraph>
<paragraph blockId="2.[136,1089,828,1570]" pageId="2" pageNumber="233">
<emphasis box="[197,305,1030,1051]" italics="true" pageId="2" pageNumber="233">Prodorsum</emphasis>
. Rostrum bidentate, with two strong teeth and semi-rectangular incision between them. Lamella (without cusp) shorter than half of prodorsum; lamellar cusp short, with long (longer than distance between insertions of lamellar setae), thin lateral tooth. Rostral seta (6573) setiform, barbed, mediodistally directed anteromediad. Lamellar (77 82) and interlamellar (102110) thick, barbed. Bothridial seta (6169) clavate, with longer stalk and shorter, elongate, slightly barbed head. Exobothridial seta not observed. Tutorial tip and genal tooth narrowly triangular. Dorsosejugal porose area not observed.
</paragraph>
<paragraph blockId="2.[136,1089,828,1570]" pageId="2" pageNumber="233">
<emphasis box="[197,304,1232,1253]" italics="true" pageId="2" pageNumber="233">Notogaster</emphasis>
. Pteromorph forming slight right angle laterally (visible in lateral aspect). All notogastral setae (
<emphasis box="[377,387,1261,1282]" italics="true" pageId="2" pageNumber="233">c</emphasis>
,
<emphasis box="[402,419,1260,1281]" italics="true" pageId="2" pageNumber="233">la</emphasis>
,
<emphasis box="[435,460,1260,1281]" italics="true" pageId="2" pageNumber="233">lm</emphasis>
,
<emphasis box="[476,488,1260,1281]" italics="true" pageId="2" pageNumber="233">h</emphasis>
3: 2832;
<emphasis box="[585,604,1260,1281]" italics="true" pageId="2" pageNumber="233">lp</emphasis>
,
<emphasis box="[619,631,1260,1281]" italics="true" pageId="2" pageNumber="233">h</emphasis>
2: 2024;
<emphasis box="[729,741,1260,1281]" italics="true" pageId="2" pageNumber="233">h</emphasis>
1,
<emphasis box="[763,775,1261,1282]" italics="true" pageId="2" pageNumber="233">p</emphasis>
1
<emphasis box="[794,806,1261,1282]" italics="true" pageId="2" pageNumber="233">p</emphasis>
3: 1416) setiform, slightly barbed. Notogastral lyrifissure
<emphasis box="[480,499,1290,1311]" italics="true" pageId="2" pageNumber="233">ip</emphasis>
not observed;
<emphasis box="[666,683,1290,1311]" italics="true" pageId="2" pageNumber="233">ia</emphasis>
,
<emphasis box="[699,724,1290,1311]" italics="true" pageId="2" pageNumber="233">im</emphasis>
,
<emphasis box="[741,760,1289,1310]" italics="true" pageId="2" pageNumber="233">ih</emphasis>
, and
<emphasis box="[826,854,1290,1311]" italics="true" pageId="2" pageNumber="233">ips</emphasis>
visible. Opisthonotal gland opening distinct.
</paragraph>
<paragraph blockId="2.[136,1089,828,1570]" pageId="2" pageNumber="233">
<taxonomicName authorityName="Krefft" authorityYear="1871" box="[197,318,1347,1368]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="233" phylum="Nematoda" rank="genus">
<emphasis box="[197,318,1347,1368]" italics="true" pageId="2" pageNumber="233">Gnathosoma</emphasis>
</taxonomicName>
. Subcapitulum size: 8290 × 6165. All subcapitular setae (
<emphasis box="[943,954,1347,1368]" italics="true" pageId="2" pageNumber="233">a</emphasis>
: 1416;
<emphasis box="[1039,1058,1347,1368]" italics="true" pageId="2" pageNumber="233">m</emphasis>
,
<emphasis box="[1071,1083,1347,1368]" italics="true" pageId="2" pageNumber="233">h</emphasis>
: 2024) setiform, slightly barbed. Adoral seta (6) setiform, roughened. Palp (length: 6165) setation: 0-2-1-3-9(+ω). Postpalpal seta (4) spiniform, smooth. Axillary saccule not observed. Chelicera (length: 8290) with two setiform, barbed setae (
<emphasis box="[835,867,1433,1454]" italics="true" pageId="2" pageNumber="233">cha</emphasis>
: 2832;
<emphasis box="[952,985,1433,1454]" italics="true" pageId="2" pageNumber="233">chb</emphasis>
: 1820).
</paragraph>
<paragraph blockId="2.[136,1089,828,1570]" pageId="2" pageNumber="233">
<emphasis box="[197,589,1462,1484]" italics="true" pageId="2" pageNumber="233">Epimeral and lateral podosomal regions</emphasis>
. Epimeral setal formula: 3123. Seta
<emphasis box="[1018,1039,1462,1483]" italics="true" pageId="2" pageNumber="233">1c</emphasis>
(32) thickened, barbed; other setae (
<emphasis box="[468,491,1491,1512]" italics="true" pageId="2" pageNumber="233">1a</emphasis>
,
<emphasis box="[503,526,1491,1512]" italics="true" pageId="2" pageNumber="233">2a</emphasis>
,
<emphasis box="[538,560,1491,1512]" italics="true" pageId="2" pageNumber="233">3a</emphasis>
: 12;
<emphasis box="[609,632,1491,1512]" italics="true" pageId="2" pageNumber="233">1b</emphasis>
,
<emphasis box="[644,668,1491,1512]" italics="true" pageId="2" pageNumber="233">3b</emphasis>
,
<emphasis box="[680,702,1491,1512]" italics="true" pageId="2" pageNumber="233">4a</emphasis>
,
<emphasis box="[715,737,1491,1512]" italics="true" pageId="2" pageNumber="233">4b</emphasis>
,
<emphasis box="[750,772,1491,1512]" italics="true" pageId="2" pageNumber="233">4c</emphasis>
: 16) setiform, slightly barbed. Custodium, discidium and circumpedal carina well developed. Humeral porose area
<emphasis box="[1059,1088,1519,1540]" italics="true" pageId="2" pageNumber="233">Ah</emphasis>
poorly visible, oval;
<emphasis box="[351,387,1549,1570]" italics="true" pageId="2" pageNumber="233">Am</emphasis>
absent.
</paragraph>
<paragraph blockId="3.[174,1127,190,356]" pageId="3" pageNumber="234">
<emphasis box="[234,412,190,212]" italics="true" pageId="3" pageNumber="234">Anogenital region</emphasis>
. Genital (
<emphasis box="[520,532,191,212]" italics="true" pageId="3" pageNumber="234">g</emphasis>
<subScript attach="left" box="[532,539,206,218]" fontSize="5" pageId="3" pageNumber="234">1</subScript>
: 1620;
<emphasis box="[627,639,191,212]" italics="true" pageId="3" pageNumber="234">g</emphasis>
<subScript attach="left" box="[639,646,206,218]" fontSize="5" pageId="3" pageNumber="234">2</subScript>
<emphasis box="[658,670,191,212]" italics="true" pageId="3" pageNumber="234">g</emphasis>
<subScript attach="left" box="[670,677,205,217]" fontSize="5" pageId="3" pageNumber="234">6</subScript>
: 16), aggenital (16), anal (16), and adanal (12) setae setiform, slightly barbed. Adanal lyrifissure located close and parallel to anal plate. Postanal porose area oval (1012 × 4).
</paragraph>
<paragraph blockId="3.[174,1127,190,356]" lastBlockId="4.[136,1089,190,327]" lastPageId="4" lastPageNumber="235" pageId="3" pageNumber="234">
<emphasis box="[234,278,277,298]" italics="true" pageId="3" pageNumber="234">Legs</emphasis>
.
<typeStatus box="[289,389,277,299]" pageId="3" pageNumber="234">Holotype</typeStatus>
and
<specimenCount box="[439,603,277,299]" pageId="3" pageNumber="234" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
monodactylous;
<specimenCount box="[783,921,277,298]" pageId="3" pageNumber="234" type="generic" typeStatus="paratype">one paratype</specimenCount>
heterotridactylous. All claws slightly barbed dorsally. Paraxial porose area on femora I-IV and on trochanters III, IV distinct. Genua I, II with ventral tooth. Formulas of leg setation and solenidia: I (153418) [122], II (153415) [112], III (231315) [110], IV (122312) [010]; homologies of setae and solenidia indicated in
<tableCitation box="[727,803,219,241]" captionStart="Table 1" captionStartId="5.[263,324,192,213]" captionText="Table 1. Leg setation and solenidia of adult Ceratobates pachiteaensis sp. n." pageId="4" pageNumber="235">Table 1</tableCitation>
. Famulus of tarsus I short, erect, slightly swollen distally, inserted before solenidion ω
<subScript attach="left" box="[766,773,263,275]" fontSize="5" pageId="4" pageNumber="235">1</subScript>
. Seta
<emphasis box="[834,843,248,269]" italics="true" pageId="4" pageNumber="235">s</emphasis>
of tarsus I eupathidial, located anterior to setae
<emphasis box="[398,408,277,298]" italics="true" pageId="4" pageNumber="235">a</emphasis>
. Tibia II and genua I, II with thick lateral (antiaxial) seta. Solenidion φ of tibia IV represented by alveolus.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160687" ID-Zenodo-Dep="7160687" httpUri="https://zenodo.org/record/7160687/files/figure.png" pageId="3" pageNumber="234" startId="3.[174,210,1523,1545]" targetBox="[176,1114,395,1493]" targetPageId="3">
<paragraph blockId="3.[174,1126,1523,1574]" pageId="3" pageNumber="234">
<emphasis bold="true" box="[174,241,1523,1545]" pageId="3" pageNumber="234">Fig. 1.</emphasis>
<taxonomicName authority="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich, 2022" authorityName="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich" authorityYear="2022" box="[248,495,1523,1544]" class="Arachnida" family="Chamobatidae" genus="Ceratobates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="pachiteaensis" status="sp. n.">
<emphasis box="[248,495,1523,1544]" italics="true" pageId="3" pageNumber="234">Ceratobates pachiteaensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[505,563,1523,1544]" pageId="3" pageNumber="234">
<taxonomicNameLabel box="[505,563,1523,1544]" pageId="3" pageNumber="234" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult, legs not shown: A = dorsal view; B = ventral view; C = right lateral view; D = posterior view, right half. Scale bar 50 μm
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160691" ID-Zenodo-Dep="7160691" httpUri="https://zenodo.org/record/7160691/files/figure.png" pageId="4" pageNumber="235" startId="4.[136,172,1467,1489]" targetBox="[139,1085,356,1440]" targetPageId="4">
<paragraph blockId="4.[136,1089,1467,1576]" pageId="4" pageNumber="235">
<emphasis bold="true" box="[136,200,1467,1489]" pageId="4" pageNumber="235">Fig. 2.</emphasis>
<taxonomicName authority="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich, 2022" authorityName="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich" authorityYear="2022" box="[206,450,1468,1489]" class="Arachnida" family="Chamobatidae" genus="Ceratobates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="pachiteaensis" status="sp. n.">
<emphasis box="[206,450,1468,1489]" italics="true" pageId="4" pageNumber="235">Ceratobates pachiteaensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[455,512,1468,1489]" pageId="4" pageNumber="235">
<taxonomicNameLabel box="[455,512,1468,1489]" pageId="4" pageNumber="235" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult: A = subcapitulum, ventral view; B = palp, right, antiaxial view; C = chelicera, left, paraxial view; D = leg I, right, antiaxial view; E = leg II, without tarsus, right, antiaxial view; F = leg III, without tarsus, left, antiaxial view; G = leg
</paragraph>
<paragraph blockId="4.[136,1089,1467,1576]" box="[407,818,1554,1576]" pageId="4" pageNumber="235">IV, left, antiaxial view. Scale bar 20 μm</paragraph>
</caption>
<caption box="[263,1036,191,213]" pageId="5" pageNumber="236" startId="5.[263,324,192,213]">
<paragraph blockId="5.[263,1036,191,213]" box="[263,1036,191,213]" pageId="5" pageNumber="236">
<emphasis bold="true" box="[263,342,191,213]" pageId="5" pageNumber="236">Table 1</emphasis>
. Leg setation and solenidia of adult
<taxonomicName authority="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich, 2022" authorityName="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich" authorityYear="2022" box="[727,972,191,212]" class="Arachnida" family="Chamobatidae" genus="Ceratobates" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="pachiteaensis" status="sp. n.">
<emphasis box="[727,972,191,212]" italics="true" pageId="5" pageNumber="236">Ceratobates pachiteaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[978,1036,192,213]" pageId="5" pageNumber="236" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="236">
<table box="[184,1105,231,466]" gridcols="6" gridrows="5" pageId="5" pageNumber="236">
<tr box="[184,1105,231,252]" gridrow="0" pageId="5" pageNumber="236">
<th box="[184,224,231,252]" gridcol="0" gridrow="0" pageId="5" pageNumber="236">Leg</th>
<th box="[250,294,231,252]" gridcol="1" gridrow="0" pageId="5" pageNumber="236">
<emphasis box="[260,284,231,252]" italics="true" pageId="5" pageNumber="236">Tr</emphasis>
</th>
<th box="[317,447,231,252]" gridcol="2" gridrow="0" pageId="5" pageNumber="236">
<emphasis box="[371,394,231,252]" italics="true" pageId="5" pageNumber="236">Fe</emphasis>
</th>
<th box="[474,554,231,252]" gridcol="3" gridrow="0" pageId="5" pageNumber="236">
<emphasis box="[501,528,231,252]" italics="true" pageId="5" pageNumber="236">Ge</emphasis>
</th>
<th box="[582,717,231,252]" gridcol="4" gridrow="0" pageId="5" pageNumber="236">
<emphasis box="[639,660,231,252]" italics="true" pageId="5" pageNumber="236">Ti</emphasis>
</th>
<th box="[749,1105,231,252]" gridcol="5" gridrow="0" pageId="5" pageNumber="236">
<emphasis box="[915,938,231,252]" italics="true" pageId="5" pageNumber="236">Ta</emphasis>
</th>
</tr>
<tr box="[184,1105,269,325]" gridrow="1" pageId="5" pageNumber="236">
<th box="[184,224,269,325]" gridcol="0" gridrow="1" pageId="5" pageNumber="236">I</th>
<td box="[250,294,269,325]" gridcol="1" gridrow="1" pageId="5" pageNumber="236">
<emphasis box="[262,281,270,291]" italics="true" pageId="5" pageNumber="236">v</emphasis>
</td>
<td box="[317,447,269,325]" gridcol="2" gridrow="1" pageId="5" pageNumber="236">
<emphasis box="[317,447,269,291]" italics="true" pageId="5" pageNumber="236">d, (l), bv”, v”</emphasis>
</td>
<td box="[474,554,269,325]" gridcol="3" gridrow="1" pageId="5" pageNumber="236">
<emphasis box="[474,533,269,291]" italics="true" pageId="5" pageNumber="236">(l), v,</emphasis>
σ
</td>
<td box="[582,717,269,325]" gridcol="4" gridrow="1" pageId="5" pageNumber="236">
<emphasis box="[582,650,269,291]" italics="true" pageId="5" pageNumber="236">(l), (v),</emphasis>
φ, φ 1 2
</td>
<td box="[749,1105,269,325]" gridcol="5" gridrow="1" pageId="5" pageNumber="236">
<emphasis italics="true" pageId="5" pageNumber="236">ft, (tc), (it), (p), (u), (a), s, (pv), (pl), v,</emphasis>
ε, ω1, ω2
</td>
</tr>
<tr box="[184,1105,337,393]" gridrow="2" pageId="5" pageNumber="236">
<th box="[184,224,337,393]" gridcol="0" gridrow="2" pageId="5" pageNumber="236">II</th>
<td box="[250,294,337,393]" gridcol="1" gridrow="2" pageId="5" pageNumber="236">
<emphasis box="[262,281,337,358]" italics="true" pageId="5" pageNumber="236">v</emphasis>
</td>
<td box="[317,447,337,393]" gridcol="2" gridrow="2" pageId="5" pageNumber="236">
<emphasis box="[317,447,337,358]" italics="true" pageId="5" pageNumber="236">d, (l), bv”, v”</emphasis>
</td>
<td box="[474,554,337,393]" gridcol="3" gridrow="2" pageId="5" pageNumber="236">
<emphasis box="[474,533,337,358]" italics="true" pageId="5" pageNumber="236">(l), v,</emphasis>
σ
</td>
<td box="[582,717,337,393]" gridcol="4" gridrow="2" pageId="5" pageNumber="236">
<emphasis box="[604,672,337,358]" italics="true" pageId="5" pageNumber="236">(l), (v),</emphasis>
φ
</td>
<td box="[749,1105,337,393]" gridcol="5" gridrow="2" pageId="5" pageNumber="236">
<emphasis box="[749,1068,337,359]" italics="true" pageId="5" pageNumber="236">(ft), (tc), (it), (p), (u), (a), s, (pv),</emphasis>
ω1, ω2
</td>
</tr>
<tr box="[184,1105,405,426]" gridrow="3" pageId="5" pageNumber="236">
<th box="[184,224,405,426]" gridcol="0" gridrow="3" pageId="5" pageNumber="236">III</th>
<td box="[250,294,405,426]" gridcol="1" gridrow="3" pageId="5" pageNumber="236">
<emphasis box="[250,294,405,426]" italics="true" pageId="5" pageNumber="236">l, v</emphasis>
</td>
<td box="[317,447,405,426]" gridcol="2" gridrow="3" pageId="5" pageNumber="236">
<emphasis box="[343,421,405,426]" italics="true" pageId="5" pageNumber="236">d, l, ev</emphasis>
</td>
<td box="[474,554,405,426]" gridcol="3" gridrow="3" pageId="5" pageNumber="236">
<emphasis box="[494,513,405,426]" italics="true" pageId="5" pageNumber="236">l,</emphasis>
σ
</td>
<td box="[582,717,405,426]" gridcol="4" gridrow="3" pageId="5" pageNumber="236">
<emphasis box="[608,667,405,426]" italics="true" pageId="5" pageNumber="236">l, (v),</emphasis>
φ
</td>
<td box="[749,1105,405,426]" gridcol="5" gridrow="3" pageId="5" pageNumber="236">
<emphasis box="[771,1083,405,426]" italics="true" pageId="5" pageNumber="236">(ft), (tc), (it), (p), (u), (a), s, (pv)</emphasis>
</td>
</tr>
<tr box="[184,1105,443,466]" gridrow="4" pageId="5" pageNumber="236">
<th box="[184,224,443,466]" gridcol="0" gridrow="4" pageId="5" pageNumber="236">IV</th>
<td box="[250,294,443,466]" gridcol="1" gridrow="4" pageId="5" pageNumber="236">
<emphasis box="[262,281,444,465]" italics="true" pageId="5" pageNumber="236">v</emphasis>
</td>
<td box="[317,447,443,466]" gridcol="2" gridrow="4" pageId="5" pageNumber="236">
<emphasis box="[356,408,443,465]" italics="true" pageId="5" pageNumber="236">d, ev</emphasis>
</td>
<td box="[474,554,443,466]" gridcol="3" gridrow="4" pageId="5" pageNumber="236">
<emphasis box="[496,533,443,464]" italics="true" pageId="5" pageNumber="236">d, l</emphasis>
</td>
<td box="[582,717,443,466]" gridcol="4" gridrow="4" pageId="5" pageNumber="236">
<emphasis box="[604,663,443,465]" italics="true" pageId="5" pageNumber="236">l, (v),</emphasis>
φ*
</td>
<td box="[749,1105,443,466]" gridcol="5" gridrow="4" pageId="5" pageNumber="236">ft”, (tc), it”, (p), (u), a, s, (pv)</td>
</tr>
</table>
</paragraph>
<paragraph blockId="5.[184,1088,482,591]" pageId="5" pageNumber="236">
<tableNote pageId="5" pageNumber="236" targetBox="[184,1105,231,466]" targetPageId="5">
Note: Roman letters refer to normal setae, Greek letters to solenidia (except ε = famulus); single quotation mark (
<emphasis box="[483,490,512,533]" italics="true" pageId="5" pageNumber="236"></emphasis>
) designates setae on the anterior and double quotation mark (
<emphasis box="[254,266,540,561]" italics="true" pageId="5" pageNumber="236"></emphasis>
) setae on the posterior side of a given leg segment; parentheses refer to a pair of setae. * As alveolus
</tableNote>
</paragraph>
<paragraph blockId="5.[174,1128,617,984]" pageId="5" pageNumber="236">
<materialsCitation collectingDate="2016-04-09" collectingDateMax="2016-04-17" collectingDateMin="2016-04-09" collectorName="District, W &amp; S. Friedrich &amp; F. Wachtel &amp; D. Hauth &amp; Department &amp; Pichcacocha Lakes &amp; Mites &amp; Winkler's" country="Peru" county="South America" elevation="2975" latitude="-10.029722" location="Andes" longLatPrecision="21" longitude="-76.14139" municipality="Central" pageId="5" pageNumber="236" specimenCount="4" specimenCount-female="2" specimenCount-male="1" stateProvince="Huanuco" typeStatus="holotype">
Material examined.
<typeStatus box="[450,550,617,639]" pageId="5" pageNumber="236">Holotype</typeStatus>
(female) and
<specimenCount box="[702,869,617,639]" pageId="5" pageNumber="236" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
(
<specimenCount box="[885,996,617,639]" pageId="5" pageNumber="236" type="male">two males</specimenCount>
and
<specimenCount pageId="5" pageNumber="236" type="female">one female</specimenCount>
):
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,
<collectingCountry box="[420,471,646,667]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
,
<collectingMunicipality box="[483,562,645,667]" pageId="5" pageNumber="236">Central</collectingMunicipality>
<collectingCountry box="[570,621,646,667]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FDBB5A8B278033AE" box="[632,703,645,667]" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="Andes" pageId="5" pageNumber="236" stateProvince="Huanuco">Andes</location>
,
<geoCoordinate box="[715,831,645,667]" degrees="09" direction="south" minutes="54" orientation="latitude" pageId="5" pageNumber="236" precision="15" seconds="30" value="-9.908333">09°5430S</geoCoordinate>
,
<geoCoordinate box="[843,963,645,667]" degrees="76" direction="west" minutes="03" orientation="longitude" pageId="5" pageNumber="236" precision="15" seconds="48" value="-76.06334">76°0348W</geoCoordinate>
,
<collectingRegion box="[974,1073,646,667]" country="Peru" name="Huanuco" pageId="5" pageNumber="236">Huánuco</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FBFB5A88216933AE" box="[1080,1110,646,667]" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="De" pageId="5" pageNumber="236" stateProvince="Huanuco">De</location>
- partment,
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FED85AAC24E6338D" box="[283,473,674,696]" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="Pachitea Province" pageId="5" pageNumber="236" stateProvince="Huanuco">Pachitea Province</location>
,
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FE215AAC27BB338D" box="[482,644,674,696]" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="Molino District" pageId="5" pageNumber="236" stateProvince="Huanuco">
Molino
<collectorName box="[563,644,674,696]" pageId="5" pageNumber="236">District</collectorName>
</location>
, W Molino,
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FD395AAD26AA338D" box="[762,917,675,696]" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="Monte Potrero" pageId="5" pageNumber="236" stateProvince="Huanuco">Monte Potrero</location>
,
<elevation box="[926,1113,674,696]" metricMagnitude="3" metricUnit="m" metricValue="2.975" metricValueMax="3.1" metricValueMin="2.85" pageId="5" pageNumber="236" unit="m" value="2975.0" valueMax="3100.0" valueMin="2850.0">
<quantity box="[926,1060,674,696]" metricMagnitude="3" metricUnit="m" metricValue="2.975" metricValueMax="3.1" metricValueMin="2.85" pageId="5" pageNumber="236" unit="m" value="2975.0" valueMax="3100.0" valueMin="2850.0">28503100 m</quantity>
a.s.l.
</elevation>
, upper soil and leaf litter in a primary mountain cloud forest,
<date box="[837,981,703,725]" pageId="5" pageNumber="236" value="2016-04-15" valueMax="2016-04-17" valueMin="2016-04-15">
<collectingDate box="[837,981,703,725]" pageId="5" pageNumber="236" value="2016-04-15" valueMax="2016-04-17" valueMin="2016-04-15">1517.IV.2016</collectingDate>
</date>
(
<collectorName box="[997,1122,703,725]" pageId="5" pageNumber="236">S. Friedrich</collectorName>
,
<collectorName box="[174,277,732,754]" pageId="5" pageNumber="236">F. Wachtel</collectorName>
and
<collectorName box="[330,432,732,754]" pageId="5" pageNumber="236">D. Hauth</collectorName>
). One
<typeStatus box="[500,595,732,753]" pageId="5" pageNumber="236">paratype</typeStatus>
(
<specimenCount box="[609,730,732,754]" pageId="5" pageNumber="236" type="female">one female</specimenCount>
):
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FD295AD226B433C7" box="[746,907,732,754]" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="South America" pageId="5" pageNumber="236" stateProvince="Huanuco">South America</location>
,
<collectingCountry box="[916,967,732,753]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FC115AD2211E33C7" box="[978,1057,732,754]" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="Central" pageId="5" pageNumber="236" stateProvince="Huanuco">Central</location>
<collectingCountry box="[1063,1114,732,753]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FF6D5AF725CA323A" box="[174,245,761,783]" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="Andes" pageId="5" pageNumber="236" stateProvince="Huanuco">Andes</location>
,
<geoCoordinate box="[261,376,761,782]" degrees="10" direction="south" minutes="01" orientation="latitude" pageId="5" pageNumber="236" precision="15" seconds="47" value="-10.029722">10°0147S</geoCoordinate>
,
<geoCoordinate box="[392,512,761,783]" degrees="76" direction="west" minutes="08" orientation="longitude" pageId="5" pageNumber="236" precision="15" seconds="29" value="-76.14139">76°0829W</geoCoordinate>
,
<collectingRegion box="[527,626,761,782]" country="Peru" name="Huanuco" pageId="5" pageNumber="236">Huánuco</collectingRegion>
<collectorName box="[638,768,761,782]" pageId="5" pageNumber="236">Department</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FCCC5AF72682323A" box="[783,957,761,783]" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="Ambo Province" pageId="5" pageNumber="236" stateProvince="Huanuco">Ambo Province</location>
,
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FC0F5AF725C0321E" country="Peru" county="South America" latitude="-10.029722" longLatPrecision="21" longitude="-76.14139" municipality="Central" name="Conchamarca District" pageId="5" pageNumber="236" stateProvince="Huanuco">Conchamarca District</location>
,
<collectorName box="[268,471,789,811]" pageId="5" pageNumber="236">Pichcacocha Lakes</collectorName>
,
<elevation box="[484,621,789,811]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="236" unit="m" value="3800.0">
<quantity box="[484,563,790,811]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="236" unit="m" value="3800.0">3800 m</quantity>
a.s.l.
</elevation>
, upper soil and leaf litter in mountain forest,
<date box="[174,270,818,840]" pageId="5" pageNumber="236" value="2016-04-09">
<collectingDate box="[174,270,818,840]" pageId="5" pageNumber="236" value="2016-04-09">9.IV.2016</collectingDate>
</date>
(
<collectorName box="[285,409,818,840]" pageId="5" pageNumber="236">S. Friedrich</collectorName>
,
<collectorName box="[419,522,818,840]" pageId="5" pageNumber="236">F. Wachtel</collectorName>
and
<collectorName box="[576,678,818,840]" pageId="5" pageNumber="236">D. Hauth</collectorName>
).
<collectorName box="[696,755,818,840]" pageId="5" pageNumber="236">Mites</collectorName>
were extracted from samples into 75% ethanol using
<collectorName box="[374,468,847,869]" pageId="5" pageNumber="236">Winklers</collectorName>
apparatus.
</materialsCitation>
</paragraph>
<paragraph blockId="5.[174,1128,617,984]" pageId="5" pageNumber="236">
Type deposition. The
<typeStatus box="[468,562,876,898]" pageId="5" pageNumber="236">holotype</typeStatus>
is deposited in the collection of the Museo de Historia Natural, Universidad Nacional Mayor de San Marcos,
<collectingRegion box="[798,855,905,927]" country="Peru" name="Lima" pageId="5" pageNumber="236">Lima</collectingRegion>
,
<collectingCountry box="[866,917,905,926]" name="Peru" pageId="5" pageNumber="236">Peru</collectingCountry>
;
<materialsCitation location="Tyumen State" pageId="5" pageNumber="236" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<specimenCount box="[928,1086,905,927]" pageId="5" pageNumber="236" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
are deposited in the collection of the
<collectingRegion box="[530,677,934,955]" country="Russia" name="Tyumen'" pageId="5" pageNumber="236">Tyumen State</collectingRegion>
</materialsCitation>
<materialsCitation country="Russia" location="University Museum of Zoology" pageId="5" pageNumber="236" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:03A9878DA2371E49FE5259D726AD3472:8EDF6040A2301E49FD6F5BAB26C1328E" box="[684,1022,933,955]" country="Russia" name="University Museum of Zoology" pageId="5" pageNumber="236" stateProvince="Tyumen'">University Museum of Zoology</location>
,
<collectingRegion box="[1032,1123,934,955]" country="Russia" name="Tyumen'" pageId="5" pageNumber="236">Tyumen</collectingRegion>
,
<collectingCountry box="[174,247,962,984]" name="Russia" pageId="5" pageNumber="236">Russia</collectingCountry>
</materialsCitation>
. All specimens are preserved in 70% ethanol solution with a drop of glycerol.
</paragraph>
<paragraph blockId="5.[175,1127,1020,1071]" box="[234,1127,1020,1042]" pageId="5" pageNumber="236">
Etymology. The species name refers to the place of origin of the
<typeStatus box="[930,1026,1020,1042]" pageId="5" pageNumber="236">holotype</typeStatus>
, Pachitea
</paragraph>
<paragraph blockId="5.[175,1127,1020,1071]" box="[175,276,1049,1071]" pageId="5" pageNumber="236">Province.</paragraph>
<paragraph blockId="5.[174,1127,1107,1234]" pageId="5" pageNumber="236">
Remarks.
<taxonomicName authority="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich, 2022" authorityName="Ermilov &amp; Subías &amp; Shtanchaeva &amp; Friedrich" authorityYear="2022" box="[357,642,1107,1133]" class="Arachnida" family="Chamobatidae" genus="Ceratobates" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="pachiteaensis" status="sp. n.">
<emphasis box="[357,642,1107,1133]" italics="true" pageId="5" pageNumber="236">Ceratobates pachiteaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[648,712,1108,1134]" pageId="5" pageNumber="236" rank="species">sp. n.</taxonomicNameLabel>
differs from all representatives of the genus by the presence of comparatively long outer tooth of lamellar cusp (longer than distance between insertions of lamellar setae versus shorter than distance between insertions of lamellar setae).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="236" type="key">
<paragraph blockId="5.[338,962,1290,1351]" pageId="5" pageNumber="236">
KEY TO KNOWN SPECIES OF
<taxonomicName authorityName="Balogh et Mahunka" authorityYear="1969" box="[752,962,1290,1316]" class="Arachnida" family="Chamobatidae" genus="Ceratobates" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">CERATOBATES</taxonomicName>
(modified after
<bibRefCitation author="ERMILOV, S. G. &amp; ANICHKIN, A. E." box="[580,906,1324,1351]" pageId="5" pageNumber="236" pagination="672 - 680" refId="ref3582" refString="ERMILOV, S. G. &amp; ANICHKIN, A. E. (2015): A new species of oribatid mites (Acari, Oribatida) from a mangrove forest of southern Vietnam. - Zoologichesky Zhurnal 94 (6): 651 - 660. [in Russian; in English: Entomological Review 95 (5): 672 - 680] https: // doi. org / 10.1134 / S 0013873815050115" type="journal article" year="2015">ERMILOV &amp; ANICHKIN 2015</bibRefCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>