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<document id="E385565986D5174F9768A7B7FA5EB64A" ID-DOI="10.5281/zenodo.214383" ID-GBIF-Dataset="39e4456d-a611-49e2-acb1-37e84cde484a" ID-ISSN="1175-5326" ID-Zenodo-Dep="214383" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460493896451" checkinUser="plazi" docAuthor="Golovatch, Sergei I., Liu, Weixin &amp; Geoffroy, Jean-Jacques" docDate="2012" docId="03A9878278604C28018AF96DFB7AFEF2" docLanguage="en" docName="zt03358p027.pdf" docOrigin="Zootaxa 3358" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hyleoglomeris heshang Golovatch, Liu &amp; Geoffroy, 2012, sp. n." docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FF90FFFA786F4C39011DFF86FFCAFFCD" masterDocTitle="Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae)" masterLastPageNumber="27" masterPageNumber="1" pageNumber="16" updateTime="1698310613439" updateUser="plazi">
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<mods:title id="F83A59E6D5B45B07D90F94C4E156EA9B">Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae)</mods:title>
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<mods:name id="00B7E8BD3CC66BC8CA984C8CFFD407E4" type="personal">
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<mods:namePart id="F1AF087937A072868A8FB1683387FBC6">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="6282F80154A05CFAD166DD6B3F477180">Liu, Weixin</mods:namePart>
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<mods:namePart id="9A7645F867501C2627CC0F34C42C3F6F">Geoffroy, Jean-Jacques</mods:namePart>
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<mods:date id="0CC4BD4FFB7A7D5675B61B76E89586C9">2012</mods:date>
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<treatment id="03A9878278604C28018AF96DFB7AFEF2" ID-DOI="http://doi.org/10.5281/zenodo.6179922" ID-GBIF-Taxon="119520265" ID-Zenodo-Dep="6179922" LSID="urn:lsid:plazi:treatment:03A9878278604C28018AF96DFB7AFEF2" httpUri="http://treatment.plazi.org/id/03A9878278604C28018AF96DFB7AFEF2" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<subSubSection id="C31A651F78604C36018AF96DFF27F8EA" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BBF369478604C36018AF96DFE24F8C9" blockId="15.[151,494,1771,1831]" box="[151,494,1771,1796]" pageId="15" pageNumber="16">
<heading id="D0F781F878604C36018AF96DFE24F8C9" bold="true" box="[151,494,1771,1796]" fontSize="10" level="2" pageId="15" pageNumber="16" reason="2">
<emphasis id="B974EA8678604C36018AF96DFE24F8C9" bold="true" box="[151,494,1771,1796]" pageId="15" pageNumber="16">
<taxonomicName id="4C004D1778604C36018AF96DFE62F8C9" ID-CoL="6MN2N" box="[151,424,1771,1796]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="heshang" status="sp. nov.">
<emphasis id="B974EA8678604C36018AF96DFE62F8C9" bold="true" box="[151,424,1771,1796]" italics="true" pageId="15" pageNumber="16">Hyleoglomeris heshang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757FD78604C3600B2F96AFE24F8C9" box="[431,494,1772,1796]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBF369478604C36018AF889FF27F8EA" blockId="15.[151,494,1771,1831]" box="[151,237,1807,1831]" pageId="15" pageNumber="16">
<figureCitation id="133B2A1178604C36018AF889FF23F8EA" box="[151,233,1807,1831]" captionStart="FIGURE 12" captionStartId="16.[151,250,1449,1472]" captionTargetBox="[163,1435,281,1415]" captionTargetId="figure@16.[151,1436,267,1426]" captionTargetPageId="16" captionText="FIGURE 12. Hyleoglomeris heshang sp. n., male paratype. A: leg 17, front view. B: leg 18, front view. C &amp; D: left half of telopods, caudal and front views, respectively. E: tip of syncoxital horn, front view. Scale bar: 0.2 mm (E drawn not to scale)." httpUri="https://zenodo.org/record/214395/files/figure.png" pageId="15" pageNumber="16">Fig. 12</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A651F78604C29018AF8D1FED9FF19" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BBF369478604C36018AF8D1FE36F811" blockId="15.[151,1436,1879,2012]" pageId="15" pageNumber="16">
<emphasis id="B974EA8678604C36018AF8D1FE49F8BD" bold="true" box="[151,387,1879,1904]" pageId="15" pageNumber="16">Material examined:</emphasis>
<typeStatus id="54BB883678604C360096F8D1FE39F8A2" box="[395,499,1879,1903]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED11AE5178604C36035CF8D1FD4EF8A2" box="[577,644,1879,1903]" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="15" pageNumber="16">IZAS</collectionCode>
),
<collectingCountry id="F317760478604C360385F8D1FD14F8A2" box="[664,734,1879,1903]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
, Guangxi Prov., Xilin County, Zhou Bang Village, Cave Zhou Bang, Cave 2, ca
<quantity id="4CF89B7178604C360077F8FAFE7EF859" box="[362,436,1916,1940]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="15" pageNumber="16" unit="m" value="890.0">890 m</quantity>
a.s.l.,
<geoCoordinate id="EE34505378604C360319F8FAFD45F859" box="[516,655,1916,1940]" direction="north" orientation="latitude" pageId="15" pageNumber="16" precision="1" value="24.55353">24.55353°N</geoCoordinate>
,
<geoCoordinate id="EE34505378604C360380F8FAFCF9F859" box="[669,819,1916,1940]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="105.11073">105.11073°E</geoCoordinate>
,
<date id="FFBE105478604C36025CF8FAFC08F859" box="[833,962,1916,1940]" pageId="15" pageNumber="16" value="2010-04-18">18.04.2010</date>
, leg. L. Deharveng et al. (CHIgx1029).
<typeStatus id="54BB883678604C36018AF827FECDF874" box="[151,263,1953,1977]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D06FD1D78604C36000FF819FEB8F87A" box="[274,370,1951,1975]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
(
<collectionCode id="ED11AE5178604C36009CF819FE0EF87A" box="[385,452,1951,1975]" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="15" pageNumber="16">IZAS</collectionCode>
),
<specimenCount id="9D06FD1D78604C3600C5F819FDECF87A" box="[472,550,1951,1975]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<specimenCount id="9D06FD1D78604C36032CF819FD58F87A" box="[561,658,1951,1975]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
(
<collectionCode id="ED11AE5178604C3603BCF819FD32F87A" LSID="urn:lsid:biocol.org:col:34988" box="[673,760,1951,1975]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
CC 177),
<specimenCount id="9D06FD1D78604C360270F819FC70F87A" box="[877,954,1951,1975]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<specimenCount id="9D06FD1D78604C3602D8F819FBECF87A" box="[965,1062,1951,1975]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
(
<collectionCode id="ED11AE5178604C360528F819FB4FF87A" box="[1077,1157,1951,1975]" pageId="15" pageNumber="16">SCAU</collectionCode>
),
<specimenCount id="9D06FD1D78604C360584F819FB2EF87A" box="[1177,1252,1951,1975]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
(
<collectionCode id="ED11AE5178604C3605EEF819FA9AF87A" box="[1267,1360,1951,1975]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="15" pageNumber="16">ZMUM</collectionCode>
), same locality, together with
<typeStatus id="54BB883678604C36008EF842FE33F811" box="[403,505,1988,2012]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BBF3694787F4C2901D4FF1EFED9FF19" blockId="16.[151,1436,151,212]" pageId="16" pageNumber="17">
<emphasis id="B974EA86787F4C2901D4FF1EFED2FF7D" bold="true" box="[201,280,152,176]" pageId="16" pageNumber="17">Name:</emphasis>
To emphasize the animal coming on a lonely cave, “
<taxonomicName id="4C004D17787F4C290279FF11FC0EFF62" box="[868,964,151,175]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="heshang">heshang</taxonomicName>
” in Chinese meaning a “monk”; a noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF7F661C787F4C29018AFA2FFAADFA12" httpUri="https://zenodo.org/record/214395/files/figure.png" pageId="16" pageNumber="17" targetBox="[163,1435,281,1415]" targetPageId="16">
<paragraph id="8BBF3694787F4C29018AFA2FFAADFA12" blockId="16.[151,1435,1449,1503]" pageId="16" pageNumber="17">
<emphasis id="B974EA86787F4C29018AFA2FFED5FA72" bold="true" box="[151,287,1449,1472]" pageId="16" pageNumber="17">FIGURE 12.</emphasis>
<taxonomicName id="4C004D17787F4C290039FA2CFDDEFA0D" box="[292,532,1450,1472]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="heshang" status="sp. nov.">
<emphasis id="B974EA86787F4C290039FA2CFDDEFA0D" box="[292,532,1450,1472]" italics="true" pageId="16" pageNumber="17">Hyleoglomeris heshang</emphasis>
</taxonomicName>
<emphasis id="B974EA86787F4C290304FA2CFD98FA72" bold="true" box="[537,594,1450,1471]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A24757FD787F4C290304FA2CFD98FA72" box="[537,594,1450,1471]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male paratype. A: leg 17, front view. B: leg 18, front view. C &amp; D: left half of telopods, caudal and front views, respectively. E: tip of syncoxital horn, front view. Scale bar: 0.2 mm (E drawn not to scale).
</paragraph>
</caption>
<subSubSection id="C31A651F787F4C2901D4F98DFB2EF9A6" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BBF3694787F4C2901D4F98DFB2EF9A6" blockId="16.[151,1436,1547,1970]" pageId="16" pageNumber="17">
<emphasis id="B974EA86787F4C2901D4F98DFE8FF9E9" bold="true" box="[201,325,1547,1572]" pageId="16" pageNumber="17">Diagnosis:</emphasis>
Differs in its residual coloration of a few ocelli, as well as by the roundly subtriangular central lobe of the telopod syncoxite, the presence of a distinct chitinized lobe from the telopod caudomedial femoral process, and the absence of a setoid filament at the tip of the telopod syncoxital horns. See also Key below.
</paragraph>
</subSubSection>
<subSubSection id="C31A651F787F4C2801D4F9F1FB18FED1" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8BBF3694787F4C2901D4F9F1FC0FF942" blockId="16.[151,1436,1547,1970]" box="[201,965,1655,1680]" pageId="16" pageNumber="17">
<emphasis id="B974EA86787F4C2901D4F9F1FE97F95D" bold="true" box="[201,349,1655,1680]" pageId="16" pageNumber="17">Description:</emphasis>
All characters like in
<taxonomicName id="4C004D17787F4C290349F9FFFD60F95D" box="[596,682,1657,1680]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tiani" status="sp. nov.">
<emphasis id="B974EA86787F4C290349F9FFFD60F95D" box="[596,682,1657,1680]" italics="true" pageId="16" pageNumber="17">H. tiani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24757FD787F4C2903ACF9FFFD24F95D" box="[689,750,1657,1680]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
, except as follows.
</paragraph>
<paragraph id="8BBF3694787F4C2901D4F91DFAE9F91A" blockId="16.[151,1436,1547,1970]" pageId="16" pageNumber="17">
Length ca 5.56.0 mm, width 2.53.0 mm in both sexes.
<typeStatus id="54BB8836787F4C290279F91DFC06F97E" box="[868,972,1691,1715]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
6.0 mm long, 3.0 mm wide. Coloration entirely pallid to sometimes very faintly yellowish, 2 or 3 uppermost ocelli sometimes faintly brownish.
</paragraph>
<paragraph id="8BBF3694787F4C2901D4F965FAE9F936" blockId="16.[151,1436,1547,1970]" box="[201,1315,1763,1788]" pageId="16" pageNumber="17">Ocelli 5+1 or 6+1, translucid, often barely visible. Antennomere 6 ca 1.71.8 times as long as high.</paragraph>
<paragraph id="8BBF3694787F4C2901D4F881FC1BF88B" blockId="16.[151,1436,1547,1970]" pageId="16" pageNumber="17">Second tergite with 8 or 9 transverse striae, three starting below, 1 or 2 level to, the remaining striae above schism, 5 or 6 striae (neither 1st nor last from below) crossing the dorsum.</paragraph>
<paragraph id="8BBF3694787F4C2901D4F8D4FE20F843" blockId="16.[151,1436,1547,1970]" pageId="16" pageNumber="17">
Male leg 17 (
<figureCitation id="133B2A11787F4C290043F8D4FE67F8A7" box="[350,429,1874,1898]" captionStart="FIGURE 12" captionStartId="16.[151,250,1449,1472]" captionTargetBox="[163,1435,281,1415]" captionTargetId="figure@16.[151,1436,267,1426]" captionTargetPageId="16" captionText="FIGURE 12. Hyleoglomeris heshang sp. n., male paratype. A: leg 17, front view. B: leg 18, front view. C &amp; D: left half of telopods, caudal and front views, respectively. E: tip of syncoxital horn, front view. Scale bar: 0.2 mm (E drawn not to scale)." httpUri="https://zenodo.org/record/214395/files/figure.png" pageId="16" pageNumber="17">Fig. 12</figureCitation>
A) with a high and rather regularly rounded outer coxal lobe; telopodite 4-segmented, tarsus with two subapical spines.
</paragraph>
<paragraph id="8BBF3694787F4C2901D4F81CFB27F87F" blockId="16.[151,1436,1547,1970]" box="[201,1261,1946,1970]" pageId="16" pageNumber="17">
Male leg 18 (
<figureCitation id="133B2A11787F4C290043F81CFE65F87F" box="[350,431,1946,1970]" captionStart="FIGURE 12" captionStartId="16.[151,250,1449,1472]" captionTargetBox="[163,1435,281,1415]" captionTargetId="figure@16.[151,1436,267,1426]" captionTargetPageId="16" captionText="FIGURE 12. Hyleoglomeris heshang sp. n., male paratype. A: leg 17, front view. B: leg 18, front view. C &amp; D: left half of telopods, caudal and front views, respectively. E: tip of syncoxital horn, front view. Scale bar: 0.2 mm (E drawn not to scale)." httpUri="https://zenodo.org/record/214395/files/figure.png" pageId="16" pageNumber="17">Fig. 12</figureCitation>
B) with a narrowly arch-shaped syncoxital notch; telopodite 4-segmented.
</paragraph>
<paragraph id="8BBF3694787E4C2801D4FF11FB18FED1" blockId="17.[151,1436,151,320]" pageId="17" pageNumber="18">
Telopods (
<figureCitation id="133B2A11787E4C280058FF11FE50FF62" box="[325,410,151,175]" captionStart="FIGURE 12" captionStartId="16.[151,250,1449,1472]" captionTargetBox="[163,1435,281,1415]" captionTargetId="figure@16.[151,1436,267,1426]" captionTargetPageId="16" captionText="FIGURE 12. Hyleoglomeris heshang sp. n., male paratype. A: leg 17, front view. B: leg 18, front view. C &amp; D: left half of telopods, caudal and front views, respectively. E: tip of syncoxital horn, front view. Scale bar: 0.2 mm (E drawn not to scale)." httpUri="https://zenodo.org/record/214395/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
CE) with a rather low, roundly subtriangular, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by a minute lobule (
<figureCitation id="133B2A11787E4C2802D5FF3AFBD6FF19" box="[968,1052,188,212]" captionStart="FIGURE 12" captionStartId="16.[151,250,1449,1472]" captionTargetBox="[163,1435,281,1415]" captionTargetId="figure@16.[151,1436,267,1426]" captionTargetPageId="16" captionText="FIGURE 12. Hyleoglomeris heshang sp. n., male paratype. A: leg 17, front view. B: leg 18, front view. C &amp; D: left half of telopods, caudal and front views, respectively. E: tip of syncoxital horn, front view. Scale bar: 0.2 mm (E drawn not to scale)." httpUri="https://zenodo.org/record/214395/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
E). Caudomedial femoral process prominent, subapically with a distinct lateral lobe on caudal face. A caudomedial process of tibia absent; tibial tubercle on caudal face evident, papillate. Tarsus rather strongly sigmoid, subacuminate apically.
</paragraph>
</subSubSection>
<subSubSection id="C31A651F787E4C2801D4FEAEFB7AFEF2" box="[201,1200,295,320]" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BBF3694787E4C2801D4FEAEFB7AFEF2" blockId="17.[151,1436,151,320]" box="[201,1200,295,320]" pageId="17" pageNumber="18">
<emphasis id="B974EA86787E4C2801D4FEAEFEFEFE8D" bold="true" box="[201,308,296,320]" pageId="17" pageNumber="18">Remark:</emphasis>
Due to its troglomorphic traits, this species might well prove to be a troglobite.
</paragraph>
</subSubSection>
</treatment>
</document>