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<document id="C4334F9ADC0F53E44D63960F05772758" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFD7BB00FF188772FDC8F94C" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia schawalleri Gompel 2020, n. sp." docType="treatment" docUuid="00E6A15E-D7F3-40BB-9A27-8919D0DAAAFF" docUuidSource="ZooBank" docVersion="8" lastPageNumber="168" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="168" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="209ECF2A8B9254D4D76597859EF6AC21">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:name id="D8E47F8C28478FA06F2C9FE461675809" type="personal">
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<mods:namePart id="05D70B99C831766F8D9536CA781157CF">Gompel, Nicolas</mods:namePart>
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<mods:date id="25F022C62BAB3E3770B391E0AAB7E492">2020</mods:date>
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<mods:number id="32EF89E11D56DBA302953B03D892F118">2020-08-26</mods:number>
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<mods:classification id="8734A1C3F961189691F5D1D42A88BF8D">journal article</mods:classification>
<mods:identifier id="87C7248684DB038A7B557D9874F3E043" type="CLB-Dataset">8744</mods:identifier>
<mods:identifier id="501E1649A1345511C200E4C62AD68C63" type="DOI">10.11646/zootaxa.4838.2.1</mods:identifier>
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<mods:identifier id="AA5DF5B4C8EB7357DFF298E48C1FDD30" type="ISSN">1175-5326</mods:identifier>
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<treatment id="039187D8FFD7BB00FF188772FDC8F94C" ID-DOI="http://doi.org/10.5281/zenodo.4404263" ID-GBIF-Taxon="177012502" ID-Zenodo-Dep="4404263" LSID="urn:lsid:zoobank.org:act:00E6A15E-D7F3-40BB-9A27-8919D0DAAAFF" httpUri="http://treatment.plazi.org/id/039187D8FFD7BB00FF188772FDC8F94C" lastPageNumber="168" pageId="17" pageNumber="168">
<subSubSection id="C3226545FFD7BB00FF188772FE40FCAD" box="[151,477,823,850]" pageId="17" pageNumber="168" type="nomenclature">
<paragraph id="8B8736CEFFD7BB00FF188772FE40FCAD" blockId="17.[151,477,823,887]" box="[151,477,823,850]" pageId="17" pageNumber="168">
<heading id="D0CF81A2FFD7BB00FF188772FE40FCAD" bold="true" box="[151,477,823,850]" fontSize="11" level="1" pageId="17" pageNumber="168" reason="1">
<emphasis id="B94CEADCFFD7BB00FF188772FE40FCAD" bold="true" box="[151,477,823,850]" pageId="17" pageNumber="168">
<taxonomicName id="4C384D4DFFD7BB00FF188772FE0EFCAD" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[151,403,823,850]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov.">
<emphasis id="B94CEADCFFD7BB00FF188772FE0EFCAD" bold="true" box="[151,403,823,850]" italics="true" pageId="17" pageNumber="168">Zarcosia schawalleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57A7FFD7BB00FE158772FE40FCAD" box="[410,477,824,850]" pageId="17" pageNumber="168" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD7BB00FF188716FE49FC88" box="[151,468,860,887]" pageId="17" pageNumber="168" type="description">
<paragraph id="8B8736CEFFD7BB00FF188716FE49FC88" blockId="17.[151,477,823,887]" box="[151,468,860,887]" pageId="17" pageNumber="168">
(
<figureCitation id="13032A4BFFD7BB00FF108716FE96FC89" box="[159,267,860,886]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="17" pageNumber="168">Figures 1</figureCitation>
;
<figureCitation id="13032A4BFFD7BB00FE978716FEDBFC89" box="[280,326,860,886]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="17" pageNumber="168">3k,l</figureCitation>
;
<figureCitation id="13032A4BFFD7BB00FEDD8716FEECFC89" box="[338,369,860,886]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="17" pageNumber="168">4n</figureCitation>
;
<figureCitation id="13032A4BFFD7BB00FEF28716FE3EFC88" box="[381,419,860,887]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="17" pageNumber="168">
<quantity id="4CC09B2BFFD7BB00FEF28716FE3EFC88" box="[381,419,860,887]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="168" unit="m" value="5.0">5m</quantity>
</figureCitation>
;
<figureCitation id="13032A4BFFD7BB00FE208716FE51FC88" box="[431,460,860,887]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="17" pageNumber="168">6h</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD7BB00FF1887EEFABAFC41" box="[151,1319,932,958]" pageId="17" pageNumber="168" type="etymology">
<paragraph id="8B8736CEFFD7BB00FF1887EEFABAFC41" blockId="17.[151,1437,932,1715]" box="[151,1319,932,958]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF1887EEFE83FC41" bold="true" box="[151,286,932,958]" pageId="17" pageNumber="168">Etymology.</emphasis>
The species is named after Wolfgang Schawaller, who collected the only known specimens.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD7BB00FF488782FC91FC1D" box="[199,780,968,994]" pageId="17" pageNumber="168" type="materials_examined">
<paragraph id="8B8736CEFFD7BB00FF488782FC91FC1D" blockId="17.[151,1437,932,1715]" box="[199,780,968,994]" pageId="17" pageNumber="168">
<materialsCitation id="3B503C93FFD7BB00FF488782FC9AFC1D" ID-GBIF-Occurrence="3020160447" box="[199,775,968,994]" country="Malaysia" location="Sepilok" municipality="Sandakan" pageId="17" pageNumber="168" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B94CEADCFFD7BB00FF488782FEFBFC1D" bold="true" box="[199,358,968,994]" pageId="17" pageNumber="168">
<typeStatus id="5483886CFFD7BB00FF488782FE9FFC1D" box="[199,258,968,994]" pageId="17" pageNumber="168">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFD7BB00FEE38782FE48FC1D" box="[364,469,968,994]" name="Malaysia" pageId="17" pageNumber="168">Malaysia</collectingCountry>
,
<collectingRegion id="49FCF82CFFD7BB00FE6E8782FDB5FC1D" box="[481,552,968,994]" country="Malaysia" name="Sabah" pageId="17" pageNumber="168">Sabah</collectingRegion>
,
<collectingMunicipality id="6BE3ACB4FFD7BB00FDBC8782FD3EFC1D" box="[563,675,968,994]" pageId="17" pageNumber="168">Sandakan</collectingMunicipality>
,
<location id="8EE76015FFD7BB00FD218782FC9AFC1D" LSID="urn:lsid:plazi:treatment:039187D8FFD7BB00FF188772FDC8F94C:8EE76015FFD7BB00FD218782FC9AFC1D" box="[686,775,968,994]" country="Malaysia" municipality="Sandakan" name="Sepilok" pageId="17" pageNumber="168" stateProvince="Sabah">Sepilok</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD7BB00FF4887A6FD3CFBF9" box="[199,673,1004,1031]" pageId="17" pageNumber="168" type="distribution">
<paragraph id="8B8736CEFFD7BB00FF4887A6FD3CFBF9" blockId="17.[151,1437,932,1715]" box="[199,673,1004,1031]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF4887A6FEFCFBF9" bold="true" box="[199,353,1004,1030]" pageId="17" pageNumber="168">Distribution.</emphasis>
Malayan Borneo (
<figureCitation id="13032A4BFFD7BB00FDBE87A6FD0DFBF9" box="[561,656,1004,1030]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="17" pageNumber="168">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD7BB00FF48805AFD09F9B9" pageId="17" pageNumber="168" type="description">
<paragraph id="8B8736CEFFD7BB00FF48805AFAF0FBB1" blockId="17.[151,1437,932,1715]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF48805AFEC4FBD5" bold="true" box="[199,345,1040,1066]" pageId="17" pageNumber="168">Description.</emphasis>
(
<figureCitation id="13032A4BFFD7BB00FEE8805AFE7BFBD4" box="[359,486,1040,1067]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="17" pageNumber="168">Figure 3k,l</figureCitation>
). Body length.
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<typeStatus id="5483886CFFD7BB00FD23805AFC8FFBD5" box="[684,786,1040,1066]" pageId="17" pageNumber="168" type="holotype">holotype</typeStatus>
:
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;
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:
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. Bicolor, head black, pronotum dark-brown and elytra orange with a broad dark transverse area, covered with short double pubescence.
</paragraph>
<paragraph id="8B8736CEFFD7BB00FF488012FB61FB68" blockId="17.[151,1437,932,1715]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF488012FE99FB8D" box="[199,260,1112,1138]" italics="true" pageId="17" pageNumber="168">Head</emphasis>
. Transverse, wider than pronotum in both sexes, covered with short white pubescence. Integument black, punctation fine and very dense, rugose. Palpi orange, last article securiform. Eyes much bigger in
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.
</paragraph>
<paragraph id="8B8736CEFFD7BB00FF4880EAFB9DFB21" blockId="17.[151,1437,932,1715]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF4880EAFEACFB45" box="[199,305,1184,1210]" italics="true" pageId="17" pageNumber="168">Antennae</emphasis>
. Orange, slender, slightly longer than elytra in
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, shorter than elytra in
<specimenCount id="9D3EFD47FFD7BB00FBD380EBFBECFB44" box="[1116,1137,1185,1211]" pageId="17" pageNumber="168" type="female">♀</specimenCount>
, antennomere 3–7 at most twice longer than wide, 8–10 shorter. Covered with hirsute blond pubescence.
</paragraph>
<paragraph id="8B8736CEFFD7BB00FF4880A3FB07FAB5" blockId="17.[151,1437,932,1715]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF4880A3FEABFAFD" box="[199,310,1257,1282]" italics="true" pageId="17" pageNumber="168">Pronotum</emphasis>
(
<figureCitation id="13032A4BFFD7BB00FECA80A2FE25FAFC" box="[325,440,1256,1283]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="17" pageNumber="168">Figure 6h</figureCitation>
). Narrow, subcylindrical in
<specimenCount id="9D3EFD47FFD7BB00FD7C80A3FC97FAFC" box="[755,778,1257,1283]" pageId="17" pageNumber="168" type="male">♂</specimenCount>
, trapezoidal in
<specimenCount id="9D3EFD47FFD7BB00FC3780A3FC50FAFC" box="[952,973,1257,1283]" pageId="17" pageNumber="168" type="female">♀</specimenCount>
, as long as wide. Lateral constrictions in the posterior half, on each side. Finely and densely punctuated, space between points much smaller than the diameters of points. Integument mat, rugose. Covered with very short, white double pubescence.
</paragraph>
<paragraph id="8B8736CEFFD7BB00FF48811EFC5BFA49" blockId="17.[151,1437,932,1715]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF48811EFE90FA91" box="[199,269,1364,1390]" italics="true" pageId="17" pageNumber="168">Elytra</emphasis>
. Length:
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<typeStatus id="5483886CFFD7BB00FE16811EFE62FA91" box="[409,511,1364,1390]" pageId="17" pageNumber="168" type="holotype">holotype</typeStatus>
:
<quantity id="4CC09B2BFFD7BB00FD84811EFDEEFA90" box="[523,627,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="168" unit="mm" value="1.1">1.10 mm</quantity>
;
<specimenCount id="9D3EFD47FFD7BB00FD0F811FFD08FA90" box="[640,661,1365,1391]" pageId="17" pageNumber="168" type="female">♀</specimenCount>
<typeStatus id="5483886CFFD7BB00FD12811FFC9FFA90" box="[669,770,1365,1391]" pageId="17" pageNumber="168" type="paratype">paratype</typeStatus>
:
<quantity id="4CC09B2BFFD7BB00FC82811EFCE8FA90" box="[781,885,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="17" pageNumber="168" unit="mm" value="1.15">1.15 mm</quantity>
. Orange, with a broad dark area with no defined boundaries, spanning half of the elytral length, from anterior fifth to posterior third. Punctation dense but shallow, covered with double pubescence. Moderately convex, depressed on disc.
</paragraph>
<paragraph id="8B8736CEFFD7BB00FF48818BFB54FA01" blockId="17.[151,1437,932,1715]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF48818BFF61FA25" box="[199,252,1473,1498]" italics="true" pageId="17" pageNumber="168">Legs</emphasis>
. Gracile. Sexual characters: forelegs with first tarsomere enlarged in male, tibia bent inward (
<figureCitation id="13032A4BFFD7BB00FA9A818AFA12FA24" box="[1301,1423,1472,1499]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="17" pageNumber="168">
Figure
<quantity id="4CC09B2BFFD7BB00FAE8818AFA12FA24" box="[1383,1423,1472,1499]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="168" unit="m" value="5.0">5m</quantity>
</figureCitation>
); mesothoracic femora with a sharp apical thorn on their inner side, curved and pointing outward.
</paragraph>
<paragraph id="8B8736CEFFD7BB00FF488242FD09F9B9" blockId="17.[151,1437,932,1715]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF488242FEA9F9DD" box="[199,308,1544,1570]" italics="true" pageId="17" pageNumber="168">Aedeagus</emphasis>
(
<figureCitation id="13032A4BFFD7BB00FECD8242FE29F9DD" box="[322,436,1544,1570]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="17" pageNumber="168">Figure 4n</figureCitation>
). Sides regularly convergent in dorsal view, with an apical button. Strongly bent in lateral view with a sudden apical deflection upward.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD7BB00FF48821AFDC8F94C" pageId="17" pageNumber="168" type="materials_examined">
<paragraph id="8B8736CEFFD7BB00FF48821AFDC8F94C" blockId="17.[151,1437,932,1715]" pageId="17" pageNumber="168">
<emphasis id="B94CEADCFFD7BB00FF48821AFEEAF995" bold="true" box="[199,375,1616,1642]" pageId="17" pageNumber="168">Type material.</emphasis>
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<typeStatus id="5483886CFFD7BB00FE0E821AFE74F995" box="[385,489,1616,1642]" pageId="17" pageNumber="168" type="holotype">Holotype</typeStatus>
(
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,
<collectionCode id="ED29AE0BFFD7BB00FDAE821BFDF9F994" box="[545,612,1617,1643]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="17" pageNumber="168">NME</collectionCode>
).
<emphasis id="B94CEADCFFD7BB00FDF6821AFD7BF995" bold="true" box="[633,742,1616,1642]" pageId="17" pageNumber="168">
<collectingCountry id="F32F765EFFD7BB00FDF6821AFD7BF995" box="[633,742,1616,1642]" name="Malaysia" pageId="17" pageNumber="168">Malaysia</collectingCountry>
</emphasis>
:
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,
<collectingMunicipality id="6BE3ACB4FFD7BB00FCC3821AFC21F995" box="[844,956,1616,1642]" pageId="17" pageNumber="168">Sandakan</collectingMunicipality>
,
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,
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(
<quantity id="4CC09B2BFFD7BB00FAAA821AFAFFF994" box="[1317,1378,1616,1643]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="168" unit="m" value="50.0">
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</quantity>
) (
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)
<date id="FF86100EFFD7BB00FEA8823EFE35F971" box="[295,424,1652,1678]" pageId="17" pageNumber="168" value="2007-03-13">
<collectingDate id="EFC2E9E6FFD7BB00FEA8823EFE35F971" box="[295,424,1652,1678]" pageId="17" pageNumber="168" value="2007-03-13">13.III.2007</collectingDate>
</date>
</materialsCitation>
.
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<typeStatus id="5483886CFFD7BB00FE39823FFD81F970" box="[438,540,1653,1679]" pageId="17" pageNumber="168" type="paratype">Paratype</typeStatus>
.
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<collectingCountry id="F32F765EFFD7BB00FDA6823EFD0BF971" box="[553,662,1652,1678]" name="Malaysia" pageId="17" pageNumber="168">Malaysia</collectingCountry>
</emphasis>
:
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,
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,
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,
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<geoCoordinate id="EE0C5009FFD7BB00FBDA823EFB5AF970" box="[1109,1223,1652,1679]" degrees="117" direction="east" minutes="56" orientation="longitude" pageId="17" pageNumber="168" precision="925" value="117.933334">117°56’ E</geoCoordinate>
(
<quantity id="4CC09B2BFFD7BB00FB55823EFA85F970" box="[1242,1304,1652,1679]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="168" unit="m" value="50.0">
<elevation id="0015D1FDFFD7BB00FB55823EFA85F970" box="[1242,1304,1652,1679]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="168" unit="m" value="50.0">50 m</elevation>
</quantity>
) (
<collectorName id="26CD5318FFD7BB00FABD823EFF42F94C" pageId="17" pageNumber="168">W. Schawaller</collectorName>
)
<date id="FF86100EFFD7BB00FF6282D2FEF1F94D" box="[237,364,1688,1714]" pageId="17" pageNumber="168" value="2007-03-13">
<collectingDate id="EFC2E9E6FFD7BB00FF6282D2FEF1F94D" box="[237,364,1688,1714]" pageId="17" pageNumber="168" value="2007-03-13">13.III.2007</collectingDate>
</date>
(
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<typeStatus id="5483886CFFD7BB00FE1782D3FE66F94C" box="[408,507,1689,1715]" pageId="17" pageNumber="168" type="paratype">paratype</typeStatus>
,
<collectionCode id="ED29AE0BFFD7BB00FD8982D3FDD4F94C" box="[518,585,1689,1715]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="17" pageNumber="168">NME</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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