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<document id="ADE0D1B133ADE748B8675A10B11FDAB1" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFC3BB14FF18811EFE35F880" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia cephalica Gompel 2020, comb. nov." docType="treatment" docVersion="7" lastPageNumber="156" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="156" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D1C15143C460161309B190CCC7F1B8A8">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:namePart id="00321D6C494C59E68E3D7669434899A7">Gompel, Nicolas</mods:namePart>
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<mods:title id="7DAAAD0B2F3E4E275FDE5D0C33014D09">Zootaxa</mods:title>
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<mods:part id="F13EB9E46C45E0C7459D71AA1D6EEF0B">
<mods:date id="8959E3A57D19B604CF82B52ADE400475">2020</mods:date>
<mods:detail id="545397C38D7AE06EB81ED404F3B66AAD" type="pubDate">
<mods:number id="54EB2D709BEB7679D3831710E7BCA0C3">2020-08-26</mods:number>
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<mods:detail id="1A9673C9D02BF530DF0363D6183CF0B0" type="volume">
<mods:number id="B716EFC601870007061CE3A55AED4901">4838</mods:number>
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<mods:number id="E133F79E3908BFC30225F04D0AB89A87">2</mods:number>
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<mods:classification id="F14C778BF76BEE48D2738778389174F1">journal article</mods:classification>
<mods:identifier id="69A74002DFBB07417BDAF431888DE607" type="CLB-Dataset">8744</mods:identifier>
<mods:identifier id="102C7BC723DB6EBC73AB030B02372188" type="DOI">10.11646/zootaxa.4838.2.1</mods:identifier>
<mods:identifier id="80AE6101CEE51A618531E0CBC01571FC" type="GBIF-Dataset">761fcec7-c79d-4e80-bb2b-23cc9e78e0b6</mods:identifier>
<mods:identifier id="3794DC66671F7CB61F79008880F41DC0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2036712D0CE60C338EC5CF28BBA2E1FC" type="Zenodo-Dep">4403783</mods:identifier>
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<treatment id="039187D8FFC3BB14FF18811EFE35F880" ID-DOI="http://doi.org/10.5281/zenodo.4453407" ID-GBIF-Taxon="177012533" ID-Zenodo-Dep="4453407" LSID="urn:lsid:plazi:treatment:039187D8FFC3BB14FF18811EFE35F880" httpUri="http://treatment.plazi.org/id/039187D8FFC3BB14FF18811EFE35F880" lastPageNumber="156" pageId="5" pageNumber="156">
<subSubSection id="C3226545FFC3BB14FF18811EFD4EFA91" box="[151,723,1363,1390]" pageId="5" pageNumber="156" type="nomenclature">
<paragraph id="8B8736CEFFC3BB14FF18811EFD4EFA91" blockId="5.[151,723,1363,1427]" box="[151,723,1363,1390]" pageId="5" pageNumber="156">
<heading id="D0CF81A2FFC3BB14FF18811EFD4EFA91" bold="true" box="[151,723,1363,1390]" fontSize="11" level="1" pageId="5" pageNumber="156" reason="1">
<emphasis id="B94CEADCFFC3BB14FF18811EFD4EFA91" bold="true" box="[151,723,1363,1390]" pageId="5" pageNumber="156">
<taxonomicName id="4C384D4DFFC3BB14FF18811EFDA2FA91" authority="(Werner, 1962)" authorityName="Gompel" authorityYear="2020" baseAuthorityName="Werner" baseAuthorityYear="1962" box="[151,575,1363,1390]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="cephalica" status="comb. nov.">
<emphasis id="B94CEADCFFC3BB14FF18811EFEE1FA91" bold="true" box="[151,380,1363,1390]" italics="true" pageId="5" pageNumber="156">Zarcosia cephalica</emphasis>
(
<bibRefCitation id="EFA94B3FFFC3BB14FE04811EFDABFA91" author="Werner, F. G." box="[395,566,1364,1390]" pageId="5" pageNumber="156" pagination="121 - 127" refId="ref18162" refString="Werner, F. G. (1962) The Aderidae of the Batu Caves, Malaya (Coleoptera). Pacific Insects, 4, 121 - 127." type="journal article" year="1962">Werner, 1962</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A27F57A7FFC3BB14FDC3811EFD4EFA91" box="[588,723,1364,1390]" pageId="5" pageNumber="156" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC3BB14FF188132FE13FA6C" box="[151,398,1400,1427]" pageId="5" pageNumber="156" type="description">
<paragraph id="8B8736CEFFC3BB14FF188132FE13FA6C" blockId="5.[151,723,1363,1427]" box="[151,398,1400,1427]" pageId="5" pageNumber="156">
(
<figureCitation id="13032A4BFFC3BB14FF108132FE96FA6D" box="[159,267,1400,1426]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="5" pageNumber="156">Figures 1</figureCitation>
;
<figureCitation id="13032A4BFFC3BB14FE978132FEAEFA6D" box="[280,307,1400,1426]" captionStart="FIGURE 2" captionStartId="8.[151,250,1960,1984]" captionTargetBox="[170,1419,212,1927]" captionTargetId="figure-30@8.[153,1434,178,1933]" captionTargetPageId="8" captionText="FIGURE 2. continued next page." figureDoi="http://doi.org/10.5281/zenodo.4403789" httpUri="https://zenodo.org/record/4403789/files/figure.png" pageId="5" pageNumber="156">2f</figureCitation>
;
<figureCitation id="13032A4BFFC3BB14FEB18132FEC3FA6D" box="[318,350,1400,1426]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="5" pageNumber="156">4d</figureCitation>
;
<figureCitation id="13032A4BFFC3BB14FEE68132FE1AFA6C" box="[361,391,1400,1427]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="5" pageNumber="156">5u</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC3BB14FF1881F6FD72FA0C" pageId="5" pageNumber="156" type="reference_group">
<paragraph id="8B8736CEFFC3BB14FF1881F6FB6DFA2B" blockId="5.[151,1264,1468,1523]" box="[151,1264,1468,1493]" pageId="5" pageNumber="156">
<treatmentCitationGroup id="AB2811E0FFC3BB14FF1881F6FB6DFA2B" box="[151,1264,1468,1493]" pageId="5" pageNumber="156">
<treatmentCitation id="0A9910DFFFC3BB14FF1881F6FDB1FA2B" author="Werner, F. G." box="[151,556,1468,1492]" page="126" pageId="5" pageNumber="156" year="1962">
<taxonomicName id="4C384D4DFFC3BB14FF1881F6FDB1FA2B" authority="Werner, 1962: 126" authorityName="Werner" authorityPageNumber="126" authorityYear="1962" box="[151,556,1468,1492]" class="Insecta" family="Aderidae" genus="Euglenes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="cephalicus">
<emphasis id="B94CEADCFFC3BB14FF1881F6FEFBFA2B" box="[151,358,1468,1492]" italics="true" pageId="5" pageNumber="156">Euglenes cephalicus</emphasis>
<materialsCitation id="3B503C93FFC3BB14FEE281F7FDB1FA2B" ID-GBIF-Occurrence="3020160391" box="[365,556,1468,1492]" collectionCode="BMNH" country="United States of America" location="Bishop Museum" pageId="5" pageNumber="156" specimenCount="1" stateProvince="Hawaii" typeStatus="holotype">
<bibRefCitation id="EFA94B3FFFC3BB14FEE281F7FDB1FA2B" author="Werner, F. G." box="[365,556,1468,1492]" pageId="5" pageNumber="156" pagination="121 - 127" refId="ref18162" refString="Werner, F. G. (1962) The Aderidae of the Batu Caves, Malaya (Coleoptera). Pacific Insects, 4, 121 - 127." type="journal article" year="1962">Werner, 1962: 126</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
;
<typeStatus id="5483886CFFC3BB14FDB981F6FD05FA2B" box="[566,664,1468,1492]" pageId="5" pageNumber="156" type="holotype">Holotype</typeStatus>
:
<location id="8EE76015FFC3BB14FD2D81F6FCD4FA2B" LSID="urn:lsid:plazi:treatment:039187D8FFC3BB14FF18811EFE35F880:8EE76015FFC3BB14FD2D81F6FCD4FA2B" box="[674,841,1468,1492]" country="United States of America" name="Bishop Museum" pageId="5" pageNumber="156" stateProvince="Hawaii">Bishop Museum</location>
(
<collectingRegion id="49FCF82CFFC3BB14FCD981F6FC3EFA2B" box="[854,931,1468,1492]" country="United States of America" name="Hawaii" pageId="5" pageNumber="156">Hawaii</collectingRegion>
,
<collectingCountry id="F32F765EFFC3BB14FC2281F7FC79FA2A" box="[941,996,1469,1493]" name="United States of America" pageId="5" pageNumber="156">USA</collectingCountry>
);
<specimenCount id="9D3EFD47FFC3BB14FC7981F7FBFDFA2A" box="[1014,1120,1469,1493]" pageId="5" pageNumber="156" type="generic" typeStatus="paratype">1 paratype</specimenCount>
at the BMNH
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8736CEFFC3BB14FF188196FD72FA0C" blockId="5.[151,1264,1468,1523]" box="[151,751,1499,1523]" pageId="5" pageNumber="156">
<taxonomicName id="4C384D4DFFC3BB14FF188196FEFEFA0C" authority="(Werner, 1962)" authorityName="Gompel" authorityYear="2020" baseAuthorityName="Werner" baseAuthorityYear="1962" box="[151,355,1499,1523]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="cephalicus" status="comb. nov.">
<emphasis id="B94CEADCFFC3BB14FF188196FEFEFA0C" box="[151,355,1499,1523]" italics="true" pageId="5" pageNumber="156">Zarcosia cephalicus</emphasis>
</taxonomicName>
[sic!] (
<bibRefCitation id="EFA94B3FFFC3BB14FE238191FDA7FA0C" author="Werner, F. G." box="[428,570,1499,1523]" pageId="5" pageNumber="156" pagination="121 - 127" refId="ref18162" refString="Werner, F. G. (1962) The Aderidae of the Batu Caves, Malaya (Coleoptera). Pacific Insects, 4, 121 - 127." type="journal article" year="1962">Werner, 1962</bibRefCitation>
):
<bibRefCitation id="EFA94B3FFFC3BB14FDC38191FD72FA0C" author="Grzymala, T. L." box="[588,751,1499,1523]" pageId="5" pageNumber="156" refId="ref17990" refString="Grzymala, T. L. (2016) Systematics of Puppet Beetles (Coleoptera, Aderidae): Classification, Molecular Phylogenetics &amp; Evolution of Sexual Dimorphism. Environmental Science, Policy, and Management. University of California, Berkeley, California, 409 pp." type="book" year="2016">Grzymala, 2016</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC3BB14FF18826BFB9CF9C3" box="[151,1025,1569,1596]" pageId="5" pageNumber="156" type="materials_examined">
<paragraph id="8B8736CEFFC3BB14FF18826BFB9CF9C3" blockId="5.[151,1437,1569,1919]" box="[151,1025,1569,1596]" pageId="5" pageNumber="156">
<materialsCitation id="3B503C93FFC3BB14FF18826BFC60F9C3" ID-GBIF-Occurrence="3020160413" box="[151,1021,1569,1596]" country="Malaysia" county="Darul Ehsan" location="Batu" municipality="Gombak District" pageId="5" pageNumber="156" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
<emphasis id="B94CEADCFFC3BB14FF18826BFEABF9C4" bold="true" box="[151,310,1569,1595]" pageId="5" pageNumber="156">
<typeStatus id="5483886CFFC3BB14FF18826BFF4FF9C4" box="[151,210,1569,1595]" pageId="5" pageNumber="156">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFC3BB14FEB3826BFE3AF9C4" box="[316,423,1569,1595]" name="Malaysia" pageId="5" pageNumber="156">Malaysia</collectingCountry>
:
<collectingRegion id="49FCF82CFFC3BB14FE3D826BFD8BF9C4" box="[434,534,1569,1595]" country="Malaysia" name="Selangor" pageId="5" pageNumber="156">Selangor</collectingRegion>
<collectingCounty id="62E64E42FFC3BB14FD93826BFD37F9C4" box="[540,682,1569,1595]" pageId="5" pageNumber="156">Darul Ehsan</collectingCounty>
,
<collectingMunicipality id="6BE3ACB4FFC3BB14FD3A826BFCE9F9C4" box="[693,884,1569,1595]" pageId="5" pageNumber="156">Gombak District</collectingMunicipality>
,
<location id="8EE76015FFC3BB14FCF08268FC29F9C3" LSID="urn:lsid:plazi:treatment:039187D8FFC3BB14FF18811EFE35F880:8EE76015FFC3BB14FCF08268FC29F9C3" box="[895,948,1570,1596]" country="Malaysia" county="Darul Ehsan" municipality="Gombak District" name="Batu" pageId="5" pageNumber="156" stateProvince="Selangor">Batu</location>
caves
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC3BB14FF48820FFB0FF9A0" box="[199,1170,1605,1632]" pageId="5" pageNumber="156" type="distribution">
<paragraph id="8B8736CEFFC3BB14FF48820FFB0FF9A0" blockId="5.[151,1437,1569,1919]" box="[199,1170,1605,1632]" pageId="5" pageNumber="156">
<emphasis id="B94CEADCFFC3BB14FF48820FFEFCF9A0" bold="true" box="[199,353,1605,1631]" pageId="5" pageNumber="156">Distribution.</emphasis>
<collectingCountry id="F32F765EFFC3BB14FEE8820FFE4FF9A0" box="[359,466,1605,1631]" name="Malaysia" pageId="5" pageNumber="156">Malaysia</collectingCountry>
: Batu caves;
<collectingCountry id="F32F765EFFC3BB14FDE9820FFD54F9A0" box="[614,713,1605,1631]" name="Malaysia" pageId="5" pageNumber="156">Sarawak</collectingCountry>
, Gunung Mulu National Park (
<figureCitation id="13032A4BFFC3BB14FBAD820FFB1CF9A0" box="[1058,1153,1605,1631]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="5" pageNumber="156">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC3BB14FF488223FE35F880" pageId="5" pageNumber="156" type="materials_examined">
<paragraph id="8B8736CEFFC3BB14FF488223FE35F880" blockId="5.[151,1437,1569,1919]" pageId="5" pageNumber="156">
<emphasis id="B94CEADCFFC3BB14FF488223FE28F97C" bold="true" box="[199,437,1641,1667]" pageId="5" pageNumber="156">Material examined.</emphasis>
<materialsCitation id="3B503C93FFC3BB14FE4F8223FE59F934" ID-GBIF-Occurrence="3020160334" collectingDate="1978-05-06" collectingDateMax="1978-05-20" collectingDateMin="1978-05-06" collectionCode="BMNH, NGPC" collectorName="P. M. Hammond &amp; J. E. Marshall" country="Malaysia" county="Division" elevation="360" latitude="4.116667" location="Deer Cave" longLatPrecision="1307" longitude="114.916664" municipality="Gunung Mulu National Park" pageId="5" pageNumber="156" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Sarawak">
<emphasis id="B94CEADCFFC3BB14FE4F8223FDB0F97C" bold="true" box="[448,557,1641,1667]" pageId="5" pageNumber="156">
<collectingCountry id="F32F765EFFC3BB14FE4F8223FDB0F97C" box="[448,557,1641,1667]" name="Malaysia" pageId="5" pageNumber="156">Malaysia</collectingCountry>
</emphasis>
:
<collectingRegion id="49FCF82CFFC3BB14FDB08223FD3FF97C" box="[575,674,1641,1667]" country="Malaysia" name="Sarawak" pageId="5" pageNumber="156">Sarawak</collectingRegion>
, 4th
<collectingCounty id="62E64E42FFC3BB14FD6E8223FCD9F97C" box="[737,836,1641,1667]" pageId="5" pageNumber="156">Division</collectingCounty>
,
<collectingMunicipality id="6BE3ACB4FFC3BB14FCDC8223FB3FF97C" box="[851,1186,1641,1667]" pageId="5" pageNumber="156">Gunung Mulu National Park</collectingMunicipality>
,
<location id="8EE76015FFC3BB14FB3D8220FAB2F97C" LSID="urn:lsid:plazi:treatment:039187D8FFC3BB14FF18811EFE35F880:8EE76015FFC3BB14FB3D8220FAB2F97C" box="[1202,1327,1641,1668]" country="Malaysia" county="Division" latitude="4.116667" longLatPrecision="1307" longitude="114.916664" municipality="Gunung Mulu National Park" name="Deer Cave" pageId="5" pageNumber="156" stateProvince="Sarawak">Deer Cave</location>
, (
<quantity id="4CC09B2BFFC3BB14FAC98223FA09F97B" box="[1350,1428,1641,1668]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="156" unit="m" value="360.0">
<elevation id="0015D1FDFFC3BB14FAC98223FA09F97B" box="[1350,1428,1641,1668]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="156" unit="m" value="360.0">360 m</elevation>
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)
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(
<collectorName id="26CD5318FFC3BB14FE0682C7FDD8F958" box="[393,581,1677,1704]" pageId="5" pageNumber="156">P. M. Hammond</collectorName>
&amp;
<collectorName id="26CD5318FFC3BB14FDE382C4FC91F958" box="[620,780,1677,1704]" pageId="5" pageNumber="156">J. E. Marshall</collectorName>
leg.)
<date id="FF86100EFFC3BB14FCC082C7FC29F958" box="[847,948,1677,1703]" pageId="5" pageNumber="156" value="1978-05-06">
<collectingDate id="EFC2E9E6FFC3BB14FCC082C7FC29F958" box="[847,948,1677,1703]" pageId="5" pageNumber="156" value="1978-05-06">6.V.1978</collectingDate>
</date>
(
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,
<specimenCount id="9D3EFD47FFC3BB14FB8E82C7FBB0F957" box="[1025,1069,1677,1704]" pageId="5" pageNumber="156" type="female">2 ♀</specimenCount>
,
<collectionCode id="ED29AE0BFFC3BB14FBB282C4FB08F957" box="[1085,1173,1678,1704]" pageId="5" pageNumber="156">BMNH</collectionCode>
),
<date id="FF86100EFFC3BB14FB2582C7FA81F958" box="[1194,1308,1677,1703]" pageId="5" pageNumber="156" value="1978-05-20">
<collectingDate id="EFC2E9E6FFC3BB14FB2582C7FA81F958" box="[1194,1308,1677,1703]" pageId="5" pageNumber="156" value="1978-05-20">20.V.1978</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC3BB14FAA282C7FAC1F957" box="[1325,1372,1677,1704]" pageId="5" pageNumber="156" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D3EFD47FFC3BB14FAE582C7FA08F957" box="[1386,1429,1677,1704]" pageId="5" pageNumber="156" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED29AE0BFFC3BB14FF1882F8FF72F933" box="[151,239,1714,1740]" pageId="5" pageNumber="156">BMNH</collectionCode>
;
<specimenCount id="9D3EFD47FFC3BB14FF7582FBFEBAF933" box="[250,295,1713,1740]" pageId="5" pageNumber="156" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D3EFD47FFC3BB14FEBC82FBFEC0F933" box="[307,349,1713,1740]" pageId="5" pageNumber="156" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED29AE0BFFC3BB14FEE382FBFE21F934" box="[364,444,1713,1739]" pageId="5" pageNumber="156">NGPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B503C93FFC3BB14FE4082FBFB8EF90F" ID-GBIF-Occurrence="3020160407" collectingDate="1978-05-20" collectionCode="BMNH" collectorName="P. M. Hammond &amp; J. E. Marshall" country="Malaysia" county="Division" elevation="360" latitude="4.116667" location="Clearwater Cave" longLatPrecision="1307" longitude="114.916664" municipality="Gunung Mulu National Park" pageId="5" pageNumber="156" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak">
<collectingRegion id="49FCF82CFFC3BB14FE4082FBFDAFF934" box="[463,562,1713,1739]" country="Malaysia" name="Sarawak" pageId="5" pageNumber="156">Sarawak</collectingRegion>
, 4th
<collectingCounty id="62E64E42FFC3BB14FDE682FBFD51F934" box="[617,716,1713,1739]" pageId="5" pageNumber="156">Division</collectingCounty>
,
<collectingMunicipality id="6BE3ACB4FFC3BB14FD5782FBFB86F934" box="[728,1051,1713,1739]" pageId="5" pageNumber="156">Gunung Mulu National Park</collectingMunicipality>
,
<location id="8EE76015FFC3BB14FBA882FBFB78F934" LSID="urn:lsid:plazi:treatment:039187D8FFC3BB14FF18811EFE35F880:8EE76015FFC3BB14FBA882FBFB78F934" box="[1063,1253,1713,1739]" country="Malaysia" county="Division" latitude="4.116667" longLatPrecision="1307" longitude="114.916664" municipality="Gunung Mulu National Park" name="Clearwater Cave" pageId="5" pageNumber="156" stateProvince="Sarawak">Clearwater Cave</location>
, (
<quantity id="4CC09B2BFFC3BB14FB7782FBFADFF933" box="[1272,1346,1713,1740]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="156" unit="m" value="360.0">
<elevation id="0015D1FDFFC3BB14FB7782FBFADFF933" box="[1272,1346,1713,1740]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="156" unit="m" value="360.0">360 m</elevation>
</quantity>
)
<geoCoordinate id="EE0C5009FFC3BB14FADD82FBFF36F90F" degrees="04" direction="north" minutes="07" orientation="latitude" pageId="5" pageNumber="156" precision="925" value="4.116667">04°07’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC3BB14FF3C829FFEB9F90F" box="[179,292,1749,1776]" degrees="114" direction="east" minutes="55" orientation="longitude" pageId="5" pageNumber="156" precision="925" value="114.916664">114°55’ E</geoCoordinate>
(
<collectorName id="26CD5318FFC3BB14FEBB829FFE73F910" box="[308,494,1749,1776]" pageId="5" pageNumber="156">P. M. Hammond</collectorName>
&amp;
<collectorName id="26CD5318FFC3BB14FD9B829CFD2FF910" box="[532,690,1749,1776]" pageId="5" pageNumber="156">J. E. Marshall</collectorName>
leg.)
<date id="FF86100EFFC3BB14FD7B829FFCFAF910" box="[756,871,1749,1775]" pageId="5" pageNumber="156" value="1978-05-20">
<collectingDate id="EFC2E9E6FFC3BB14FD7B829FFCFAF910" box="[756,871,1749,1775]" pageId="5" pageNumber="156" value="1978-05-20">20.V.1978</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC3BB14FCF5829FFC38F90F" box="[890,933,1749,1776]" pageId="5" pageNumber="156" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED29AE0BFFC3BB14FC3D829CFB96F90F" box="[946,1035,1750,1776]" pageId="5" pageNumber="156">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B503C93FFC3BB14FB91829FFA81F8C7" ID-GBIF-Occurrence="3020160428" collectingDate="1998-07-12" collectionCode="A, MNHN, MZB" collectorName="L. Deharveng &amp; A. Bedos" country="Indonesia" county="Selatan" elevation="30" latitude="3.3930554" location="Tring Meuduro Tunong" longLatPrecision="21" longitude="97.12389" municipality="Sawang" pageId="5" pageNumber="156" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Aceh">
<emphasis id="B94CEADCFFC3BB14FB91829FFABBF910" bold="true" box="[1054,1318,1749,1775]" pageId="5" pageNumber="156">
<collectingCountry id="F32F765EFFC3BB14FB91829FFABBF910" box="[1054,1318,1749,1775]" name="Indonesia" pageId="5" pageNumber="156">Republic of Indonesia</collectingCountry>
</emphasis>
:
<collectingRegion id="49FCF82CFFC3BB14FABB829FFAF2F910" box="[1332,1391,1749,1775]" country="Indonesia" name="Aceh" pageId="5" pageNumber="156">Aceh</collectingRegion>
<collectingCounty id="62E64E42FFC3BB14FAF8829FFF4DF8EC" pageId="5" pageNumber="156">Selatan</collectingCounty>
,
<collectingMunicipality id="6BE3ACB4FFC3BB14FF5282B3FEA5F8EC" box="[221,312,1785,1811]" pageId="5" pageNumber="156">Sawang</collectingMunicipality>
,
<location id="8EE76015FFC3BB14FECB82B3FDCDF8EB" LSID="urn:lsid:plazi:treatment:039187D8FFC3BB14FF18811EFE35F880:8EE76015FFC3BB14FECB82B3FDCDF8EB" box="[324,592,1785,1812]" country="Indonesia" county="Selatan" latitude="3.3930554" longLatPrecision="21" longitude="97.12389" municipality="Sawang" name="Tring Meuduro Tunong" pageId="5" pageNumber="156" stateProvince="Aceh">Tring Meuduro Tunong</location>
,
<location id="8EE76015FFC3BB14FDD382B3FC9EF8EC" LSID="urn:lsid:plazi:treatment:039187D8FFC3BB14FF18811EFE35F880:8EE76015FFC3BB14FDD382B3FC9EF8EC" box="[604,771,1785,1811]" country="Indonesia" county="Selatan" latitude="3.3930554" longLatPrecision="21" longitude="97.12389" municipality="Sawang" name="Gua Kiciarean" pageId="5" pageNumber="156" stateProvince="Aceh">Gua Kiciarean</location>
,
<location id="8EE76015FFC3BB14FC8182B3FCD7F8EC" LSID="urn:lsid:plazi:treatment:039187D8FFC3BB14FF18811EFE35F880:8EE76015FFC3BB14FC8182B3FCD7F8EC" box="[782,842,1785,1811]" country="Indonesia" county="Selatan" latitude="3.3930554" longLatPrecision="21" longitude="97.12389" municipality="Sawang" name="Cave" pageId="5" pageNumber="156" stateProvince="Aceh">Cave</location>
, (
<quantity id="4CC09B2BFFC3BB14FCD082B3FC06F8EB" box="[863,923,1785,1812]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="156" unit="m" value="30.0">
<elevation id="0015D1FDFFC3BB14FCD082B3FC06F8EB" box="[863,923,1785,1812]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="156" unit="m" value="30.0">30 m</elevation>
</quantity>
)
<geoCoordinate id="EE0C5009FFC3BB14FC2482B3FBA1F8EB" box="[939,1084,1785,1812]" degrees="03" direction="north" minutes="23" orientation="latitude" pageId="5" pageNumber="156" precision="15" seconds="35" value="3.3930554">03°23’35” N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC3BB14FBCB82B3FB4FF8EB" box="[1092,1234,1785,1812]" degrees="97" direction="east" minutes="07" orientation="longitude" pageId="5" pageNumber="156" precision="15" seconds="26" value="97.12389">97°07’26” E</geoCoordinate>
(
<collectorName id="26CD5318FFC3BB14FB6C82B3FAE2F8EC" box="[1251,1407,1785,1811]" pageId="5" pageNumber="156">L. Deharveng</collectorName>
&amp;
<collectionCode id="ED29AE0BFFC3BB14FF188354FF33F8C7" box="[151,174,1822,1848]" pageId="5" pageNumber="156">A</collectionCode>
.
<collectorName id="26CD5318FFC3BB14FF348357FE9CF8C8" box="[187,257,1821,1847]" pageId="5" pageNumber="156">Bedos</collectorName>
leg.)
<date id="FF86100EFFC3BB14FECA8357FE53F8C8" box="[325,462,1821,1847]" pageId="5" pageNumber="156" value="1998-07-12">
<collectingDate id="EFC2E9E6FFC3BB14FECA8357FE53F8C8" box="[325,462,1821,1847]" pageId="5" pageNumber="156" value="1998-07-12">12.VII.1998</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC3BB14FE6E8357FD90F8C7" box="[481,525,1821,1848]" pageId="5" pageNumber="156" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D3EFD47FFC3BB14FD948357FDDAF8C7" box="[539,583,1821,1848]" pageId="5" pageNumber="156" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED29AE0BFFC3BB14FDD98354FD32F8C7" box="[598,687,1822,1848]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="156">MNHN</collectionCode>
;
<specimenCount id="9D3EFD47FFC3BB14FD338357FD75F8C7" box="[700,744,1821,1848]" pageId="5" pageNumber="156" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED29AE0BFFC3BB14FD778354FCA4F8C7" box="[760,825,1822,1848]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="156" type="Museum">MZB</collectionCode>
), (
<collectorName id="26CD5318FFC3BB14FCD88357FC24F8C8" box="[855,953,1821,1847]" pageId="5" pageNumber="156">J. Gibert</collectorName>
leg.)
<date id="FF86100EFFC3BB14FC728357FB18F8C8" box="[1021,1157,1821,1847]" pageId="5" pageNumber="156" value="1998-07-12">
<collectingDate id="EFC2E9E6FFC3BB14FC728357FB18F8C8" box="[1021,1157,1821,1847]" pageId="5" pageNumber="156" value="1998-07-12">12.VII.1998</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC3BB14FB168357FB5EF8C7" box="[1177,1219,1821,1848]" pageId="5" pageNumber="156" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED29AE0BFFC3BB14FB5C8354FA89F8C7" box="[1235,1300,1822,1848]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="156" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B503C93FFC3BB14FAA68354FE39F880" ID-GBIF-Occurrence="3020160432" collectingDate="2011-02-17" collectionCode="A, NGPC" collectorName="L. Deharveng &amp; A. Bedos" country="Indonesia" county="Cilacap" elevation="70" latitude="-7.7166667" location="Goa Ratu" municipality="Nusakambangan Island" pageId="5" pageNumber="156" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Timur">
<collectingRegion id="49FCF82CFFC3BB14FAA68354FAFCF8C7" box="[1321,1377,1822,1848]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="156">Jawa</collectingRegion>
Tengah,
<collectingCounty id="62E64E42FFC3BB14FF40830BFEBAF8A4" box="[207,295,1857,1883]" pageId="5" pageNumber="156">Cilacap</collectingCounty>
,
<collectingMunicipality id="6BE3ACB4FFC3BB14FEBB830BFDA2F8A4" box="[308,575,1857,1883]" pageId="5" pageNumber="156">Nusakambangan Island</collectingMunicipality>
,
<location id="8EE76015FFC3BB14FDC3830BFD26F8A3" LSID="urn:lsid:plazi:treatment:039187D8FFC3BB14FF18811EFE35F880:8EE76015FFC3BB14FDC3830BFD26F8A3" box="[588,699,1857,1884]" country="Indonesia" county="Cilacap" latitude="-7.7166667" municipality="Nusakambangan Island" name="Goa Ratu" pageId="5" pageNumber="156" stateProvince="Jawa Timur">Goa Ratu</location>
, (
<quantity id="4CC09B2BFFC3BB14FD5F830BFC90F8A3" box="[720,781,1857,1884]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="156" unit="m" value="70.0">
<elevation id="0015D1FDFFC3BB14FD5F830BFC90F8A3" box="[720,781,1857,1884]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="156" unit="m" value="70.0">70 m</elevation>
</quantity>
) cave,
<geoCoordinate id="EE0C5009FFC3BB14FCEF830BFC5EF8A4" box="[864,963,1857,1883]" degrees="07" direction="south" minutes="43" orientation="latitude" pageId="5" pageNumber="156" precision="925" value="-7.7166667">07°43’ S</geoCoordinate>
<geoCoordinate id="EE0C5009FFC3BB14FC43830BFBB9F8A3" box="[972,1060,1857,1884]" pageId="5" pageNumber="156">108°57’</geoCoordinate>
(
<collectorName id="26CD5318FFC3BB14FBB9830BFB4FF8A4" box="[1078,1234,1857,1883]" pageId="5" pageNumber="156">L. Deharveng</collectorName>
&amp;
<collectionCode id="ED29AE0BFFC3BB14FB788308FA93F8A3" box="[1271,1294,1858,1884]" pageId="5" pageNumber="156">A</collectionCode>
.
<collectorName id="26CD5318FFC3BB14FA94830BFAFCF8A4" box="[1307,1377,1857,1883]" pageId="5" pageNumber="156">Bedos</collectorName>
leg.)
<date id="FF86100EFFC3BB14FF18832FFE96F880" box="[151,267,1893,1919]" pageId="5" pageNumber="156" value="2011-02-17">
<collectingDate id="EFC2E9E6FFC3BB14FF18832FFE96F880" box="[151,267,1893,1919]" pageId="5" pageNumber="156" value="2011-02-17">17.II.2011</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC3BB14FE91832FFEA2F880" box="[286,319,1893,1919]" pageId="5" pageNumber="156" type="female">1♀</specimenCount>
,
<collectionCode id="ED29AE0BFFC3BB14FEC3832FFE01F880" box="[332,412,1893,1919]" pageId="5" pageNumber="156">NGPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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