256 lines
28 KiB
XML
256 lines
28 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6210779" ID-GBIF-Dataset="7485373c-0219-4ed7-bb5c-ac664acf6775" ID-GBIF-Taxon="114923682" ID-Zenodo-Dep="6210779" LSID="urn:lsid:zoobank.org:act:EADB4ADB-ED02-4F5F-A1EC-C0AEB4F53371" checkinTime="1427905504904" checkinUser="donat" docAuthor="Alberto Ballerio & Munetoshi Maruyama" docDate="2010" docId="038951095C043D0C66B6FE59ABB6FCED" docLanguage="en" docName="ZK_article_2140.pdf" docOrigin="Zookeys 34" docTitle="Madrasostes hashimi Ballerio & Maruyama, 2010, sp. n." docType="treatment" docUuid="EADB4ADB-ED02-4F5F-A1EC-C0AEB4F53371" docUuidSource="ZooBank" docVersion="9" lastPageId="16" lastPageNumber="93" masterDocId="FFB029715C083D1C663AFFC4A802FFD6" masterDocTitle="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="104" masterPageNumber="77" pageId="12" pageNumber="89" updateTime="1645477292090" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Alberto Ballerio</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Munetoshi Maruyama</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zookeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>34</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>77</mods:start>
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<mods:end>104</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.34.268</mods:identifier>
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<mods:identifier type="ZooBank">553C91CE-FEA1-4867-929B-2865647E4B2E</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6210779" ID-GBIF-Taxon="114923682" ID-Zenodo-Dep="6210779" LSID="urn:lsid:zoobank.org:act:EADB4ADB-ED02-4F5F-A1EC-C0AEB4F53371" httpUri="http://treatment.plazi.org/id/038951095C043D0C66B6FE59ABB6FCED" lastPageId="16" lastPageNumber="93" pageId="12" pageNumber="89">
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<subSubSection box="[140,459,413,439]" pageId="12" pageNumber="89" type="nomenclature">
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<paragraph box="[140,459,413,439]" pageId="12" pageNumber="89">
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<heading bold="true" box="[140,459,413,439]" fontSize="11" level="1" pageId="12" pageNumber="89" reason="6">
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<emphasis bold="true" box="[140,459,413,439]" pageId="12" pageNumber="89">
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<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:EADB4ADB-ED02-4F5F-A1EC-C0AEB4F53371" box="[140,388,413,439]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="89" phylum="Arthropoda" rank="species" species="hashimi" status="sp. n.">
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<emphasis bold="true" box="[140,388,413,439]" italics="true" pageId="12" pageNumber="89">Madrasostes hashimi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[395,459,413,439]" pageId="12" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="89" type="description">
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<paragraph pageId="12" pageNumber="89">
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urn:lsid:zoobank.org:act:EADB4ADB-ED02-4F5F-A1EC-C0AEB4F53371 (
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<figureCitation box="[148,210,483,510]" captionStart="Figure 1" captionStartId="13.[140,211,1590,1612]" captionTargetBox="[140,1107,211,1470]" captionTargetId="figure@13.[140,1108,204,1474]" captionTargetPageId="13" captionText="Figure 1. Ceratocanthinae of Ulu Gombak. A Madrasostes variolosum habitus dorsal B Madrasostes simplex habitus dorsal C Madrasostes hashimi sp. n. habitus dorsal D idem habitus lateral E Madrasostes sculpturatum habitus dorsal." pageId="12" pageNumber="89">Fig. 1</figureCitation>
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C, D)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="89" type="materials_examined">
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<paragraph box="[140,990,553,580]" pageId="12" pageNumber="89">
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<emphasis bold="true" box="[140,301,554,580]" pageId="12" pageNumber="89">Type locality:</emphasis>
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West Malaysia: Ulu Gombak (Selangor), 3°19
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<emphasis box="[827,833,554,580]" italics="true" pageId="12" pageNumber="89">'</emphasis>
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N 101°45
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<emphasis box="[952,959,554,580]" italics="true" pageId="12" pageNumber="89">'</emphasis>
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E.
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</paragraph>
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<paragraph pageId="12" pageNumber="89">
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<emphasis bold="true" box="[185,417,589,615]" pageId="12" pageNumber="89">Material examined.</emphasis>
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<emphasis box="[426,521,589,615]" italics="true" pageId="12" pageNumber="89">Holotype</emphasis>
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, male: West Malaysia, Selangor, Ulu Gombak, 250 mt., 24.IV.2007, Leg. M. Maruyama /
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<taxonomicName box="[572,792,624,651]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="89" phylum="Arthropoda" rank="species" species="hashimi" status="sp. n.">
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<emphasis box="[572,792,624,651]" italics="true" pageId="12" pageNumber="89">Madrasostes hashimi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[798,858,625,651]" pageId="12" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
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det. Ballerio & Maruyama 2009 Holotypus (KUM) [distended specimen glued on a card, dissected, genitalia mounted in DMHF resin on a separate card, same pin].
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<emphasis box="[800,900,695,721]" italics="true" pageId="12" pageNumber="89">Paratypes</emphasis>
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[all dissected]: 19. IV. 2007, M. Maruyama et al. (1 male); 29. IV. 2007, M. Maruyama (1 female); 5. V. 2007, M. Maruyama (1 male). All from termite nest(s). (2 KUM, 1 ABCB)
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="93" pageId="12" pageNumber="89" type="description">
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<paragraph pageId="12" pageNumber="89">
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<emphasis bold="true" box="[185,335,800,826]" pageId="12" pageNumber="89">Description.</emphasis>
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HL = 1.21 mm; HW = 2 mm; PL = 1.28 mm; PW = 3.42 mm; EL = 3.57 mm; EW = 3.5 mm.
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</paragraph>
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<paragraph pageId="12" pageNumber="89">Medium sized Ceratocanthinae, surface shiny, finely setose (45x); “rolling up” coaptations perfect; wings present and fairly developed.</paragraph>
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<paragraph box="[185,1044,941,967]" pageId="12" pageNumber="89">Dorsum dark brown with bronze sheen, sternum reddish, antennae reddish.</paragraph>
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<paragraph pageId="12" pageNumber="89">
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<emphasis box="[185,251,976,1003]" italics="true" pageId="12" pageNumber="89">Head:</emphasis>
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wide (W/L ratio = 1.64) pentagonal, fore portion triangular, apex forming an obtuse angle (about 120°), both sides of the angle slightly convex, irregularly finely serrated, not reflexed upward, tip of triangle blunt; genae almost aligned with fore margin, slightly protruding outwards, while forming a rounded acute angle, genal canthus normally developed, reaching the occipital area of head, dorsal ocular area small, dorsal interocular area about 18 times the maximum width of the dorsal ocular area, ventral ocular area relatively large; head dorsal surface slightly convex, with dense puncturation, punctures deeply impressed, their distance being inferior than their diameter, intervals between punctures irregularly raised, giving a granulose appearance to head sculpturing, punctures ocellate, bearing a short fine seta, fore margin with two deep, fine, transverse striae.
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</paragraph>
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<paragraph lastPageId="14" lastPageNumber="91" pageId="12" pageNumber="89">
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<emphasis box="[185,301,1364,1390]" italics="true" pageId="12" pageNumber="89">Pronotum:</emphasis>
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subtrapezoidal, wider than long (W/L ratio = 2.66), almost as wide as elytra; fore margin feebly bisinuate; fore angles slightly, but distinctly protrudent forward, broadly subtruncate at apex, outer apex of truncation acute and distinctly protruding, a distinct sinuature at the outer side of the apex; fore edge continuously finely margined, edges of sides without any visible margin (dorsal view), base strongly margined; base at middle very slightly protruding backwards, basal edge neither swollen nor raised; pronotal surface regularly convex with one shallow depression at each side of disc (paradiscal depressions). Surface shiny, smooth, with dense strong irregular puncturation; punctures deep, ocellate, their distance being less than their diameter, intervals between punctures irregularly raised, giving a granulose appearance to pronotal sculpturing, punctures bearing often a fine short seta.
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</paragraph>
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<caption pageId="13" pageNumber="90" targetBox="[140,1107,211,1470]" targetPageId="13">
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<paragraph pageId="13" pageNumber="90">
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<emphasis bold="true" box="[140,238,1590,1612]" pageId="13" pageNumber="90">Figure 1.</emphasis>
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Ceratocanthinae of Ulu Gombak.
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<emphasis bold="true" box="[563,581,1590,1612]" pageId="13" pageNumber="90">A</emphasis>
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<emphasis box="[587,795,1590,1612]" italics="true" pageId="13" pageNumber="90">
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<taxonomicName box="[587,692,1590,1612]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
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variolosum
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</emphasis>
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habitus dorsal
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<emphasis bold="true" box="[936,952,1590,1612]" pageId="13" pageNumber="90">B</emphasis>
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<emphasis italics="true" pageId="13" pageNumber="90">
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<taxonomicName class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="simplex">Madrasostes simplex</taxonomicName>
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</emphasis>
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habitus dorsal
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<emphasis bold="true" box="[326,344,1622,1644]" pageId="13" pageNumber="90">C</emphasis>
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<emphasis box="[353,538,1622,1644]" italics="true" pageId="13" pageNumber="90">
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<taxonomicName box="[353,538,1622,1644]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="hashimi" status="sp. n.">Madrasostes hashimi</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[547,601,1623,1644]" pageId="13" pageNumber="90" rank="species">sp. n.</taxonomicNameLabel>
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habitus dorsal
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<emphasis bold="true" box="[751,771,1622,1644]" pageId="13" pageNumber="90">D</emphasis>
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idem habitus lateral
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<emphasis bold="true" box="[978,993,1622,1644]" pageId="13" pageNumber="90">E</emphasis>
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<taxonomicName class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="90" phylum="Arthropoda" rank="species" species="sculpturatum">
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<emphasis italics="true" pageId="13" pageNumber="90">Madrasostes sculpturatum</emphasis>
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</taxonomicName>
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habitus dorsal.
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</paragraph>
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</caption>
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<paragraph pageId="14" pageNumber="91">
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<emphasis box="[185,300,237,263]" italics="true" pageId="14" pageNumber="91">Scutellum:</emphasis>
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wider than long, sides proximally subparallel and distinctly notched by elytral articular process, then convergent to form a triangle with elongate acute apex and sides slightly curved inward. Surface slightly depressed in the middle. Apical portion of mesepisterna visible from above. Scutellum uniformly densely punctured; punctures about as large and shaped as on head.
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</paragraph>
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<paragraph pageId="14" pageNumber="91">
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<emphasis box="[185,257,413,439]" italics="true" pageId="14" pageNumber="91">Elytra:</emphasis>
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slightly longer than wide (W/L ratio = 0.98), apical fourth regularly round- ed (dorsal view), apex slightly re-entering inward (lateral view); elytra regularly convex, although slightly flattened at disc; elytral suture very finely raised; inferior sutural stria present, ending just before humeral area, delimiting a small marginal elytral area; marginal area poorly developed, smooth, articular area with striation, visible in lateral view; humeral callus small; elytral articular process small, smooth and shiny. Elytra strongly densely punctured, basal and median third covered by a mix of horseshoeshaped impressed punctures, their distance from each other being shorter than their diameter, each one bearing a very short fine seta and simple impressed puncures, distal third covered by dense ocellate punctures, discal surface with sparse longitudinal short carinae, distal third and sides with sparse, short tubercles. Wings present.
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</paragraph>
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<paragraph pageId="14" pageNumber="91">Clypeopleuron very short and transversely slightly grooved at each side. Labrum wide and short, semicircular, distally fringed by long fine setae directed forward. Distal epipharynx semicircular, longitudinally divided by a strong anterior median process; pariae distinctly raised with respect to the haptolachus; median brush and corypha absent; apical fringe made of long fine setae, absent in the middle. Mentum ventrally flat, widely emarginated in the middle, emargination regularly wide-U-shaped; labial palpi (including palpiger) four jointed, first joint very short and transverse, joint two short, joint three longer and plumper than joint two, joint four subconical, apically bearing some short sensilla, all joints, except four, fringed with long setae. Maxillae with an elongate single membranous lacinia, covered with fine long setae, monolobed galea proximally sclerotized and distally clothed with very coarse, long, fine setae, with distinctive comb-like tip (galeal brush), maxillary palpi (including palpiger) four jointed, palpiger very small, joint two wide and relatively short, joint three relatively short, joint four long and subconical, about as long as two and three together, apically bearing some short sensilla. Mandibles short, regularly curved, apicais with pointed apical tooth short and blunt, not protruding over mesal brush, lateral sclerite of apicalis bearing a distinct large pore, conjunctive present, mesal brush wide and well developed, basalis with molar lobe relatively strong. Antennae 10-segmented, scape long (about half the total length of antenna), securiform, pedicellus plump and rounded, flagellum short, with articles distinctly wider than long, antennal club with three uniformly setose articles.</paragraph>
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<paragraph pageId="14" pageNumber="91">
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<emphasis box="[185,400,1504,1531]" italics="true" pageId="14" pageNumber="91">Sexual dimorphism:</emphasis>
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males with protibiae ending with an apical spur plumper than in females and with distal third more dramatically bent downwards, mestotibiae ending with a straight apical spur and an inner apical spur bent inwards at a right angle, almost fused with the apex of mesotibia. Females with apical spur of protibiae slender and more gently bent downwards, mesotibiae ending with two straight apical spurs.
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</paragraph>
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<caption pageId="15" pageNumber="92" targetBox="[140,1107,172,1515]" targetPageId="15">
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<paragraph pageId="15" pageNumber="92">
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<emphasis bold="true" box="[140,237,1558,1580]" pageId="15" pageNumber="92">Figure 2.</emphasis>
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Ceratocanthinae of Ulu Gombak.
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<emphasis bold="true" box="[557,576,1558,1580]" pageId="15" pageNumber="92">A</emphasis>
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<emphasis box="[581,779,1558,1580]" italics="true" pageId="15" pageNumber="92">
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<taxonomicName box="[581,779,1558,1580]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="depressum">Madrasostes depressum</taxonomicName>
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</emphasis>
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habitus dorsal
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<emphasis bold="true" box="[917,934,1558,1580]" pageId="15" pageNumber="92">B</emphasis>
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<emphasis italics="true" pageId="15" pageNumber="92">
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<taxonomicName class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="boucomonti">Madrasostes boucomonti</taxonomicName>
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</emphasis>
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habitus dorsal
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<emphasis bold="true" box="[333,352,1590,1612]" pageId="15" pageNumber="92">C</emphasis>
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<emphasis box="[357,524,1590,1612]" italics="true" pageId="15" pageNumber="92">
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<taxonomicName box="[357,524,1590,1612]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="agostii" status="sp. n.">Madrasostes agostii</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[530,581,1591,1612]" pageId="15" pageNumber="92" rank="species">sp. n.</taxonomicNameLabel>
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habitus dorsal
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<emphasis bold="true" box="[720,740,1590,1612]" pageId="15" pageNumber="92">D</emphasis>
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<emphasis box="[746,923,1590,1612]" italics="true" pageId="15" pageNumber="92">
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<taxonomicName box="[746,850,1590,1612]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
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clypeale
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</emphasis>
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male habitus dorsal
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<emphasis bold="true" box="[140,155,1622,1644]" pageId="15" pageNumber="92">E</emphasis>
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<emphasis box="[161,367,1622,1644]" italics="true" pageId="15" pageNumber="92">
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<taxonomicName box="[161,266,1622,1644]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
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malyanum
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</emphasis>
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habitus dorsal
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<emphasis bold="true" box="[509,523,1622,1644]" pageId="15" pageNumber="92">F</emphasis>
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<taxonomicName box="[529,729,1622,1644]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="92" phylum="Arthropoda" rank="species" species="mirificum" status="sp. n.">
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<emphasis box="[529,729,1622,1644]" italics="true" pageId="15" pageNumber="92">Madrasostes mirificum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[735,786,1623,1644]" pageId="15" pageNumber="92" rank="species">sp. n.</taxonomicNameLabel>
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habitus dorsal
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<emphasis bold="true" box="[927,946,1622,1644]" pageId="15" pageNumber="92">G</emphasis>
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habitus lateral of male
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<emphasis bold="true" box="[190,210,1654,1676]" pageId="15" pageNumber="92">H</emphasis>
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fore body of female
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<emphasis bold="true" box="[401,409,1654,1676]" pageId="15" pageNumber="92">I</emphasis>
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male protibia dorsal view
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<emphasis bold="true" box="[654,661,1654,1676]" pageId="15" pageNumber="92">J</emphasis>
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male protibia lateral view.
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</paragraph>
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</caption>
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<paragraph pageId="16" pageNumber="93">
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<emphasis box="[185,353,166,193]" italics="true" pageId="16" pageNumber="93">Male genitalia:</emphasis>
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genital segment fairly sclerotized, V-shaped (
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<figureCitation box="[882,951,166,193]" captionStart="Figure 8" captionStartId="23.[140,211,1558,1580]" captionTargetBox="[145,1078,415,1513]" captionTargetId="figure@23.[140,1108,394,1541]" captionTargetPageId="23" captionText="Figure 8. Madrasostes mirificum sp. n.: A male genital segment (scale bar: 0, 2 mm) B aedeagus (scale bar: 0, 1 mm) C parameres in dorsal view (scale bar: 0, 0 5 mm) D Madrasostes hasimi sp. n.: male genital segment (scale bar: 0, 1 mm) E parameres (scale bar: 0, 1 mm) F sclerite of internal sac of aedeagus (scale bar: 0, 1 mm) G Madrasostes depressum: sclerite of internal sac of aedeagus (scale bar: 0, 1 mm)." pageId="16" pageNumber="93">Fig. 8</figureCitation>
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D). Aedeagus with basal piece twisted, about three times longer than parameres; internal sac relatively small, containing a large very thick asymmetrical spiraliform sclerite (
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<figureCitation box="[1008,1074,237,263]" captionStart="Figure 8" captionStartId="23.[140,211,1558,1580]" captionTargetBox="[145,1078,415,1513]" captionTargetId="figure@23.[140,1108,394,1541]" captionTargetPageId="23" captionText="Figure 8. Madrasostes mirificum sp. n.: A male genital segment (scale bar: 0, 2 mm) B aedeagus (scale bar: 0, 1 mm) C parameres in dorsal view (scale bar: 0, 0 5 mm) D Madrasostes hasimi sp. n.: male genital segment (scale bar: 0, 1 mm) E parameres (scale bar: 0, 1 mm) F sclerite of internal sac of aedeagus (scale bar: 0, 1 mm) G Madrasostes depressum: sclerite of internal sac of aedeagus (scale bar: 0, 1 mm)." pageId="16" pageNumber="93">Fig. 8</figureCitation>
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F); parameres asymmetrical (
|
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<figureCitation box="[426,490,272,299]" captionStart="Figure 8" captionStartId="23.[140,211,1558,1580]" captionTargetBox="[145,1078,415,1513]" captionTargetId="figure@23.[140,1108,394,1541]" captionTargetPageId="23" captionText="Figure 8. Madrasostes mirificum sp. n.: A male genital segment (scale bar: 0, 2 mm) B aedeagus (scale bar: 0, 1 mm) C parameres in dorsal view (scale bar: 0, 0 5 mm) D Madrasostes hasimi sp. n.: male genital segment (scale bar: 0, 1 mm) E parameres (scale bar: 0, 1 mm) F sclerite of internal sac of aedeagus (scale bar: 0, 1 mm) G Madrasostes depressum: sclerite of internal sac of aedeagus (scale bar: 0, 1 mm)." pageId="16" pageNumber="93">Fig. 8</figureCitation>
|
||
E), laterally flattened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="93" type="discussion">
|
||
<paragraph pageId="16" pageNumber="93">
|
||
<emphasis bold="true" box="[185,315,307,334]" pageId="16" pageNumber="93">Variability.</emphasis>
|
||
The type series shows variability mainly in the development of longitudinal carinae of elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="93" type="diagnosis">
|
||
<paragraph pageId="16" pageNumber="93">
|
||
<emphasis bold="true" box="[185,356,378,404]" pageId="16" pageNumber="93">Identification.</emphasis>
|
||
<taxonomicName box="[365,588,377,404]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="hashimi" status="sp. n.">
|
||
<emphasis box="[365,588,377,404]" italics="true" pageId="16" pageNumber="93">Madrasostes hashimi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[597,660,378,404]" pageId="16" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
can be mistaked only with
|
||
<taxonomicName authority="Paulian, 1989" authorityName="Paulian" authorityYear="1989" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="thoracicum">
|
||
<emphasis italics="true" pageId="16" pageNumber="93">M. thoracicum</emphasis>
|
||
Paulian, 1989
|
||
</taxonomicName>
|
||
known from Borneo (Sabah). The new species however differs from the former because of a) the puncturation of elytra which in
|
||
<taxonomicName box="[832,993,448,475]" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="thoracicum">
|
||
<emphasis box="[832,993,448,475]" italics="true" pageId="16" pageNumber="93">M. thoracicum</emphasis>
|
||
</taxonomicName>
|
||
is slightly larger and denser, b) the shape of elytral carinae that in
|
||
<emphasis box="[780,908,483,510]" italics="true" pageId="16" pageNumber="93">
|
||
<taxonomicName box="[780,908,483,510]" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.bestMatchDistance="180" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="hashimi" status="sp. n.">M. hashimi</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[916,979,484,510]" pageId="16" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
are usually longer than in
|
||
<taxonomicName box="[307,468,518,545]" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="thoracicum">
|
||
<emphasis box="[307,468,518,545]" italics="true" pageId="16" pageNumber="93">M. thoracicum</emphasis>
|
||
</taxonomicName>
|
||
, c) the sculpturing of pronotum which is much stronger in
|
||
<taxonomicName box="[173,336,553,580]" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="thoracicum">
|
||
<emphasis box="[173,336,553,580]" italics="true" pageId="16" pageNumber="93">M. thoracicum</emphasis>
|
||
</taxonomicName>
|
||
, with puncturation deeper and more transverse, and finally d) the shape of the sclerite of the internal sac of aedeagus, which in the new species is larger and distinctly spiraliform (
|
||
<figureCitation box="[441,506,624,651]" captionStart="Figure 8" captionStartId="23.[140,211,1558,1580]" captionTargetBox="[145,1078,415,1513]" captionTargetId="figure@23.[140,1108,394,1541]" captionTargetPageId="23" captionText="Figure 8. Madrasostes mirificum sp. n.: A male genital segment (scale bar: 0, 2 mm) B aedeagus (scale bar: 0, 1 mm) C parameres in dorsal view (scale bar: 0, 0 5 mm) D Madrasostes hasimi sp. n.: male genital segment (scale bar: 0, 1 mm) E parameres (scale bar: 0, 1 mm) F sclerite of internal sac of aedeagus (scale bar: 0, 1 mm) G Madrasostes depressum: sclerite of internal sac of aedeagus (scale bar: 0, 1 mm)." pageId="16" pageNumber="93">Fig. 8</figureCitation>
|
||
F), whereas in
|
||
<taxonomicName box="[673,834,624,651]" class="Insecta" family="Hybosoridae" genus="Madrasostes" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="93" phylum="Arthropoda" rank="species" species="thoracicum">
|
||
<emphasis box="[673,834,624,651]" italics="true" pageId="16" pageNumber="93">M. thoracicum</emphasis>
|
||
</taxonomicName>
|
||
is short and shaped in a different way (
|
||
<figureCitation box="[303,367,659,686]" captionStart="Figure 8" captionStartId="23.[140,211,1558,1580]" captionTargetBox="[145,1078,415,1513]" captionTargetId="figure@23.[140,1108,394,1541]" captionTargetPageId="23" captionText="Figure 8. Madrasostes mirificum sp. n.: A male genital segment (scale bar: 0, 2 mm) B aedeagus (scale bar: 0, 1 mm) C parameres in dorsal view (scale bar: 0, 0 5 mm) D Madrasostes hasimi sp. n.: male genital segment (scale bar: 0, 1 mm) E parameres (scale bar: 0, 1 mm) F sclerite of internal sac of aedeagus (scale bar: 0, 1 mm) G Madrasostes depressum: sclerite of internal sac of aedeagus (scale bar: 0, 1 mm)." pageId="16" pageNumber="93">Fig. 8</figureCitation>
|
||
G)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="93" type="etymology">
|
||
<paragraph pageId="16" pageNumber="93">
|
||
<emphasis bold="true" box="[185,320,695,721]" pageId="16" pageNumber="93">Etymology.</emphasis>
|
||
Dedicated to Dr. Rosli Hashim, who always helps Munetoshi Maruyama’s research in Malaysia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="93" type="distribution">
|
||
<paragraph pageId="16" pageNumber="93">
|
||
<emphasis bold="true" box="[185,491,765,791]" pageId="16" pageNumber="93">Distribution and habitat.</emphasis>
|
||
This new species is know only from the type locality. For details on collecting circumstances see the introductory paragraphs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |