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<mods:title id="CC15FC6F3CDF80BB02B4CF54FE076F30">Revision of Gymnosophistis Meyrick, 1934 (Lepidoptera: Sesiidae) from East Africa with description of a new species</mods:title>
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<treatment id="03891674FF8C9176E99606DCFD655968" ID-DOI="http://doi.org/10.5281/zenodo.5968933" ID-GBIF-Taxon="154521114" ID-Zenodo-Dep="5968933" LSID="urn:lsid:plazi:treatment:03891674FF8C9176E99606DCFD655968" httpUri="http://treatment.plazi.org/id/03891674FF8C9176E99606DCFD655968" lastPageId="3" lastPageNumber="148" pageId="1" pageNumber="146">
<subSubSection id="C33AF4E9FF8C9174E99606DCFDF45A9A" box="[151,530,725,751]" pageId="1" pageNumber="146" type="nomenclature">
<paragraph id="8B9FA762FF8C9174E99606DCFDF45A9A" blockId="1.[151,530,725,751]" box="[151,530,725,751]" pageId="1" pageNumber="146">
<heading id="D0D7100EFF8C9174E99606DCFDF45A9A" bold="true" box="[151,530,725,751]" fontSize="11" level="1" pageId="1" pageNumber="146" reason="1">
<taxonomicName id="4C20DCE1FF8C9174E99606DCFDF45A9A" ID-CoL="929Z7" authority="Meyrick, 1934" authorityName="Meyrick" authorityYear="1934" box="[151,530,725,751]" class="Insecta" family="Sesiidae" genus="Gymnosophistis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8C9174E99606DCFDF45A9A" bold="true" box="[151,530,725,751]" pageId="1" pageNumber="146">
<emphasis id="B9547B70FF8C9174E99606DCFEB15A9A" bold="true" box="[151,343,725,751]" italics="true" pageId="1" pageNumber="146">Gymnosophistis</emphasis>
<bibRefCitation id="EFB1DA93FF8C9174E85F06DCFDF45A9A" author="Meyrick, E." box="[350,530,725,751]" pageId="1" pageNumber="146" pagination="454 - 456" refId="ref3189" refString="Meyrick, E. (1934) Aegeriadae. Exotic Microlepidoptera, 4, 454 - 456." type="journal article" year="1934">Meyrick, 1934</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33AF4E9FF8C9174E9960712FA915B3A" pageId="1" pageNumber="146" type="materials_examined">
<paragraph id="8B9FA762FF8C9174E9960712FC405B45" blockId="1.[151,1399,794,847]" box="[151,934,794,816]" pageId="1" pageNumber="146">
<typeStatus id="549B19C0FF8C9174E9960712FF2F5B45" box="[151,201,795,816]" pageId="1" pageNumber="146">Type</typeStatus>
species:
<taxonomicName id="4C20DCE1FF8C9174E8270713FCE95B45" authority="Meyrick, 1934: 454" authorityName="Meyrick" authorityPageNumber="454" authorityYear="1934" box="[294,783,794,816]" class="Insecta" family="Sesiidae" genus="Gymnosophistis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="thyrsodoxa">
<emphasis id="B9547B70FF8C9174E8270713FDD95B45" box="[294,575,794,816]" italics="true" pageId="1" pageNumber="146">Gymnosophistis thyrsodoxa</emphasis>
<bibRefCitation id="EFB1DA93FF8C9174EB450713FD3B5B45" author="Meyrick, E." box="[580,733,794,816]" pageId="1" pageNumber="146" pagination="454 - 456" refId="ref3189" refString="Meyrick, E. (1934) Aegeriadae. Exotic Microlepidoptera, 4, 454 - 456." type="journal article" year="1934">Meyrick, 1934</bibRefCitation>
: 454
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="8B9FA762FF8C9174E9960730FA915B3A" blockId="1.[151,1399,794,847]" box="[151,1399,825,847]" pageId="1" pageNumber="146">
References:
<bibRefCitation id="EFB1DA93FF8C9174E8150733FE555B3A" author="Naumann, C. M." box="[276,435,825,847]" pageId="1" pageNumber="146" pagination="1 - 190" refId="ref3207" refString="Naumann, C. M. (1971) Untersuchungen zur Systematik und Phylogenese der holarktischen Sesiiden (Insecta, Lepidoptera). Bonner Zoologische Monographien, 1, 1 - 190." type="journal article" year="1971">Naumann 1971</bibRefCitation>
: 17;
<bibRefCitation id="EFB1DA93FF8C9174E8E50730FD585B3A" author="Fletcher, D. S." box="[484,702,825,847]" pageId="1" pageNumber="146" refId="ref3022" refString="Fletcher, D. S. (1982) Bombycoidea, Castnioidea, Cossoidea, Mimallonoidea. Sesioidea, Sphingoidea, Zygaenoidea. In: Nye, I. W. B. (Ed.), The Generic names of Moths of the World, Uol. 4. British Museum (Natural History) Publication No. 848. British Museum (Natural History), London, 192 pp." type="book" year="1982">Fletcher &amp; Nye 1982</bibRefCitation>
: 72; Heppner &amp; Duckworth 1981: 43;
<bibRefCitation id="EFB1DA93FF8C9174ED440730FAA85B3A" author="Puhringer, F. &amp; Kallies, A." box="[1093,1358,825,847]" pageId="1" pageNumber="146" pagination="1 - 85" refId="ref3240" refString="Puhringer, F. &amp; Kallies, A. (2004) Provisional checklist of the Sesiidae of the world (Lepidoptera: Ditrysia). Mitteilungen der Entomologischen Arbeitsgemeinschaft Salzkammergu t, 4, 1 - 85. [http: // www. sesiidae. net / Checklst. htm (last updated 2017)]" type="journal article" year="2004">Pühringer &amp; Kallies 2004</bibRefCitation>
: 44.
</paragraph>
</subSubSection>
<subSubSection id="C33AF4E9FF8C9174E9960775FC3E5EA5" pageId="1" pageNumber="146" type="description">
<paragraph id="8B9FA762FF8C9174E9960775FD805BE1" blockId="1.[151,1437,892,2033]" box="[151,614,892,917]" pageId="1" pageNumber="146">
<emphasis id="B9547B70FF8C9174E9960775FEA25BE0" bold="true" box="[151,324,892,917]" pageId="1" pageNumber="146">Redescription.</emphasis>
Alar expanse
<quantity id="4CD80A87FF8C9174E8E40775FD875BE1" box="[485,609,892,917]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="146" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">2025 mm</quantity>
.
</paragraph>
<paragraph id="8B9FA762FF8C9174E9C60796FE285C18" blockId="1.[151,1437,892,2033]" pageId="1" pageNumber="146">
<emphasis id="B9547B70FF8C9174E9C60796FEE25BCD" box="[199,260,927,952]" italics="true" pageId="1" pageNumber="146">Head</emphasis>
: haustellum well developed; labial palpus strongly upturned, covering frons, tip between bases of antennae, second palpomere somewhat longer than first and third; dorsally smooth scaled, ventrally rough, somewhat appressed, laterally protruding; width of frons about twice the diameter of an eye; scales of frons and vertex long, hair-like; pericephalic scales dorsally long, hair-like, laterally short; antennae strikingly short, about half as long as forewing, strongly clavate, flagellum proximally somewhat serrate, ciliate, with proximal ciliae longest, club without ciliae.
</paragraph>
<paragraph id="8B9FA762FF8C9174E9C6007EFE315D1D" blockId="1.[151,1437,892,2033]" pageId="1" pageNumber="146">
<emphasis id="B9547B70FF8C9174E9C6007EFEFE5CE5" box="[199,280,1143,1168]" italics="true" pageId="1" pageNumber="146">Thorax</emphasis>
: strong, densely covered with hair-like scales. Legs strong; mid- and hind tibia as well as first tarsomere of hind leg tufted with rough long, hair-like scales; all spur pairs with lateral spurs somewhat less than half as long as mesal ones. Wings: largely transparent; apical area absent; margins very narrow, except for broad costal margin; discal spot of forewing extremely narrow; discal spot of hind wing absent; venation in forewing with R1 to R3 closely approximated, R4 and R5 stalked to more than half length; all veins from stalk of R4/R5 to M3 arise almost equally spaced from cross vein; CuA1 and CuA2 coincident; hindwing venation typical of
<taxonomicName id="4C20DCE1FF8C9174EC190125FA825D30" authorityName="Boisduval" authorityYear="1828" box="[1304,1380,1324,1349]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="tribe" tribe="Sesiini">Sesiini</taxonomicName>
with veins M3 and CuA1 stalked.
</paragraph>
<paragraph id="8B9FA762FF8C9174E9C6017DFD845DC5" blockId="1.[151,1437,892,2033]" pageId="1" pageNumber="146">
<emphasis id="B9547B70FF8C9174E9C6017DFED75DF8" box="[199,305,1396,1421]" italics="true" pageId="1" pageNumber="146">Abdomen</emphasis>
: rather short and broad, spindle-shaped, first tergites densely, other tergites scarcely covered with hair-like scales, anal tuft well developed.
</paragraph>
<paragraph id="8B9FA762FF8C9174E9C601B5FE7D5DA0" blockId="1.[151,1437,892,2033]" box="[199,411,1468,1493]" pageId="1" pageNumber="146">Females unknown.</paragraph>
<paragraph id="8B9FA762FF8C9174E9C601D6FC3E5EA5" blockId="1.[151,1437,892,2033]" pageId="1" pageNumber="146">
<emphasis id="B9547B70FF8C9174E9C601D6FE975D8D" box="[199,369,1503,1528]" italics="true" pageId="1" pageNumber="146">Male genitalia</emphasis>
. (
<figureCitation id="131BBBE7FF8C9174E88F01D6FDE65D8D" box="[398,512,1503,1528]" captionStart="FIGURES 78" captionStartId="4.[151,264,1928,1951]" captionTargetBox="[151,1436,347,1897]" captionTargetId="figure@4.[151,1436,347,1897]" captionTargetPageId="4" captionText="FIGURES 78. Male genitalia of Gymnosophistis Meyrick, 1934: 7. G. thyrsodoxa Meyrick, 1934, lectotype; 8. G. staudei sp. nov., paratype. Scales 1 mm." httpUri="https://zenodo.org/record/2615011/files/figure.png" pageId="1" pageNumber="146">Figs. 78</figureCitation>
). Tegumen broad; gnathos small, slightly asymmetric, distally pointed; uncus narrow, ventro-lateral margins densely covered with long, hair- to bristle-like, proximad pointing setae, dorsally seamlessly fused, ventrally connected to tegumen by a membrane; valva broad, ovoid to rectangular, without crista sacculi or other raised structures, dorso-medially and distally densely covered with proximad pointing, bristle-like, apically mostly bifurcate setae; juxta narrow, posteriorly wide protruding; vinculum forming a short to moderately long saccus; phallus long and slender, almost straight, longer than valva; coecum penis present, short, slightly curved, proximally rounded; vesica basally with numerous teeth or thorns.
</paragraph>
</subSubSection>
<subSubSection id="C33AF4E9FF8C9176E9C602D5FB7A58A0" lastPageId="3" lastPageNumber="148" pageId="1" pageNumber="146" type="diagnosis">
<paragraph id="8B9FA762FF8C9177E9C602D5FD085935" blockId="1.[151,1437,892,2033]" lastBlockId="2.[151,1436,151,320]" lastPageId="2" lastPageNumber="147" pageId="1" pageNumber="146">
<emphasis id="B9547B70FF8C9174E9C602D5FEA75E80" bold="true" box="[199,321,1756,1781]" pageId="1" pageNumber="146">Diagnosis.</emphasis>
Medium sized, powerfully built clearwing moths. The fusion of forewing veins CuA1 and CuA2 is here considered a synapomorphy of the genus; it is unique within the
<taxonomicName id="4C20DCE1FF8C9174EAA002F6FBE45F6D" authorityName="Boisduval" authorityYear="1828" box="[929,1026,1791,1816]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="subFamily" subFamily="Sesiinae">Sesiinae</taxonomicName>
. Antennae extraordinarily short, that of the male clavate, ciliate, rather than pectinate or simple as in most other African
<taxonomicName id="4C20DCE1FF8C9174ED34032DFB635F48" authorityName="Boisduval" authorityYear="1828" box="[1077,1157,1828,1853]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="tribe" tribe="Sesiini">Sesiini</taxonomicName>
.
<taxonomicName id="4C20DCE1FF8C9174ED8E032DFA655F48" authority="Walker, 1856" authorityName="Walker" authorityYear="1856" box="[1167,1411,1828,1853]" class="Insecta" family="Sesiidae" genus="Alonina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8C9174ED8E032DFB0E5F48" box="[1167,1256,1828,1853]" italics="true" pageId="1" pageNumber="146">Alonina</emphasis>
<bibRefCitation id="EFB1DA93FF8C9174EDEF032DFA655F48" author="Walker, F." box="[1262,1411,1828,1853]" pageId="1" pageNumber="146" refId="ref3456" refString="Walker, F. (1856) Sphingidae. List of the Specimens of Lepidopterous Insects in the Collection of the British Museum, 8, 1 - 271. [8 - 72, 83 - 84, 260, 270 (index)]" type="journal volume" year="1856">Walker, 1856</bibRefCitation>
</taxonomicName>
is most probably the sister group of
<taxonomicName id="4C20DCE1FF8C9174EB1D034EFD295F15" authorityName="Meyrick" authorityYear="1934" box="[540,719,1863,1888]" class="Insecta" family="Sesiidae" genus="Gymnosophistis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8C9174EB1D034EFD295F15" box="[540,719,1863,1888]" italics="true" pageId="1" pageNumber="146">Gymnosophistis</emphasis>
</taxonomicName>
. Both genera have a functional haustellum; very similar shape and covering with scales of the labial palpus; coverage of large parts of the body and legs with hair-like scales; in the male genitalia similar shape of tegumen, uncus, valva, juxta and phallus. Most importantly, both genera display a very similar bristle-like shape and proximad pointing arrangement of setae of the uncus and dense, bristle-like, medially and distally located proximad pointing, setae of the valva. Both genera also have clavate and ciliate antennae; however, they are distinctly longer and narrower in
<taxonomicName id="4C20DCE1FF8F9177EA51049EFC4F58C5" authorityName="Walker" authorityYear="1856" box="[848,937,151,176]" class="Insecta" family="Sesiidae" genus="Alonina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8F9177EA51049EFC4F58C5" box="[848,937,151,176]" italics="true" pageId="2" pageNumber="147">Alonina</emphasis>
</taxonomicName>
. Another clear difference of the two genera, beside the fused forewing veins, is the shape of the gnathos, which is symmetric in all species of
<taxonomicName id="4C20DCE1FF8F9177EDD204B5FACA58A0" authorityName="Walker" authorityYear="1856" box="[1235,1324,188,213]" class="Insecta" family="Sesiidae" genus="Alonina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8F9177EDD204B5FACA58A0" box="[1235,1324,188,213]" italics="true" pageId="2" pageNumber="147">Alonina</emphasis>
</taxonomicName>
, with two short, down curved hooks (asymmetric, a single distally slightly pointed protrusion in
<taxonomicName id="4C20DCE1FF8F9177EDD804D6FA6A588D" authorityName="Meyrick" authorityYear="1934" box="[1241,1420,223,248]" class="Insecta" family="Sesiidae" genus="Gymnosophistis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8F9177EDD804D6FA6A588D" box="[1241,1420,223,248]" italics="true" pageId="2" pageNumber="147">Gymnosophistis</emphasis>
</taxonomicName>
). Superficially similar species of
<taxonomicName id="4C20DCE1FF8F9177E8F6050DFD8B5968" authorityName="Naumann" authorityYear="1971" box="[503,621,260,285]" class="Insecta" family="Sesiidae" genus="Felderiola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8F9177E8F6050DFD8B5968" box="[503,621,260,285]" italics="true" pageId="2" pageNumber="147">Felderiola</emphasis>
</taxonomicName>
and
<taxonomicName id="4C20DCE1FF8F9177EBA2050DFCB95968" authorityName="Wallengren" authorityYear="1858" box="[675,863,260,285]" class="Insecta" family="Sesiidae" genus="Monopetalotaxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8F9177EBA2050DFCB95968" box="[675,863,260,285]" italics="true" pageId="2" pageNumber="147">Monopetalotaxis</emphasis>
</taxonomicName>
differ by the lack of a functional haustellum, males moreover by the simple, thorn-like setae of the valva.
</paragraph>
<caption id="DF5FF7EAFF8F9177E9960391FC505FB8" httpUri="https://zenodo.org/record/2615007/files/figure.png" pageId="2" pageNumber="147" startId="2.[151,264,1944,1967]" targetBox="[151,1427,373,1892]" targetPageId="2">
<paragraph id="8B9FA762FF8F9177E9960391FC505FB8" blockId="2.[151,1437,1944,1997]" pageId="2" pageNumber="147">
<emphasis id="B9547B70FF8F9177E9960391FEDD5FD8" bold="true" box="[151,315,1944,1967]" pageId="2" pageNumber="147">FIGURES 16.</emphasis>
Specimens of
<taxonomicName id="4C20DCE1FF8F9177E8D30391FCC15FDB" authority="Meyrick, 1934: 1" authorityName="Meyrick" authorityPageNumber="1" authorityYear="1934" box="[466,807,1944,1966]" class="Insecta" family="Sesiidae" genus="Gymnosophistis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8F9177E8D30391FD925FDB" box="[466,628,1944,1966]" italics="true" pageId="2" pageNumber="147">Gymnosophistis</emphasis>
Meyrick, 1934: 1
</taxonomicName>
2.
<taxonomicName id="4C20DCE1FF8F9177EA530391FB985FDB" authority="Meyrick, 1934" authorityName="Meyrick" authorityYear="1934" box="[850,1150,1944,1966]" class="Insecta" family="Sesiidae" genus="Gymnosophistis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="thyrsodoxa">G. thyrsodoxa Meyrick, 1934</taxonomicName>
, paralectotype; 3. lectotype. 46.
<taxonomicName id="4C20DCE1FF8F9177E9C803B1FEC85FB8" authority="Bartsch, 2018" authorityName="Bartsch" authorityYear="2018" box="[201,302,1975,1997]" class="Insecta" family="Sesiidae" genus="Gymnosophistis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="staudei" status="sp. nov.">
<emphasis id="B9547B70FF8F9177E9C803B1FEC85FB8" box="[201,302,1975,1997]" italics="true" pageId="2" pageNumber="147">G. staudei</emphasis>
</taxonomicName>
<emphasis id="B9547B70FF8F9177E83503BEFE635FB9" bold="true" box="[308,389,1975,1996]" pageId="2" pageNumber="147">
<taxonomicNameLabel id="A267C60BFF8F9177E83503BEFE635FB9" box="[308,389,1975,1996]" pageId="2" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype. Scales 10 mm. (4) head lateral. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="8B9FA762FF8E9176E9C60491FB7A58A0" blockId="3.[151,1436,151,285]" pageId="3" pageNumber="148">
Heppner &amp; Duckworth (1981) list
<taxonomicName id="4C20DCE1FF8E9176EB65049EFCF158C5" authorityName="Meyrick" authorityYear="1934" box="[612,791,151,176]" class="Insecta" family="Sesiidae" genus="Gymnosophistis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B9547B70FF8E9176EB65049EFCF158C5" box="[612,791,151,176]" italics="true" pageId="3" pageNumber="148">Gymnosophistis</emphasis>
</taxonomicName>
under “genera unassigned to subfamily”;
<bibRefCitation id="EFB1DA93FF8E9176EC0D049EFEDE58A1" author="Puhringer, F. &amp; Kallies, A." pageId="3" pageNumber="148" pagination="1 - 85" refId="ref3240" refString="Puhringer, F. &amp; Kallies, A. (2004) Provisional checklist of the Sesiidae of the world (Lepidoptera: Ditrysia). Mitteilungen der Entomologischen Arbeitsgemeinschaft Salzkammergu t, 4, 1 - 85. [http: // www. sesiidae. net / Checklst. htm (last updated 2017)]" type="journal article" year="2004">Pühringer &amp; Kallies (2004)</bibRefCitation>
placed it in “
<taxonomicName id="4C20DCE1FF8E9176E8CD04B5FDCA58A0" authorityName="Boisduval" authorityYear="1828" box="[460,556,188,213]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="subFamily" subFamily="Sesiinae">Sesiinae</taxonomicName>
genera unassigned to tribe”. It is here placed in
<taxonomicName id="4C20DCE1FF8E9176ED4804B5FB7F58A0" authorityName="Boisduval" authorityYear="1828" box="[1097,1177,188,213]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="tribe" tribe="Sesiini">Sesiini</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33AF4E9FF8E9176E9C604D6FD655968" pageId="3" pageNumber="148" type="distribution">
<paragraph id="8B9FA762FF8E9176E9C604D6FCBE588D" blockId="3.[151,1436,151,285]" box="[199,856,223,248]" pageId="3" pageNumber="148">
<emphasis id="B9547B70FF8E9176E9C604D6FE86588D" bold="true" box="[199,352,223,248]" pageId="3" pageNumber="148">Distribution.</emphasis>
So far only known from
<collectingCountry id="F337E7F2FF8E9176EB7A04E9FD21588D" box="[635,711,224,248]" name="Kenya" pageId="3" pageNumber="148">Kenya</collectingCountry>
, East Africa.
</paragraph>
<paragraph id="8B9FA762FF8E9176E9C6050DFD655968" blockId="3.[151,1436,151,285]" box="[199,643,260,285]" pageId="3" pageNumber="148">Biology and host plant(s) are unknown.</paragraph>
</subSubSection>
</treatment>
</document>