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<document id="886BFE37501A450C0E55271E0950F7AA" ID-ISSN="1175-5326" ID-ZooBank="59F6B2D8-D444-468B-943E-DB4C8DE09259" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629821200530" checkinUser="felipe" docAuthor="Akkari, Nesrine &amp; Enghoff, Henrik" docDate="2012" docId="03F987992526FFC6FF4BFBAFA73CF8C9" docLanguage="en" docName="zt03538p053.pdf" docOrigin="Zootaxa 3538" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ommatoiulus pseudoflagellatus Akkari &amp; Enghoff 2012, n. sp." docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="FFC0FFE12506FFE4FFDCFF99A674FFCF" masterDocTitle="Review of the genus Ommatoiulus in Andalusia, Spain (Diplopoda: Julida) with description of ten new species and notes on a remarkable gonopod structure, the fovea" masterLastPageNumber="53" masterPageNumber="1" pageNumber="33" updateTime="1699116717231" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C4E00B9F4F8751348D81508288D340D1">Review of the genus Ommatoiulus in Andalusia, Spain (Diplopoda: Julida) with description of ten new species and notes on a remarkable gonopod structure, the fovea</mods:title>
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<mods:namePart id="662DFACAACA8584EB540D430CC54FECE">Akkari, Nesrine</mods:namePart>
<mods:affiliation id="C4FCF7DAB2032AC14F31113E8AD023EC">Nesrine Akkari (nes. akkari @ gmail. com, nakkari @ snm. ku. dk)</mods:affiliation>
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<mods:namePart id="85062481A1B05E58B1140A738724BA00">Enghoff, Henrik</mods:namePart>
<mods:affiliation id="2846EDD993B991C3B0EF3DFBAFD705A4">Henrik Enghoff (henghoff @ snm. ku. dk)</mods:affiliation>
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<mods:date id="121110F840703AD51B9152D66B2EEF22">2012</mods:date>
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<mods:number id="C6636301FA5D8B5F4E62ADD0D344DE95">2012-11-05</mods:number>
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<treatment id="03F987992526FFC6FF4BFBAFA73CF8C9" ID-DOI="http://doi.org/10.5281/zenodo.5258112" ID-Zenodo-Dep="5258112" LSID="urn:lsid:plazi:treatment:03F987992526FFC6FF4BFBAFA73CF8C9" httpUri="http://treatment.plazi.org/id/03F987992526FFC6FF4BFBAFA73CF8C9" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
<subSubSection id="C34A65042526FFC4FF4BFBAFA53FFB9F" box="[151,843,1078,1104]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BEF368F2526FFC4FF4BFBAFA53FFB9F" blockId="32.[151,843,1078,1137]" box="[151,843,1078,1104]" pageId="32" pageNumber="33">
<heading id="D0A781E32526FFC4FF4BFBAFA53FFB9F" bold="true" box="[151,843,1078,1104]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="B924EA9D2526FFC4FF4BFBAFA53FFB9F" bold="true" box="[151,843,1078,1104]" pageId="32" pageNumber="33">
<taxonomicName id="4C504D0C2526FFC4FF4BFBAFA577FB9F" ID-CoL="49CR8" authority="Akkari &amp; Enghoff" authorityName="Akkari &amp; Enghoff" authorityYear="2012" box="[151,771,1078,1104]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pseudoflagellatus" status="sp. nov.">
<emphasis id="B924EA9D2526FFC4FF4BFBAFA463FB9F" bold="true" box="[151,535,1078,1104]" italics="true" pageId="32" pageNumber="33">Ommatoiulus pseudoflagellatus</emphasis>
Akkari &amp; Enghoff
</taxonomicName>
<taxonomicNameLabel id="A21757E62526FFC4FCD6FBAFA53FFB9F" box="[778,843,1078,1104]" pageId="32" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A65042526FFC4FF4BFBC0A760FBBE" box="[151,276,1113,1137]" pageId="32" pageNumber="33" type="description">
<paragraph id="8BEF368F2526FFC4FF4BFBC0A760FBBE" blockId="32.[151,843,1078,1137]" box="[151,276,1113,1137]" pageId="32" pageNumber="33">
<figureCitation id="136B2A0A2526FFC4FF4BFBC0A760FBBE" box="[151,276,1113,1137]" captionStart-0="FIGURES 7275" captionStart-1="FIGURES 7677" captionStartId-0="33.[151,263,1879,1902]" captionStartId-1="34.[151,264,1372,1395]" captionTargetBox-0="[167,1430,185,1846]" captionTargetBox-1="[161,1427,502,1345]" captionTargetId-0="figure-99@33.[158,1443,143,1882]" captionTargetId-1="figure-406@34.[151,1436,487,1358]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="FIGURES 7275. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 72: Right promerite, posterior view, Fig. 73: Left gonopod, posterior view, Fig. 74: Right posterior gonopod, anterior view, Fig. 75: Left gonopod, mesal view.: Abbreviations: F: Fovea, g: Seminal groove, Jp: Jagged process Lap: Lateral apical process of solenomerite, Lm: Lateral margin, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Mr: Mesal ridge, Ms: Mesomerite, P: Promerite, Px: Paracoxite, S: Solenomerite." captionText-1="FIGURES 7677. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 76: detail of the distal part of the right posterior gonopod, Fig. 77: Close-up of furrow of mesal process of solenomerite and apical part of paracoxite. Abbreviations: If: Internal fringe of solenomerite, Lap: Lateral apical process of solenomrite, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Ms: Mesomerite, Px: Paracoxite, S: Solenomerite." pageId="32" pageNumber="33">Figs 7277</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A65042526FFC4FF4BFB38A402FACE" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BEF368F2526FFC4FF4BFB38A402FACE" blockId="32.[151,1436,1185,2038]" pageId="32" pageNumber="33">
<emphasis id="B924EA9D2526FFC4FF4BFB38A772FB76" box="[151,262,1185,1210]" inLineHeading="true" pageId="32" pageNumber="33" reason="1">
<emphasis id="B924EA9D2526FFC4FF4BFB38A68BFB75" bold="true" box="[151,255,1185,1210]" pageId="32" pageNumber="33">Material</emphasis>
.
</emphasis>
<materialsCitation id="3B383CD22526FFC4FED3FB38A366FB12" collectingDate="2008-02-05" collectionCode="ZMUC" collectorName="H. Reip &amp; K. Voigtlander" county="Cadiz Province" latitude="36.75" location="Riverbank Rio del Bosque" longLatPrecision="1185" longitude="-5.4666667" municipality="Grazalema Benamahoma" pageId="32" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Andalucia" typeStatus="holotype">
<emphasis id="B924EA9D2526FFC4FED3FB38A708FB76" bold="true" box="[271,380,1185,1209]" pageId="32" pageNumber="33">
<typeStatus id="54EB882D2526FFC4FED3FB38A708FB76" box="[271,380,1185,1209]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D56FD062526FFC4FE51FB38A7C8FB76" box="[397,444,1185,1209]" pageId="32" pageNumber="33" type="male">1 ♂</specimenCount>
,
<collectingRegion id="4994F86D2526FFC4FE16FB38A44AFB76" box="[458,574,1185,1209]" country="Spain" name="Andalucia" pageId="32" pageNumber="33">Andalusia</collectingRegion>
,
<collectingCounty id="628E4E032526FFC4FD90FB38A488FB76" box="[588,764,1185,1209]" pageId="32" pageNumber="33">Cádiz Province</collectingCounty>
, west side of
<collectingMunicipality id="6B8BACF52526FFC4FC42FB38A2C8FB76" box="[926,1212,1185,1209]" pageId="32" pageNumber="33">Grazalema Benamahoma</collectingMunicipality>
,
<location id="8E8F60542526FFC4FB16FB38A69AFB12" LSID="urn:lsid:plazi:treatment:03F987992526FFC6FF4BFBAFA73CF8C9:8E8F60542526FFC4FB16FB38A69AFB12" county="Cadiz Province" latitude="36.75" longLatPrecision="1185" longitude="-5.4666667" municipality="Grazalema Benamahoma" name="Riverbank Rio del Bosque" pageId="32" pageNumber="33" stateProvince="Andalucia">Riverbank Rio del Bosque</location>
,
<geoCoordinate id="EE6450482526FFC4FF25FB5CA721FB12" box="[249,341,1221,1245]" degrees="36" direction="north" minutes="45" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="36.75">36º45'N</geoCoordinate>
,
<geoCoordinate id="EE6450482526FFC4FEBCFB5CA7C7FB12" box="[352,435,1221,1245]" degrees="5" direction="west" minutes="28" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="-5.4666667">5º28'W</geoCoordinate>
, under bark of
<taxonomicName id="4C504D0C2526FFC4FD87FB5CA4A3FB12" box="[603,727,1221,1245]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9D2526FFC4FD87FB5CA4A3FB12" box="[603,727,1221,1245]" italics="true" pageId="32" pageNumber="33">Eucalyptus</emphasis>
</taxonomicName>
,
<date id="FFEE104F2526FFC4FD39FB5CA53FFB12" box="[741,843,1221,1245]" pageId="32" pageNumber="33" value="2008-02-05">
<collectingDate id="EFAAE9A72526FFC4FD39FB5CA53FFB12" box="[741,843,1221,1245]" pageId="32" pageNumber="33" value="2008-02-05">5.ii.2008</collectingDate>
</date>
,
<collectorName id="26A553592526FFC4FC8AFB5FA5D9FB12" box="[854,941,1221,1245]" pageId="32" pageNumber="33">H. Reip</collectorName>
&amp;
<collectorName id="26A553592526FFC4FC0CFB5FA200FB12" box="[976,1140,1221,1245]" pageId="32" pageNumber="33">K. Voigtländer</collectorName>
leg. (
<collectionCode id="ED41AE4A2526FFC4FB6FFB5CA37EFB12" box="[1203,1290,1221,1245]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="32" pageNumber="33">ZMUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B383CD22526FFC4FAC0FB5CA406FACE" collectingDate="2008-02-05" collectionCode="ZMUC" collectorName="H. Reip &amp; K. Voigtlander" county="Cadiz Province" latitude="36.75" location="Riverbank Rio del Bosque" longLatPrecision="1185" longitude="-5.4666667" municipality="Grazalema Benamahoma" pageId="32" pageNumber="33" specimenCount="2" specimenCount-female="2" stateProvince="Andalucia" typeStatus="paratype">
<emphasis id="B924EA9D2526FFC4FAC0FB5CA3E0FB12" bold="true" box="[1308,1428,1221,1245]" pageId="32" pageNumber="33">
<typeStatus id="54EB882D2526FFC4FAC0FB5CA3E0FB12" box="[1308,1428,1221,1245]" pageId="32" pageNumber="33" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D56FD062526FFC4FF4BFB70A6A2FACE" box="[151,214,1257,1281]" pageId="32" pageNumber="33" type="female">2 ♀♀</specimenCount>
, same data as the holotype (
<collectionCode id="ED41AE4A2526FFC4FDCFFB70A41EFACE" box="[531,618,1257,1281]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="32" pageNumber="33">ZMUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A65042526FFC4FF1BFA94A2BAFA5E" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BEF368F2526FFC4FF1BFA94A2BAFA5E" blockId="32.[151,1436,1185,2038]" pageId="32" pageNumber="33">
<emphasis id="B924EA9D2526FFC4FF1BFA94A735FAEA" box="[199,321,1293,1318]" inLineHeading="true" pageId="32" pageNumber="33" reason="1">
<emphasis id="B924EA9D2526FFC4FF1BFA94A74EFAE9" bold="true" box="[199,314,1293,1318]" pageId="32" pageNumber="33">Diagnosis</emphasis>
.
</emphasis>
A most peculiar species, somehow comparable with
<taxonomicName id="4C504D0C2526FFC4FC4DFA94A27BFAEA" baseAuthorityName="Verhoeff" baseAuthorityYear="1910" box="[913,1039,1293,1317]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B924EA9D2526FFC4FC4DFA94A27BFAEA" box="[913,1039,1293,1317]" italics="true" pageId="32" pageNumber="33">O. armatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C504D0C2526FFC4FBC0FA94A2FEFAEA" authorityName="Akkari &amp; Enghoff" authorityYear="2012" box="[1052,1162,1293,1317]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="baileyi">
<emphasis id="B924EA9D2526FFC4FBC0FA94A2FEFAEA" box="[1052,1162,1293,1317]" italics="true" pageId="32" pageNumber="33">O. baileyi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C504D0C2526FFC4FB4BFA94A33FFAEA" authorityName="Verhoeff" authorityYear="1893" box="[1175,1355,1293,1317]" class="Diplopoda" family="Julidae" genus="Julus" kingdom="Animalia" order="Julida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="dorsovittatus">
<emphasis id="B924EA9D2526FFC4FB4BFA94A33FFAEA" box="[1175,1355,1293,1317]" italics="true" pageId="32" pageNumber="33">O. dorsovittatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D0C2526FFC4FA5DFA94A697FA86" authorityName=": Mauries" authorityYear="1978" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fuentei">
<emphasis id="B924EA9D2526FFC4FA5DFA94A697FA86" italics="true" pageId="32" pageNumber="33">O. fuentei</emphasis>
</taxonomicName>
in the presence of a voluminous solenomerite with two main lobes separated by a serrated furrow; however, differing from all other
<taxonomicName id="4C504D0C2526FFC4FE75FACCA4EEFAA2" box="[425,666,1365,1389]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B924EA9D2526FFC4FE75FACCA434FAA2" box="[425,576,1365,1389]" italics="true" pageId="32" pageNumber="33">Ommatoiulus</emphasis>
species
</taxonomicName>
by the combination of a mesal solenomertial process strongly bent distally and protruding into a pointed process, a reduced mesomerite and a flagelloid paracoxite.
</paragraph>
</subSubSection>
<subSubSection id="C34A65042526FFC4FF1BFA04A768FA16" pageId="32" pageNumber="33" type="etymology">
<paragraph id="8BEF368F2526FFC4FF1BFA04A768FA16" blockId="32.[151,1436,1185,2038]" pageId="32" pageNumber="33">
<emphasis id="B924EA9D2526FFC4FF1BFA04A739FA7A" box="[199,333,1437,1461]" inLineHeading="true" pageId="32" pageNumber="33" reason="1">
<emphasis id="B924EA9D2526FFC4FF1BFA04A732FA7A" bold="true" box="[199,326,1437,1461]" pageId="32" pageNumber="33">Etymology</emphasis>
.
</emphasis>
The species epithet refers to the particular configuration of the paracoxite, resembling a flagellum.
</paragraph>
</subSubSection>
<subSubSection id="C34A65042526FFC6FF1BFA7CA2E8FE0A" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
<paragraph id="8BEF368F2526FFC4FF1BFA7CA7FDF91A" blockId="32.[151,1436,1185,2038]" pageId="32" pageNumber="33">
<emphasis id="B924EA9D2526FFC4FF1BFA7CA72CFA32" box="[199,344,1509,1534]" inLineHeading="true" pageId="32" pageNumber="33" reason="1">
<emphasis id="B924EA9D2526FFC4FF1BFA7CA725FA31" bold="true" box="[199,337,1509,1534]" pageId="32" pageNumber="33">Description</emphasis>
.
</emphasis>
Male: L: 26.4 mm, H: 2.5 mm, 46 PR+2 AR+T. Females: L: 21.22.5 mm, H: 2.52.9 mm, 4245 PR+ 1 AR+ T. Eyes composed of 35 black ocelli arranged in 8 vertical rows. General colour with greyish to light purple prozonites and light yellowish metazonites, becoming brownish dorsally. Frontal part of the head and collum dark tawny-brown, labral zone yellowish, mouthparts bright yellow. Antennae dark brown, legs tawnybrown; preanal ring dark brown, anal valves blackish. Ozopores small, circular, openings at ca. their diameter behind suture, suture complete, curving at ozopore level; prozonites with irregular oblique striation; metazonites with regular striation.
</paragraph>
<paragraph id="8BEF368F2526FFC4FF1BF978A2AAF88E" blockId="32.[151,1436,1185,2038]" pageId="32" pageNumber="33">Telson: Preanal ring with a hyaline upturned process and a horizontal caudal projection bearing 5+5 setae on the tip and 2+2 on the sides; subanal scale with a triangular apex and 5+5 setae; anal valves with 4+4 long setae on the surface, a marginal row of several short setae and a submarginal row of ca.10+10 longer ones.</paragraph>
<paragraph id="8BEF368F2526FFC6FF1BF8D4A2E8FE0A" blockId="32.[151,1436,1185,2038]" lastBlockId="34.[151,1436,151,453]" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
Gonopods. Promerite (
<emphasis id="B924EA9D2526FFC4FE18F8D4A7A1F8AA" bold="true" box="[452,469,1869,1893]" pageId="32" pageNumber="33">P</emphasis>
) narrow basally, broadest at midlength, lateral margin (
<emphasis id="B924EA9D2526FFC4FBE0F8D4A212F8AA" bold="true" box="[1084,1126,1869,1893]" pageId="32" pageNumber="33">Lm</emphasis>
) expanded in a big rounded lobe (
<figureCitation id="136B2A0A2526FFC4FF0AF8E8A75CF846" box="[214,296,1905,1929]" captionStart="FIGURES 7275" captionStartId="33.[151,263,1879,1902]" captionTargetBox="[167,1430,185,1846]" captionTargetId="figure-99@33.[158,1443,143,1882]" captionTargetPageId="33" captionText="FIGURES 7275. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 72: Right promerite, posterior view, Fig. 73: Left gonopod, posterior view, Fig. 74: Right posterior gonopod, anterior view, Fig. 75: Left gonopod, mesal view.: Abbreviations: F: Fovea, g: Seminal groove, Jp: Jagged process Lap: Lateral apical process of solenomerite, Lm: Lateral margin, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Mr: Mesal ridge, Ms: Mesomerite, P: Promerite, Px: Paracoxite, S: Solenomerite." pageId="32" pageNumber="33">Fig. 72</figureCitation>
), gradually narrowing distad, apically with a small lateral jagged process (
<emphasis id="B924EA9D2526FFC4FBBDF8E8A20AF846" bold="true" box="[1121,1150,1905,1929]" pageId="32" pageNumber="33">Jp</emphasis>
); mesal ridge (
<emphasis id="B924EA9D2526FFC4FAFAF8E8A339F846" bold="true" box="[1318,1357,1905,1929]" pageId="32" pageNumber="33">Mr</emphasis>
) broad and protruding distally in a rounded lobe, rudimentary telopodite absent or indistinct. Posterior gonopod: Mesomerite (
<emphasis id="B924EA9D2526FFC4FEF1F820A726F81E" bold="true" box="[301,338,1977,2001]" pageId="32" pageNumber="33">Ms</emphasis>
) broadest at the basis, gradually narrowing distad, mesal margin bent at blunt angles at about 2/3 length (
<figureCitation id="136B2A0A2526FFC4FF37F844A735F83A" box="[235,321,2013,2037]" captionStart="FIGURES 7275" captionStartId="33.[151,263,1879,1902]" captionTargetBox="[167,1430,185,1846]" captionTargetId="figure-99@33.[158,1443,143,1882]" captionTargetPageId="33" captionText="FIGURES 7275. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 72: Right promerite, posterior view, Fig. 73: Left gonopod, posterior view, Fig. 74: Right posterior gonopod, anterior view, Fig. 75: Left gonopod, mesal view.: Abbreviations: F: Fovea, g: Seminal groove, Jp: Jagged process Lap: Lateral apical process of solenomerite, Lm: Lateral margin, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Mr: Mesal ridge, Ms: Mesomerite, P: Promerite, Px: Paracoxite, S: Solenomerite." pageId="32" pageNumber="33">Figs 74</figureCitation>
,
<figureCitation id="136B2A0A2526FFC4FE91F844A718F83A" box="[333,364,2013,2037]" captionStart="FIGURES 7677" captionStartId="34.[151,264,1372,1395]" captionTargetBox="[161,1427,502,1345]" captionTargetId="figure-406@34.[151,1436,487,1358]" captionTargetPageId="34" captionText="FIGURES 7677. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 76: detail of the distal part of the right posterior gonopod, Fig. 77: Close-up of furrow of mesal process of solenomerite and apical part of paracoxite. Abbreviations: If: Internal fringe of solenomerite, Lap: Lateral apical process of solenomrite, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Ms: Mesomerite, Px: Paracoxite, S: Solenomerite." pageId="32" pageNumber="33">76</figureCitation>
), distal third curved, tapering apically and pointing anteromesad. Solenomerite (
<emphasis id="B924EA9D2526FFC4FB36F844A28DF839" bold="true" box="[1258,1273,2013,2038]" pageId="32" pageNumber="33">S</emphasis>
) large, extending beyond the other branches, composed of a uniformly broad, folded lateral process (
<emphasis id="B924EA9D2524FFC6FB67FF01A2A9FF7F" bold="true" box="[1211,1245,152,176]" pageId="34" pageNumber="35">Lp</emphasis>
), apically with a rounded lobe bearing scattered serrations, and a long mesal process (
<emphasis id="B924EA9D2524FFC6FC6CFF22A5ADFF1C" bold="true" box="[944,985,187,211]" pageId="34" pageNumber="35">Mp</emphasis>
) with a folded, lateral margin showing numerous rows of spikes on the internal fringe (
<emphasis id="B924EA9D2524FFC6FD71FF44A4B5FF39" bold="true" box="[685,705,221,246]" pageId="34" pageNumber="35">If</emphasis>
) (
<figureCitation id="136B2A0A2524FFC6FD06FF44A558FF3A" box="[730,812,221,245]" captionStart="FIGURES 7677" captionStartId="34.[151,264,1372,1395]" captionTargetBox="[161,1427,502,1345]" captionTargetId="figure-406@34.[151,1436,487,1358]" captionTargetPageId="34" captionText="FIGURES 7677. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 76: detail of the distal part of the right posterior gonopod, Fig. 77: Close-up of furrow of mesal process of solenomerite and apical part of paracoxite. Abbreviations: If: Internal fringe of solenomerite, Lap: Lateral apical process of solenomrite, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Ms: Mesomerite, Px: Paracoxite, S: Solenomerite." pageId="34" pageNumber="35">Fig. 77</figureCitation>
), apically bent 90° in a rounded angle, protruding into a transparent process (
<emphasis id="B924EA9D2524FFC6FEA1FE99A7D8FED7" bold="true" box="[381,428,256,280]" pageId="34" pageNumber="35">Lap</emphasis>
) covered with numerous spikes on the surface and the margin, pointing laterad (
<figureCitation id="136B2A0A2524FFC6FACEFF66A312FED8" box="[1298,1382,255,279]" captionStart="FIGURES 7275" captionStartId="33.[151,263,1879,1902]" captionTargetBox="[167,1430,185,1846]" captionTargetId="figure-99@33.[158,1443,143,1882]" captionTargetPageId="33" captionText="FIGURES 7275. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 72: Right promerite, posterior view, Fig. 73: Left gonopod, posterior view, Fig. 74: Right posterior gonopod, anterior view, Fig. 75: Left gonopod, mesal view.: Abbreviations: F: Fovea, g: Seminal groove, Jp: Jagged process Lap: Lateral apical process of solenomerite, Lm: Lateral margin, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Mr: Mesal ridge, Ms: Mesomerite, P: Promerite, Px: Paracoxite, S: Solenomerite." pageId="34" pageNumber="35">Figs 73</figureCitation>
,
<figureCitation id="136B2A0A2524FFC6FAADFF66A3E4FED8" box="[1393,1424,255,279]" captionStart="FIGURES 7677" captionStartId="34.[151,264,1372,1395]" captionTargetBox="[161,1427,502,1345]" captionTargetId="figure-406@34.[151,1436,487,1358]" captionTargetPageId="34" captionText="FIGURES 7677. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 76: detail of the distal part of the right posterior gonopod, Fig. 77: Close-up of furrow of mesal process of solenomerite and apical part of paracoxite. Abbreviations: If: Internal fringe of solenomerite, Lap: Lateral apical process of solenomrite, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Ms: Mesomerite, Px: Paracoxite, S: Solenomerite." pageId="34" pageNumber="35">76</figureCitation>
). Both lateral and mesal processes furrowed and connected distally by a serrated membrane covered by several spikes; fovea (
<emphasis id="B924EA9D2524FFC6FF38FEDCA681FE92" bold="true" box="[228,245,325,349]" pageId="34" pageNumber="35">F</emphasis>
) located at the basis of the mesal process (
<figureCitation id="136B2A0A2524FFC6FD16FEDCA568FE92" box="[714,796,325,349]" captionStart="FIGURES 7275" captionStartId="33.[151,263,1879,1902]" captionTargetBox="[167,1430,185,1846]" captionTargetId="figure-99@33.[158,1443,143,1882]" captionTargetPageId="33" captionText="FIGURES 7275. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 72: Right promerite, posterior view, Fig. 73: Left gonopod, posterior view, Fig. 74: Right posterior gonopod, anterior view, Fig. 75: Left gonopod, mesal view.: Abbreviations: F: Fovea, g: Seminal groove, Jp: Jagged process Lap: Lateral apical process of solenomerite, Lm: Lateral margin, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Mr: Mesal ridge, Ms: Mesomerite, P: Promerite, Px: Paracoxite, S: Solenomerite." pageId="34" pageNumber="35">Fig. 74</figureCitation>
), seminal groove (
<emphasis id="B924EA9D2524FFC6FC3BFEDCA581FE92" bold="true" box="[999,1013,325,349]" pageId="34" pageNumber="35">g</emphasis>
) running from the fovea posterodistad up to the extremity of the mesal process of the solenomerite. From the lateral basis of the mesal process emerges a flagelloid process which we interpret as the paracoxite (
<emphasis id="B924EA9D2524FFC6FCC5FE12A54CFE6C" bold="true" box="[793,824,395,419]" pageId="34" pageNumber="35">Px</emphasis>
), running along the fold, between the jagged internal fringes of the mesal process, ending subapically at the level of the apical flexion (
<figureCitation id="136B2A0A2524FFC6FBCCFE34A212FE0A" box="[1040,1126,429,453]" captionStart="FIGURES 7275" captionStartId="33.[151,263,1879,1902]" captionTargetBox="[167,1430,185,1846]" captionTargetId="figure-99@33.[158,1443,143,1882]" captionTargetPageId="33" captionText="FIGURES 7275. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 72: Right promerite, posterior view, Fig. 73: Left gonopod, posterior view, Fig. 74: Right posterior gonopod, anterior view, Fig. 75: Left gonopod, mesal view.: Abbreviations: F: Fovea, g: Seminal groove, Jp: Jagged process Lap: Lateral apical process of solenomerite, Lm: Lateral margin, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Mr: Mesal ridge, Ms: Mesomerite, P: Promerite, Px: Paracoxite, S: Solenomerite." pageId="34" pageNumber="35">Figs 73</figureCitation>
,
<figureCitation id="136B2A0A2524FFC6FBADFE34A2E4FE0A" box="[1137,1168,429,453]" captionStart="FIGURES 7677" captionStartId="34.[151,264,1372,1395]" captionTargetBox="[161,1427,502,1345]" captionTargetId="figure-406@34.[151,1436,487,1358]" captionTargetPageId="34" captionText="FIGURES 7677. Ommatoiulus pseudoflagellatus n. sp., holotype: Fig. 76: detail of the distal part of the right posterior gonopod, Fig. 77: Close-up of furrow of mesal process of solenomerite and apical part of paracoxite. Abbreviations: If: Internal fringe of solenomerite, Lap: Lateral apical process of solenomrite, Lp: Lateral process of solenomerite, Mp: Mesal process of solenomerite, Ms: Mesomerite, Px: Paracoxite, S: Solenomerite." pageId="34" pageNumber="35">77</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2F66072527FFC5FF4BF8CEA735F83B" pageId="33" pageNumber="34" startId="33.[151,263,1879,1902]" targetBox="[167,1430,185,1846]" targetPageId="33">
<paragraph id="8BEF368F2527FFC5FF4BF8CEA735F83B" blockId="33.[151,1436,1879,2037]" pageId="33" pageNumber="34">
<emphasis id="B924EA9D2527FFC5FF4BF8CEA723F8A1" bold="true" box="[151,343,1879,1902]" pageId="33" pageNumber="34">FIGURES 7275.</emphasis>
<taxonomicName id="4C504D0C2527FFC5FEBCF8CEA4D1F8A2" authority="Akkari &amp; Enghoff, 2012" authorityName="Akkari &amp; Enghoff" authorityYear="2012" box="[352,677,1879,1901]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pseudoflagellatus" status="sp. nov.">
<emphasis id="B924EA9D2527FFC5FEBCF8CEA4D1F8A2" box="[352,677,1879,1901]" italics="true" pageId="33" pageNumber="34">Ommatoiulus pseudoflagellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757E62527FFC5FD73F8C1A49EF8A2" box="[687,746,1880,1901]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
, holotype: Fig. 72: Right promerite, posterior view, Fig. 73: Left gonopod, posterior view, Fig. 74: Right posterior gonopod, anterior view, Fig. 75: Left gonopod, mesal view.: Abbreviations:
<emphasis id="B924EA9D2527FFC5FA59F8E0A3E0F841" bold="true" box="[1413,1428,1913,1934]" pageId="33" pageNumber="34">F</emphasis>
: Fovea,
<emphasis id="B924EA9D2527FFC5FF3FF803A684F860" bold="true" box="[227,240,1946,1967]" pageId="33" pageNumber="34">g</emphasis>
: Seminal groove,
<emphasis id="B924EA9D2527FFC5FE72F803A7BDF860" bold="true" box="[430,457,1946,1967]" pageId="33" pageNumber="34">Jp</emphasis>
: Jagged process
<emphasis id="B924EA9D2527FFC5FDA7F803A4D2F860" bold="true" box="[635,678,1946,1967]" pageId="33" pageNumber="34">Lap</emphasis>
: Lateral apical process of solenomerite,
<emphasis id="B924EA9D2527FFC5FB90F803A206F860" bold="true" box="[1100,1138,1946,1967]" pageId="33" pageNumber="34">Lm</emphasis>
: Lateral margin,
<emphasis id="B924EA9D2527FFC5FAFAF803A331F860" bold="true" box="[1318,1349,1946,1967]" pageId="33" pageNumber="34">Lp</emphasis>
: Lateral process of solenomerite,
<emphasis id="B924EA9D2527FFC5FE48F825A7CEF81E" bold="true" box="[404,442,1980,2001]" pageId="33" pageNumber="34">Mp</emphasis>
: Mesal process of solenomerite,
<emphasis id="B924EA9D2527FFC5FCD5F825A558F81E" bold="true" box="[777,812,1980,2001]" pageId="33" pageNumber="34">Mr</emphasis>
: Mesal ridge,
<emphasis id="B924EA9D2527FFC5FC63F825A595F81E" bold="true" box="[959,993,1980,2001]" pageId="33" pageNumber="34">Ms</emphasis>
: Mesomerite,
<emphasis id="B924EA9D2527FFC5FBA8F825A2F7F81E" bold="true" box="[1140,1155,1980,2001]" pageId="33" pageNumber="34">P</emphasis>
: Promerite,
<emphasis id="B924EA9D2527FFC5FADDF825A369F81E" bold="true" box="[1281,1309,1980,2001]" pageId="33" pageNumber="34">Px</emphasis>
: Paracoxite,
<emphasis id="B924EA9D2527FFC5FF4BF847A6D1F83A" bold="true" box="[151,165,2014,2037]" pageId="33" pageNumber="34">S</emphasis>
: Solenomerite.
</paragraph>
</caption>
<caption id="DF2F66072524FFC6FF4BFAC5A552FA00" pageId="34" pageNumber="35" startId="34.[151,264,1372,1395]" targetBox="[161,1427,502,1345]" targetPageId="34">
<paragraph id="8BEF368F2524FFC6FF4BFAC5A552FA00" blockId="34.[151,1436,1372,1487]" pageId="34" pageNumber="35">
<emphasis id="B924EA9D2524FFC6FF4BFAC5A72CFABC" bold="true" box="[151,344,1372,1395]" pageId="34" pageNumber="35">FIGURES 7677.</emphasis>
<taxonomicName id="4C504D0C2524FFC6FEBDFAC4A4D3FABC" authority="Akkari &amp; Enghoff, 2012" authorityName="Akkari &amp; Enghoff" authorityYear="2012" box="[353,679,1373,1395]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pseudoflagellatus" status="sp. nov.">
<emphasis id="B924EA9D2524FFC6FEBDFAC4A4D3FABC" box="[353,679,1373,1395]" italics="true" pageId="34" pageNumber="35">Ommatoiulus pseudoflagellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757E62524FFC6FD6DFAC4A499FABD" box="[689,749,1373,1394]" pageId="34" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
, holotype: Fig. 76: detail of the distal part of the right posterior gonopod, Fig. 77: Close-up of furrow of mesal process of solenomerite and apical part of paracoxite. Abbreviations:
<emphasis id="B924EA9D2524FFC6FAF2FAE2A334FA5D" bold="true" box="[1326,1344,1403,1426]" pageId="34" pageNumber="35">If</emphasis>
: Internal fringe of solenomerite,
<emphasis id="B924EA9D2524FFC6FE58FA03A7C4FA60" bold="true" box="[388,432,1434,1455]" pageId="34" pageNumber="35">Lap</emphasis>
: Lateral apical process of solenomrite,
<emphasis id="B924EA9D2524FFC6FC9CFA03A52BFA60" bold="true" box="[832,863,1434,1455]" pageId="34" pageNumber="35">Lp</emphasis>
: Lateral process of solenomerite,
<emphasis id="B924EA9D2524FFC6FB61FA03A297FA60" bold="true" box="[1213,1251,1434,1455]" pageId="34" pageNumber="35">Mp</emphasis>
: Mesal process of solenomerite,
<emphasis id="B924EA9D2524FFC6FEFAFA20A73CFA01" bold="true" box="[294,328,1465,1486]" pageId="34" pageNumber="35">Ms</emphasis>
: Mesomerite,
<emphasis id="B924EA9D2524FFC6FE07FA20A783FA01" bold="true" box="[475,503,1465,1486]" pageId="34" pageNumber="35">Px</emphasis>
: Paracoxite,
<emphasis id="B924EA9D2524FFC6FDA0FA21A4FEFA00" bold="true" box="[636,650,1464,1487]" pageId="34" pageNumber="35">S</emphasis>
: Solenomerite.
</paragraph>
</caption>
<subSubSection id="C34A65042524FFC6FF4BFA62A4E1F9F9" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BEF368F2524FFC6FF4BFA62A59DF9DC" blockId="34.[151,1436,1531,1798]" box="[151,1001,1531,1556]" pageId="34" pageNumber="35">
<emphasis id="B924EA9D2524FFC6FF4BFA62A758F9DC" box="[151,300,1531,1556]" inLineHeading="true" pageId="34" pageNumber="35" reason="1">
<emphasis id="B924EA9D2524FFC6FF4BFA62A752F9DB" bold="true" box="[151,294,1531,1556]" pageId="34" pageNumber="35">Distribution</emphasis>
.
</emphasis>
Known only from
<typeStatus id="54EB882D2524FFC6FE22FA65A459F9DB" box="[510,557,1532,1556]" pageId="34" pageNumber="35">type</typeStatus>
locality in Cádiz Province in
<collectingRegion id="4994F86D2524FFC6FCAEFA62A591F9DC" box="[882,997,1531,1555]" country="Spain" name="Andalucia" pageId="34" pageNumber="35">Andalusia</collectingRegion>
.
</paragraph>
<paragraph id="8BEF368F2524FFC6FF1BF984A4E1F9F9" blockId="34.[151,1436,1531,1798]" box="[199,661,1565,1590]" pageId="34" pageNumber="35">
<emphasis id="B924EA9D2524FFC6FF1BF984A75CF9F9" box="[199,296,1565,1590]" inLineHeading="true" pageId="34" pageNumber="35" reason="1">
<emphasis id="B924EA9D2524FFC6FF1BF984A755F9F9" bold="true" box="[199,289,1565,1590]" pageId="34" pageNumber="35">Habitat</emphasis>
.
</emphasis>
Found under bark of
<taxonomicName id="4C504D0C2524FFC6FDC9F987A4FBF9F9" box="[533,655,1566,1590]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B924EA9D2524FFC6FDC9F987A4FBF9F9" box="[533,655,1566,1590]" italics="true" pageId="34" pageNumber="35">Eucalyptus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A65042524FFC6FF1BF9D9A73CF8C9" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BEF368F2524FFC6FF1BF9D9A73CF8C9" blockId="34.[151,1436,1531,1798]" pageId="34" pageNumber="35">
<emphasis id="B924EA9D2524FFC6FF1BF9D9A738F997" box="[199,332,1600,1625]" inLineHeading="true" pageId="34" pageNumber="35" reason="1">
<emphasis id="B924EA9D2524FFC6FF1BF9D9A732F996" bold="true" box="[199,326,1600,1625]" pageId="34" pageNumber="35">Comments</emphasis>
.
</emphasis>
The gonopods of
<taxonomicName id="4C504D0C2524FFC6FDF4F9D8A57BF996" authorityName="Akkari &amp; Enghoff" authorityYear="2012" box="[552,783,1601,1625]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pseudoflagellatus">
<emphasis id="B924EA9D2524FFC6FDF4F9D8A57BF996" box="[552,783,1601,1625]" italics="true" pageId="34" pageNumber="35">O. pseudoflagellatus</emphasis>
</taxonomicName>
are quite different from those of all other
<taxonomicName id="4C504D0C2524FFC6FAD4F9D8A69DF9B4" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B924EA9D2524FFC6FAD4F9D8A3E8F996" box="[1288,1436,1601,1625]" italics="true" pageId="34" pageNumber="35">Ommatoiulus</emphasis>
species
</taxonomicName>
, and we had great difficulties homologising the various processes of the posterior gonopod with the normal components of an
<taxonomicName id="4C504D0C2524FFC6FEBEF91FA782F951" box="[354,502,1670,1694]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B924EA9D2524FFC6FEBEF91FA782F951" box="[354,502,1670,1694]" italics="true" pageId="34" pageNumber="35">Ommatoiulus</emphasis>
</taxonomicName>
gonopod. One might argue that the species is so deviating that it should be placed in is own new genus. However, awaiting further ongoing revisionary work on
<taxonomicName id="4C504D0C2524FFC6FC39F930A20DF90E" box="[997,1145,1705,1729]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B924EA9D2524FFC6FC39F930A20DF90E" box="[997,1145,1705,1729]" italics="true" pageId="34" pageNumber="35">Ommatoiulus</emphasis>
</taxonomicName>
and, hopefully, additional material of
<taxonomicName id="4C504D0C2524FFC6FEC3F955A472F92B" box="[287,518,1740,1764]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pesudoflagellatus">
<emphasis id="B924EA9D2524FFC6FEC3F955A472F92B" box="[287,518,1740,1764]" italics="true" pageId="34" pageNumber="35">O. pesudoflagellatus</emphasis>
</taxonomicName>
, we classify it, for the time being, in the same genus as the other Andalusian schizophyllines.
</paragraph>
</subSubSection>
</treatment>
</document>