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<document id="84168019AA6E5ED0830FBEEC7323DBC6" ID-DOI="10.5281/zenodo.5353544" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353544" ID-ZooBank="772BDDF2-AD14-4950-B3B3-D6A2C9E403C2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630210667410" checkinUser="felipe" docAuthor="Gordon, Robert D., F, Guillermo González &amp; Hanley, Guy A." docDate="2020" docId="03D687B4FF9B6A16FF0AFF54FC4CFBC4" docLanguage="en" docName="0766.pdf" docOrigin="Insecta Mundi 2020 (766)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Calloeneis roxanne Gordon and Hanley 2020, new species" docType="treatment" docVersion="4" lastPageNumber="11" masterDocId="FFEFFFCCFF976A1AFFCAFF83FFB6FF8F" masterDocTitle="South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="11" updateTime="1699187120983" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="B3961D208A7AA91D8A6AB73CC0373A13" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="10CD96797A6A484DC3328BC56E2216A8">
<mods:title id="8AC89BB7E3CE841D5C5A35F3BC92C6DD">South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini)</mods:title>
</mods:titleInfo>
<mods:name id="B8CC5AE50C38DD324C535EB3F1203053" type="personal">
<mods:role id="96A95C351F737B4BD14FDE47CABE57C4">
<mods:roleTerm id="F67946199FDB520DB4F20181F3D0AF3F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="703881BE3AD322ADE9A8D97007AB2C35">Gordon, Robert D.</mods:namePart>
<mods:affiliation id="6CFF041DF69C46EA00DEAACEEB5B17FB">Northern Plains Entomology PO Box 65 Willow City, ND 58384, USA</mods:affiliation>
</mods:name>
<mods:name id="28D014124E31802CCACB2ACB8CD57FA3" type="personal">
<mods:role id="2B2AC22816D7DBC0299A96FD970E2248">
<mods:roleTerm id="C3EFEB15FA761B71C313DE8D4C8DBF38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F4A0A0EE909F805C51702F08575AF98">F, Guillermo González</mods:namePart>
<mods:affiliation id="E087D46622AC016362938EBEC127C966">Nocedal 6455 La Reina Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name id="393694356306D65B07BA817F56E6A3F9" type="personal">
<mods:role id="9EE57BEB4636EC5585A6577414287863">
<mods:roleTerm id="B75DB909366825709EBBE4B90286025A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="102EE07010BA01C86E5D9351A263F87F">Hanley, Guy A.</mods:namePart>
<mods:affiliation id="DBE96AF637304EF21FDE33E4A2C5ED36">Northern Plains Entomology Minot, ND</mods:affiliation>
</mods:name>
<mods:typeOfResource id="BC6C5D3AF4B9D6DE49B9F94D0206AB62">text</mods:typeOfResource>
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<mods:titleInfo id="0C3991AFE5E1AC6ABDBBF4BAEFA69937">
<mods:title id="F30061331640FBF89DB0E2ABC22C1AEB">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="A2EDAFF5881313A75D4B5094C734B0A9">
<mods:date id="4FFEC155DB23A2C4EC68E7BEE31D1814">2020</mods:date>
<mods:detail id="F673FB97681A6C931EF08F0D6DE956D6" type="pubDate">
<mods:number id="BB20A5BE415393850E08790F5957E672">2020-05-29</mods:number>
</mods:detail>
<mods:detail id="B1A181381315EEEA335E8668D22C081A" type="volume">
<mods:number id="9F368883F97A64AD46EC66F04D84101A">2020</mods:number>
</mods:detail>
<mods:detail id="AE33503E219C2ECBD8D21C0A66640AC3" type="issue">
<mods:number id="CB52046172BE6A957A0F7F108AE6B6F4">766</mods:number>
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<mods:start id="4C7F065C16CCE5685A0617D78BF97022">1</mods:start>
<mods:end id="65988611D9AA4AD6199C41EFA1FD04A4">25</mods:end>
</mods:extent>
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<mods:classification id="577FCFD1E61CDD4B1753708C368B0BCC">journal article</mods:classification>
<mods:identifier id="95F6B0DE09AC1DD2BB2DD853710990F6" type="DOI">10.5281/zenodo.5353544</mods:identifier>
<mods:identifier id="6FADF217E9687D9489109F360C442A53" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="5BC2BE873A5A96BADACC734C61B30B83" type="Zenodo-Dep">5353544</mods:identifier>
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</mods:mods>
<treatment id="03D687B4FF9B6A16FF0AFF54FC4CFBC4" ID-DOI="http://doi.org/10.5281/zenodo.5451060" ID-Zenodo-Dep="5451060" LSID="urn:lsid:plazi:treatment:03D687B4FF9B6A16FF0AFF54FC4CFBC4" httpUri="http://treatment.plazi.org/id/03D687B4FF9B6A16FF0AFF54FC4CFBC4" lastPageNumber="11" pageId="12" pageNumber="11">
<subSubSection id="C3656529FF9B6A16FF0AFF54FC0AFF60" box="[192,956,214,239]" pageId="12" pageNumber="11" type="nomenclature">
<paragraph id="8BC036A2FF9B6A16FF0AFF54FC0AFF60" blockId="12.[192,956,214,239]" box="[192,956,214,239]" pageId="12" pageNumber="11">
<heading id="D08881CEFF9B6A16FF0AFF54FC0AFF60" bold="true" box="[192,956,214,239]" fontSize="10" level="2" pageId="12" pageNumber="11" reason="3">
<emphasis id="B90BEAB0FF9B6A16FF0AFF54FC0AFF60" bold="true" box="[192,956,214,239]" pageId="12" pageNumber="11">
11.
<taxonomicName id="4C7F4D21FF9B6A16FF24FF55FCBDFF60" authority="Gordon and Hanley" authorityName="Gordon and Hanley" authorityYear="2020" box="[238,779,214,239]" class="Insecta" family="Coccinellidae" genus="Calloeneis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="roxanne" status="sp. nov.">
<emphasis id="B90BEAB0FF9B6A16FF24FF55FE42FF61" bold="true" box="[238,500,214,238]" italics="true" pageId="12" pageNumber="11">Calloeneis roxanne</emphasis>
Gordon and Hanley
</taxonomicName>
,
<taxonomicNameLabel id="A23857CBFF9B6A16FCD2FF54FC0AFF60" box="[792,956,215,239]" pageId="12" pageNumber="11" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF9B6A16FF0AFE84FBACFC26" pageId="12" pageNumber="11" type="description">
<paragraph id="8BC036A2FF9B6A16FF0AFE84FA85FC88" blockId="12.[190,1442,263,775]" pageId="12" pageNumber="11">
<emphasis id="B90BEAB0FF9B6A16FF0AFE84FDF6FEAF" bold="true" box="[192,576,263,288]" pageId="12" pageNumber="11">
Description. Male
<typeStatus id="54C48800FF9B6A16FE0AFE8BFD8CFEAF" box="[448,570,264,288]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
.
</emphasis>
Length
<quantity id="4C879B47FF9B6A16FD66FE84FCBAFE90" box="[684,780,263,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="11" unit="mm" value="1.6">1.6 mm</quantity>
, width
<quantity id="4C879B47FF9B6A16FCA6FE8BFC7AFE90" box="[876,972,264,288]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="11" unit="mm" value="1.3">1.3 mm</quantity>
; dorsal surface shiny, head distinctly alutaceous. Color yellow; elytron with narrow, oval dark brown macula on sutural margin in basal 1/2, additional black macula present on lateral margin in basal 1/2 (
<figureCitation id="13442A27FF9B6A16FC2AFECBFB8FFEEE" box="[992,1081,328,353]" captionStart="Figures 38–53" captionStartId="23.[192,287,1678,1700]" captionTargetBox="[205,1432,233,1600]" captionTargetId="figure-89@23.[192,1440,217,1650]" captionTargetPageId="23" captionText="Figures 38–53. Calloeneis spp.38–42) Calloeneis krista. 38) Habitus.39) Frons.40) Penis guide ventral.41) Penis guide lateral. 42) Penis. 43–48) Calloeneis roxanne. 43) Habitus. 44) Frons. 45) Penis guide ventral. 46) Penis guide lateral. 47) Penis. 48) Spermathecal capsule. 49–53) Calloeneis angelica. 49) Habitus. 50) Frons. 51) Penis guide ventral. 52) Penis guide lateral. 53) Penis." figureDoi="http://doi.org/10.5281/zenodo.5353554" httpUri="https://zenodo.org/record/5353554/files/figure.png" pageId="12" pageNumber="11">Fig. 43</figureCitation>
); meso,- metasternum black; abdomen with basal ventrite dark brown. Head punctures separated by a diameter or less; pronotal punctures small, separated by up to twice a diameter; elytral punctures larger than on pronotum, separated by up to three times a diameter; prosternal punctures small, separated by about a diameter; mesosternal punctures larger than on prosternum, separated by less than a diameter, metasternal punctures large, separated by a diameter or less medially, absent in lateral 1/3; abdominal ventrites 1, 2 with fine punctures separated by up to 3 times diameter medially, ventrites 2–4 finely, densely punctured medially; ventrite 5 finely, densely punctured. Head with frons parallel sided, 1.2 times as wide as eye, medially depressed. clypeal apex truncate (
<figureCitation id="13442A27FF9B6A16FCADFDCFFC08FDEB" box="[871,958,588,612]" captionStart="Figures 38–53" captionStartId="23.[192,287,1678,1700]" captionTargetBox="[205,1432,233,1600]" captionTargetId="figure-89@23.[192,1440,217,1650]" captionTargetPageId="23" captionText="Figures 38–53. Calloeneis spp.38–42) Calloeneis krista. 38) Habitus.39) Frons.40) Penis guide ventral.41) Penis guide lateral. 42) Penis. 43–48) Calloeneis roxanne. 43) Habitus. 44) Frons. 45) Penis guide ventral. 46) Penis guide lateral. 47) Penis. 48) Spermathecal capsule. 49–53) Calloeneis angelica. 49) Habitus. 50) Frons. 51) Penis guide ventral. 52) Penis guide lateral. 53) Penis." figureDoi="http://doi.org/10.5281/zenodo.5353554" httpUri="https://zenodo.org/record/5353554/files/figure.png" pageId="12" pageNumber="11">Fig. 44</figureCitation>
); eye canthus short, about half width of eye. Prosternum with lateral carina on each side curved, extended from apex of intercoxal process to lateral margin of prosternum. Epipleuron not descending externally. Postcoxal line on basal abdominal ventrite curved, extended 3/4 distance to rear margin of ventrite. Genitalia with penis guide shorter than paramere, sides parallel in basal 3/4, rounded to acute apex; paramere slender, curved, apex abruptly rounded (
<figureCitation id="13442A27FF9B6A16FEFCFD6CFE0FFC88" box="[310,441,751,775]" captionStart="Figures 38–53" captionStartId="23.[192,287,1678,1700]" captionTargetBox="[205,1432,233,1600]" captionTargetId="figure-89@23.[192,1440,217,1650]" captionTargetPageId="23" captionText="Figures 38–53. Calloeneis spp.38–42) Calloeneis krista. 38) Habitus.39) Frons.40) Penis guide ventral.41) Penis guide lateral. 42) Penis. 43–48) Calloeneis roxanne. 43) Habitus. 44) Frons. 45) Penis guide ventral. 46) Penis guide lateral. 47) Penis. 48) Spermathecal capsule. 49–53) Calloeneis angelica. 49) Habitus. 50) Frons. 51) Penis guide ventral. 52) Penis guide lateral. 53) Penis." figureDoi="http://doi.org/10.5281/zenodo.5353554" httpUri="https://zenodo.org/record/5353554/files/figure.png" pageId="12" pageNumber="11">Fig. 45, 46</figureCitation>
); penis long, slender, apex shortly attenuate, basal capsule lost (
<figureCitation id="13442A27FF9B6A16FB04FD6CFA90FC88" box="[1230,1318,751,775]" captionStart="Figures 38–53" captionStartId="23.[192,287,1678,1700]" captionTargetBox="[205,1432,233,1600]" captionTargetId="figure-89@23.[192,1440,217,1650]" captionTargetPageId="23" captionText="Figures 38–53. Calloeneis spp.38–42) Calloeneis krista. 38) Habitus.39) Frons.40) Penis guide ventral.41) Penis guide lateral. 42) Penis. 43–48) Calloeneis roxanne. 43) Habitus. 44) Frons. 45) Penis guide ventral. 46) Penis guide lateral. 47) Penis. 48) Spermathecal capsule. 49–53) Calloeneis angelica. 49) Habitus. 50) Frons. 51) Penis guide ventral. 52) Penis guide lateral. 53) Penis." figureDoi="http://doi.org/10.5281/zenodo.5353554" httpUri="https://zenodo.org/record/5353554/files/figure.png" pageId="12" pageNumber="11">Fig. 47</figureCitation>
).
</paragraph>
<paragraph id="8BC036A2FF9B6A16FF0AFC9CFE67FCD7" blockId="12.[192,1440,799,856]" pageId="12" pageNumber="11">
<emphasis id="B90BEAB0FF9B6A16FF0AFC9CFE9BFCB8" bold="true" box="[192,301,799,823]" pageId="12" pageNumber="11">Female.</emphasis>
Similar to male except genitalia with ramus bulbous, cornu strongly narrowed from base to slender apex (
<figureCitation id="13442A27FF9B6A16FEA6FCC3FE72FCD7" box="[364,452,832,856]" captionStart="Figures 38–53" captionStartId="23.[192,287,1678,1700]" captionTargetBox="[205,1432,233,1600]" captionTargetId="figure-89@23.[192,1440,217,1650]" captionTargetPageId="23" captionText="Figures 38–53. Calloeneis spp.38–42) Calloeneis krista. 38) Habitus.39) Frons.40) Penis guide ventral.41) Penis guide lateral. 42) Penis. 43–48) Calloeneis roxanne. 43) Habitus. 44) Frons. 45) Penis guide ventral. 46) Penis guide lateral. 47) Penis. 48) Spermathecal capsule. 49–53) Calloeneis angelica. 49) Habitus. 50) Frons. 51) Penis guide ventral. 52) Penis guide lateral. 53) Penis." figureDoi="http://doi.org/10.5281/zenodo.5353554" httpUri="https://zenodo.org/record/5353554/files/figure.png" pageId="12" pageNumber="11">Fig. 48</figureCitation>
).
</paragraph>
<paragraph id="8BC036A2FF9B6A16FF0AFCF3FBACFC26" blockId="12.[191,1440,880,937]" pageId="12" pageNumber="11">
<emphasis id="B90BEAB0FF9B6A16FF0AFCF3FEFAFC07" bold="true" box="[192,332,880,904]" pageId="12" pageNumber="11">Variation.</emphasis>
Length 1.5 to 2.0 mm, width
<quantity id="4C879B47FF9B6A16FD65FCF3FCE0FC07" box="[687,854,880,904]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="12" pageNumber="11" unit="mm" value="1.5" valueMax="1.7" valueMin="1.3">1.3 to 1.7 mm</quantity>
. Elytral color pattern varies from that described above to having lateral macula absent or having both maculae absent.
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF9B6A16FF0AFC42FB16FC75" pageId="12" pageNumber="11" type="materials_examined">
<paragraph id="8BC036A2FF9B6A16FF0AFC42FB16FC75" blockId="12.[190,1440,961,1018]" pageId="12" pageNumber="11">
<emphasis id="B90BEAB0FF9B6A16FF0AFC42FE31FC56" bold="true" box="[192,391,961,985]" pageId="12" pageNumber="11">Type material.</emphasis>
<materialsCitation id="3B173CFFFF9B6A16FE46FC42FBD9FC56" box="[396,1135,961,985]" collectionCode="CMNH" collectorName="Acc No." country="Brazil" location="Brazil" pageId="12" pageNumber="11" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C48800FF9B6A16FE46FC42FE4CFC56" box="[396,506,961,985]" pageId="12" pageNumber="11" type="holotype">Holotype</typeStatus>
male: Chapada,
<collectingCountry id="F3687632FF9B6A16FD09FC42FCA7FC56" box="[707,785,961,985]" name="Brazil" pageId="12" pageNumber="11">Brazil</collectingCountry>
,
<collectorName id="268A5374FF9B6A16FCD1FC42FCC0FC56" box="[795,886,961,985]" pageId="12" pageNumber="11">Acc No.</collectorName>
2966, Aug. (
<collectionCode id="ED6EAE67FF9B6A16FBC3FC42FBD0FC56" box="[1033,1126,961,985]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="12" pageNumber="11" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54C48800FF9B6A16FBB2FC41FB42FC55" box="[1144,1268,962,986]" pageId="12" pageNumber="11" type="paratype">Paratypes</typeStatus>
29, same data as
<typeStatus id="54C48800FF9B6A16FF2AFC61FEFEFC75" box="[224,328,994,1018]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
except months April, June, September, October, November. (
<collectionCode id="ED6EAE67FF9B6A16FBFDFC61FB22FC75" box="[1079,1172,994,1018]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="12" pageNumber="11" type="Museum">CMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF9B6A16FF0AFB91FC4CFBC4" pageId="12" pageNumber="11" type="discussion">
<paragraph id="8BC036A2FF9B6A16FF0AFB91FC4CFBC4" blockId="12.[192,1440,1042,1099]" pageId="12" pageNumber="11">
<emphasis id="B90BEAB0FF9B6A16FF0AFB91FEF5FBA5" bold="true" box="[192,323,1042,1066]" pageId="12" pageNumber="11">Remarks.</emphasis>
This species has a recognizable dorsal color pattern, although the pattern is somewhat variable. Otherwise male genitalia must be examined for identification.
</paragraph>
</subSubSection>
</treatment>
</document>
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