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<document id="F9827AB92DCEF0DF9A5E95963E683B04" ID-DOI="10.1649/0010-065x-67.4.587" ID-ISSN="1938-4394" ID-Zenodo-Dep="10104109" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630166095551" checkinUser="carolina" docAuthor="Kanda, Kojun" docDate="2013" docId="03CAF87A9567BD6AFF41815E1EB4FB14" docLanguage="en" docName="0010-065X-67.4.587.pdf.imf" docOrigin="The Coleopterists Bulletin 67 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-67.4.587" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Madreallecula mcclevei Kanda, new species" docType="treatment" docVersion="3" lastPageNumber="590" masterDocId="FFF380029566BD69FFED823A1A20FF9D" masterDocTitle="Madreallecula Mcclevei Kanda (Coleoptera: Tenebrionidae: Alleculinae: Alleculini), New Genus and New Species of Comb-Clawed Beetle from Arizona, USA" masterLastPageNumber="590" masterPageNumber="587" pageNumber="588" updateTime="1699628455198" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="76FBB305C5B8AE0C9A9B765B96C59AFF">Madreallecula Mcclevei Kanda (Coleoptera: Tenebrionidae: Alleculinae: Alleculini), New Genus and New Species of Comb-Clawed Beetle from Arizona, USA</mods:title>
</mods:titleInfo>
<mods:name id="287B2B27DCF6051C134D15159E6B58FE" type="personal">
<mods:role id="4A7A648AB53F83E9C97B2BAA12F1E58E">
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<mods:namePart id="F179930A8D4BE08A5A23F945F788DAB9">Kanda, Kojun</mods:namePart>
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<mods:typeOfResource id="3507C3AF5BCEA0A7B746B1CC7E195BC9">text</mods:typeOfResource>
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<mods:titleInfo id="0AC81F69F435DC5565AA44A31B3B7A4D">
<mods:title id="24E7C601960140D053E7EA93438007A6">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="D37111B560B9C76EBB9C7657F083004C">
<mods:date id="D498C74945F6CAB48A7FD3D14F0D5C67">2013</mods:date>
<mods:detail id="5D7E6BE389B0656A7FD6F1981BCB5AF4" type="pubDate">
<mods:number id="A4DA074F2F632444294828C492702DF7">2013-12-31</mods:number>
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<mods:number id="F5B989FF457AB4176D801005E9F39B3D">67</mods:number>
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<mods:number id="E5139719D718444007B58BE78086FDC9">4</mods:number>
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<mods:url id="2DFB55187E82D9A826B84C744735D12C">http://dx.doi.org/10.1649/0010-065x-67.4.587</mods:url>
</mods:location>
<mods:classification id="8BD043524C21EDB8DBE43508DC2597A1">journal article</mods:classification>
<mods:identifier id="F89D584B4F015879CF74D540BA37CCC3" type="DOI">10.1649/0010-065x-67.4.587</mods:identifier>
<mods:identifier id="CB47588E288984750B82650B6D981B31" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="8D251F4A6BB7B3F16BB5915CAF656243" type="Zenodo-Dep">10104109</mods:identifier>
</mods:mods>
<treatment id="03CAF87A9567BD6AFF41815E1EB4FB14" LSID="urn:lsid:plazi:treatment:03CAF87A9567BD6AFF41815E1EB4FB14" httpUri="http://treatment.plazi.org/id/03CAF87A9567BD6AFF41815E1EB4FB14" lastPageId="3" lastPageNumber="590" pageId="1" pageNumber="588">
<subSubSection id="C3791AE79567BD68FF41815E1848FCE7" box="[172,616,868,890]" pageId="1" pageNumber="588" type="nomenclature">
<paragraph id="8BDC496C9567BD68FF41815E1848FCE7" blockId="1.[172,616,868,890]" box="[172,616,868,890]" pageId="1" pageNumber="588">
<heading id="D094FE009567BD68FF41815E1848FCE7" box="[172,616,868,890]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="588" reason="2">
<emphasis id="B917957E9567BD68FF41815E1848FCE7" bold="true" box="[172,616,868,890]" pageId="1" pageNumber="588">
<taxonomicName id="4C6332EF9567BD68FF41815E1BC6FCE7" authority="Kanda" authorityName="Kanda" box="[172,486,868,890]" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="mcclevei" status="sp. nov.">
<emphasis id="B917957E9567BD68FF41815E1BB6FCE7" bold="true" box="[172,406,868,890]" italics="true" pageId="1" pageNumber="588">Madreallecula mcclevei</emphasis>
Kanda
</taxonomicName>
,
<taxonomicNameLabel id="A22428059567BD68FE1F815F1848FCE7" box="[498,616,869,890]" pageId="1" pageNumber="588" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3791AE79567BD68FF4281B51872F99A" pageId="1" pageNumber="588" type="materials_examined">
<paragraph id="8BDC496C9567BD68FF4281B5185FFAF5" blockId="1.[151,639,911,1729]" pageId="1" pageNumber="588">
<emphasis id="B917957E9567BD68FF4281B51B66FC39" bold="true" box="[175,326,911,932]" pageId="1" pageNumber="588">Type Material.</emphasis>
<materialsCitation id="3B0B43319567BD68FEB981B51B1AFBC3" collectionCode="OSAC" collectorName="S. McCleve" location="Galiuro Mts." pageId="1" pageNumber="588" specimenCount="2" stateProvince="Oregon" typeStatus="holotype">
<emphasis id="B917957E9567BD68FEB981B51B9BFC39" bold="true" box="[340,443,911,932]" pageId="1" pageNumber="588">
<typeStatus id="54D8F7CE9567BD68FEB981B51B9BFC39" box="[340,443,911,932]" pageId="1" pageNumber="588" type="holotype">Holotype</typeStatus>
</emphasis>
: male with three labels: “AZ: GRAHAM CO.\
<location id="8EBC1FB79567BD68FE2F8190181EFC22" LSID="urn:lsid:plazi:treatment:03CAF87A9567BD6AFF41815E1EB4FB14:8EBC1FB79567BD68FE2F8190181EFC22" box="[450,574,938,959]" name="Galiuro Mts." pageId="1" pageNumber="588" stateProvince="Oregon">Galiuro Mts.</location>
\ High Cr. 1660M\ vii-20-78 at\ light
<collectorName id="26962CBA9567BD68FE3D81FE181CFC44" box="[464,572,964,985]" pageId="1" pageNumber="588">S.McCleve</collectorName>
”; [red
<collectionCode id="ED72D1A99567BD68FF7A81E51AF8FC69" box="[151,216,991,1012]" country="USA" httpUri="http://grbio.org/cool/vn0n-jrfa" name="Oregon State Arthropod Collection" pageId="1" pageNumber="588" type="University or college">OSAC</collectionCode>
barcode label, barcode and identification number 0000529577 on top, “
<collectingRegion id="49A7878E9567BD68FE5C81C0180AFB92" box="[433,554,1018,1039]" country="United States of America" name="Oregon" pageId="1" pageNumber="588">Oregon State</collectingRegion>
\ University\
<collectionCode id="ED72D1A99567BD68FF2C862E1ADEFBB4" box="[193,254,1044,1065]" country="USA" httpUri="http://grbio.org/cool/vn0n-jrfa" name="Oregon State Arthropod Collection" pageId="1" pageNumber="588" type="University or college">OSAC</collectionCode>
” on bottom]; “
<typeStatus id="54D8F7CE9567BD68FE6C862E1BDAFBB4" box="[385,506,1044,1065]" pageId="1" pageNumber="588" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C6332EF9567BD68FE10862E1B4CFBD9" authority="Kojun Kanda" authorityName="Kojun Kanda" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="mcclevei">Madreallecula mcclevei Kojun Kanda</taxonomicName>
” [red paper, taxonomic name written by hand]
</materialsCitation>
.
<materialsCitation id="3B0B43319567BD68FEAA8673185BFAF5" collectingDate="1987-08-12" collectingDateMax="1987-08-13" collectingDateMin="1987-08-12" collectionCode="MV, PARA-, TYPE" collectorName="S. McCleve &amp; C. B. &amp; J. E. Barr" county="Santa Cruz Co." elevation="1926" location="One" municipality="One" pageId="1" pageNumber="588" specimenCount="1" stateProvince="Oregon" typeStatus="paratype">
<emphasis id="B917957E9567BD68FEAA86731B91FBC3" bold="true" box="[327,433,1097,1118]" pageId="1" pageNumber="588">
<typeStatus id="54D8F7CE9567BD68FEAA86731B8DFBC3" box="[327,429,1097,1118]" pageId="1" pageNumber="588" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D6582E59567BD68FE5786731863FBC3" box="[442,579,1097,1118]" pageId="1" pageNumber="588" type="generic">one specimen</specimenCount>
, male labeled “AZ: GRAHAM CO.\ Galiuro Mts.\ High Cr. 1660M\ vii-20-78 at\ light
<collectorName id="26962CBA9567BD68FE228644181BFB0E" box="[463,571,1150,1171]" pageId="1" pageNumber="588">S.McCleve</collectorName>
”.
<location id="8EBC1FB79567BD68FDB88644185DFB0E" LSID="urn:lsid:plazi:treatment:03CAF87A9567BD6AFF41815E1EB4FB14:8EBC1FB79567BD68FDB88644185DFB0E" box="[597,637,1150,1171]" county="Santa Cruz Co." municipality="One" name="One" pageId="1" pageNumber="588" stateProvince="Oregon">One</location>
specimen, female labeled “AZ: COCHISE CO.\ Dragoon Mts. \Sorin Cn
<quantity id="4C9BE4899567BD68FE6286891BF8FB55" box="[399,472,1203,1224]" metricMagnitude="3" metricUnit="m" metricValue="1.9259999999999997" pageId="1" pageNumber="588" unit="m" value="1926.0">
<elevation id="004EAE5F9567BD68FE6286891BF8FB55" box="[399,472,1203,1224]" metricMagnitude="3" metricUnit="m" metricValue="1.9259999999999997" pageId="1" pageNumber="588" unit="m" value="1926.0">1926m</elevation>
</quantity>
. vii-23, 24-79 at\ light
<collectorName id="26962CBA9567BD68FF2A86F41B0DFB7E" box="[199,301,1230,1251]" pageId="1" pageNumber="588">S.McCleve</collectorName>
”.
<collectingMunicipality id="6BB8D3169567BD68FEA986F41B4BFB7E" box="[324,363,1230,1251]" pageId="1" pageNumber="588">One</collectingMunicipality>
specimen, male labeled “AZ:
<collectingCounty id="62BD31E09567BD68FF7A86D31B08FB63" box="[151,296,1257,1278]" pageId="1" pageNumber="588">Santa Cruz Co.</collectingCounty>
\ Coronado N. F., Santa\ Rita Mtns, Madera\ Cyn.,
<date id="FFDD6FAC9567BD68FECA87391B9FFA85" box="[295,447,1283,1304]" pageId="1" pageNumber="588" value="1987-08-12" valueMax="1987-08-13" valueMin="1987-08-12">
<collectingDate id="EF9996449567BD68FECA87391B9FFA85" box="[295,447,1283,1304]" pageId="1" pageNumber="588" value="1987-08-12" valueMax="1987-08-13" valueMin="1987-08-12">12-13-VIII-1987</collectingDate>
</date>
\
<collectorName id="26962CBA9567BD68FE228739185AFA85" box="[463,634,1283,1304]" pageId="1" pageNumber="588">C. B.&amp; J. E. Barr</collectorName>
,
<collectionCode id="ED72D1A99567BD68FF7A87241A9CFAAE" box="[151,188,1310,1331]" country="0" name="University of Montana Museum" pageId="1" pageNumber="588">MV</collectionCode>
/BL”. All
<typeStatus id="54D8F7CE9567BD68FEF487241B51FAAE" box="[281,369,1310,1331]" pageId="1" pageNumber="588" type="paratype">paratypes</typeStatus>
with a second label, “
<collectionCode id="ED72D1A99567BD68FDD787241AF4FAD0" pageId="1" pageNumber="588">PARA- TYPE</collectionCode>
<taxonomicName id="4C6332EF9567BD68FF0F87021853FAD0" authority="Kojun Kanda" authorityName="Kojun Kanda" box="[226,627,1336,1357]" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="mcclevei">Madreallecula mcclevei Kojun Kanda</taxonomicName>
” [yellow paper, taxonomic name written by hand]
</materialsCitation>
.
</paragraph>
<caption id="DF1C19E49567BD68FD28862B1981FBA1" ID-DOI="http://doi.org/10.5281/zenodo.10104111" ID-Zenodo-Dep="10104111" httpUri="https://zenodo.org/record/10104111/files/figure.png" pageId="1" pageNumber="588" startId="1.[709,741,1041,1060]" targetBox="[685,1171,194,1017]" targetPageId="1">
<paragraph id="8BDC496C9567BD68FD28862B1981FBA1" blockId="1.[685,1171,1041,1084]" pageId="1" pageNumber="588">
<emphasis id="B917957E9567BD68FD28862B1920FBB9" bold="true" box="[709,768,1041,1060]" pageId="1" pageNumber="588">Fig. 1.</emphasis>
Dorsal habitus of a paratype of
<taxonomicName id="4C6332EF9567BD68FBF1862B18D4FBA6" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="mcclevei">
<emphasis id="B917957E9567BD68FBF1862B18D4FBA6" italics="true" pageId="1" pageNumber="588">Madreallecula mcclevei</emphasis>
</taxonomicName>
. Scale bar = 2 mm.
</paragraph>
</caption>
<paragraph id="8BDC496C9567BD68FF4287571872F99A" blockId="1.[151,639,911,1729]" pageId="1" pageNumber="588">
<materialsCitation id="3B0B43319567BD68FF428757186EF99A" collectionCode="OSAC, R" collectorName="S. McCleve &amp; University &amp; Insect Collection &amp; C. B. and J. E. Barr &amp; L. Aalbu" location="Arthropod Collection" municipality="The" pageId="1" pageNumber="588" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
<collectingMunicipality id="6BB8D3169567BD68FF4287571AF5FA1F" box="[175,213,1389,1410]" pageId="1" pageNumber="588">The</collectingMunicipality>
specimens are deposited in the following collections:
<typeStatus id="54D8F7CE9567BD68FEE587B21B79FA00" box="[264,345,1416,1437]" pageId="1" pageNumber="588" type="holotype">holotype</typeStatus>
in the
<collectingRegion id="49A7878E9567BD68FE7187B21836FA00" box="[412,534,1416,1437]" country="United States of America" name="Oregon" pageId="1" pageNumber="588">Oregon State</collectingRegion>
<location id="8EBC1FB79567BD68FDF087B21AD6FA25" LSID="urn:lsid:plazi:treatment:03CAF87A9567BD6AFF41815E1EB4FB14:8EBC1FB79567BD68FDF087B21AD6FA25" municipality="The" name="Arthropod Collection" pageId="1" pageNumber="588" stateProvince="Oregon">Arthropod Collection</location>
(
<collectionCode id="ED72D1A99567BD68FEEE87991B61FA25" box="[259,321,1443,1464]" country="USA" httpUri="http://grbio.org/cool/vn0n-jrfa" name="Oregon State Arthropod Collection" pageId="1" pageNumber="588" type="University or college">OSAC</collectionCode>
_0000529577),
<specimenCount id="9D6582E59567BD68FE3987991875FA25" box="[468,597,1443,1464]" pageId="1" pageNumber="588" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
collected by
<collectorName id="26962CBA9567BD68FF1B87871B47FA4F" box="[246,359,1469,1490]" pageId="1" pageNumber="588">S. McCleve</collectorName>
in the
<collectorName id="26962CBA9567BD68FE408787182FFA4F" box="[429,527,1469,1490]" pageId="1" pageNumber="588">University</collectorName>
of
<collectingRegion id="49A7878E9567BD68FDDE8787185EFA4F" box="[563,638,1469,1490]" country="United States of America" name="Arizona" pageId="1" pageNumber="588">Arizona</collectingRegion>
<collectorName id="26962CBA9567BD68FF7A87E21B1FFA70" box="[151,319,1496,1517]" pageId="1" pageNumber="588">Insect Collection</collectorName>
, the
<typeStatus id="54D8F7CE9567BD68FE9D87E21BE2FA70" box="[368,450,1496,1517]" pageId="1" pageNumber="588" type="paratype">paratype</typeStatus>
collected by
<collectorName id="26962CBA9567BD68FDA487E21B3AF99A" pageId="1" pageNumber="588">C. B. and J. E. Barr</collectorName>
in the collection of
<collectionCode id="ED72D1A99567BD68FE3787C81BCDF99A" box="[474,493,1522,1543]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="588">R</collectionCode>
.
<collectorName id="26962CBA9567BD68FE1A87C8186EF99A" box="[503,590,1522,1543]" pageId="1" pageNumber="588">L. Aalbu</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3791AE79567BD6AFF4284371B4BFAA2" lastPageId="3" lastPageNumber="590" pageId="1" pageNumber="588" type="description">
<paragraph id="8BDC496C9567BD6AFF4284371847FB18" blockId="1.[151,639,911,1729]" lastBlockId="3.[151,639,817,1555]" lastPageId="3" lastPageNumber="590" pageId="1" pageNumber="588">
<emphasis id="B917957E9567BD68FF4284371B8BF9BF" bold="true" box="[175,427,1549,1570]" pageId="1" pageNumber="588">
Description of
<typeStatus id="54D8F7CE9567BD68FEA584371B87F9BF" box="[328,423,1549,1570]" pageId="1" pageNumber="588" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Length from anterior margin of pronotum to apex of elytra:
<quantity id="4C9BE4899567BD68FF7A84781AC7F9CA" box="[151,231,1602,1623]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="588" unit="mm" value="6.8">6.8 mm</quantity>
; width of posterior margin of pronotum:
<quantity id="4C9BE4899567BD68FF7A84671ACAF9EF" box="[151,234,1629,1650]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="588" unit="mm" value="2.5">2.5 mm</quantity>
. Integument rufo-testaceous, polished. Body clothed in semi-erect, golden setae.
<emphasis id="B917957E9567BD68FDAD844D1857F911" bold="true" box="[576,631,1655,1676]" pageId="1" pageNumber="588">Head</emphasis>
: Small, at widest half the width of basal margin of pronotum. Dorsal surface sparsely covered in shallow setate punctures. Ventral surface of head much less hirsute and punctate than dorsal surface. Gula smooth, impunctate. Eyes prominent, separated dorsally by 1/2 the width of one eye. Frontoclypeal suture complete. Antennae with terminal 8 segments moniliform, 3
<superScript id="7C16E4249567BD68FBEC86D91E30FB6C" attach="left" box="[1025,1040,1251,1265]" fontSize="6" pageId="1" pageNumber="588">rd</superScript>
antennomere as long as scape and pedicel combined, 4
<superScript id="7C16E4249567BD68FBB486C41E47FA91" attach="left" box="[1113,1127,1278,1292]" fontSize="6" pageId="1" pageNumber="588">th</superScript>
and 11
<superScript id="7C16E4249567BD68FD28872218F3FABB" attach="left" box="[709,723,1304,1318]" fontSize="6" pageId="1" pageNumber="588">th</superScript>
antennomere slightly longer than wide, width of antennomeres 5 to 10 equal to or slightly greater than length (
<figureCitation id="135855E99567BD68FCC18769194AFAF5" box="[812,874,1363,1384]" captionStart="Figs" captionStartId="2.[197,238,1686,1705]" captionTargetBox="[214,1101,969,1653]" captionTargetId="figure-20@2.[188,1178,959,1664]" captionTargetPageId="2" captionText="Figs. 35. Antennae of Alleculina. 3) Madreallecula mcclevei; 4) Hymenorus sp.; 5) Knausia crassicornis. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10104115" httpUri="https://zenodo.org/record/10104115/files/figure.png" pageId="1" pageNumber="588">Fig. 3</figureCitation>
). Terminal segment of maxillary palpi large, broadly triangular.
<emphasis id="B917957E9567BD68FBD187541EACFA1E" bold="true" box="[1084,1164,1390,1411]" pageId="1" pageNumber="588">Thorax</emphasis>
: Pronotum convex anteriorly and flat posteriorly, posterior margin slightly produced medially, anterior margin rounded. Posterior angles obtuse and rounded. Dorsal surface of pronotum coarsely punctate, ventral surface slightly rugose. Prosternum flat, declivous. Distance between anterior margin of procoxal cavity and anterior margin of prosternum shorter than length of prosternal process. Mesosternum with slight keel antero-medially, coarsely punctate. Mesepisternum smooth towards medial line of body. Metasternum coarsely punctate, more densely punctate medially. Metepisternum coarsely punctate.
<emphasis id="B917957E9565BD6AFF15810B1B07FCDB" bold="true" box="[248,295,817,838]" pageId="3" pageNumber="590">Legs</emphasis>
: Femora barely extending beyond lateral margin of elytra when held perpendicular to midline of body. Pro- and mesotarsi with membranous lobes on penultimate and antepenultimate tarsomeres. Metatarsi with membranous lobe on penultimate tarsomere only. Protarsal claws with 11 teeth, mesotarsal claws with 10 teeth, and metatarsal claws with 8 teeth.
<emphasis id="B917957E9565BD6AFE2281D1182FFB9D" bold="true" box="[463,527,1003,1024]" pageId="3" pageNumber="590">Elytra</emphasis>
: Distinctly striate, strial punctures becoming shallower distally. Epipleura slightly tuberculate.
<emphasis id="B917957E9565BD6AFDFB861A1856FBA8" bold="true" box="[534,630,1056,1077]" pageId="3" pageNumber="590">Genitalia</emphasis>
: Aedeagus elongate with parameres narrow, slightly acuminate and apically rounded, without setae, about 1/4 the length of the phallobase (
<figureCitation id="135855E99565BD6AFDF2864A187CFB18" box="[543,604,1136,1157]" captionStart="Fig" captionStartId="3.[175,207,698,717]" captionTargetBox="[326,462,190,675]" captionTargetId="figure-423@3.[151,638,183,683]" captionTargetPageId="3" captionText="Fig. 6. Ventral view of the aedeagus of Madreallecula mcclevei (parameres and portion of the phallobase). Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10104117" httpUri="https://zenodo.org/record/10104117/files/figure.png" pageId="3" pageNumber="590">Fig. 6</figureCitation>
).
</paragraph>
<caption id="DF1C19E49564BD6BFF2881441E63FC0C" ID-DOI="http://doi.org/10.5281/zenodo.10104113" ID-Zenodo-Dep="10104113" box="[197,1091,894,913]" httpUri="https://zenodo.org/record/10104113/files/figure.png" pageId="2" pageNumber="589" startId="2.[197,229,894,913]" targetBox="[183,1183,194,870]" targetPageId="2">
<paragraph id="8BDC496C9564BD6BFF2881441E63FC0C" blockId="2.[197,1091,894,913]" box="[197,1091,894,913]" pageId="2" pageNumber="589">
<emphasis id="B917957E9564BD6BFF2881441B21FC0C" bold="true" box="[197,257,894,913]" pageId="2" pageNumber="589">Fig. 2.</emphasis>
Head of a paratype of
<taxonomicName id="4C6332EF9564BD6BFE3681441885FC0D" box="[475,677,894,912]" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="mcclevei">
<emphasis id="B917957E9564BD6BFE3681441885FC0D" box="[475,677,894,912]" italics="true" pageId="2" pageNumber="589">Madreallecula mcclevei</emphasis>
</taxonomicName>
with clypeus emphasized. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="DF1C19E49564BD6BFF2884AC1B79F95C" ID-DOI="http://doi.org/10.5281/zenodo.10104115" ID-Zenodo-Dep="10104115" httpUri="https://zenodo.org/record/10104115/files/figure.png" pageId="2" pageNumber="589" startId="2.[197,238,1686,1705]" targetBox="[214,1101,969,1653]" targetPageId="2">
<paragraph id="8BDC496C9564BD6BFF2884AC1E89F934" blockId="2.[173,1193,1686,1729]" box="[197,1193,1686,1705]" pageId="2" pageNumber="589">
<emphasis id="B917957E9564BD6BFF2884AC1B03F934" bold="true" box="[197,291,1686,1705]" pageId="2" pageNumber="589">Figs. 35.</emphasis>
Antennae of
<taxonomicName id="4C6332EF9564BD6BFE5C84AC182DF934" box="[433,525,1686,1705]" class="Insecta" family="Tenebrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="subTribe" subTribe="Alleculina">Alleculina</taxonomicName>
.
<emphasis id="B917957E9564BD6BFDF784AC1805F934" bold="true" box="[538,549,1686,1705]" pageId="2" pageNumber="589">3</emphasis>
)
<taxonomicName id="4C6332EF9564BD6BFDDA84AC1929F935" box="[567,777,1686,1704]" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="mcclevei">
<emphasis id="B917957E9564BD6BFDDA84AC1929F935" box="[567,777,1686,1704]" italics="true" pageId="2" pageNumber="589">Madreallecula mcclevei</emphasis>
</taxonomicName>
;
<emphasis id="B917957E9564BD6BFCF484AC1904F934" bold="true" box="[793,804,1686,1705]" pageId="2" pageNumber="589">4</emphasis>
)
<taxonomicName id="4C6332EF9564BD6BFCD884AC199EF934" box="[821,958,1686,1705]" class="Insecta" family="Tenebrionidae" genus="Hymenorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B917957E9564BD6BFCD884AC19B8F935" box="[821,920,1686,1704]" italics="true" pageId="2" pageNumber="589">Hymenorus</emphasis>
sp.
</taxonomicName>
;
<emphasis id="B917957E9564BD6BFC2184AC19F7F934" bold="true" box="[972,983,1686,1705]" pageId="2" pageNumber="589">5</emphasis>
)
<taxonomicName id="4C6332EF9564BD6BFC0584AC1E84F935" authorityName="Fall" authorityYear="1931" box="[1000,1188,1686,1704]" class="Insecta" family="Tenebrionidae" genus="Knausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="B917957E9564BD6BFC0584AC1E84F935" box="[1000,1188,1686,1704]" italics="true" pageId="2" pageNumber="589">Knausia crassicornis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDC496C9564BD6BFF4084941B79F95C" blockId="2.[173,1193,1686,1729]" box="[173,345,1710,1729]" pageId="2" pageNumber="589">Scale bar = 1 mm.</paragraph>
</caption>
<caption id="DF1C19E49565BD6AFF4280801B3CFD60" ID-DOI="http://doi.org/10.5281/zenodo.10104117" ID-Zenodo-Dep="10104117" httpUri="https://zenodo.org/record/10104117/files/figure.png" pageId="3" pageNumber="590" startId="3.[175,207,698,717]" targetBox="[326,462,190,675]" targetPageId="3">
<paragraph id="8BDC496C9565BD6AFF4280801B3CFD60" blockId="3.[151,638,698,765]" pageId="3" pageNumber="590">
<emphasis id="B917957E9565BD6AFF4280801ACBFD50" bold="true" box="[175,235,698,717]" pageId="3" pageNumber="590">Fig. 6.</emphasis>
Ventral view of the aedeagus of
<taxonomicName id="4C6332EF9565BD6AFDE580811AFEFD79" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="mcclevei">
<emphasis id="B917957E9565BD6AFDE580811AFEFD79" italics="true" pageId="3" pageNumber="590">Madreallecula mcclevei</emphasis>
</taxonomicName>
(parameres and portion of the phallobase). Scale bar = 0.25 mm.
</paragraph>
</caption>
<caption id="DF1C19E49565BD6AFD28812019DEFCC0" ID-DOI="http://doi.org/10.5281/zenodo.10104119" ID-Zenodo-Dep="10104119" httpUri="https://zenodo.org/record/10104119/files/figure.png" pageId="3" pageNumber="590" startId="3.[709,741,794,813]" targetBox="[685,1171,194,769]" targetPageId="3">
<paragraph id="8BDC496C9565BD6AFD28812019DEFCC0" blockId="3.[685,1173,794,861]" pageId="3" pageNumber="590">
<emphasis id="B917957E9565BD6AFD2881201923FCB0" bold="true" box="[709,771,794,813]" pageId="3" pageNumber="590">Fig. 7.</emphasis>
Map of Arizona showing known localities for
<emphasis id="B917957E9565BD6AFD3C81081995FCD9" box="[721,949,818,836]" italics="true" pageId="3" pageNumber="590">
<taxonomicName id="4C6332EF9565BD6AFD3C81081990FCD9" box="[721,944,818,836]" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="mcclevei">Madreallecula mcclevei</taxonomicName>
.
</emphasis>
Scale bar = 100 miles. (Map courtesy of Digital Vector Maps).
</paragraph>
</caption>
<paragraph id="8BDC496C9565BD6AFF4286B01B4BFAA2" blockId="3.[151,639,817,1555]" pageId="3" pageNumber="590">
<emphasis id="B917957E9565BD6AFF4286B01B34FB02" bold="true" box="[175,276,1162,1183]" pageId="3" pageNumber="590">Variation.</emphasis>
No external morphological differences are apparent between the male and female specimens. Length from anterior margin of pronotum to apex of elytra is
<quantity id="4C9BE4899565BD6AFE4A86E01839FB72" box="[423,537,1241,1263]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="6.9" metricValueMin="6.1" pageId="3" pageNumber="590" unit="mm" value="6.5" valueMax="6.9" valueMin="6.1">6.16.9 mm</quantity>
, and width of posterior margin of pronotum is
<quantity id="4C9BE4899565BD6AFDEC86CE185BFA94" box="[513,635,1268,1290]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="3" pageNumber="590" unit="mm" value="2.5" valueMax="2.6" valueMin="2.4">2.42.6 mm</quantity>
. Mesotarsal claws have 1011 teeth, metatarsal claws have 89 teeth.
</paragraph>
</subSubSection>
<subSubSection id="C3791AE79565BD6AFF42877E1B74FA59" pageId="3" pageNumber="590" type="distribution">
<paragraph id="8BDC496C9565BD6AFF42877E1B74FA59" blockId="3.[151,639,817,1555]" pageId="3" pageNumber="590">
<emphasis id="B917957E9565BD6AFF42877E1B12FAC4" bold="true" box="[175,306,1348,1369]" pageId="3" pageNumber="590">Distribution.</emphasis>
All known specimens were collected in the Madrean Sky Islands in Southeast
<collectingRegion id="49A7878E9565BD6AFF7A87431AC7FA13" box="[151,231,1401,1422]" country="United States of America" name="Arizona" pageId="3" pageNumber="590">Arizona</collectingRegion>
. The species is known to occur specifically in the Galiuro, Dragoon, and Santa Rita mountains (
<figureCitation id="135855E99565BD6AFEE687951B68FA59" box="[267,328,1455,1476]" captionStart="Fig" captionStartId="3.[709,741,794,813]" captionTargetBox="[685,1171,194,769]" captionTargetId="figure-446@3.[685,1172,193,770]" captionTargetPageId="3" captionText="Fig. 7. Map of Arizona showing known localities for Madreallecula mcclevei. Scale bar = 100 miles. (Map courtesy of Digital Vector Maps)." figureDoi="http://doi.org/10.5281/zenodo.10104119" httpUri="https://zenodo.org/record/10104119/files/figure.png" pageId="3" pageNumber="590">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3791AE79565BD6AFF4287F31EB4FB14" pageId="3" pageNumber="590" type="etymology">
<paragraph id="8BDC496C9565BD6AFF4287F31BD7F98E" blockId="3.[151,639,817,1555]" pageId="3" pageNumber="590">
<emphasis id="B917957E9565BD6AFF4287F31B02FA43" bold="true" box="[175,290,1481,1502]" pageId="3" pageNumber="590">Etymology.</emphasis>
This species is named in honor of Scott McCleve who collected the
<typeStatus id="54D8F7CE9565BD6AFE1487DE1870FA64" box="[505,592,1508,1529]" pageId="3" pageNumber="590" type="holotype">holotype</typeStatus>
and two of the
<typeStatus id="54D8F7CE9565BD6AFEE987C41B7FF98E" box="[260,351,1534,1555]" pageId="3" pageNumber="590" type="paratype">paratypes</typeStatus>
of this species.
</paragraph>
<paragraph id="8BDC496C9565BD6AFF4284091840F9FE" blockId="3.[151,638,1587,1635]" pageId="3" pageNumber="590">
The following modification to key AA of Aalbu
<emphasis id="B917957E9565BD6AFF7A84741AEAF9FE" box="[151,202,1614,1635]" italics="true" pageId="3" pageNumber="590">et al.</emphasis>
(2002) will accommodate
<taxonomicName id="4C6332EF9565BD6AFE3D8474187AF9FE" authorityName="Kanda" authorityYear="2013" box="[464,602,1614,1635]" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B917957E9565BD6AFE3D8474187AF9FE" box="[464,602,1614,1635]" italics="true" pageId="3" pageNumber="590">Madreallecula</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph id="8BDC496C9565BD6AFF7A844D185DF95C" blockId="3.[151,638,1655,1729]" pageId="3" pageNumber="590">
2(1). Wings greatly reduced or absent, much shorter than elytra; one species,
<collectingRegion id="49A7878E9565BD6AFDC284A8185EF93A" box="[559,638,1682,1703]" country="United States of America" name="Arizona" pageId="3" pageNumber="590">Arizona</collectingRegion>
...................................................
<taxonomicName id="4C6332EF9565BD6AFDC18491185DF95C" authorityName="Champion" authorityYear="1888" box="[556,637,1707,1729]" class="Insecta" family="Tenebrionidae" genus="Phedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B917957E9565BD6AFDC18491185DF95C" bold="true" box="[556,637,1707,1729]" italics="true" pageId="3" pageNumber="590">Phedius</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BDC496C9565BD6AFD4081B51EB4FC22" blockId="3.[685,1172,911,1161]" pageId="3" pageNumber="590">–—. Wings fully developed, when unfolded distinctly longer than elytra .......................... 2a</paragraph>
<paragraph id="8BDC496C9565BD6AFD4081F61EB4FB8A" blockId="3.[685,1172,911,1161]" pageId="3" pageNumber="590">
2a(2). Antennae moniliform, head declivous, clypeus with front angles projecting laterally; one species,
<collectingRegion id="49A7878E9565BD6AFC87863B1994FB8B" box="[874,948,1025,1046]" country="United States of America" name="Arizona" pageId="3" pageNumber="590">Arizona</collectingRegion>
...............
<taxonomicName id="4C6332EF9565BD6AFBE6863B1EB4FB8A" authorityName="Kanda" authorityYear="2013" box="[1035,1172,1025,1047]" class="Insecta" family="Tenebrionidae" genus="Madreallecula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B917957E9565BD6AFBE6863B1EB4FB8A" bold="true" box="[1035,1172,1025,1047]" italics="true" pageId="3" pageNumber="590">Madreallecula</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BDC496C9565BD6AFD40861E1EB4FB14" blockId="3.[685,1172,911,1161]" pageId="3" pageNumber="590">
–—. Antennae serrate with antennomeres 410 strongly compressed (
<taxonomicName id="4C6332EF9565BD6AFC0E86051E18FBC9" authorityName="Fall" authorityYear="1931" box="[995,1080,1087,1108]" class="Insecta" family="Tenebrionidae" genus="Knausia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B917957E9565BD6AFC0E86051E18FBC9" box="[995,1080,1087,1108]" italics="true" pageId="3" pageNumber="590">Knausia</emphasis>
</taxonomicName>
) or filiform, clypeus with lateral margins straight or converging .............................................. 3
</paragraph>
</subSubSection>
</treatment>
</document>