267 lines
34 KiB
XML
267 lines
34 KiB
XML
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<mods:title id="0C7AC40B57D776EF8948F6A8403377FC">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group</mods:title>
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<mods:namePart id="BB88CE8F619D09952FDE67688A16A976">Shavrin, Alexey V.</mods:namePart>
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<mods:namePart id="21ED60992B42FFD1B014BDC9ED37F810">Smetana, Aleš</mods:namePart>
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<mods:date id="891FD31DAD568B32783636030F637C33">2018</mods:date>
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<treatment id="03EB87F8F15FFF9AFF31F91278664BAA" ID-DOI="http://doi.org/10.5281/zenodo.6495060" ID-GBIF-Taxon="154496012" ID-Zenodo-Dep="6495060" LSID="urn:lsid:plazi:treatment:03EB87F8F15FFF9AFF31F91278664BAA" httpUri="http://treatment.plazi.org/id/03EB87F8F15FFF9AFF31F91278664BAA" lastPageId="47" lastPageNumber="498" pageId="44" pageNumber="495">
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<subSubSection id="C3586565F15FFF99FF31F9127ED34F17" box="[151,715,1790,1817]" pageId="44" pageNumber="495" type="nomenclature">
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<paragraph id="8BFD36EEF15FFF99FF31F9127ED34F17" blockId="44.[151,715,1790,1850]" box="[151,715,1790,1817]" pageId="44" pageNumber="495">
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<heading id="D0B58182F15FFF99FF31F9127ED34F17" bold="true" box="[151,715,1790,1817]" fontSize="11" level="1" pageId="44" pageNumber="495" reason="1">
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<emphasis id="B936EAFCF15FFF99FF31F9127ED34F17" bold="true" box="[151,715,1790,1817]" pageId="44" pageNumber="495">
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<taxonomicName id="4C424D6DF15FFF99FF31F9127E5C4F17" ID-CoL="5VCBF" authority="(Coiffait, 1977)" authorityName="Shavrin & Smetana" authorityYear="2018" baseAuthorityName="Coiffait" baseAuthorityYear="1977" box="[151,580,1790,1817]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="alticola" status="comb. nov.">
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<emphasis id="B936EAFCF15FFF99FF31F9127D984F16" bold="true" box="[151,384,1790,1816]" italics="true" pageId="44" pageNumber="495">Anthobium alticola</emphasis>
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(
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<bibRefCitation id="EFD34B1FF15FFF99FE29F9137E244F17" author="Coiffait, H." box="[399,572,1791,1817]" pageId="44" pageNumber="495" pagination="243 - 275" refId="ref35437" refString="Coiffait, H. (1977) Staphilinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112 (1976), 243 - 275." type="journal article" year="1977">Coiffait, 1977</bibRefCitation>
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||
)
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||
</taxonomicName>
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<taxonomicNameLabel id="A2055787F15FFF99FDEAF9137ED34F17" box="[588,715,1791,1817]" pageId="44" pageNumber="495" rank="species">comb.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3586565F15FFF99FF31F8CD7D884F34" box="[151,400,1825,1850]" pageId="44" pageNumber="495" type="description">
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<paragraph id="8BFD36EEF15FFF99FF31F8CD7D884F34" blockId="44.[151,715,1790,1850]" box="[151,400,1825,1850]" pageId="44" pageNumber="495">
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||
(
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<figureCitation id="13792A6BF15FFF99FF39F8CD7CE44F34" box="[159,252,1825,1850]" captionStart="FIGURES 94–95" captionStartId="45.[151,264,1980,2003]" captionTargetBox="[229,1383,1092,1951]" captionTargetId="figure@45.[151,1437,1087,1958]" captionTargetPageId="45" captionText="FIGURES 94–95. Habitus of Anthobium nigrum group: 94—A. alticola (NEPAL: Annapurna Mts.) and 95—A. flaveolum (paratype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607378/files/figure.png" pageId="44" pageNumber="495">Figs. 94</figureCitation>
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,
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<figureCitation id="13792A6BF15FFF99FEAEF8CD7D494F34" box="[264,337,1825,1850]" captionStart="FIGURES 96–99" captionStartId="46.[151,264,1972,1995]" captionTargetBox="[325,1319,213,1913]" captionTargetId="figure@46.[228,1365,193,1952]" captionTargetPageId="46" captionText="FIGURES 96–99. Aedeagus of Anthobium alticola (96–97: NEPAL: Annapurna Mts.) and A. flaveolum (98–99: holotype): 96, 98—parameral view, 97, 99—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607380/files/figure.png" pageId="44" pageNumber="495">96–97</figureCitation>
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,
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<figureCitation id="13792A6BF15FFF99FEFBF8CE7D904F34" box="[349,392,1826,1850]" captionStart="FIGURE 100" captionStartId="48.[151,250,1958,1980]" captionTargetBox="[178,1399,1201,1918]" captionTargetId="figure@48.[178,1399,1201,1918]" captionTargetPageId="48" captionText="FIGURE 100. Distribution of Anthobium alticola (black circles) and A. flaveolum (black square) in the Himalayan Region." httpUri="https://zenodo.org/record/2607384/files/figure.png" pageId="44" pageNumber="495">100</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3586565F15FFF99FF31F88578524F90" pageId="44" pageNumber="495" type="reference_group">
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<paragraph id="8BFD36EEF15FFF99FF31F8857E074F71" blockId="44.[151,1098,1897,1950]" box="[151,543,1897,1919]" pageId="44" pageNumber="495">
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<treatmentCitationGroup id="AB5211C0F15FFF99FF31F8857E074F71" box="[151,543,1897,1919]" pageId="44" pageNumber="495">
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<taxonomicName id="4C424D6DF15FFF99FF31F8857E074F71" ID-CoL="34GNG" authority="Coiffait, 1977: 265" authorityName="Coiffait" authorityPageNumber="265" authorityYear="1977" box="[151,543,1897,1919]" class="Insecta" family="Staphylinidae" genus="Deliphrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="alticola">
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<emphasis id="B936EAFCF15FFF99FF31F8857D404F71" box="[151,344,1897,1919]" italics="true" pageId="44" pageNumber="495">Deliphrum alticola</emphasis>
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<treatmentCitation id="0AE310FFF15FFF99FEF8F8857E074F71" author="Coiffait, H." box="[350,543,1897,1919]" page="265" pageId="44" pageNumber="495" year="1977">
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<bibRefCitation id="EFD34B1FF15FFF99FEF8F8857DE84F71" author="Coiffait, H." box="[350,496,1897,1919]" pageId="44" pageNumber="495" pagination="243 - 275" refId="ref35437" refString="Coiffait, H. (1977) Staphilinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112 (1976), 243 - 275." type="journal article" year="1977">Coiffait, 1977</bibRefCitation>
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: 265
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</treatmentCitation>
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</taxonomicName>
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BFD36EEF15FFF99FF31F86478524F90" blockId="44.[151,1098,1897,1950]" box="[151,1098,1928,1950]" pageId="44" pageNumber="495">
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<treatmentCitationGroup id="AB5211C0F15FFF99FF31F86478524F90" box="[151,1098,1928,1950]" pageId="44" pageNumber="495">
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<taxonomicName id="4C424D6DF15FFF99FF31F8647E344F90" ID-CoL="34GNG" authority=": Herman, 2001: 281" authorityName="Herman" authorityPageNumber="281" authorityYear="2001" box="[151,556,1928,1950]" class="Insecta" family="Staphylinidae" genus="Deliphrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="alticola">
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<emphasis id="B936EAFCF15FFF99FF31F8647D404F90" box="[151,344,1928,1950]" italics="true" pageId="44" pageNumber="495">Deliphrum alticola</emphasis>
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: Herman, 2001: 281
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</taxonomicName>
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;
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<treatmentCitation id="0AE310FFF15FFF99FD91F8647F1C4F90" author="Smetana, A." box="[567,772,1928,1950]" page="242" pageId="44" pageNumber="495" year="2004">
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||
<bibRefCitation id="EFD34B1FF15FFF99FD91F8647ECA4F90" author="Smetana, A." box="[567,722,1928,1950]" pageId="44" pageNumber="495" pagination="268" refId="ref40333" refString="Smetana, A. (2004) Staphylinidae, subfamilies Omaliinae-Dasycerinae, Phloeocharinae-Apateticinae, Piestinae-Staphylininae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. II. Hydrophiloidea-Histeroidea- Staphylinoidea. Appolo Books, Stenstrup, pp. 237 ‾ 268." type="journal article" year="2004">Smetana, 2004</bibRefCitation>
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||
: 242
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="0AE310FFF15FFF99FCA8F86478524F90" author="Schulke, M. & Smetana, A." box="[782,1098,1928,1950]" page="312" pageId="44" pageNumber="495" year="2015">
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||
<bibRefCitation id="EFD34B1FF15FFF99FCA8F86478024F90" author="Schulke, M. & Smetana, A." box="[782,1050,1928,1950]" pageId="44" pageNumber="495" pagination="304 - 1134" refId="ref39989" refString="Schulke, M. & Smetana, A. (2015) Staphylinidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea-Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book chapter" year="2015">Schülke & Smetana, 2015</bibRefCitation>
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||
: 312
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||
</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3586565F15FFF98FF31F8267DBC494E" lastPageId="45" lastPageNumber="496" pageId="44" pageNumber="495" type="materials_examined">
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<paragraph id="8BFD36EEF15FFF99FF31F82679844FED" blockId="44.[151,1436,1994,2019]" box="[151,1436,1994,2019]" pageId="44" pageNumber="495">
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<emphasis id="B936EAFCF15FFF99FF31F8267DD94FED" bold="true" box="[151,449,1994,2019]" pageId="44" pageNumber="495">Type material examined:</emphasis>
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<typeStatus id="54F9884CF15FFF99FE61F8267E374FED" box="[455,559,1994,2019]" pageId="44" pageNumber="495" type="holotype">Holotype</typeStatus>
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by monotypy of
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<taxonomicName id="4C424D6DF15FFF99FD57F82778724FED" authority="Coiffait, 1977" authorityName="Coiffait" authorityYear="1977" box="[753,1130,1994,2019]" class="Insecta" family="Staphylinidae" genus="Deliphrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="alticola">
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<emphasis id="B936EAFCF15FFF99FD57F8277FDF4FED" box="[753,967,1995,2019]" italics="true" pageId="44" pageNumber="495">Deliphrum alticola</emphasis>
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<bibRefCitation id="EFD34B1FF15FFF99FC68F82678724FED" author="Coiffait, H." box="[974,1130,1994,2019]" pageId="44" pageNumber="495" pagination="243 - 275" refId="ref35437" refString="Coiffait, H. (1977) Staphilinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112 (1976), 243 - 275." type="journal article" year="1977">Coiffait, 1977</bibRefCitation>
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</taxonomicName>
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<specimenCount id="9D44FD67F15FFF99FBD7F82778904FEC" box="[1137,1160,1995,2018]" pageId="44" pageNumber="495" type="female">♀</specimenCount>
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: ‘Umg. Mahidoela-Pass |
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</paragraph>
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<paragraph id="8BFD36EEF15EFF98FF31FF7B7F8248F6" blockId="45.[151,1437,151,1040]" pageId="45" pageNumber="496">
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<quantity id="4CBA9B0BF15EFF98FF31FF7B7CF048BE" box="[151,232,151,176]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="45" pageNumber="496" unit="m" value="5000.0">5000m</quantity>
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,b.Maharigaon’ <rectangular label, printed>, ‘Gebiet von Jumla | Westnepal, lg.H. Franz’ <rectangular label, printed>, ‘TYPE’ <red rectangular label, printed>, ‘
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<taxonomicName id="4C424D6DF15EFF98FD46FF507F4648DB" authorityName="Erichson" authorityYear="1839" box="[736,862,188,213]" class="Insecta" family="Staphylinidae" genus="Deliphrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="498" phylum="Arthropoda" rank="genus">Deliphrum</taxonomicName>
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||
|
|
||
<taxonomicName id="4C424D6DF15EFF98FCD2FF50781C48DA" authority="Cam" authorityName="Cam" box="[884,1028,188,213]" class="Insecta" family="Staphylinidae" genus="Deliphrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="496" phylum="Arthropoda" rank="species" species="alticola">alticola Cam</taxonomicName>
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| H.Coiffait det. 1976’ <rectangular label, printed, except handwritten “76”> (NHMW; Franz collection).
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||
</paragraph>
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||
<paragraph id="8BFD36EEF15EFF98FF61FEE87DBC494E" blockId="45.[151,1437,151,1040]" pageId="45" pageNumber="496">
|
||
<emphasis id="B936EAFCF15EFF98FF61FEE87DA44913" bold="true" box="[199,444,260,285]" pageId="45" pageNumber="496">Additional material:</emphasis>
|
||
<materialsCitation id="3B2A3CB3F15EFF98FE64FEE87DB8494E" ID-GBIF-Occurrence="2028139685" collectionCode="SNSD" elevation="3700" location="Telbrung Danda" pageId="45" pageNumber="496" specimenCount="1" specimenCount-male="1" stateProvince="Eger">
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||
<specimenCount id="9D44FD67F15EFF98FE64FEE87DF74912" box="[450,495,260,284]" pageId="45" pageNumber="496" type="male">1 ♂</specimenCount>
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||
: ‘NEPAL-HIMALAYA | SE-Annapurna mts. | leg. O.
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||
<collectingRegion id="4986F80CF15EFF98FBEBFEE978904913" box="[1101,1160,261,285]" country="Hungary" name="Eger" pageId="45" pageNumber="496">Jäger</collectingRegion>
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||
1997’, ‘
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||
<location id="8E9D6035F15EFF98FB44FEE879844913" LSID="urn:lsid:plazi:treatment:03EB87F8F15FFF9AFF31F91278664BAA:8E9D6035F15EFF98FB44FEE879844913" box="[1250,1436,260,285]" name="Telbrung Danda" pageId="45" pageNumber="496" stateProvince="Eger">Telbrung Danda</location>
|
||
|
|
||
<quantity id="4CBA9B0BF15EFF98FF02FEC47CEC494E" box="[164,244,296,320]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="45" pageNumber="496" unit="m" value="3700.0">
|
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<elevation id="006FD1DDF15EFF98FF02FEC47CEC494E" box="[164,244,296,320]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="45" pageNumber="496" unit="m" value="3700.0">3700m</elevation>
|
||
</quantity>
|
||
, 9.VI’ (
|
||
<collectionCode id="ED53AE2BF15EFF98FEECFEC47D8F494E" box="[330,407,296,320]" pageId="45" pageNumber="496">SNSD</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3586565F15EFF9AFF61FEA0790C4AFE" lastPageId="47" lastPageNumber="498" pageId="45" pageNumber="496" type="description">
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<paragraph id="8BFD36EEF15EFF98FF61FEA07EAF49A3" blockId="45.[151,1437,151,1040]" pageId="45" pageNumber="496">
|
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<emphasis id="B936EAFCF15EFF98FF61FEA07D6C496B" bold="true" box="[199,372,332,357]" pageId="45" pageNumber="496">Redescription.</emphasis>
|
||
Measurements (n=2): HW: 0.67–0.68; HL: 0.40; AL(
|
||
<typeStatus id="54F9884CF15EFF98FC6AFEA0782B496B" box="[972,1075,332,357]" pageId="45" pageNumber="496" type="holotype">holotype</typeStatus>
|
||
): 1.11; OL: 0.17–0.19; PL: 0.60; PW: 1.20; ESL: 1.52–1.55; EW: 1.42–1.48; AW: 1.15–1.45; MTbL(
|
||
<typeStatus id="54F9884CF15EFF98FC31FE837FE54986" box="[919,1021,367,392]" pageId="45" pageNumber="496" type="holotype">holotype</typeStatus>
|
||
): 0.75, MTrL(
|
||
<typeStatus id="54F9884CF15EFF98FB39FE83791C4986" box="[1183,1284,367,392]" pageId="45" pageNumber="496" type="holotype">holotype</typeStatus>
|
||
): 0.40 (MTrL 1–4: 0.25; MTrL 5: 0.15); AedL: 0.70; TL: 3.50.
|
||
</paragraph>
|
||
<paragraph id="8BFD36EEF15EFF98FF61FE5B7EF44B1A" blockId="45.[151,1437,151,1040]" pageId="45" pageNumber="496">
|
||
Body moderately elongate, convex. Body and antennomeres 4–11 brown (
|
||
<typeStatus id="54F9884CF15EFF98FBB4FE5B786E49DE" box="[1042,1142,439,464]" pageId="45" pageNumber="496" type="holotype">holotype</typeStatus>
|
||
paler, with yellow-brown elytra); lateral, basal and apical parts of pronotum yellow-brown; mouthparts, antennomeres 1–3 and legs yellow. Body very shiny, forebody with bronze sheen; anterior portion of head between antennal insertion and anterior margin of eye with fine transverse meshes, posterior parts of head between ocelli and posterior margins of eyes with traces of fine transverse sculpture; abdomen with distinct isodiametric microsculpture. Head with irregular small punctation, slightly denser on infraorbital ridges; middle part of neck with irregular small punctation; pronotum with punctation markedly denser, larger and deeper than that of head, sparser on basal third and lateral portions; scutellum without or with several small punctures; punctation of elytra markedly larger and deeper than that on pronotum, dense, distinctly smaller on parascutellar area, each elytron with six, vague and tangled longitudinal rows of punctures. Habitus as in
|
||
<figureCitation id="13792A6BF15EFF98FD33FD107EFF4B1A" box="[661,743,764,789]" captionStart="FIGURES 94–95" captionStartId="45.[151,264,1980,2003]" captionTargetBox="[229,1383,1092,1951]" captionTargetId="figure@45.[151,1437,1087,1958]" captionTargetPageId="45" captionText="FIGURES 94–95. Habitus of Anthobium nigrum group: 94—A. alticola (NEPAL: Annapurna Mts.) and 95—A. flaveolum (paratype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607378/files/figure.png" pageId="45" pageNumber="496">Fig. 94</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFD36EEF15EFF98FF61FCF37F104C1E" blockId="45.[151,1437,151,1040]" pageId="45" pageNumber="496">
|
||
Head 1.6 times as wide as long; middle portion of head somewhat flattened, without transverse impression in middle; occipital furrow between ocelli wide and deep; grooves in front of ocelli long and deep, reaching apical third of length of eye; postocular ridges distinct, smooth; anterior portion between antennal insertion and anterior margin of eye with wide and moderately deep semicircular notch. Ocelli large and very convex, situated at level of postocular ridges, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Length × width of antennomeres (
|
||
<typeStatus id="54F9884CF15EFF98FDB5FC387E644BE3" box="[531,636,980,1005]" pageId="45" pageNumber="496" type="holotype">holotype</typeStatus>
|
||
): 1: 0.15 × 0.05; 2: 0.10 × 0.05; 3: 0.12 × 0.04; 4: 0.07 × 0.04; 5: 0.10 × 0.05; 6: 0.10 × 0.06; 7–10: 0.08 × 0.07; 11: 0.15 × 0.07.
|
||
</paragraph>
|
||
<caption id="DF3D6666F15EFF98FF31F8507DD34FFF" httpUri="https://zenodo.org/record/2607378/files/figure.png" pageId="45" pageNumber="496" startId="45.[151,264,1980,2003]" targetBox="[229,1383,1092,1951]" targetPageId="45">
|
||
<paragraph id="8BFD36EEF15EFF98FF31F8507DD34FFF" blockId="45.[151,1436,1980,2033]" pageId="45" pageNumber="496">
|
||
<emphasis id="B936EAFCF15EFF98FF31F8507D4E4FDF" bold="true" box="[151,342,1980,2003]" pageId="45" pageNumber="496">FIGURES 94–95.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C424D6DF15EFF98FE71F8507E804FDC" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[471,664,1980,2002]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="496" phylum="Arthropoda" rank="species" species="nigrum">
|
||
<emphasis id="B936EAFCF15EFF98FE71F8507E804FDC" box="[471,664,1980,2002]" italics="true" pageId="45" pageNumber="496">Anthobium nigrum</emphasis>
|
||
</taxonomicName>
|
||
group: 94—
|
||
<taxonomicName id="4C424D6DF15EFF98FC86F8517F944FDC" box="[800,908,1980,2002]" class="Insecta" family="Staphylinidae" genus="Deliphrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="496" phylum="Arthropoda" rank="species" species="alticola">
|
||
<emphasis id="B936EAFCF15EFF98FC86F8517F944FDC" box="[800,908,1980,2002]" italics="true" pageId="45" pageNumber="496">A. alticola</emphasis>
|
||
</taxonomicName>
|
||
(NEPAL: Annapurna Mts.) and 95—
|
||
<taxonomicName id="4C424D6DF15EFF98FABFF85179844FDC" authorityName="Shavrin & Smetana" authorityYear="2018" box="[1305,1436,1980,2002]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="496" phylum="Arthropoda" rank="species" species="flaveolum">
|
||
<emphasis id="B936EAFCF15EFF98FABFF85179844FDC" box="[1305,1436,1980,2002]" italics="true" pageId="45" pageNumber="496">A. flaveolum</emphasis>
|
||
</taxonomicName>
|
||
(paratype). Scale bar: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3D6666F15DFF9BFF31F8587F134FE2" httpUri="https://zenodo.org/record/2607380/files/figure.png" pageId="46" pageNumber="497" startId="46.[151,264,1972,1995]" targetBox="[325,1319,213,1913]" targetPageId="46">
|
||
<paragraph id="8BFD36EEF15DFF9BFF31F8587F134FE2" blockId="46.[151,1436,1972,2028]" pageId="46" pageNumber="497">
|
||
<emphasis id="B936EAFCF15DFF9BFF31F8587D4B4FC4" bold="true" box="[151,339,1972,1995]" pageId="46" pageNumber="497">FIGURES 96–99.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="4C424D6DF15DFF9BFE7BF8587EB84FC4" authorityName="Shavrin & Smetana" authorityYear="2018" baseAuthorityName="Coiffait" baseAuthorityYear="1977" box="[477,672,1972,1994]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="alticola">
|
||
<emphasis id="B936EAFCF15DFF9BFE7BF8587EB84FC4" box="[477,672,1972,1994]" italics="true" pageId="46" pageNumber="497">Anthobium alticola</emphasis>
|
||
</taxonomicName>
|
||
(96–97: NEPAL: Annapurna Mts.) and
|
||
<taxonomicName id="4C424D6DF15DFF9BFB92F85978AC4FC4" authorityName="Shavrin & Smetana" authorityYear="2018" box="[1076,1204,1972,1994]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="497" phylum="Arthropoda" rank="species" species="flaveolum">
|
||
<emphasis id="B936EAFCF15DFF9BFB92F85978AC4FC4" box="[1076,1204,1972,1994]" italics="true" pageId="46" pageNumber="497">A. flaveolum</emphasis>
|
||
</taxonomicName>
|
||
(98–99: holotype): 96, 98—parameral view, 97, 99—lateral view. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD36EEF15CFF9AFF61FF747E97494E" blockId="47.[151,1436,151,933]" pageId="47" pageNumber="498">Pronotum twice as wide as long, 1.7 times as wide as head, widest in middle, evenly narrowed both posteriad and anteriad; anterior angles widely rounded, not protruded anteriad; posterior angles obtuse or rounded; lateral edges of pronotum with irregular smooth crenulation; disc of pronotum with wide and markedly convex middle elevation, with pair of indistinct transverse impression on mediobasal third; lateral portions wide and slightly explanate, each with depp oval pit in middle.</paragraph>
|
||
<paragraph id="8BFD36EEF15CFF9AFF61FEA078B349A3" blockId="47.[151,1436,151,933]" pageId="47" pageNumber="498">Elytra slightly longer than wide, slightly widened apicad, reaching apical margin of abdominal tergite V, about two and half times as long as pronotum, with widely rounded apical margins truncated at suture; lateral portions moderately narrow, slightly explanate; latero-apical margins with small flattened crenulation.</paragraph>
|
||
<paragraph id="8BFD36EEF15CFF9AFF61FE5B7D0449FA" blockId="47.[151,1436,151,933]" pageId="47" pageNumber="498">Abdomen distinctly narrower than elytra, with a pair of small rounded tomentose wing-folding spots in middle of tergite V.</paragraph>
|
||
<paragraph id="8BFD36EEF15CFF9AFF61FE1378D24A6E" blockId="47.[151,1436,151,933]" pageId="47" pageNumber="498">
|
||
Male. Apical margins of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (
|
||
<figureCitation id="13792A6BF15CFF9AFAA5FE1379404A16" box="[1283,1368,511,536]" captionStart="FIGURES 96–99" captionStartId="46.[151,264,1972,1995]" captionTargetBox="[325,1319,213,1913]" captionTargetId="figure@46.[228,1365,193,1952]" captionTargetPageId="46" captionText="FIGURES 96–99. Aedeagus of Anthobium alticola (96–97: NEPAL: Annapurna Mts.) and A. flaveolum (98–99: holotype): 96, 98—parameral view, 97, 99—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607380/files/figure.png" pageId="47" pageNumber="498">Fig. 96</figureCitation>
|
||
) with median lobe wide basally, gradually narrowed toward truncate apex; parameres markedly exceeding apex of median lobe, with two pairs of short apical and preapical setae. Aedeagus laterally as in
|
||
<figureCitation id="13792A6BF15CFF9AFBD5FDAB78DE4A6E" box="[1139,1222,583,608]" captionStart="FIGURES 96–99" captionStartId="46.[151,264,1972,1995]" captionTargetBox="[325,1319,213,1913]" captionTargetId="figure@46.[228,1365,193,1952]" captionTargetPageId="46" captionText="FIGURES 96–99. Aedeagus of Anthobium alticola (96–97: NEPAL: Annapurna Mts.) and A. flaveolum (98–99: holotype): 96, 98—parameral view, 97, 99—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607380/files/figure.png" pageId="47" pageNumber="498">Fig. 97</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFD36EEF15CFF9AFF61FD8079914A8B" blockId="47.[151,1436,151,933]" box="[199,1417,620,645]" pageId="47" pageNumber="498">Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</paragraph>
|
||
<paragraph id="8BFD36EEF15CFF9AFF61FD63790C4AFE" blockId="47.[151,1436,151,933]" pageId="47" pageNumber="498">
|
||
<emphasis id="B936EAFCF15CFF9AFF61FD637DAA4AA6" bold="true" box="[199,434,655,680]" pageId="47" pageNumber="498">Comparative notes.</emphasis>
|
||
Based on the shape of the convex body, the flattened head and the sparse punctation of the forebody,
|
||
<taxonomicName id="4C424D6DF15CFF9AFE94FD587DBF4AC2" authorityName="Shavrin & Smetana" authorityYear="2018" baseAuthorityName="Coiffait" baseAuthorityYear="1977" box="[306,423,692,716]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="alticola">
|
||
<emphasis id="B936EAFCF15CFF9AFE94FD587DBF4AC2" box="[306,423,692,716]" italics="true" pageId="47" pageNumber="498">A. alticola</emphasis>
|
||
</taxonomicName>
|
||
is most similar to the Himalayan
|
||
<taxonomicName id="4C424D6DF15CFF9AFC82FD5878BF4AC2" authority="Shavrin & Smetana, 2017" authorityName="Shavrin & Smetana" authorityYear="2017" box="[804,1191,692,717]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="ivani">
|
||
<emphasis id="B936EAFCF15CFF9AFC82FD587F634AC2" box="[804,891,692,716]" italics="true" pageId="47" pageNumber="498">A. ivani</emphasis>
|
||
<bibRefCitation id="EFD34B1FF15CFF9AFC25FD5878BF4AC2" author="Shavrin, A. V. & Smetana, A." box="[899,1191,692,717]" pageId="47" pageNumber="498" pagination="1 - 39" refId="ref40158" refString="Shavrin, A. V. & Smetana, A. (2017) A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups. Zootaxa, 4365 (1), 1 - 39. https: // doi. org / 10.11646 / zootaxa. 4365.1.1" type="journal article" year="2017">Shavrin & Smetana, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
, from which it differs by shorter body, paler coloration, longer elytra and by the shape and internal structure of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3586565F15CFF9AFF61FD107D284B53" pageId="47" pageNumber="498" type="distribution">
|
||
<paragraph id="8BFD36EEF15CFF9AFF61FD10797B4B1A" blockId="47.[151,1436,151,933]" box="[199,1379,764,789]" pageId="47" pageNumber="498">
|
||
<emphasis id="B936EAFCF15CFF9AFF61FD107D784B1B" bold="true" box="[199,352,764,789]" pageId="47" pageNumber="498">Distribution.</emphasis>
|
||
The species is at present known from two locations in central and eastern
|
||
<collectingCountry id="F355767EF15CFF9AFB04FD1078FD4B1B" box="[1186,1253,764,789]" name="Nepal" pageId="47" pageNumber="498">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="13792A6BF15CFF9AFB52FD10794E4B1A" box="[1268,1366,764,789]" captionStart="FIGURE 100" captionStartId="48.[151,250,1958,1980]" captionTargetBox="[178,1399,1201,1918]" captionTargetId="figure@48.[178,1399,1201,1918]" captionTargetPageId="48" captionText="FIGURE 100. Distribution of Anthobium alticola (black circles) and A. flaveolum (black square) in the Himalayan Region." httpUri="https://zenodo.org/record/2607384/files/figure.png" pageId="47" pageNumber="498">Fig. 100</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFD36EEF15CFF9AFF61FCF37D284B53" blockId="47.[151,1436,151,933]" pageId="47" pageNumber="498">
|
||
<emphasis id="B936EAFCF15CFF9AFF61FCF37D524B36" bold="true" box="[199,330,799,824]" pageId="47" pageNumber="498">Bionomics.</emphasis>
|
||
Specimens were collected at elevations from 3700 and
|
||
<quantity id="4CBA9B0BF15CFF9AFC62FCF378024B39" box="[964,1050,799,824]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="47" pageNumber="498" unit="m" value="5000.0">5000 m</quantity>
|
||
a.s.l. The detailed ecological data are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3586565F15CFF9AFF61FC8B78664BAA" pageId="47" pageNumber="498" type="discussion">
|
||
<paragraph id="8BFD36EEF15CFF9AFF61FC8B78664BAA" blockId="47.[151,1436,151,933]" pageId="47" pageNumber="498">
|
||
<emphasis id="B936EAFCF15CFF9AFF61FC8B7D234B8E" bold="true" box="[199,315,871,896]" pageId="47" pageNumber="498">Remarks.</emphasis>
|
||
The study of the
|
||
<typeStatus id="54F9884CF15CFF9AFDB5FC8B7E6D4B8E" box="[531,629,871,896]" pageId="47" pageNumber="498" type="holotype">holotype</typeStatus>
|
||
and the additional male from central
|
||
<collectingCountry id="F355767EF15CFF9AFB90FC8B78614B8E" box="[1078,1145,871,896]" name="Nepal" pageId="47" pageNumber="498">Nepal</collectingCountry>
|
||
revealed that
|
||
<taxonomicName id="4C424D6DF15CFF9AFA85FC847CF54BAA" class="Insecta" family="Staphylinidae" genus="Deliphrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="alticola">
|
||
<emphasis id="B936EAFCF15CFF9AFA85FC847CF54BAA" italics="true" pageId="47" pageNumber="498">Deliphrum alticola</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C424D6DF15CFF9AFE31FC617DF04BAA" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[407,488,909,932]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="species" species="nigrum">
|
||
<emphasis id="B936EAFCF15CFF9AFE31FC617DF04BAA" box="[407,488,909,932]" italics="true" pageId="47" pageNumber="498">nigrum</emphasis>
|
||
</taxonomicName>
|
||
group of the genus
|
||
<taxonomicName id="4C424D6DF15CFF9AFD61FC6078624BAA" authority="(Shavrin & Smetana 2017)" baseAuthorityName="Shavrin & Smetana" baseAuthorityYear="2017" box="[711,1146,908,933]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="498" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B936EAFCF15CFF9AFD61FC607F5A4BAA" box="[711,834,908,932]" italics="true" pageId="47" pageNumber="498">Anthobium</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD34B1FF15CFF9AFCF7FC60786A4BAA" author="Shavrin, A. V. & Smetana, A." box="[849,1138,908,933]" pageId="47" pageNumber="498" pagination="1 - 39" refId="ref40158" refString="Shavrin, A. V. & Smetana, A. (2017) A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups. Zootaxa, 4365 (1), 1 - 39. https: // doi. org / 10.11646 / zootaxa. 4365.1.1" type="journal article" year="2017">Shavrin & Smetana 2017</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |