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<document ID-DOI="http://doi.org/10.5281/zenodo.4619827" ID-GBIF-Dataset="a3b1a6f6-898c-4c71-afb7-b3cc5b59ce85" ID-GBIF-Taxon="119316521" ID-ISSN="1175­5326" ID-Zenodo-Dep="4619827" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1458923791862" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="03EB87A5FFCFC10FFE8AFC42A824F96D" docLanguage="en" docName="zt00030.pdf.imf" docOrigin="Zootaxa 30" docStyle="DocumentStyle:83E4DCE73C651764D97D3683740715A6.1:zzz_test.0000.journal_article" docStyleId="83E4DCE73C651764D97D3683740715A6" docStyleName="zzz_test.0000.journal_article" docStyleVersion="1" docTitle="Tomoglossa Kraatz 1856" docType="treatment" docVersion="9" lastPageId="4" lastPageNumber="5" masterDocId="FFD2FFDDFFCDC10BFF82FFB6AB33FFEB" masterDocTitle="A revision of Nearctic species of the genus Tomoglossa Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="19" masterPageNumber="1" pageId="2" pageNumber="3" updateTime="1643593573506" updateUser="ExternalLinkService">
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<mods:title>A revision of Nearctic species of the genus Tomoglossa Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart>Gusarov, Vladimir I.</mods:namePart>
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<mods:date>2002</mods:date>
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<subSubSection box="[264,723,1011,1038]" pageId="2" pageNumber="3" type="nomenclature">
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<heading bold="true" box="[264,723,1011,1038]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<taxonomicName authority="Kraatz, 1856" authorityName="Kraatz" authorityYear="1856" box="[264,576,1011,1038]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[264,406,1012,1038]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
<bibRefCitation author="Kraatz, G." box="[414,576,1011,1037]" pageId="2" pageNumber="3" refId="ref7346" refString="Kraatz, G. (1856 - 1858) Naturgeschichte der Insecten Deutschlands. Abt. 1. Coleoptera. Band 2. Verlag der Nicolaischen Buchhandlung, Berlin, 1080 pp." type="book" year="1856">
Kraatz,
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</taxonomicName>
(
<figureCitation box="[591,714,1011,1037]" captionStart-0="FIGURES 1­12" captionStart-1="FIGURES 13­20" captionStart-2="FIGURES 21­28" captionStart-3="FIGURES 29­33" captionStart-4="FIGURES 34­37" captionStart-5="FIGURES 38­42" captionStart-6="FIGURES 43­46" captionStartId-0="6.[264,383,1719,1743]" captionStartId-1="7.[264,383,1766,1790]" captionStartId-2="8.[264,383,1724,1748]" captionStartId-3="10.[264,383,1748,1772]" captionStartId-4="11.[264,383,1741,1765]" captionStartId-5="13.[264,383,1771,1795]" captionStartId-6="15.[264,383,1451,1475]" captionTargetBox-0="[340,1243,331,1659]" captionTargetBox-1="[351,1233,333,1743]" captionTargetBox-2="[379,1211,497,1686]" captionTargetBox-3="[422,1164,577,1689]" captionTargetBox-4="[344,1236,722,1679]" captionTargetBox-5="[384,1200,958,1742]" captionTargetBox-6="[450,1136,732,1414]" captionTargetId-0="figure@6.[337,1250,329,1666]" captionTargetId-1="figure@7.[345,1241,326,1749]" captionTargetId-2="figure@8.[370,1217,489,1697]" captionTargetId-3="figure@10.[416,1171,569,1698]" captionTargetId-4="figure@11.[340,1247,719,1683]" captionTargetId-5="figure@13.[380,1207,943,1745]" captionTargetId-6="figure@15.[447,1140,729,1424]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="13" captionTargetPageId-6="15" captionText-0="FIGURES 1­12. Mouthparts of Tomoglossa decora (Casey) (Vermont). 1, left mandible, ventral view; 2, left mandible, dorsal view; 3, right mandible, dorsal view; 4, left maxilla, ventral view; 5, right galea, ventral view; 6, right galea, dorsal view; 7, left lacinia, dorsal view; 8, labrum; 9, epipharynx; 10, prementum; 11, hypopharynx; 12, mentum. Scale bar 0.1 mm (1­9, 12), 0.07 mm (11), 0.05 mm (10)." captionText-1="FIGURES 13­20. Details of Tomoglossa decora (Casey) (Vermont (13­15, 20), Pennsylvania (16­ 18), lectotype (19)). 13, pronotum; 14, right metatarsus, posterior view; 15, mesometathorax, ventral view; 16, apical margin of female sternum VIII; 17, female tergum VIII; 18, female sternum VIII; 19, male tergum VIII; 20, male sternum VIII. Scale bar 0.2 mm (13­15, 17­20), 0.1 mm (16)." captionText-2="FIGURES 21­28. Genitalia and terminalia of Tomoglossa decora (Casey) (Vermont (21­25), Pennsylvania (26­28)). 21, median lobe of aedeagus, lateral view; 22, median lobe of aedeagus, ventral view; 23, internal sac of aedeagus, in packed state; 24, deeply concave sclerite of internal sac; 25, copulatory piece; 26, spermatheca; 27, left portion of female sternum IX; 28, female accessory sclerite. U ­ umbilicus. Scale bar 0.1 mm." captionText-3="FIGURES 29­33. Antennae of Nearctic Tomoglossa. 29, T. ornatella (Casey) (lectotype); 30, T. improvisa (Casey) (lectotype); 31, T. decora (Casey) (Pennsylvania); 32, T. arizonica Gusarov, sp. nov. (holotype); 33, T. floridana Gusarov, sp. nov. (holotype). Scale bar 0.1 mm." captionText-4="FIGURES 34­37. Details of Tomoglossa ornatella (Casey) (lectotype). 34, female tergum VIII; 35, female sternum VIII; 36, spermatheca; 37, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (34­35), 0.1 mm (36­37)." captionText-5="FIGURES 38­42. Details of Tomoglossa arizonica Gusarov, sp. nov. (holotype). 38, male tergum VIII; 39, male sternum VIII; 40, median lobe of aedeagus, lateral view; 41, median lobe of aedeagus, ventral view; 42, internal sac of aedeagus, in packed state. Scale bar 0.2 mm (38­39), 0.1 mm (40­42)." captionText-6="FIGURES 43­46. Details of Tomoglossa floridana Gusarov, sp. nov. (holotype). 43, female tergum VIII; 44, female sternum VIII; 45, spermatheca; 46, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (43­44), 0.1 mm (45­46)." httpUri-0="https://zenodo.org/record/155537/files/figure.png" httpUri-1="https://zenodo.org/record/155538/files/figure.png" httpUri-2="https://zenodo.org/record/155539/files/figure.png" httpUri-3="https://zenodo.org/record/155540/files/figure.png" httpUri-4="https://zenodo.org/record/155541/files/figure.png" httpUri-5="https://zenodo.org/record/155542/files/figure.png" httpUri-6="https://zenodo.org/record/155543/files/figure.png" pageId="2" pageNumber="3">
Figs.
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)
</emphasis>
</heading>
</paragraph>
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<paragraph blockId="2.[264,881,1081,1556]" box="[264,603,1081,1104]" pageId="2" pageNumber="3">
<taxonomicName authority="Kraatz, 1856: 342" authorityName="Kraatz" authorityPageNumber="342" authorityYear="1856" box="[264,599,1081,1104]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,388,1081,1104]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
<bibRefCitation author="Kraatz, G." box="[397,599,1081,1104]" pageId="2" pageNumber="3" refId="ref7346" refString="Kraatz, G. (1856 - 1858) Naturgeschichte der Insecten Deutschlands. Abt. 1. Coleoptera. Band 2. Verlag der Nicolaischen Buchhandlung, Berlin, 1080 pp." type="book" year="1856">
Kraatz,
<number box="[482,539,1081,1104]" pageId="2" pageNumber="3" value="1856.0">1856</number>
:
<number box="[556,599,1081,1104]" pageId="2" pageNumber="3" value="342.0">342</number>
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</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,662,1116,1139]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[264,388,1116,1139]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,388,1116,1139]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
</taxonomicName>
;
<bibRefCitation author="Bernhauer, M." box="[404,658,1116,1139]" pageId="2" pageNumber="3" pagination="381 - 405" refId="ref7085" refString="Bernhauer, M. (1907 b) Neue Aleocharini aus Nordamerika. (Col.) (3. Stuck.). Deutsche Entomologische Zeitschrift, 1907 (4), 381 - 405." type="journal article" year="1907">
Bernhauer,
<number box="[528,584,1116,1139]" pageId="2" pageNumber="3" value="1907.0">1907</number>
b:
<number box="[615,658,1116,1139]" pageId="2" pageNumber="3" value="401.0">401</number>
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.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="description">
<paragraph blockId="2.[264,881,1081,1556]" box="[264,657,1149,1174]" pageId="2" pageNumber="3">
<taxonomicName box="[264,502,1150,1174]" pageId="2" pageNumber="3">
<emphasis box="[264,362,1151,1174]" italics="true" pageId="2" pageNumber="3">Noverota</emphasis>
<bibRefCitation author="Casey, T. L." box="[369,548,1150,1173]" pageId="2" pageNumber="3" pagination="1 - 183" refId="ref7205" refString="Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera, 1, 1 - 183." type="journal article" year="1910">
Casey,
<number box="[445,502,1150,1173]" pageId="2" pageNumber="3" value="1910.0">1910</number>
:
<number box="[519,548,1150,1173]" pageId="2" pageNumber="3" value="90.0">90</number>
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</taxonomicName>
,
<emphasis bold="true" box="[559,657,1149,1173]" pageId="2" pageNumber="3">syn. nov.</emphasis>
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,614,1185,1208]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[264,388,1185,1208]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,388,1185,1208]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
</taxonomicName>
;
<bibRefCitation author="Fenyes, A." box="[404,610,1185,1208]" pageId="2" pageNumber="3" pagination="111 - 414" refId="ref7266" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.) Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Brussels, pp. 111 - 414." type="book chapter" year="1920">
Fenyes,
<number box="[493,550,1185,1208]" pageId="2" pageNumber="3" value="1920.0">1920</number>
:
<number box="[567,610,1185,1208]" pageId="2" pageNumber="3" value="255.0">255</number>
</bibRefCitation>
.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,679,1220,1244]" pageId="2" pageNumber="3">
<taxonomicName authority="(Noverota)" authorityName="Noverota" box="[264,457,1220,1244]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,333,1220,1243]" italics="true" pageId="2" pageNumber="3">Atheta</emphasis>
(
<emphasis box="[349,446,1221,1244]" italics="true" pageId="2" pageNumber="3">Noverota</emphasis>
)
</taxonomicName>
;
<bibRefCitation author="Fenyes, A." box="[470,675,1220,1243]" pageId="2" pageNumber="3" pagination="111 - 414" refId="ref7266" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.) Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Brussels, pp. 111 - 414." type="book chapter" year="1920">
Fenyes,
<number box="[558,615,1220,1243]" pageId="2" pageNumber="3" value="1920.0">1920</number>
:
<number box="[632,675,1220,1243]" pageId="2" pageNumber="3" value="199.0">199</number>
</bibRefCitation>
.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,808,1254,1278]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[264,388,1255,1278]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,388,1255,1278]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
</taxonomicName>
;
<bibRefCitation author="Bernhauer, M. &amp; Scheerpeltz, O." box="[404,804,1254,1277]" pageId="2" pageNumber="3" pagination="499 - 988" refId="ref7122" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, 499 - 988." type="journal article" year="1926">
Bernhauer &amp; Scheerpeltz,
<number box="[687,744,1254,1277]" pageId="2" pageNumber="3" value="1926.0">1926</number>
:
<number box="[761,804,1254,1277]" pageId="2" pageNumber="3" value="598.0">598</number>
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.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,874,1289,1313]" pageId="2" pageNumber="3">
<taxonomicName authority="(Noverota)" authorityName="Noverota" box="[264,457,1289,1313]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,333,1289,1312]" italics="true" pageId="2" pageNumber="3">Atheta</emphasis>
(
<emphasis box="[349,446,1290,1313]" italics="true" pageId="2" pageNumber="3">Noverota</emphasis>
)
</taxonomicName>
;
<bibRefCitation author="Bernhauer, M. &amp; Scheerpeltz, O." box="[470,869,1289,1312]" pageId="2" pageNumber="3" pagination="499 - 988" refId="ref7122" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.) Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, 499 - 988." type="journal article" year="1926">
Bernhauer &amp; Scheerpeltz,
<number box="[753,810,1289,1312]" pageId="2" pageNumber="3" value="1926.0">1926</number>
:
<number box="[826,869,1289,1312]" pageId="2" pageNumber="3" value="606.0">606</number>
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.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,663,1324,1347]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[264,388,1324,1347]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,388,1324,1347]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
</taxonomicName>
;
<bibRefCitation author="Scheerpeltz, O." box="[404,659,1324,1347]" pageId="2" pageNumber="3" pagination="123 - 132" refId="ref7999" refString="Scheerpeltz, O. (1963) Neue Arten der Gattung Tomoglossa Kraatz, mit einer Bestimmingstabelle der bis jetzt bekannt gewordenen Arten dieser Gattung (Col. Staph.). Entomologische Berichten, 23 (7), 123 - 132." type="journal article" year="1963">
Scheerpeltz,
<number box="[543,600,1324,1347]" pageId="2" pageNumber="3" value="1963.0">1963</number>
:
<number box="[616,659,1324,1347]" pageId="2" pageNumber="3" value="123.0">123</number>
</bibRefCitation>
.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,714,1358,1382]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[264,388,1359,1382]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,388,1359,1382]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
</taxonomicName>
;
<bibRefCitation author="Benick, G. &amp; Lohse, G. A." box="[404,710,1358,1381]" pageId="2" pageNumber="3" pagination="72 - 220" refId="ref6982" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.) Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">
Benick &amp; Lohse,
<number box="[593,650,1358,1381]" pageId="2" pageNumber="3" value="1974.0">1974</number>
:
<number box="[667,710,1358,1381]" pageId="2" pageNumber="3" value="107.0">107</number>
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.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,621,1393,1416]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[264,388,1393,1416]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,388,1393,1416]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
</taxonomicName>
;
<bibRefCitation author="Sawada, K." box="[404,617,1393,1416]" pageId="2" pageNumber="3" pagination="171 - 222" refId="ref7901" refString="Sawada, K. (1977) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). III: Japanese Species described by the previous Authors. Contributions from the Biological Laboratory, Kyoto University, 25 (2), 171 - 222." type="journal article" year="1977">
Sawada,
<number box="[500,557,1393,1416]" pageId="2" pageNumber="3" value="1977.0">1977</number>
:
<number box="[574,617,1393,1416]" pageId="2" pageNumber="3" value="192.0">192</number>
</bibRefCitation>
.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,593,1428,1451]" pageId="2" pageNumber="3">
<taxonomicName box="[264,529,1428,1451]" pageId="2" pageNumber="3">
<emphasis box="[264,362,1428,1451]" italics="true" pageId="2" pageNumber="3">Noverota</emphasis>
;
<bibRefCitation author="Seevers, C. H." box="[376,589,1428,1451]" pageId="2" pageNumber="3" pagination="1 - 275" refId="ref8043" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, i-vi + 1 - 275." type="journal article" year="1978">
Seevers,
<number box="[472,529,1428,1451]" pageId="2" pageNumber="3" value="1978.0">1978</number>
:
<number box="[546,589,1428,1451]" pageId="2" pageNumber="3" value="122.0">122</number>
</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,604,1462,1486]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[264,388,1463,1486]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[264,388,1463,1486]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
</taxonomicName>
;
<bibRefCitation author="Lohse, G. A." box="[404,600,1462,1485]" pageId="2" pageNumber="3" pagination="185 - 243" refId="ref7412" refString="Lohse, G. A. (1989) Erganzungen und Berichtigungen zu Freude-Harde-Lohse &quot; Die Kafer Mitteleuropas &quot; Band 5 (1974). In: Lohse, G. A. &amp; Lucht, W. H. (Eds.) Die Kafer Mitteleuropas. 1. Supplementband mit Katalogteil. Goecke &amp; Evers Verlag, Krefeld, pp. 185 - 243." type="journal article" year="1989">
Lohse,
<number box="[483,540,1462,1485]" pageId="2" pageNumber="3" value="1989.0">1989</number>
:
<number box="[557,600,1462,1485]" pageId="2" pageNumber="3" value="208.0">208</number>
</bibRefCitation>
.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,881,1497,1520]" pageId="2" pageNumber="3">
<taxonomicName box="[264,817,1497,1520]" pageId="2" pageNumber="3">
<emphasis box="[264,362,1497,1520]" italics="true" pageId="2" pageNumber="3">Noverota</emphasis>
;
<bibRefCitation author="Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[377,877,1497,1520]" editor="Arnett, R. H. &amp; Thomas, M. C." journalOrPublisher="CRC Press, Boca Raton" pageId="2" pageNumber="3" pagination="272 - 418" refId="ref7542" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.) American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, pp. 272 - 418." title="Staphylinidae Latreille, 1802" type="book chapter" volumeTitle="American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia" year="2000">
Newton, Thayer, Ashe &amp; Chandler,
<number box="[760,817,1497,1520]" pageId="2" pageNumber="3" value="2000.0">2000</number>
:
<number box="[834,877,1497,1520]" pageId="2" pageNumber="3" value="369.0">369</number>
</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[264,881,1081,1556]" box="[264,880,1532,1556]" pageId="2" pageNumber="3">
(Other references for Palaearctic
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[618,744,1533,1556]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[618,744,1533,1556]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
</taxonomicName>
are omitted)
</paragraph>
</subSubSection>
<subSubSection box="[264,1056,1601,1627]" pageId="2" pageNumber="3" type="materials_examined">
<paragraph blockId="2.[264,1324,1601,1867]" box="[264,1056,1601,1627]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[264,431,1601,1627]" pageId="2" pageNumber="3">
<typeStatus box="[264,325,1601,1627]" pageId="2" pageNumber="3">Type</typeStatus>
Species.
</emphasis>
<taxonomicName authority="Erichson, 1837" authorityName="Erichson" authorityYear="1837" baseAuthorityName="Erichson" baseAuthorityYear="1837" box="[446,883,1601,1627]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="luteicornis">
<emphasis box="[446,696,1602,1627]" italics="true" pageId="2" pageNumber="3">Homalota luteicornis</emphasis>
<bibRefCitation author="Erichson, W. F." box="[703,883,1601,1627]" pageId="2" pageNumber="3" refId="ref7233" refString="Erichson, W. F. (1837 - 1839) Die Kafer der Mark Brandenburg. Band 1. F. H. Morin, Berlin, viii + 740 pp." type="book" year="1837">
Erichson,
<number box="[822,883,1601,1627]" pageId="2" pageNumber="3" value="1837.0">1837</number>
</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="diagnosis">
<paragraph blockId="2.[264,1324,1601,1867]" lastBlockId="3.[264,1324,284,1870]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis bold="true" box="[312,440,1641,1667]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[456,594,1642,1667]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[456,594,1642,1667]" italics="true" pageId="2" pageNumber="3">Tomoglossa</emphasis>
</taxonomicName>
can be distinguished from other athetine genera by the combination of the following characters: parallel­sided body; slender sickle­shaped mandibles (
<figureCitation box="[350,457,1721,1747]" captionStart="FIGURES 1­12" captionStartId="6.[264,383,1719,1743]" captionTargetBox="[340,1243,331,1659]" captionTargetId="figure@6.[337,1250,329,1666]" captionTargetPageId="6" captionText="FIGURES 1­12. Mouthparts of Tomoglossa decora (Casey) (Vermont). 1, left mandible, ventral view; 2, left mandible, dorsal view; 3, right mandible, dorsal view; 4, left maxilla, ventral view; 5, right galea, ventral view; 6, right galea, dorsal view; 7, left lacinia, dorsal view; 8, labrum; 9, epipharynx; 10, prementum; 11, hypopharynx; 12, mentum. Scale bar 0.1 mm (1­9, 12), 0.07 mm (11), 0.05 mm (10)." httpUri="https://zenodo.org/record/155537/files/figure.png" pageId="2" pageNumber="3">
Figs.
<number box="[417,457,1721,1747]" pageId="2" pageNumber="3" value="2.0" valueMax="3.0" valueMin="1.0">1­3</number>
</figureCitation>
); ligula with two completely separate lobes (
<figureCitation box="[986,1074,1721,1747]" captionStart="FIGURES 1­12" captionStartId="6.[264,383,1719,1743]" captionTargetBox="[340,1243,331,1659]" captionTargetId="figure@6.[337,1250,329,1666]" captionTargetPageId="6" captionText="FIGURES 1­12. Mouthparts of Tomoglossa decora (Casey) (Vermont). 1, left mandible, ventral view; 2, left mandible, dorsal view; 3, right mandible, dorsal view; 4, left maxilla, ventral view; 5, right galea, ventral view; 6, right galea, dorsal view; 7, left lacinia, dorsal view; 8, labrum; 9, epipharynx; 10, prementum; 11, hypopharynx; 12, mentum. Scale bar 0.1 mm (1­9, 12), 0.07 mm (11), 0.05 mm (10)." httpUri="https://zenodo.org/record/155537/files/figure.png" pageId="2" pageNumber="3">
Fig.
<number box="[1041,1074,1721,1747]" pageId="2" pageNumber="3" value="10.0">10</number>
</figureCitation>
); mentum fused with submentum (
<figureCitation box="[419,508,1761,1787]" captionStart="FIGURES 1­12" captionStartId="6.[264,383,1719,1743]" captionTargetBox="[340,1243,331,1659]" captionTargetId="figure@6.[337,1250,329,1666]" captionTargetPageId="6" captionText="FIGURES 1­12. Mouthparts of Tomoglossa decora (Casey) (Vermont). 1, left mandible, ventral view; 2, left mandible, dorsal view; 3, right mandible, dorsal view; 4, left maxilla, ventral view; 5, right galea, ventral view; 6, right galea, dorsal view; 7, left lacinia, dorsal view; 8, labrum; 9, epipharynx; 10, prementum; 11, hypopharynx; 12, mentum. Scale bar 0.1 mm (1­9, 12), 0.07 mm (11), 0.05 mm (10)." httpUri="https://zenodo.org/record/155537/files/figure.png" pageId="2" pageNumber="3">
Fig.
<number box="[476,508,1761,1787]" pageId="2" pageNumber="3" value="12.0">12</number>
</figureCitation>
); pronotum with microsetae directed posteriorly throughout the disc (
<typeStatus box="[272,332,1801,1827]" pageId="2" pageNumber="3">Type</typeStatus>
VI,
<bibRefCitation author="Benick, G. &amp; Lohse, G. A." box="[385,664,1801,1827]" pageId="2" pageNumber="3" pagination="72 - 220" refId="ref6982" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.) Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">
Benick and Lohse
<number box="[604,664,1801,1827]" pageId="2" pageNumber="3" value="1974.0">1974</number>
</bibRefCitation>
) (
<figureCitation box="[688,775,1801,1827]" captionStart="FIGURES 13 ­ 20" captionStartId="7.[264,383,1766,1790]" captionTargetBox="[351,1233,333,1743]" captionTargetId="figure@7.[345,1241,326,1749]" captionTargetPageId="7" captionText="FIGURES 13 ­ 20. Details of Tomoglossa decora (Casey) (Vermont (13 ­ 15, 20), Pennsylvania (16 ­ 18), lectotype (19 )). 13, pronotum; 14, right metatarsus, posterior view; 15, mesometathorax, ventral view; 16, apical margin of female sternum VIII; 17, female tergum VIII; 18, female sternum VIII; 19, male tergum VIII; 20, male sternum VIII. Scale bar 0.2 mm (13 ­ 15, 17 ­ 20), 0.1 mm (16)." httpUri="https://zenodo.org/record/155538/files/figure.png" pageId="2" pageNumber="3">
Fig.
<number box="[743,775,1801,1827]" pageId="2" pageNumber="3" value="13.0">13</number>
</figureCitation>
); pronotal macrosetae inconspicuous; pronotal hypomera fully visible in lateral aspect; metasternal process extremely short and wide, virtually non­existent, perhaps better described as convexity of the anterior margin of metasternum (
<figureCitation box="[381,473,324,350]" captionStart="FIGURES 13 ­ 20" captionStartId="7.[264,383,1766,1790]" captionTargetBox="[351,1233,333,1743]" captionTargetId="figure@7.[345,1241,326,1749]" captionTargetPageId="7" captionText="FIGURES 13 ­ 20. Details of Tomoglossa decora (Casey) (Vermont (13 ­ 15, 20), Pennsylvania (16 ­ 18), lectotype (19 )). 13, pronotum; 14, right metatarsus, posterior view; 15, mesometathorax, ventral view; 16, apical margin of female sternum VIII; 17, female tergum VIII; 18, female sternum VIII; 19, male tergum VIII; 20, male sternum VIII. Scale bar 0.2 mm (13 ­ 15, 17 ­ 20), 0.1 mm (16)." httpUri="https://zenodo.org/record/155538/files/figure.png" pageId="3" pageNumber="4">
Fig.
<number box="[441,473,324,350]" pageId="3" pageNumber="4" value="15.0">15</number>
</figureCitation>
); mesotibia with short median macroseta (shorter than tibial width); metatarsal segment
<number box="[510,525,364,390]" pageId="3" pageNumber="4" value="1.0">1</number>
longer than segment
<number box="[803,818,364,390]" pageId="3" pageNumber="4" value="2.0">2</number>
(
<figureCitation box="[841,934,364,390]" captionStart="FIGURES 13 ­ 20" captionStartId="7.[264,383,1766,1790]" captionTargetBox="[351,1233,333,1743]" captionTargetId="figure@7.[345,1241,326,1749]" captionTargetPageId="7" captionText="FIGURES 13 ­ 20. Details of Tomoglossa decora (Casey) (Vermont (13 ­ 15, 20), Pennsylvania (16 ­ 18), lectotype (19 )). 13, pronotum; 14, right metatarsus, posterior view; 15, mesometathorax, ventral view; 16, apical margin of female sternum VIII; 17, female tergum VIII; 18, female sternum VIII; 19, male tergum VIII; 20, male sternum VIII. Scale bar 0.2 mm (13 ­ 15, 17 ­ 20), 0.1 mm (16)." httpUri="https://zenodo.org/record/155538/files/figure.png" pageId="3" pageNumber="4">
Fig.
<number box="[902,934,364,390]" pageId="3" pageNumber="4" value="14.0">14</number>
</figureCitation>
); without empodial setae; with numerous erect macrosetae and long semi­erect microsetae on abdominal terga VII­VIII.
</paragraph>
<paragraph blockId="3.[264,1324,284,1870]" pageId="3" pageNumber="4">
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[312,451,444,469]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[312,451,444,469]" italics="true" pageId="3" pageNumber="4">Tomoglossa</emphasis>
</taxonomicName>
shares with
<taxonomicName authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[624,988,444,470]" class="Insecta" family="Staphylinidae" genus="Hydrosmecta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[624,779,444,469]" italics="true" pageId="3" pageNumber="4">Hydrosmecta</emphasis>
Thomson,
<number box="[929,988,444,470]" pageId="3" pageNumber="4" value="1858.0">1858</number>
</taxonomicName>
slender mandibles, inconspicuous process of the metasternum, and the lack of empodial setae.
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[1098,1237,484,509]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[1098,1237,484,509]" italics="true" pageId="3" pageNumber="4">Tomoglossa</emphasis>
</taxonomicName>
can be easily distinguished from
<taxonomicName authorityName="Thomson" authorityYear="1858" box="[579,734,524,549]" class="Insecta" family="Staphylinidae" genus="Hydrosmecta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[579,734,524,549]" italics="true" pageId="3" pageNumber="4">Hydrosmecta</emphasis>
</taxonomicName>
by its less slender body, more widely separated lobes of the ligula, less transverse mentum fused with submentum, pubescence directed posteriorly everywhere on the pronotal disc (
<typeStatus box="[784,844,604,630]" pageId="3" pageNumber="4">Type</typeStatus>
VI), shorter metasternum, and long erect setation of the apical abdominal segments.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph blockId="3.[264,1324,284,1870]" pageId="3" pageNumber="4">
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[312,451,684,709]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[312,451,684,709]" italics="true" pageId="3" pageNumber="4">Tomoglossa</emphasis>
</taxonomicName>
is similar to
<taxonomicName authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[613,915,684,710]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[613,719,684,709]" italics="true" pageId="3" pageNumber="4">Geostiba</emphasis>
Thomson,
<quantity box="[856,915,684,710]" metricMagnitude="1" metricUnit="m" metricValue="4.71932" pageId="3" pageNumber="4" unit="in" value="1858.0">1858</quantity>
</taxonomicName>
in having pronotal pubescence of
<typeStatus box="[264,322,724,750]" pageId="3" pageNumber="4">Type</typeStatus>
VI; however,
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[487,626,724,749]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[487,626,724,749]" italics="true" pageId="3" pageNumber="4">Tomoglossa</emphasis>
</taxonomicName>
differs in having slender mandibles, more widely separated lobes of ligula, less transverse mentum fused with submentum, inconspicuous metasternal process, long and erect setation of the apical abdominal segments and in lacking empodial setae.
</paragraph>
<paragraph blockId="3.[264,1324,284,1870]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[312,464,883,909]" pageId="3" pageNumber="4">Description.</emphasis>
Length
<quantity box="[565,708,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" pageId="3" pageNumber="4" unit="mm" value="1.85" valueMax="2.1" valueMin="1.6">1.6­2.1 mm</quantity>
. Head reddish brown to dark brown; pronotum yellow to reddish yellow, lighter than head; elytra from brownish yellow to brown, with yellow apical (
<number box="[404,415,964,990]" pageId="3" pageNumber="4" value="1.0">1</number>
/
<number box="[427,442,964,990]" pageId="3" pageNumber="4" value="3.0">3</number>
to
<number box="[482,493,964,990]" pageId="3" pageNumber="4" value="1.0">1</number>
/
<number box="[504,519,964,990]" pageId="3" pageNumber="4" value="5.0">5</number>
) portion; abdominal segments III­V and VII­VIII yellow to reddish yellow; abdominal tergum VI darker, reddish brown to dark brown; antennae uniformly yellow or with darker articles
<number box="[619,676,1044,1070]" pageId="3" pageNumber="4" value="6.5" valueMax="11.0" valueMin="2.0">2­11</number>
; maxillar palpi uniformly yellow or with darker article
<number box="[264,281,1084,1110]" pageId="3" pageNumber="4" value="3.0">3</number>
; legs yellow to reddish yellow. Body form parallel­sided.
</paragraph>
<paragraph blockId="3.[264,1324,284,1870]" pageId="3" pageNumber="4">
Head transverse, eyes as long as temples (seen from above); infraorbital carina complete. Antennal articles
<number box="[545,560,1164,1190]" pageId="3" pageNumber="4" value="2.0">2</number>
and
<number box="[619,634,1164,1190]" pageId="3" pageNumber="4" value="3.0">3</number>
of equal length,
<number box="[835,874,1164,1190]" pageId="3" pageNumber="4" value="4.5" valueMax="5.0" valueMin="4.0">4­5</number>
subquadrate,
<number box="[1040,1093,1164,1190]" pageId="3" pageNumber="4" value="8.0" valueMax="10.0" valueMin="6.0">6­10</number>
slightly transverse, apical article as long as
<number box="[542,557,1204,1230]" pageId="3" pageNumber="4" value="9.0">9</number>
and
<number box="[613,642,1204,1230]" pageId="3" pageNumber="4" value="10.0">10</number>
combined, without coeloconic sensilla. Labrum (
<figureCitation box="[1220,1314,1204,1230]" captionStart="FIGURES 1­12" captionStartId="6.[264,383,1719,1743]" captionTargetBox="[340,1243,331,1659]" captionTargetId="figure@6.[337,1250,329,1666]" captionTargetPageId="6" captionText="FIGURES 1­12. Mouthparts of Tomoglossa decora (Casey) (Vermont). 1, left mandible, ventral view; 2, left mandible, dorsal view; 3, right mandible, dorsal view; 4, left maxilla, ventral view; 5, right galea, ventral view; 6, right galea, dorsal view; 7, left lacinia, dorsal view; 8, labrum; 9, epipharynx; 10, prementum; 11, hypopharynx; 12, mentum. Scale bar 0.1 mm (1­9, 12), 0.07 mm (11), 0.05 mm (10)." httpUri="https://zenodo.org/record/155537/files/figure.png" pageId="3" pageNumber="4">
Fig.
<number box="[1274,1314,1204,1230]" pageId="3" pageNumber="4" value="8.5" valueMax="9.0" valueMin="8.0">8­9</number>
</figureCitation>
) transverse and narrow, with straight anterior margin. Mandibles (
<figureCitation box="[1044,1150,1244,1270]" captionStart="FIGURES 1­12" captionStartId="6.[264,383,1719,1743]" captionTargetBox="[340,1243,331,1659]" captionTargetId="figure@6.[337,1250,329,1666]" captionTargetPageId="6" captionText="FIGURES 1­12. Mouthparts of Tomoglossa decora (Casey) (Vermont). 1, left mandible, ventral view; 2, left mandible, dorsal view; 3, right mandible, dorsal view; 4, left maxilla, ventral view; 5, right galea, ventral view; 6, right galea, dorsal view; 7, left lacinia, dorsal view; 8, labrum; 9, epipharynx; 10, prementum; 11, hypopharynx; 12, mentum. Scale bar 0.1 mm (1­9, 12), 0.07 mm (11), 0.05 mm (10)." httpUri="https://zenodo.org/record/155537/files/figure.png" pageId="3" pageNumber="4">
Figs.
<number box="[1112,1150,1244,1270]" pageId="3" pageNumber="4" value="2.0" valueMax="3.0" valueMin="1.0">1­3</number>
</figureCitation>
) symmetrical, slender, sickle­shaped, with a small medial tooth; ventral molar area without patches of denticles, dorsal molar region without visible “velvety patch” (
<number box="[1000,1047,1324,1350]" pageId="3" pageNumber="4" value="400.0">400</number>
x) (Figs.
<quantity metricMagnitude="0" metricUnit="m" metricValue="5.822188" metricValueMax="5.872988" metricValueMin="5.771388" pageId="3" pageNumber="4" unit="in" value="229.22" valueMax="231.22" valueMin="227.22">227.22­231.22 in</quantity>
<bibRefCitation author="Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[293,579,1364,1390]" editor="Arnett, R. H. &amp; Thomas, M. C." journalOrPublisher="CRC Press, Boca Raton" pageId="3" pageNumber="4" pagination="272 - 418" refId="ref7542" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.) American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, pp. 272 - 418." title="Staphylinidae Latreille, 1802" type="book chapter" volumeTitle="American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia" year="2000">
Newton
<emphasis box="[392,450,1364,1389]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
<number box="[457,520,1364,1390]" pageId="3" pageNumber="4" value="2000.0">2000</number>
:
<number box="[530,579,1364,1390]" pageId="3" pageNumber="4" value="309.0">309</number>
</bibRefCitation>
). Maxilla (
<figureCitation box="[705,810,1364,1390]" captionStart="FIGURES 1­12" captionStartId="6.[264,383,1719,1743]" captionTargetBox="[340,1243,331,1659]" captionTargetId="figure@6.[337,1250,329,1666]" captionTargetPageId="6" captionText="FIGURES 1­12. Mouthparts of Tomoglossa decora (Casey) (Vermont). 1, left mandible, ventral view; 2, left mandible, dorsal view; 3, right mandible, dorsal view; 4, left maxilla, ventral view; 5, right galea, ventral view; 6, right galea, dorsal view; 7, left lacinia, dorsal view; 8, labrum; 9, epipharynx; 10, prementum; 11, hypopharynx; 12, mentum. Scale bar 0.1 mm (1­9, 12), 0.07 mm (11), 0.05 mm (10)." httpUri="https://zenodo.org/record/155537/files/figure.png" pageId="3" pageNumber="4">
Figs.
<number box="[770,810,1364,1390]" pageId="3" pageNumber="4" value="5.5" valueMax="7.0" valueMin="4.0">4­7</number>
</figureCitation>
) with galea as long as lacinia, apical lobe of galea covered with numerous fine and short setae; interior margin of galea with row of setae subapically; apical third of lacinia with row of closely spaced spines, middle third produced medially and covered with numerous setae. Labial palpi with three articles; ligula divided into two completely separate lobes; medial pseudopore field of prementum with
<number box="[326,365,1564,1590]" pageId="3" pageNumber="4" value="4.5" valueMax="5.0" valueMin="4.0">4­5</number>
pseudopores (
<bibRefCitation author="Sawada, K." box="[540,703,1564,1590]" pageId="3" pageNumber="4" pagination="23 - 64" refId="ref7825" refString="Sawada, K. (1970) Aleocharinae (Staphylinidae, Coleoptera) of the IBP-Station in the Shiga Heights, Central Japan (I). Bulletin of the National Science Museum, 13 (1), 23 - 64." type="journal article" year="1970">Sawada 1970</bibRefCitation>
,
<bibRefCitation author="Sawada, K." box="[718,780,1564,1590]" pageId="3" pageNumber="4" pagination="31 - 59" refId="ref7868" refString="Sawada, K. (1972) Methodological Research in the Taxonomy of Aleocharinae. Contributions from the Biological Laboratory, Kyoto University, 24 (1), 31 - 59." type="journal article" year="1972">1972</bibRefCitation>
), lateral areas with pair of twin pores, single spinose pore and
<number box="[468,507,1604,1630]" pageId="3" pageNumber="4" value="5.0" valueMax="6.0" valueMin="4.0">4­6</number>
irregular pseudopores (
<figureCitation box="[785,872,1604,1630]" captionStart="FIGURES 1­12" captionStartId="6.[264,383,1719,1743]" captionTargetBox="[340,1243,331,1659]" captionTargetId="figure@6.[337,1250,329,1666]" captionTargetPageId="6" captionText="FIGURES 1­12. Mouthparts of Tomoglossa decora (Casey) (Vermont). 1, left mandible, ventral view; 2, left mandible, dorsal view; 3, right mandible, dorsal view; 4, left maxilla, ventral view; 5, right galea, ventral view; 6, right galea, dorsal view; 7, left lacinia, dorsal view; 8, labrum; 9, epipharynx; 10, prementum; 11, hypopharynx; 12, mentum. Scale bar 0.1 mm (1­9, 12), 0.07 mm (11), 0.05 mm (10)." httpUri="https://zenodo.org/record/155537/files/figure.png" pageId="3" pageNumber="4">
Fig.
<number box="[839,872,1604,1630]" pageId="3" pageNumber="4" value="10.0">10</number>
</figureCitation>
). Mentum (
<figureCitation box="[1009,1093,1604,1630]" captionStart="FIGURES 1­12" captionStartId="6.[264,383,1719,1743]" captionTargetBox="[340,1243,331,1659]" captionTargetId="figure@6.[337,1250,329,1666]" captionTargetPageId="6" captionText="FIGURES 1­12. Mouthparts of Tomoglossa decora (Casey) (Vermont). 1, left mandible, ventral view; 2, left mandible, dorsal view; 3, right mandible, dorsal view; 4, left maxilla, ventral view; 5, right galea, ventral view; 6, right galea, dorsal view; 7, left lacinia, dorsal view; 8, labrum; 9, epipharynx; 10, prementum; 11, hypopharynx; 12, mentum. Scale bar 0.1 mm (1­9, 12), 0.07 mm (11), 0.05 mm (10)." httpUri="https://zenodo.org/record/155537/files/figure.png" pageId="3" pageNumber="4">
Fig.
<number box="[1063,1093,1604,1630]" pageId="3" pageNumber="4" value="12.0">12</number>
</figureCitation>
) with anterior margin concave, posteriorly fused with submentum.
</paragraph>
<paragraph blockId="3.[264,1324,284,1870]" lastBlockId="4.[264,1324,284,1670]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Pronotum transverse, broadest slightly in front of middle, sides broadly rounded, anterior margin straight, anterior and posterior angles rounded; posterior margin convex; surface covered with short microsetae directed posteriorly in both midline and lateral areas (
<typeStatus box="[272,332,1804,1830]" pageId="3" pageNumber="4">Type</typeStatus>
VI,
<bibRefCitation author="Benick, G. &amp; Lohse, G. A." box="[385,646,1804,1830]" pageId="3" pageNumber="4" pagination="72 - 220" refId="ref6982" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.) Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">
Benick &amp; Lohse
<number box="[586,646,1804,1830]" pageId="3" pageNumber="4" value="1974.0">1974</number>
</bibRefCitation>
); macrosetae very short, limited to one macroseta at each of anterior and posterior angles and one on each lateral margin; hypomera fully visible in lateral aspect. Posterior margin of elytra straight. Wings present, fully developed. Mesosternal process short and wide, extending
<number box="[776,786,324,350]" pageId="4" pageNumber="5" value="1.0">1</number>
/
<number box="[797,814,324,350]" pageId="4" pageNumber="5" value="4.0">4</number>
length of mesocoxal cavities, metasternal process almost non­existent, anterior margin of metasternum only slightly convex medially (
<figureCitation box="[273,361,404,430]" captionStart="FIGURES 13 ­ 20" captionStartId="7.[264,383,1766,1790]" captionTargetBox="[351,1233,333,1743]" captionTargetId="figure@7.[345,1241,326,1749]" captionTargetPageId="7" captionText="FIGURES 13 ­ 20. Details of Tomoglossa decora (Casey) (Vermont (13 ­ 15, 20), Pennsylvania (16 ­ 18), lectotype (19 )). 13, pronotum; 14, right metatarsus, posterior view; 15, mesometathorax, ventral view; 16, apical margin of female sternum VIII; 17, female tergum VIII; 18, female sternum VIII; 19, male tergum VIII; 20, male sternum VIII. Scale bar 0.2 mm (13 ­ 15, 17 ­ 20), 0.1 mm (16)." httpUri="https://zenodo.org/record/155538/files/figure.png" pageId="4" pageNumber="5">
Fig.
<number box="[329,361,404,430]" pageId="4" pageNumber="5" value="15.0">15</number>
</figureCitation>
); mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about
<number box="[1062,1081,444,470]" pageId="4" pageNumber="5" value="6.0">6</number>
:
<number box="[1086,1123,444,470]" pageId="4" pageNumber="5" value="19.0">19</number>
:
<number box="[1127,1139,444,470]" pageId="4" pageNumber="5" value="1.0">1</number>
; mesocoxal cavities margined posteriorly; mesocoxae narrowly separated. Mesotibia with very short median macroseta (shorter than tibia width). Tarsal segmentation
<number box="[1061,1102,524,550]" pageId="4" pageNumber="5" value="4.5" valueMax="5.0" valueMin="4.0">4­5</number>
­
<number box="[1112,1127,524,550]" pageId="4" pageNumber="5" value="5.0">5</number>
; metatarsal segment
<number box="[330,344,564,590]" pageId="4" pageNumber="5" value="1.0">1</number>
longer than segment
<number box="[597,612,564,590]" pageId="4" pageNumber="5" value="2.0">2</number>
(
<figureCitation box="[627,715,564,590]" captionStart="FIGURES 13 ­ 20" captionStartId="7.[264,383,1766,1790]" captionTargetBox="[351,1233,333,1743]" captionTargetId="figure@7.[345,1241,326,1749]" captionTargetPageId="7" captionText="FIGURES 13 ­ 20. Details of Tomoglossa decora (Casey) (Vermont (13 ­ 15, 20), Pennsylvania (16 ­ 18), lectotype (19 )). 13, pronotum; 14, right metatarsus, posterior view; 15, mesometathorax, ventral view; 16, apical margin of female sternum VIII; 17, female tergum VIII; 18, female sternum VIII; 19, male tergum VIII; 20, male sternum VIII. Scale bar 0.2 mm (13 ­ 15, 17 ­ 20), 0.1 mm (16)." httpUri="https://zenodo.org/record/155538/files/figure.png" pageId="4" pageNumber="5">
Fig.
<number box="[682,715,564,590]" pageId="4" pageNumber="5" value="14.0">14</number>
</figureCitation>
). No empodial setae present.
</paragraph>
<paragraph blockId="4.[264,1324,284,1670]" pageId="4" pageNumber="5">
Abdominal terga III­V with moderate transverse basal impressions. Tergum VII
<number box="[1287,1324,604,630]" pageId="4" pageNumber="5" value="1.6">1.6</number>
times longer than VI. Puncturation of terga III­V equally sparse, puncturation of terga VI­ VII sparser. Terga VII­VIII with numerous erect macrosetae and long semi­erect microsetae. Tergum VIII (
<figureCitation box="[476,616,724,750]" captionStart="FIGURES 13 ­ 20" captionStartId="7.[264,383,1766,1790]" captionTargetBox="[351,1233,333,1743]" captionTargetId="figure@7.[345,1241,326,1749]" captionTargetPageId="7" captionText="FIGURES 13 ­ 20. Details of Tomoglossa decora (Casey) (Vermont (13 ­ 15, 20), Pennsylvania (16 ­ 18), lectotype (19 )). 13, pronotum; 14, right metatarsus, posterior view; 15, mesometathorax, ventral view; 16, apical margin of female sternum VIII; 17, female tergum VIII; 18, female sternum VIII; 19, male tergum VIII; 20, male sternum VIII. Scale bar 0.2 mm (13 ­ 15, 17 ­ 20), 0.1 mm (16)." httpUri="https://zenodo.org/record/155538/files/figure.png" pageId="4" pageNumber="5">Figs. 17, 19</figureCitation>
,
<figureCitation box="[629,661,724,750]" captionStart="FIGURES 34 ­ 37" captionStartId="11.[264,383,1741,1765]" captionTargetBox="[344,1236,722,1679]" captionTargetId="figure@11.[340,1247,719,1683]" captionTargetPageId="11" captionText="FIGURES 34 ­ 37. Details of Tomoglossa ornatella (Casey) (lectotype). 34, female tergum VIII; 35, female sternum VIII; 36, spermatheca; 37, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (34 ­ 35), 0.1 mm (36 ­ 37)." httpUri="https://zenodo.org/record/155541/files/figure.png" pageId="4" pageNumber="5">34</figureCitation>
,
<figureCitation box="[672,704,724,750]" captionStart="FIGURES 38 ­ 42" captionStartId="13.[264,383,1771,1795]" captionTargetBox="[384,1200,958,1742]" captionTargetId="figure@13.[380,1207,943,1745]" captionTargetPageId="13" captionText="FIGURES 38 ­ 42. Details of Tomoglossa arizonica Gusarov, sp. nov. (holotype). 38, male tergum VIII; 39, male sternum VIII; 40, median lobe of aedeagus, lateral view; 41, median lobe of aedeagus, ventral view; 42, internal sac of aedeagus, in packed state. Scale bar 0.2 mm (38 ­ 39), 0.1 mm (40 ­ 42)." httpUri="https://zenodo.org/record/155542/files/figure.png" pageId="4" pageNumber="5">38</figureCitation>
,
<figureCitation box="[716,747,724,750]" captionStart="FIGURES 43 ­ 46" captionStartId="15.[264,383,1451,1475]" captionTargetBox="[450,1136,732,1414]" captionTargetId="figure@15.[447,1140,729,1424]" captionTargetPageId="15" captionText="FIGURES 43 ­ 46. Details of Tomoglossa floridana Gusarov, sp. nov. (holotype). 43, female tergum VIII; 44, female sternum VIII; 45, spermatheca; 46, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (43 ­ 44), 0.1 mm (45 ­ 46)." httpUri="https://zenodo.org/record/155543/files/figure.png" pageId="4" pageNumber="5">
<number box="[716,747,724,750]" pageId="4" pageNumber="5" value="43.0">43</number>
</figureCitation>
) in both sexes with basal row of long microsetae. Female sternum VIII (
<figureCitation box="[524,665,764,790]" captionStart="FIGURES 13 ­ 20" captionStartId="7.[264,383,1766,1790]" captionTargetBox="[351,1233,333,1743]" captionTargetId="figure@7.[345,1241,326,1749]" captionTargetPageId="7" captionText="FIGURES 13 ­ 20. Details of Tomoglossa decora (Casey) (Vermont (13 ­ 15, 20), Pennsylvania (16 ­ 18), lectotype (19 )). 13, pronotum; 14, right metatarsus, posterior view; 15, mesometathorax, ventral view; 16, apical margin of female sternum VIII; 17, female tergum VIII; 18, female sternum VIII; 19, male tergum VIII; 20, male sternum VIII. Scale bar 0.2 mm (13 ­ 15, 17 ­ 20), 0.1 mm (16)." httpUri="https://zenodo.org/record/155538/files/figure.png" pageId="4" pageNumber="5">Figs. 16, 18</figureCitation>
,
<figureCitation box="[677,709,764,790]" captionStart="FIGURES 34 ­ 37" captionStartId="11.[264,383,1741,1765]" captionTargetBox="[344,1236,722,1679]" captionTargetId="figure@11.[340,1247,719,1683]" captionTargetPageId="11" captionText="FIGURES 34 ­ 37. Details of Tomoglossa ornatella (Casey) (lectotype). 34, female tergum VIII; 35, female sternum VIII; 36, spermatheca; 37, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (34 ­ 35), 0.1 mm (36 ­ 37)." httpUri="https://zenodo.org/record/155541/files/figure.png" pageId="4" pageNumber="5">35</figureCitation>
,
<figureCitation box="[721,752,764,790]" captionStart="FIGURES 43 ­ 46" captionStartId="15.[264,383,1451,1475]" captionTargetBox="[450,1136,732,1414]" captionTargetId="figure@15.[447,1140,729,1424]" captionTargetPageId="15" captionText="FIGURES 43 ­ 46. Details of Tomoglossa floridana Gusarov, sp. nov. (holotype). 43, female tergum VIII; 44, female sternum VIII; 45, spermatheca; 46, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (43 ­ 44), 0.1 mm (45 ­ 46)." httpUri="https://zenodo.org/record/155543/files/figure.png" pageId="4" pageNumber="5">44</figureCitation>
) with row of apical microsetae (as in other Athetini).
</paragraph>
<paragraph blockId="4.[264,1324,284,1670]" pageId="4" pageNumber="5">
Median lobe of aedeagus ventrally with very characteristic structure in the area of basal pore (
<figureCitation box="[404,543,884,910]" captionStart="FIGURES 21 ­ 28" captionStartId="8.[264,383,1724,1748]" captionTargetBox="[379,1211,497,1686]" captionTargetId="figure@8.[370,1217,489,1697]" captionTargetPageId="8" captionText="FIGURES 21 ­ 28. Genitalia and terminalia of Tomoglossa decora (Casey) (Vermont (21 ­ 25), Pennsylvania (26 ­ 28 )). 21, median lobe of aedeagus, lateral view; 22, median lobe of aedeagus, ventral view; 23, internal sac of aedeagus, in packed state; 24, deeply concave sclerite of internal sac; 25, copulatory piece; 26, spermatheca; 27, left portion of female sternum IX; 28, female accessory sclerite. U ­ umbilicus. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155539/files/figure.png" pageId="4" pageNumber="5">
Figs.
<number box="[472,502,884,910]" pageId="4" pageNumber="5" value="21.0">21</number>
­22
</figureCitation>
,
<figureCitation box="[557,630,884,910]" captionStart="FIGURES 38 ­ 42" captionStartId="13.[264,383,1771,1795]" captionTargetBox="[384,1200,958,1742]" captionTargetId="figure@13.[380,1207,943,1745]" captionTargetPageId="13" captionText="FIGURES 38 ­ 42. Details of Tomoglossa arizonica Gusarov, sp. nov. (holotype). 38, male tergum VIII; 39, male sternum VIII; 40, median lobe of aedeagus, lateral view; 41, median lobe of aedeagus, ventral view; 42, internal sac of aedeagus, in packed state. Scale bar 0.2 mm (38 ­ 39), 0.1 mm (40 ­ 42)." httpUri="https://zenodo.org/record/155542/files/figure.png" pageId="4" pageNumber="5">
40­
<number box="[599,630,884,910]" pageId="4" pageNumber="5" value="41.0">41</number>
</figureCitation>
). Internal sac with a deeply concave sclerite (
<figureCitation box="[1179,1318,884,910]" captionStart="FIGURES 21 ­ 28" captionStartId="8.[264,383,1724,1748]" captionTargetBox="[379,1211,497,1686]" captionTargetId="figure@8.[370,1217,489,1697]" captionTargetPageId="8" captionText="FIGURES 21 ­ 28. Genitalia and terminalia of Tomoglossa decora (Casey) (Vermont (21 ­ 25), Pennsylvania (26 ­ 28 )). 21, median lobe of aedeagus, lateral view; 22, median lobe of aedeagus, ventral view; 23, internal sac of aedeagus, in packed state; 24, deeply concave sclerite of internal sac; 25, copulatory piece; 26, spermatheca; 27, left portion of female sternum IX; 28, female accessory sclerite. U ­ umbilicus. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155539/files/figure.png" pageId="4" pageNumber="5">
Figs.
<number box="[1247,1278,884,910]" pageId="4" pageNumber="5" value="23.0">23</number>
­24
</figureCitation>
,
<figureCitation box="[264,297,924,950]" captionStart="FIGURES 38 ­ 42" captionStartId="13.[264,383,1771,1795]" captionTargetBox="[384,1200,958,1742]" captionTargetId="figure@13.[380,1207,943,1745]" captionTargetPageId="13" captionText="FIGURES 38 ­ 42. Details of Tomoglossa arizonica Gusarov, sp. nov. (holotype). 38, male tergum VIII; 39, male sternum VIII; 40, median lobe of aedeagus, lateral view; 41, median lobe of aedeagus, ventral view; 42, internal sac of aedeagus, in packed state. Scale bar 0.2 mm (38 ­ 39), 0.1 mm (40 ­ 42)." httpUri="https://zenodo.org/record/155542/files/figure.png" pageId="4" pageNumber="5">42</figureCitation>
). Copulatory piece (
<bibRefCitation author="Sawada, K." box="[534,694,924,950]" pageId="4" pageNumber="5" pagination="31 - 59" refId="ref7868" refString="Sawada, K. (1972) Methodological Research in the Taxonomy of Aleocharinae. Contributions from the Biological Laboratory, Kyoto University, 24 (1), 31 - 59." type="journal article" year="1972">
Sawada
<number box="[634,694,924,950]" pageId="4" pageNumber="5" value="1972.0">1972</number>
</bibRefCitation>
) without apical process (
<figureCitation box="[988,1087,924,950]" captionStart="FIGURES 21 ­ 28" captionStartId="8.[264,383,1724,1748]" captionTargetBox="[379,1211,497,1686]" captionTargetId="figure@8.[370,1217,489,1697]" captionTargetPageId="8" captionText="FIGURES 21 ­ 28. Genitalia and terminalia of Tomoglossa decora (Casey) (Vermont (21 ­ 25), Pennsylvania (26 ­ 28 )). 21, median lobe of aedeagus, lateral view; 22, median lobe of aedeagus, ventral view; 23, internal sac of aedeagus, in packed state; 24, deeply concave sclerite of internal sac; 25, copulatory piece; 26, spermatheca; 27, left portion of female sternum IX; 28, female accessory sclerite. U ­ umbilicus. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155539/files/figure.png" pageId="4" pageNumber="5">Figs. 25</figureCitation>
,
<figureCitation box="[1099,1133,924,950]" captionStart="FIGURES 38 ­ 42" captionStartId="13.[264,383,1771,1795]" captionTargetBox="[384,1200,958,1742]" captionTargetId="figure@13.[380,1207,943,1745]" captionTargetPageId="13" captionText="FIGURES 38 ­ 42. Details of Tomoglossa arizonica Gusarov, sp. nov. (holotype). 38, male tergum VIII; 39, male sternum VIII; 40, median lobe of aedeagus, lateral view; 41, median lobe of aedeagus, ventral view; 42, internal sac of aedeagus, in packed state. Scale bar 0.2 mm (38 ­ 39), 0.1 mm (40 ­ 42)." httpUri="https://zenodo.org/record/155542/files/figure.png" pageId="4" pageNumber="5">42</figureCitation>
).
</paragraph>
<paragraph blockId="4.[264,1324,284,1670]" box="[312,1103,964,990]" pageId="4" pageNumber="5">
Spermatheca with wide umbilicus (
<bibRefCitation author="Muona, J." box="[725,878,964,990]" pageId="4" pageNumber="5" pagination="17 - 24" refId="ref7510" refString="Muona, J. (1990) The Fennoscandian and Danish species of the genus Amischa Thomson (Coleoptera, Staphylinidae). Entomologisk Tidskrift, 111, 17 - 24." type="journal article" year="1990">
Muona
<number box="[818,878,964,990]" pageId="4" pageNumber="5" value="1990.0">1990</number>
</bibRefCitation>
) (
<figureCitation box="[902,1000,964,990]" captionStart="FIGURES 21 ­ 28" captionStartId="8.[264,383,1724,1748]" captionTargetBox="[379,1211,497,1686]" captionTargetId="figure@8.[370,1217,489,1697]" captionTargetPageId="8" captionText="FIGURES 21 ­ 28. Genitalia and terminalia of Tomoglossa decora (Casey) (Vermont (21 ­ 25), Pennsylvania (26 ­ 28 )). 21, median lobe of aedeagus, lateral view; 22, median lobe of aedeagus, ventral view; 23, internal sac of aedeagus, in packed state; 24, deeply concave sclerite of internal sac; 25, copulatory piece; 26, spermatheca; 27, left portion of female sternum IX; 28, female accessory sclerite. U ­ umbilicus. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155539/files/figure.png" pageId="4" pageNumber="5">Figs. 26</figureCitation>
,
<figureCitation box="[1012,1044,964,990]" captionStart="FIGURES 34 ­ 37" captionStartId="11.[264,383,1741,1765]" captionTargetBox="[344,1236,722,1679]" captionTargetId="figure@11.[340,1247,719,1683]" captionTargetPageId="11" captionText="FIGURES 34 ­ 37. Details of Tomoglossa ornatella (Casey) (lectotype). 34, female tergum VIII; 35, female sternum VIII; 36, spermatheca; 37, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (34 ­ 35), 0.1 mm (36 ­ 37)." httpUri="https://zenodo.org/record/155541/files/figure.png" pageId="4" pageNumber="5">36</figureCitation>
,
<figureCitation box="[1056,1089,964,990]" captionStart="FIGURES 43 ­ 46" captionStartId="15.[264,383,1451,1475]" captionTargetBox="[450,1136,732,1414]" captionTargetId="figure@15.[447,1140,729,1424]" captionTargetPageId="15" captionText="FIGURES 43 ­ 46. Details of Tomoglossa floridana Gusarov, sp. nov. (holotype). 43, female tergum VIII; 44, female sternum VIII; 45, spermatheca; 46, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (43 ­ 44), 0.1 mm (45 ­ 46)." httpUri="https://zenodo.org/record/155543/files/figure.png" pageId="4" pageNumber="5">
<number box="[1056,1089,964,990]" pageId="4" pageNumber="5" value="45.0">45</number>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="discussion">
<paragraph blockId="4.[264,1324,284,1670]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[312,452,1003,1029]" pageId="4" pageNumber="5">Discussion.</emphasis>
The
<typeStatus box="[535,585,1004,1030]" pageId="4" pageNumber="5">type</typeStatus>
species of
<emphasis box="[733,840,1005,1030]" italics="true" pageId="4" pageNumber="5">Noverota</emphasis>
(
<emphasis box="[862,1009,1005,1030]" italics="true" pageId="4" pageNumber="5">
N.
<taxonomicName authority="Casey, 1910" authorityName="Gusarov" authorityYear="2002" baseAuthorityName="Casey" baseAuthorityYear="1910" box="[902,1009,1005,1030]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ornatella">ornatella</taxonomicName>
</emphasis>
<bibRefCitation author="Casey, T. L." box="[1022,1172,1004,1030]" pageId="4" pageNumber="5" pagination="1 - 183" refId="ref7205" refString="Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera, 1, 1 - 183." type="journal article" year="1910">
Casey,
<number box="[1111,1172,1004,1030]" pageId="4" pageNumber="5" value="1910.0">1910</number>
</bibRefCitation>
, by original designation) and
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[469,607,1045,1070]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[469,607,1045,1070]" italics="true" pageId="4" pageNumber="5">Tomoglossa</emphasis>
</taxonomicName>
(
<taxonomicName authority="(Erichson, 1837))" authorityName="Erichson" authorityYear="1837" baseAuthorityName="Erichson" baseAuthorityYear="1837" box="[626,998,1044,1070]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="luteicornis">
<emphasis box="[626,782,1045,1070]" italics="true" pageId="4" pageNumber="5">T. luteicornis</emphasis>
(
<bibRefCitation author="Erichson, W. F." box="[798,979,1044,1070]" pageId="4" pageNumber="5" refId="ref7233" refString="Erichson, W. F. (1837 - 1839) Die Kafer der Mark Brandenburg. Band 1. F. H. Morin, Berlin, viii + 740 pp." type="book" year="1837">
Erichson,
<number box="[920,979,1044,1070]" pageId="4" pageNumber="5" value="1837.0">1837</number>
</bibRefCitation>
))
</taxonomicName>
differ only in body coloration (
<taxonomicName authorityName="Erichson" authorityYear="1837" baseAuthorityName="Erichson" baseAuthorityYear="1837" box="[327,481,1085,1110]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="luteicornis">
<emphasis box="[327,481,1085,1110]" italics="true" pageId="4" pageNumber="5">T. luteicornis</emphasis>
</taxonomicName>
being darker) and in minor details of genitalia (
<figureCitation box="[1038,1172,1084,1110]" captionStart="FIGURES 21 ­ 28" captionStartId="8.[264,383,1724,1748]" captionTargetBox="[379,1211,497,1686]" captionTargetId="figure@8.[370,1217,489,1697]" captionTargetPageId="8" captionText="FIGURES 21 ­ 28. Genitalia and terminalia of Tomoglossa decora (Casey) (Vermont (21 ­ 25), Pennsylvania (26 ­ 28 )). 21, median lobe of aedeagus, lateral view; 22, median lobe of aedeagus, ventral view; 23, internal sac of aedeagus, in packed state; 24, deeply concave sclerite of internal sac; 25, copulatory piece; 26, spermatheca; 27, left portion of female sternum IX; 28, female accessory sclerite. U ­ umbilicus. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155539/files/figure.png" pageId="4" pageNumber="5">
Figs.
<quantity box="[1104,1172,1084,1110]" metricMagnitude="-1" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.604000000000001" metricValueMin="5.3340000000000005" pageId="4" pageNumber="5" unit="in" value="23.5" valueMax="26.0" valueMin="21.0">21­26</quantity>
</figureCitation>
in this paper and Fig.
<quantity box="[375,453,1124,1150]" metricMagnitude="0" metricUnit="m" metricValue="3.1242" pageId="4" pageNumber="5" unit="in" value="123.0">123 in</quantity>
<bibRefCitation author="Lohse, G. A." box="[464,609,1124,1150]" pageId="4" pageNumber="5" pagination="185 - 243" refId="ref7412" refString="Lohse, G. A. (1989) Erganzungen und Berichtigungen zu Freude-Harde-Lohse &quot; Die Kafer Mitteleuropas &quot; Band 5 (1974). In: Lohse, G. A. &amp; Lucht, W. H. (Eds.) Die Kafer Mitteleuropas. 1. Supplementband mit Katalogteil. Goecke &amp; Evers Verlag, Krefeld, pp. 185 - 243." type="journal article" year="1989">
Lohse
<number box="[546,609,1124,1150]" pageId="4" pageNumber="5" value="1989.0">1989</number>
</bibRefCitation>
).
<collectingCountry box="[633,666,1124,1150]" name="American Samoa" pageId="4" pageNumber="5">As</collectingCountry>
a result,
<emphasis box="[784,892,1125,1150]" italics="true" pageId="4" pageNumber="5">Noverota</emphasis>
is placed in synonymy with
<taxonomicName authorityName="Kraatz" authorityYear="1856" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="5">Tomoglossa</emphasis>
</taxonomicName>
. Apparently Casey was not familiar with
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[823,962,1165,1190]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[823,962,1165,1190]" italics="true" pageId="4" pageNumber="5">Tomoglossa</emphasis>
</taxonomicName>
and when describing
<emphasis box="[1215,1322,1165,1190]" italics="true" pageId="4" pageNumber="5">Noverota</emphasis>
he compared it only with
<taxonomicName authorityName="Thomson" authorityYear="1858" box="[569,724,1205,1230]" class="Insecta" family="Staphylinidae" genus="Hydrosmecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[569,724,1205,1230]" italics="true" pageId="4" pageNumber="5">Hydrosmecta</emphasis>
</taxonomicName>
.
<bibRefCitation author="Casey, T. L." box="[739,896,1204,1230]" pageId="4" pageNumber="5" pagination="1 - 183" refId="ref7205" refString="Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera, 1, 1 - 183." type="journal article" year="1910">
Casey (
<number box="[827,888,1204,1230]" pageId="4" pageNumber="5" value="1910.0">1910</number>
)
</bibRefCitation>
included seven species in the genus
<emphasis box="[264,371,1245,1270]" italics="true" pageId="4" pageNumber="5">Noverota</emphasis>
. Three of these species belong to
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[815,954,1245,1270]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[815,954,1245,1270]" italics="true" pageId="4" pageNumber="5">Tomoglossa</emphasis>
</taxonomicName>
. They are redescribed below.
<emphasis box="[264,479,1285,1310]" italics="true" pageId="4" pageNumber="5">Noverota clemens</emphasis>
<bibRefCitation author="Casey, T. L." box="[493,645,1284,1310]" pageId="4" pageNumber="5" pagination="1 - 183" refId="ref7205" refString="Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera, 1, 1 - 183." type="journal article" year="1910">
Casey,
<number box="[583,645,1284,1310]" pageId="4" pageNumber="5" value="1910.0">1910</number>
</bibRefCitation>
,
<emphasis box="[663,794,1285,1310]" italics="true" pageId="4" pageNumber="5">N. finitima</emphasis>
<bibRefCitation author="Casey, T. L." box="[807,959,1284,1310]" pageId="4" pageNumber="5" pagination="1 - 183" refId="ref7205" refString="Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera, 1, 1 - 183." type="journal article" year="1910">
Casey,
<number box="[897,959,1284,1310]" pageId="4" pageNumber="5" value="1910.0">1910</number>
</bibRefCitation>
, and
<emphasis box="[1032,1161,1285,1310]" italics="true" pageId="4" pageNumber="5">N. scenica</emphasis>
<bibRefCitation author="Casey, T. L." box="[1174,1323,1284,1310]" pageId="4" pageNumber="5" pagination="1 - 183" refId="ref7205" refString="Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera, 1, 1 - 183." type="journal article" year="1910">
Casey,
<number box="[1264,1323,1284,1310]" pageId="4" pageNumber="5" value="1910.0">1910</number>
</bibRefCitation>
belong to
<taxonomicName authority="Mulsant &amp; Rey, 1873" authorityName="Mulsant &amp; Rey" authorityYear="1873" box="[387,773,1324,1350]" class="Insecta" family="Staphylinidae" genus="Philhygra" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[387,504,1325,1350]" italics="true" pageId="4" pageNumber="5">Philhygra</emphasis>
Mulsant &amp; Rey,
<number box="[714,773,1324,1350]" pageId="4" pageNumber="5" value="1873.0">1873</number>
</taxonomicName>
and will be redescribed elsewhere. The taxonomic position of
<emphasis box="[484,637,1365,1390]" italics="true" pageId="4" pageNumber="5">N. personata</emphasis>
<bibRefCitation author="Casey, T. L." box="[646,791,1364,1390]" pageId="4" pageNumber="5" pagination="1 - 183" refId="ref7205" refString="Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. Memoirs on the Coleoptera, 1, 1 - 183." type="journal article" year="1910">
Casey,
<number box="[732,791,1364,1390]" pageId="4" pageNumber="5" value="1910.0">1910</number>
</bibRefCitation>
is not clear, but it does not belong to
<taxonomicName authorityName="Kraatz" authorityYear="1856" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="5">Tomoglossa</emphasis>
</taxonomicName>
. The species is known from a single female
<typeStatus box="[868,919,1404,1430]" pageId="4" pageNumber="5">type</typeStatus>
and a detailed study of its mouthparts is not possible. When additional specimens of this species are found the position of the species can be clarified. Two new species of
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[848,986,1485,1510]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[848,986,1485,1510]" italics="true" pageId="4" pageNumber="5">Tomoglossa</emphasis>
</taxonomicName>
are described below. Thus, five valid Nearctic species of
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[610,749,1525,1550]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[610,749,1525,1550]" italics="true" pageId="4" pageNumber="5">Tomoglossa</emphasis>
</taxonomicName>
are currently known.
</paragraph>
<paragraph blockId="4.[264,1324,284,1670]" pageId="4" pageNumber="5">
In the key by Ashe (
<bibRefCitation author="Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[555,788,1564,1590]" editor="Arnett, R. H. &amp; Thomas, M. C." journalOrPublisher="CRC Press, Boca Raton" pageId="4" pageNumber="5" pagination="272 - 418" refId="ref7542" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Eds.) American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, pp. 272 - 418." title="Staphylinidae Latreille, 1802" type="book chapter" volumeTitle="American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia" year="2000">
Newton
<emphasis box="[658,719,1565,1590]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
<number box="[727,788,1564,1590]" pageId="4" pageNumber="5" value="2000.0">2000</number>
</bibRefCitation>
) the specimens belonging to
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[1137,1275,1565,1590]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[1137,1275,1565,1590]" italics="true" pageId="4" pageNumber="5">Tomoglossa</emphasis>
</taxonomicName>
run to the second half of the couplet
<number box="[650,683,1604,1630]" pageId="4" pageNumber="5" value="58.0">58</number>
. In the key by
<bibRefCitation author="Benick, G. &amp; Lohse, G. A." box="[857,1155,1604,1630]" pageId="4" pageNumber="5" pagination="72 - 220" refId="ref6982" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.) Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">
Benick and Lohse (
<number box="[1085,1146,1604,1630]" pageId="4" pageNumber="5" value="1974.0">1974</number>
)
</bibRefCitation>
the same specimens easily reach
<taxonomicName authorityName="Kraatz" authorityYear="1856" box="[501,640,1645,1670]" class="Insecta" family="Staphylinidae" genus="Tomoglossa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[501,640,1645,1670]" italics="true" pageId="4" pageNumber="5">Tomoglossa</emphasis>
</taxonomicName>
at couplet
<number box="[769,786,1644,1670]" pageId="4" pageNumber="5" value="8.0">8</number>
.
</paragraph>
</subSubSection>
</treatment>
</document>