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<document id="0952015703B950DD1D3CF89738658E4E" ID-CLB-Dataset="31260" ID-DOI="10.11646/zootaxa.4359.1.1" ID-GBIF-Dataset="a5cbd5ef-7f49-4a94-a2f4-61adcc8c8d01" ID-ISSN="1175-5326" ID-Zenodo-Dep="1069168" ID-ZooBank="8D06174D-B19F-4B5C-B9B0-DA74E6D43C75" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512122415347" checkinUser="plazi" docAuthor="Tibiriçá, Yara, Pola, Marta &amp; Cervera, Juan Lucas" docDate="2017" docId="03EB87A2FFDBFF8F9790FDFAFD18FAC5" docLanguage="en" docName="zootaxa.4359.1.1.pdf.imf" docOrigin="Zootaxa 4359 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pleurobranchus forskalii Ruppel &amp; Leuckart 1828" docType="treatment" docVersion="8" lastPageNumber="12" masterDocId="FFD2FFDAFFD0FF849707FF83FFB7FFC0" masterDocTitle="Astonishing diversity revealed: an annotated and illustrated inventory of Nudipleura (Gastropoda: Heterobranchia) from Mozambique" masterLastPageNumber="133" masterPageNumber="1" pageNumber="12" updateTime="1698506029818" updateUser="ExternalLinkService">
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<mods:title id="29A6F1F312134D29506232C46CA12A19">Astonishing diversity revealed: an annotated and illustrated inventory of Nudipleura (Gastropoda: Heterobranchia) from Mozambique</mods:title>
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<mods:namePart id="79930387E4EEB6C291BA0C892269A5FA">Tibiriçá, Yara</mods:namePart>
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<mods:namePart id="C7707E7CB328CAC2294AEE47B67FCDFF">Pola, Marta</mods:namePart>
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<mods:namePart id="F84A1CE94504F54B4F8BF4FC720CFDB6">Cervera, Juan Lucas</mods:namePart>
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<mods:date id="0C9B747FF6EC2DA2E31B07591E35E7AD">2017</mods:date>
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<treatment id="03EB87A2FFDBFF8F9790FDFAFD18FAC5" ID-DOI="http://doi.org/10.5281/zenodo.6025740" ID-GBIF-Taxon="137581732" ID-Zenodo-Dep="6025740" LSID="urn:lsid:plazi:treatment:03EB87A2FFDBFF8F9790FDFAFD18FAC5" httpUri="http://treatment.plazi.org/id/03EB87A2FFDBFF8F9790FDFAFD18FAC5" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="C358653FFFDBFF8F9790FDFAFE8DFD74" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BFD36B4FFDBFF8F9790FDFAFCBAFD53" blockId="11.[151,781,633,693]" box="[151,781,633,659]" pageId="11" pageNumber="12">
<heading id="D0B581D8FFDBFF8F9790FDFAFCBAFD53" bold="true" box="[151,781,633,659]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="4C424D37FFDBFF8F9790FDFAFCBAFD53" authority="Ruppel &amp; Leuckart, 1828" authorityName="Ruppel &amp; Leuckart" authorityYear="1828" box="[151,781,633,659]" class="Gastropoda" family="Pleurobranchidae" genus="Pleurobranchus" kingdom="Animalia" order="Pleurobranchomorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="forskalii">
<emphasis id="B936EAA6FFDBFF8F9790FDFAFCBAFD53" bold="true" box="[151,781,633,659]" pageId="11" pageNumber="12">
<emphasis id="B936EAA6FFDBFF8F9790FDFAFE70FD53" bold="true" box="[151,455,633,659]" italics="true" pageId="11" pageNumber="12">Pleurobranchus forskalii</emphasis>
<bibRefCitation id="EFD34B45FFDBFF8F96C9FDFAFCBAFD53" author="Ruppel" box="[462,781,633,659]" pageId="11" pageNumber="12" refString="Ruppell, W. P. E. S. &amp; Leuckart, F. S. (1828) Mollusca. In: Atlas zu der Reise im nordlichen Afrika. v on Eduard Ruppell. 1. Abth. Zoologie. 5. Neue wirbellose. Thiere des Rothen Meers, Frankfurt, pp. 15 - 47, pls. 1 - 12." type="book chapter" year="1828">Rüppel &amp; Leuckart, 1828</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFD36B4FFDBFF8F9790FD1FFE8DFD74" blockId="11.[151,781,633,693]" box="[151,314,668,693]" pageId="11" pageNumber="12">
(
<figureCitation id="13792A31FFDBFF8F9799FD1FFE85FD74" box="[158,306,668,693]" captionStart="FIGURE 2" captionStartId="10.[151,250,1879,1901]" captionTargetBox="[218,1354,195,1865]" captionTargetId="figure@10.[218,1354,195,1865]" captionTargetPageId="10" captionText="FIGURE 2. A, Berthella martensi (MB28-004607). BD, Berthellina delicata species complex (MB28-005057, MB28- 004578, MB28-005060, respectively). E, Berthellina sp. (MB28-005016). F, Pleurobranchaea brockii (in situ). G, Pleurobranchus albiguttatus (MB28-004603). HI, Pleurobranchus forskalii (MB28-004434 and MHN-YT1655, respectively). J, Pleurobranchus grandis (in situ). KL, Pleurobranchus peronii (MHN-YT276 and MB28-005042, respectively)." httpUri="https://zenodo.org/record/1069172/files/figure.png" pageId="11" pageNumber="12">Figure 2 HI</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C358653FFFDBFF8F9790FD67FA91FCA5" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BFD36B4FFDBFF8F9790FD67FE7AFC82" blockId="11.[151,1437,740,1285]" pageId="11" pageNumber="12">
<emphasis id="B936EAA6FFDBFF8F9790FD67FE36FD3D" bold="true" box="[151,385,740,765]" pageId="11" pageNumber="12">Material examined.</emphasis>
Four specimens. MB28-004434, ZRP,
<date id="FFFC1074FFDBFF8F943AFD67FC61FD3C" box="[829,982,740,764]" pageId="11" pageNumber="12" value="2011-12-10">
10 Dec.
<quantity id="4CBA9B51FFDBFF8F949AFD67FC61FD3C" box="[925,982,740,764]" metricMagnitude="3" metricUnit="m" metricValue="2.0111" pageId="11" pageNumber="12" unit="m" value="2011.1">2011</quantity>
</date>
, 1m,
<quantity id="4CBA9B51FFDBFF8F9311FD67FBD6FD3D" box="[1046,1121,740,765]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="mm" value="35.0">35mm</quantity>
; MHN-YT596, BL,
<date id="FFFC1074FFDBFF8F924EFD67FF65FCDF" pageId="11" pageNumber="12" value="2012-10-12">
12 Oct.
<quantity id="4CBA9B51FFDBFF8F9790FC84FF65FCDF" box="[151,210,775,799]" metricMagnitude="3" metricUnit="m" metricValue="2.0122" pageId="11" pageNumber="12" unit="m" value="2012.2">2012</quantity>
</date>
, 2m,
<quantity id="4CBA9B51FFDBFF8F961FFC84FEC6FCDF" box="[280,369,775,799]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="12" unit="mm" value="120.0">120mm</quantity>
; MB28-004904, MIF,
<date id="FFFC1074FFDBFF8F957BFC84FCAFFCDF" box="[636,792,774,799]" pageId="11" pageNumber="12" value="2016-06-11">
11 Jun.
<quantity id="4CBA9B51FFDBFF8F95DAFC85FCAFFCDF" box="[733,792,774,799]" metricMagnitude="3" metricUnit="m" metricValue="2.0163" pageId="11" pageNumber="12" unit="m" value="2016.3">2016</quantity>
</date>
, 3m,
<quantity id="4CBA9B51FFDBFF8F9458FC84FC1DFCDF" box="[863,938,775,799]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="11" pageNumber="12" unit="mm" value="88.0">88mm</quantity>
; MHN-YT1655,
<collectionCode id="ED53AE71FFDBFF8F9372FC84FB04FCDE" box="[1141,1203,775,798]" country="Rwanda" httpUri="http://grbio.org/cool/jbr1-cqb0" name="Institute of Scientific and Technological Research (IRST)" pageId="11" pageNumber="12">NHR</collectionCode>
,
<date id="FFFC1074FFDBFF8F93C4FC84FAD6FCDF" box="[1219,1377,774,799]" pageId="11" pageNumber="12" value="2016-10-23">
23 Oct.
<quantity id="4CBA9B51FFDBFF8F9221FC85FAD6FCDF" box="[1318,1377,774,799]" metricMagnitude="3" metricUnit="m" metricValue="2.0168" pageId="11" pageNumber="12" unit="m" value="2016.8">2016</quantity>
</date>
, 8m,
<quantity id="4CBA9B51FFDBFF8F9790FCA9FF5BFC82" box="[151,236,810,834]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="mm" value="200.0">200mm</quantity>
(crawling on sand).
</paragraph>
<paragraph id="8BFD36B4FFDBFF8F97C0FCCFFA91FCA5" blockId="11.[151,1437,740,1285]" box="[199,1318,844,869]" pageId="11" pageNumber="12">
<emphasis id="B936EAA6FFDBFF8F97C0FCCFFE82FCA5" bold="true" box="[199,309,844,869]" pageId="11" pageNumber="12">Habitats.</emphasis>
Subtropical tidal reefs, rocky reefs and seagrass and tropical coral bommies and seagrass.
</paragraph>
</subSubSection>
<subSubSection id="C358653FFFDBFF8F97C0FCEDFB57FC2F" pageId="11" pageNumber="12" type="description">
<paragraph id="8BFD36B4FFDBFF8F97C0FCEDFC36FC46" blockId="11.[151,1437,740,1285]" box="[199,897,878,903]" pageId="11" pageNumber="12">
<emphasis id="B936EAA6FFDBFF8F97C0FCEDFED3FC47" bold="true" box="[199,356,878,903]" pageId="11" pageNumber="12">Occurrences.</emphasis>
Zavora, Barra, Mozambique Island and Nuarro.
</paragraph>
<paragraph id="8BFD36B4FFDBFF8F97C0FC12FB57FC2F" blockId="11.[151,1437,740,1285]" pageId="11" pageNumber="12">
<emphasis id="B936EAA6FFDBFF8F97C0FC12FE59FC6A" bold="true" box="[199,494,913,938]" pageId="11" pageNumber="12">Geographic distribution.</emphasis>
Indo-west Pacific.
<collectingCountry id="F3557624FFDBFF8F95C0FC11FCBDFC69" box="[711,778,914,937]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
,
<collectingCountry id="F3557624FFDBFF8F9412FC11FC47FC6A" box="[789,1008,913,938]" name="Papua New Guinea" pageId="11" pageNumber="12">
<collectingRegion id="4986F856FFDBFF8F9412FC11FCEEFC69" box="[789,857,914,937]" country="Indonesia" name="Papua Barat" pageId="11" pageNumber="12">Papua</collectingRegion>
New Guinea
</collectingCountry>
,
<collectingCountry id="F3557624FFDBFF8F94FDFC12FBD4FC6A" box="[1018,1123,913,938]" name="Australia" pageId="11" pageNumber="12">Australia</collectingCountry>
,
<collectingCountry id="F3557624FFDBFF8F9368FC12FB2EFC6A" box="[1135,1177,913,938]" name="Fiji" pageId="11" pageNumber="12">Fiji</collectingCountry>
,
<collectingCountry id="F3557624FFDBFF8F93A4FC12FAA4FC6A" box="[1187,1299,913,938]" name="Indonesia" pageId="11" pageNumber="12">Indonesia</collectingCountry>
,
<collectingCountry id="F3557624FFDBFF8F9219FC12FA2BFC6A" box="[1310,1436,913,938]" name="Philippines" pageId="11" pageNumber="12">Philippines</collectingCountry>
(Gosliner
<emphasis id="B936EAA6FFDBFF8F9600FC36FE88FC0D" box="[263,319,948,973]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2008;
<bibRefCitation id="EFD34B45FFDBFF8F968AFC37FD31FC0D" author="Goodheart" box="[397,646,948,973]" pageId="11" pageNumber="12" refString="Goodheart, J., Camacho-Garcia, Y., Padula, V., Schrodl, M., Cervera, J. L., Gosliner, T. M. &amp; Valdes, A. (2015) Systematics and biogeography of Pleurobranchus Cuvier, 1804, sea slugs (Heterobranchia: Nudipleura: Pleurobranchidae). Zoological Journal of the Linnean Society, 174, 322 - 362. https: // doi. org / 10.1111 / zoj. 12237" type="journal article" year="2015">
Goodheart
<emphasis id="B936EAA6FFDBFF8F950CFC36FDF4FC0D" box="[523,579,948,973]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2015
</bibRefCitation>
),
<collectingRegion id="4986F856FFDBFF8F959EFC37FD7AFC0D" box="[665,717,948,973]" country="Papua New Guinea" name="Gulf" pageId="11" pageNumber="12">Gulf</collectingRegion>
of
<collectingCountry id="F3557624FFDBFF8F95F5FC37FC81FC0C" box="[754,822,948,972]" name="Oman" pageId="11" pageNumber="12">Oman</collectingCountry>
(
<bibRefCitation id="EFD34B45FFDBFF8F9442FC37FB01FC0D" author="Fatemi" box="[837,1206,948,973]" pageId="11" pageNumber="12" refString="Fatemi, Y. &amp; Attaran-Fariman, G. (2015) Checklist of the opisthobranchs (Heterobranchia: Gastropoda) along the Iranian Coasts of the Gulf of Oman. Journal of Biodiversity Environmental Science, 6, 1 - 7." type="journal article" year="2015">Fatemi &amp; Attaran-Fariman 2015</bibRefCitation>
),
<collectingCountry id="F3557624FFDBFF8F93CEFC37FA9AFC0D" box="[1225,1325,948,973]" name="Thailand" pageId="11" pageNumber="12">Thailand</collectingCountry>
(
<bibRefCitation id="EFD34B45FFDBFF8F923CFC37FE87FC2F" author="Mehrota" pageId="11" pageNumber="12" refString="Mehrotra, R. &amp; Scott, C. (2015) Species inventory of sea slugs (Gastropoda: Heterobranchia) for Koh Tao, Thailand, with 25 first records fo Thai waters. Marine Biodiversity, 46, 761 - 771. https: // doi. org / 10.1007 / s 12526 - 015 - 0424 - 7" type="journal article" year="2015">Mehrota &amp; Scott 2015</bibRefCitation>
), Red Sea,
<collectingCountry id="F3557624FFDBFF8F96A9FC55FDA5FC2F" box="[430,530,982,1007]" name="Tanzania" pageId="11" pageNumber="12">Tanzania</collectingCountry>
(Gosliner
<emphasis id="B936EAA6FFDBFF8F958DFC54FD74FC2F" box="[650,707,982,1007]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2008;
<bibRefCitation id="EFD34B45FFDBFF8F9417FC55FBB1FC2F" author="Goodheart" box="[784,1030,982,1007]" pageId="11" pageNumber="12" refString="Goodheart, J., Camacho-Garcia, Y., Padula, V., Schrodl, M., Cervera, J. L., Gosliner, T. M. &amp; Valdes, A. (2015) Systematics and biogeography of Pleurobranchus Cuvier, 1804, sea slugs (Heterobranchia: Nudipleura: Pleurobranchidae). Zoological Journal of the Linnean Society, 174, 322 - 362. https: // doi. org / 10.1111 / zoj. 12237" type="journal article" year="2015">
Goodheart
<emphasis id="B936EAA6FFDBFF8F948AFC54FC72FC2F" box="[909,965,982,1007]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2015
</bibRefCitation>
) and
<collectingCountry id="F3557624FFDBFF8F9343FC55FB6CFC2F" box="[1092,1243,982,1007]" name="Mozambique" pageId="11" pageNumber="12">Mozambique</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C358653FFFDBFF8F97C0FC7AFD18FAC5" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BFD36B4FFDBFF8F97C0FC7AFD18FAC5" blockId="11.[151,1437,740,1285]" pageId="11" pageNumber="12">
<emphasis id="B936EAA6FFDBFF8F97C0FC7AFE8CFBD2" bold="true" box="[199,315,1017,1042]" pageId="11" pageNumber="12">Remarks.</emphasis>
<bibRefCitation id="EFD34B45FFDBFF8F9643FC7AFDE7FBD2" box="[324,592,1017,1042]" pageId="11" pageNumber="12" refString="Goodheart, J., Camacho-Garcia, Y., Padula, V., Schrodl, M., Cervera, J. L., Gosliner, T. M. &amp; Valdes, A. (2015) Systematics and biogeography of Pleurobranchus Cuvier, 1804, sea slugs (Heterobranchia: Nudipleura: Pleurobranchidae). Zoological Journal of the Linnean Society, 174, 322 - 362. https: // doi. org / 10.1111 / zoj. 12237" type="journal article">
Goodheart
<emphasis id="B936EAA6FFDBFF8F96C4FC79FE41FBD2" box="[451,502,1017,1042]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. (2015)
</bibRefCitation>
reviewed the genus
<taxonomicName id="4C424D37FFDBFF8F9439FC7AFC45FBD2" authorityName="Cuvier" authorityYear="1804" box="[830,1010,1017,1042]" class="Gastropoda" family="Pleurobranchidae" genus="Pleurobranchus" kingdom="Animalia" order="Pleurobranchomorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="B936EAA6FFDBFF8F9439FC7AFC45FBD2" box="[830,1010,1017,1042]" italics="true" pageId="11" pageNumber="12">Pleurobranchus</emphasis>
</taxonomicName>
based on molecular and morphology and observed that colour variation in this species is related to the ontogeny, where juveniles are lighter in colour than adults. Small specimens from
<collectingCountry id="F3557624FFDBFF8F9518FBBDFD04FB97" box="[543,691,1086,1111]" name="Mozambique" pageId="11" pageNumber="12">Mozambique</collectingCountry>
show complex colouration with several hues of brown mixed with small amounts of grey and white. The large and small tubercles often have a reddish dot at the top. A dark brown ring subtended by a whitish semi ring surrounds each large compound tubercle (
<figureCitation id="13792A31FFDBFF8F9322FB07FB3EFB5C" box="[1061,1161,1156,1181]" captionStart="FIGURE 2" captionStartId="10.[151,250,1879,1901]" captionTargetBox="[218,1354,195,1865]" captionTargetId="figure@10.[218,1354,195,1865]" captionTargetPageId="10" captionText="FIGURE 2. A, Berthella martensi (MB28-004607). BD, Berthellina delicata species complex (MB28-005057, MB28- 004578, MB28-005060, respectively). E, Berthellina sp. (MB28-005016). F, Pleurobranchaea brockii (in situ). G, Pleurobranchus albiguttatus (MB28-004603). HI, Pleurobranchus forskalii (MB28-004434 and MHN-YT1655, respectively). J, Pleurobranchus grandis (in situ). KL, Pleurobranchus peronii (MHN-YT276 and MB28-005042, respectively)." httpUri="https://zenodo.org/record/1069172/files/figure.png" pageId="11" pageNumber="12">Fig. 2 G</figureCitation>
). The base of the foot is translucent light-brown spotted with dark-brown, irregular marks. The larger specimen had a dark-purple mantle with partly open white rings surrounding each cluster of tubercles (
<figureCitation id="13792A31FFDBFF8F9492FB4AFC4EFB22" box="[917,1017,1225,1250]" captionStart="FIGURE 2" captionStartId="10.[151,250,1879,1901]" captionTargetBox="[218,1354,195,1865]" captionTargetId="figure@10.[218,1354,195,1865]" captionTargetPageId="10" captionText="FIGURE 2. A, Berthella martensi (MB28-004607). BD, Berthellina delicata species complex (MB28-005057, MB28- 004578, MB28-005060, respectively). E, Berthellina sp. (MB28-005016). F, Pleurobranchaea brockii (in situ). G, Pleurobranchus albiguttatus (MB28-004603). HI, Pleurobranchus forskalii (MB28-004434 and MHN-YT1655, respectively). J, Pleurobranchus grandis (in situ). KL, Pleurobranchus peronii (MHN-YT276 and MB28-005042, respectively)." httpUri="https://zenodo.org/record/1069172/files/figure.png" pageId="11" pageNumber="12">Fig. 2 H</figureCitation>
). In this morphotype, the base of the foot is the same colour as the rest of the mantle.
</paragraph>
</subSubSection>
</treatment>
</document>