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<mods:title id="305725A790EABFDE5B9224CFAA8057AA">Glaciambulata neumayeri gen. et sp. nov., a new Antarctic trachymedusa (Cnidaria: Hydrozoa), with a revision of the family Ptychogastriidae</mods:title>
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<mods:namePart id="6E5D938FD1995911A003AA63FF664D4D">Horia R. Galea</mods:namePart>
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<mods:namePart id="9A2265EBA30E776177A647BCD6CC2F07">Cornelia Roder</mods:namePart>
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<mods:namePart id="384ABB3C5858990A19D0AE6673472187">Christoph Walcher</mods:namePart>
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<mods:namePart id="356A1A1AF25E680CFA49AC1DF05A64F0">Marco Warmuth</mods:namePart>
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<mods:namePart id="E97E54C0FDA071C451DF7CFFFA60AC0E">Eberhard Kohlberg</mods:namePart>
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<mods:namePart id="BC5BF99778FAB38857FA7AB01F669894">Philipp F. Fischer</mods:namePart>
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<treatment id="03EB879BFF91FFA2FDFD0043FAFBE4DB" ID-DOI="http://doi.org/10.5281/zenodo.5628722" ID-GBIF-Taxon="127636786" ID-Zenodo-Dep="5628722" LSID="urn:lsid:plazi:treatment:03EB879BFF91FFA2FDFD0043FAFBE4DB" httpUri="http://treatment.plazi.org/id/03EB879BFF91FFA2FDFD0043FAFBE4DB" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="C3586506FF91FFA3FDFD0043FC72E570" box="[565,1022,1614,1641]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BFD368DFF91FFA3FDFD0043FC72E570" blockId="2.[565,1022,1614,1641]" box="[565,1022,1614,1641]" pageId="2" pageNumber="3">
<heading id="D0B581E1FF91FFA3FDFD0043FC72E570" box="[565,1022,1614,1641]" centered="true" fontSize="11" level="3" pageId="2" pageNumber="3" reason="4">
Family
<taxonomicName id="4C424D0EFF91FFA3FD580043FC72E570" ID-CoL="7NKRH" authority="Mayer, 1910" authorityName="Mayer" authorityYear="1910" box="[656,1022,1614,1641]" class="Hydrozoa" family="Ptychogastriidae" kingdom="Animalia" order="Trachymedusae" pageId="2" pageNumber="3" phylum="Cnidaria" rank="family">
<emphasis id="B936EA9FFF91FFA3FD580043FCEEE571" bold="true" box="[656,866,1614,1640]" pageId="2" pageNumber="3">Ptychogastriidae</emphasis>
<bibRefCitation id="EFD34B7CFF91FFA3FCA10042FC72E570" author="Mayer A. G." box="[873,1022,1615,1641]" journalOrPublisher="Carnegie Institution of Washington Publication" pageId="2" pageNumber="3" pagination="231 - 498" part="109" publicationUrl="http://dx.doi.org/10.5962/bhl.title.5996" refString="Mayer A. G. 1910. Medusae of the world. Vol. II. The hydromedusae. Carnegie Institution of Washington Publication 109: 231 - 498. http: // dx. doi. org / 10.5962 / bhl. title. 5996" title="Medusae of the world. Vol. II. The hydromedusae" type="journal article" year="1910">Mayer, 1910</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BFD368DFF91FFA3FF75009DFEBAE5B3" blockId="2.[189,1399,1680,1964]" box="[189,310,1680,1706]" pageId="2" pageNumber="3">
<heading id="D0B581E1FF91FFA3FF75009DFEBAE5B3" bold="true" box="[189,310,1680,1706]" fontSize="11" level="2" pageId="2" pageNumber="3" reason="6">
<emphasis id="B936EA9FFF91FFA3FF75009DFEBAE5B3" bold="true" box="[189,310,1680,1706]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFD368DFF91FFA2FF7500B3FCF4E250" blockId="2.[189,1399,1680,1964]" lastBlockId="3.[189,1398,267,329]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<taxonomicName id="4C424D0EFF91FFA3FF7500B3FEFAE5C1" box="[189,374,1726,1752]" class="Hydrozoa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trachymedusae" pageId="2" pageNumber="3" phylum="Cnidaria" rank="order">Trachymedusae</taxonomicName>
with either Fat or dome-shaped umbrella; with four-lobed manubrium, not giving rise to mesenteries, or with eight-lobed manubrium, giving rise to eight mesenterial partitions of the subumbrella; without gastric peduncle; mouth with four simple lips; tentacles marginal, either Fliform, not arranged in groups and inserted at varied levels on the umbrellar margin, or Fliform and adhesive occurring simultaneously, and grouped into more or less well-deFned clusters composed of several superimposed rows; eight radial canals joining the circular canal; centripetal canals either absent or present; gonads either simple, elongated and attached to the body of manubrium, or in pairs resulting from the mesenterial partitions and developing on either side of the manubrial lobes; with or without a velum; with either 8 or 16 free, ecto-endodermal statocysts.
</paragraph>
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<subSubSection id="C3586506FF90FFA2FF75077DFD01E1EF" pageId="3" pageNumber="4" type="discussion">
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<emphasis id="B936EA9FFF90FFA2FF75077DFEA3E293" bold="true" box="[189,303,368,394]" pageId="3" pageNumber="4">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFD368DFF90FFA2FF750793FD01E1EF" blockId="3.[189,1399,414,758]" pageId="3" pageNumber="4">
Earlier diagnoses (
<bibRefCitation id="EFD34B7CFF90FFA2FE5D0792FDA4E2A0" author="Mayer A. G." box="[405,552,415,441]" journalOrPublisher="Carnegie Institution of Washington Publication" pageId="3" pageNumber="4" pagination="231 - 498" part="109" publicationUrl="http://dx.doi.org/10.5962/bhl.title.5996" refString="Mayer A. G. 1910. Medusae of the world. Vol. II. The hydromedusae. Carnegie Institution of Washington Publication 109: 231 - 498. http: // dx. doi. org / 10.5962 / bhl. title. 5996" title="Medusae of the world. Vol. II. The hydromedusae" type="journal article" year="1910">Mayer 1910</bibRefCitation>
;
<bibRefCitation id="EFD34B7CFF90FFA2FDFB0792FD4BE2A0" author="Kramp P. L." box="[563,711,414,441]" journalOrPublisher="Dana Report 46" pageId="3" pageNumber="4" pagination="1 - 283" refString="Kramp P. L. 1959. The hydromedusae of the Atlantic Ocean and adjacent waters. Dana Report 46: 1 - 283." title="The hydromedusae of the Atlantic Ocean and adjacent waters" type="book chapter" year="1959">Kramp 1959</bibRefCitation>
,
<bibRefCitation id="EFD34B7CFF90FFA2FD1B0793FC9CE2A1" author="Kramp P. L." box="[723,784,414,440]" journalOrPublisher="Journal of the Marine Biological Association of the United Kingdom" pageId="3" pageNumber="4" pagination="1 - 469" part="40" publicationUrl="http://dx.doi.org/10.1017/S0025315400007347" refString="Kramp P. L. 1961. Synopsis of the medusae of the world. Journal of the Marine Biological Association of the United Kingdom 40: 1 - 469. http: // dx. doi. org / 10.1017 / S 0025315400007347" title="Synopsis of the medusae of the world" type="journal article" year="1961">1961</bibRefCitation>
,
<bibRefCitation id="EFD34B7CFF90FFA2FCD30793FCD5E2A1" author="Kramp P. L." box="[795,857,414,440]" journalOrPublisher="Dana Report 72" pageId="3" pageNumber="4" pagination="1 - 200" refString="Kramp P. L. 1968. The hydromedusae of the PaciFc and Indian Oceans. Sections II and III. Dana Report 72: 1 - 200." title="The hydromedusae of the PaciFc and Indian Oceans. Sections II and III" type="book chapter" year="1968">1968</bibRefCitation>
;
<bibRefCitation id="EFD34B7CFF90FFA2FCAC0793FBF6E2A0" author="Bouillon J. &amp; Boero F." box="[868,1146,414,441]" journalOrPublisher="Thalassia Salentina" pageId="3" pageNumber="4" pagination="47 - 296" part="24" refString="Bouillon J. &amp; Boero F. 2000. Phylogeny and classiFcation of Hydroidomedusae. Synopsis of the families and genera of the Hydroidomedusae of the world, with a list of the worldwide species. Thalassia Salentina 24: 47 - 296." title="Phylogeny and classiFcation of Hydroidomedusae. Synopsis of the families and genera of the Hydroidomedusae of the world, with a list of the worldwide species" type="journal article" year="2000">Bouillon &amp; Boero 2000</bibRefCitation>
;
<bibRefCitation id="EFD34B7CFF90FFA2FB4D0793FAE2E2A0" author="Bouillon J. &amp; Gravili C. &amp; Pages F. &amp; Gili J. M. &amp; Boero F." box="[1157,1390,414,441]" journalOrPublisher="Museum national d'Histoire naturelle, Paris" pageId="3" pageNumber="4" pagination="194" publicationUrl="http://dx.doi" refString="Bouillon J., Gravili C., Pages F., Gili J. M. &amp; Boero F. 2006. An introduction to Hydrozoa. Memoires du Museum national d'Histoire naturelle 194, Museum national d'Histoire naturelle, Paris. http: // dx. doi. org / 10.1086 / 586983" title="An introduction to Hydrozoa. Memoires du Museum national d'Histoire naturelle" type="book chapter" year="2006">
Bouillon
<emphasis id="B936EA9FFF90FFA2FB3A0792FAA9E2A1" box="[1266,1317,414,440]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
. 2006
</bibRefCitation>
) are broadened by taking into consideration the shape of the umbrella, the presence/absence of a velum, and of centripetal canals. Indeed, unlike both
<taxonomicName id="4C424D0EFF90FFA2FD2F07E8FBB4E2E6" authority="Allman, 1878" authorityName="Allman" authorityYear="1878" box="[743,1080,485,511]" class="Hydrozoa" family="Ptychogastriidae" genus="Ptychogastria" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus">
<emphasis id="B936EA9FFF90FFA2FD2F07E8FC00E2E6" box="[743,908,485,511]" italics="true" pageId="3" pageNumber="4">Ptychogastria</emphasis>
<bibRefCitation id="EFD34B7CFF90FFA2FC5B07E8FBB4E2E6" author="Allman G. J." box="[915,1080,485,511]" editor="Feilden H. W." journalOrPublisher="G. S. Nares" pageId="3" pageNumber="4" pagination="290 - 292" part="2" publicationUrl="http://dx.doi.org/10.5962/bhl.title.6828" refString="Allman G. J. 1878. Appendix No. XI. Hydrozoa. In: Feilden H. W. (ed.) Narrative of a voyage to the Polar Sea during 1875 - 6 in H. M. ships &quot; Alert &quot; and &quot; Discovery &quot; by Capt. G. S. Nares 2: 290 - 292. Sampson Low, Marston, Searle &amp; Rivington, London. http: // dx. doi. org / 10.5962 / bhl. title. 6828" title="Appendix No. XI. Hydrozoa" type="journal article" volumeTitle="Narrative of a voyage to the Polar Sea during 1875 - 6 in H. M. ships &quot; Alert &quot; and &quot; Discovery &quot; by Capt" year="1878">Allman, 1878</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C424D0EFF90FFA2FBBE07EBFF76E13A" authority="Beyer, 1959" authorityName="Beyer" authorityYear="1959" class="Hydrozoa" family="Ptychogastriidae" genus="Tesserogastria" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus">
<emphasis id="B936EA9FFF90FFA2FBBE07EBFAADE2E6" box="[1142,1313,486,511]" italics="true" pageId="3" pageNumber="4">Tesserogastria</emphasis>
<bibRefCitation id="EFD34B7CFF90FFA2FAE207E8FF76E13A" author="Beyer F." journalOrPublisher="Nytt Magasin for Zoologi" pageId="3" pageNumber="4" pagination="121 - 143" part="6" refString="Beyer F. 1959. A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps. Nytt Magasin for Zoologi 6: 121 - 143." title="A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps" type="journal article" year="1959">Beyer, 1959</bibRefCitation>
</taxonomicName>
, the medusae belonging to the so far monotypic genus
<taxonomicName id="4C424D0EFF90FFA2FC4C0405FBB8E13B" box="[900,1076,520,546]" class="Hydrozoa" family="Ptychogastriidae" genus="Glaciambulata" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus" status="gen. nov.">
<emphasis id="B936EA9FFF90FFA2FC4C0405FBB8E13B" box="[900,1076,520,546]" italics="true" pageId="3" pageNumber="4">Glaciambulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557E4FF90FFA2FBF40404FB2BE13A" box="[1084,1191,521,547]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
(see below) have Fattened rather than dome-shaped umbrellas, and they also lack a velum. The presence of centripetal canals was curiously excluded from earlier diagnoses of the family, though they are present in
<taxonomicName id="4C424D0EFF90FFA2FACF0442FEE8E195" authority="Allman, 1878" authorityName="Allman" authorityYear="1878" class="Hydrozoa" family="Ptychogastriidae" genus="Ptychogastria" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="species" species="polaris">
<emphasis id="B936EA9FFF90FFA2FACF0442FAFAE171" box="[1287,1398,590,616]" italics="true" pageId="3" pageNumber="4">P. polaris</emphasis>
<bibRefCitation id="EFD34B7CFF90FFA2FF75047FFEE8E195" author="Allman G. J." box="[189,356,626,652]" editor="Feilden H. W." journalOrPublisher="G. S. Nares" pageId="3" pageNumber="4" pagination="290 - 292" part="2" publicationUrl="http://dx.doi.org/10.5962/bhl.title.6828" refString="Allman G. J. 1878. Appendix No. XI. Hydrozoa. In: Feilden H. W. (ed.) Narrative of a voyage to the Polar Sea during 1875 - 6 in H. M. ships &quot; Alert &quot; and &quot; Discovery &quot; by Capt. G. S. Nares 2: 290 - 292. Sampson Low, Marston, Searle &amp; Rivington, London. http: // dx. doi. org / 10.5962 / bhl. title. 6828" title="Appendix No. XI. Hydrozoa" type="journal article" volumeTitle="Narrative of a voyage to the Polar Sea during 1875 - 6 in H. M. ships &quot; Alert &quot; and &quot; Discovery &quot; by Capt" year="1878">Allman, 1878</bibRefCitation>
</taxonomicName>
. The occurrence of the gonads is henceforth restricted to the manubrium only, because
<taxonomicName id="4C424D0EFF90FFA2FF75049BFDBBE1B6" authority="Haeckel, 1879" authorityName="Haeckel" authorityYear="1879" box="[189,567,661,687]" class="Magnoliopsida" family="Asteraceae" genus="Pectis" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="antarctica">
<emphasis id="B936EA9FFF90FFA2FF75049BFE0FE1B6" box="[189,387,662,687]" italics="true" pageId="3" pageNumber="4">Pectis antarctica</emphasis>
<bibRefCitation id="EFD34B7CFF90FFA2FE430498FDBBE1B6" author="Haeckel E." box="[395,567,661,687]" journalOrPublisher="Gustav Fisher, Jena" pageId="3" pageNumber="4" publicationUrl="http://dx.doi.org/10.5962/bhl.title.46856" refString="Haeckel E. 1879. Das System der Medusen. Erster Theil einer Monographie der Medusen. Mit einem Atlas von vierzig Tafeln. Gustav Fisher, Jena. http: // dx. doi. org / 10.5962 / bhl. title. 46856" title="Das System der Medusen. Erster Theil einer Monographie der Medusen. Mit einem Atlas von vierzig Tafeln" type="book" year="1879">Haeckel, 1879</bibRefCitation>
</taxonomicName>
, formerly assigned to the genus
<taxonomicName id="4C424D0EFF90FFA2FC7B0498FBD4E1B6" box="[947,1112,661,687]" class="Hydrozoa" family="Ptychogastriidae" genus="Ptychogastria" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus">
<emphasis id="B936EA9FFF90FFA2FC7B0498FBD4E1B6" box="[947,1112,661,687]" italics="true" pageId="3" pageNumber="4">Ptychogastria</emphasis>
</taxonomicName>
by a number of authors, has the gonads on the radial canals and, for this reason, combined with additional characters, does not belong to
<taxonomicName id="4C424D0EFF90FFA2FEFA04D6FE7AE1EC" box="[306,502,731,757]" class="Hydrozoa" family="Ptychogastriidae" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="family">Ptychogastriidae</taxonomicName>
(see below).
</paragraph>
</subSubSection>
<subSubSection id="C3586506FF90FFA2FF750510FBFEE02E" box="[189,1138,797,823]" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BFD368DFF90FFA2FF750510FBFEE02E" blockId="3.[189,1138,797,823]" box="[189,1138,797,823]" pageId="3" pageNumber="4">
The main distinguishing characters of ptychogastriid genera are listed in
<tableCitation id="C6C00336FF90FFA2FBDC0510FBE0E02E" box="[1044,1132,797,823]" captionStart="Table 1" captionStartId="3.[189,258,900,926]" captionTargetBox="[189,1398,986,1673]" captionTargetPageId="3" captionText="Table 1. Main distinguishing characters of the genera belonging to the family Ptychogastriidae Mayer, 1900." httpUri="http://table.plazi.org/id/DF3D6605FF90FFA2FF750589FF73E0D8" pageId="3" pageNumber="4" tableUuid="DF3D6605FF90FFA2FF750589FF73E0D8">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF3D6605FF90FFA2FF750589FF73E0D8" ID-Table-UUID="DF3D6605FF90FFA2FF750589FF73E0D8" httpUri="http://table.plazi.org/id/DF3D6605FF90FFA2FF750589FF73E0D8" pageId="3" pageNumber="4" targetBox="[189,1398,986,1673]" targetIsTable="true" targetPageId="3">
<paragraph id="8BFD368DFF90FFA2FF750589FF73E0D8" blockId="3.[189,1398,900,961]" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF750589FEACE087" bold="true" box="[189,288,900,926]" pageId="3" pageNumber="4">Table 1.</emphasis>
Main distinguishing characters of the genera belonging to the family
<taxonomicName id="4C424D0EFF90FFA2FB900589FF77E0D8" authority="Mayer, 1900" authorityName="Mayer" authorityYear="1900" class="Hydrozoa" family="Ptychogastriidae" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="family">Ptychogastriidae Mayer, 1900</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BFD368DFF90FFA2FF0D0225FBD2E590" pageId="3" pageNumber="4">
<table id="F942C42DFF90005EFF7505D7FAFAE590" box="[189,1398,986,1673]" gridcols="4" gridrows="9" pageId="3" pageNumber="4">
<tr id="357234CFFF90005EFF7505D7FAFAE75B" box="[189,1398,986,1090]" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF0D0225FD50E71E" bold="true" box="[197,732,1004,1090]" pageId="3" pageNumber="4">
<th id="76A35DB3FF90005EFF7505D7FE2FE75B" box="[189,419,986,1090]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">Umbrella</th>
<th id="76A35DB3FF90005EFE6505D7FD6EE75B" box="[429,738,986,1090]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="4C424D0EFF90FFA2FE7C05E1FDEBE71E" box="[436,615,1004,1031]" class="Hydrozoa" family="Ptychogastriidae" genus="Glaciambulata" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus" status="gen. nov.">
<emphasis id="B936EA9FFF90FFA2FE7C05E1FDEBE71E" bold="true" box="[436,615,1004,1031]" italics="true" pageId="3" pageNumber="4">Glaciambulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557E4FF90FFA2FDA605E0FD50E71E" box="[622,732,1005,1031]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
Flat
</th>
</emphasis>
<th id="76A35DB3FF90005EFD3A05D7FBABE75B" box="[754,1063,986,1090]" gridcol="2" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="4C424D0EFF90FFA2FCF205D7FC6CE701" authority="Allman, 1878" authorityName="Allman" authorityYear="1878" box="[826,992,986,1048]" class="Hydrozoa" family="Ptychogastriidae" genus="Ptychogastria" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus">
<emphasis id="B936EA9FFF90FFA2FCF205D7FC6CE701" bold="true" box="[826,992,986,1048]" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FCF205D7FC6CE0EC" bold="true" box="[826,992,986,1013]" italics="true" pageId="3" pageNumber="4">Ptychogastria</emphasis>
Allman, 1878
</emphasis>
</taxonomicName>
Dome-shaped
</th>
<th id="76A35DB3FF90005EFBFF05D7FAFAE75B" box="[1079,1398,986,1090]" gridcol="3" gridrow="0" pageId="3" pageNumber="4">
<taxonomicName id="4C424D0EFF90FFA2FBB305D6FA96E701" authority="Beyer, 1959" authorityName="Beyer" authorityYear="1959" box="[1147,1320,987,1048]" class="Hydrozoa" family="Ptychogastriidae" genus="Tesserogastria" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus">
<emphasis id="B936EA9FFF90FFA2FBB305D6FA96E701" bold="true" box="[1147,1320,987,1048]" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FBB305D6FAA4E0EC" bold="true" box="[1147,1320,987,1013]" italics="true" pageId="3" pageNumber="4">Tesserogastria</emphasis>
Beyer, 1959
</emphasis>
</taxonomicName>
Dome-shaped
</th>
</tr>
<tr id="357234CFFF90005EFF750258FAFAE769" box="[189,1398,1109,1136]" gridrow="1" pageId="3" pageNumber="4">
<th id="76A35DB3FF90005EFF750258FE2FE769" box="[189,419,1109,1136]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF0D0258FED6E776" bold="true" box="[197,346,1109,1135]" pageId="3" pageNumber="4">Manubrium</emphasis>
</th>
<td id="76A35DB3FF90005EFE650258FD6EE769" box="[429,738,1109,1136]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">Eight-lobed</td>
<td id="76A35DB3FF90005EFD3A0258FBABE769" box="[754,1063,1109,1136]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">Eight-lobed</td>
<td id="76A35DB3FF90005EFBFF0258FAFAE769" box="[1079,1398,1109,1136]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">Four-lobed</td>
</tr>
<tr id="357234CFFF90005EFF75028FFAFAE784" box="[189,1398,1154,1181]" gridrow="2" pageId="3" pageNumber="4">
<th id="76A35DB3FF90005EFF75028FFE2FE784" box="[189,419,1154,1181]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF0D028FFEDBE785" bold="true" box="[197,343,1154,1180]" pageId="3" pageNumber="4">Mesenteries</emphasis>
</th>
<td id="76A35DB3FF90005EFE65028FFD6EE784" box="[429,738,1154,1181]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">8</td>
<td id="76A35DB3FF90005EFD3A028FFBABE784" box="[754,1063,1154,1181]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">8</td>
<td id="76A35DB3FF90005EFBFF028FFAFAE784" box="[1079,1398,1154,1181]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">Absent</td>
</tr>
<tr id="357234CFFF90005EFF7502BDFAFAE64A" box="[189,1398,1200,1363]" gridrow="3" pageId="3" pageNumber="4">
<th id="76A35DB3FF90005EFF7502BDFE2FE64A" box="[189,419,1200,1363]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF0D02BDFEBAE7D3" bold="true" box="[197,310,1200,1226]" pageId="3" pageNumber="4">Tentacles</emphasis>
</th>
<td id="76A35DB3FF90005EFE6502BDFD6EE64A" box="[429,738,1200,1363]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">Filiform + adhesive; forming contiguous groups; inserted in superimposed rows on umbrella margin</td>
<td id="76A35DB3FF90005EFD3A02BDFBABE64A" box="[754,1063,1200,1363]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">Filiform + adhesive; forming contiguous groups; inserted in superimposed rows on umbrella margin</td>
<td id="76A35DB3FF90005EFBFF02BDFAFAE64A" box="[1079,1398,1200,1363]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">Filiform; reportedly not arranged in groups; inserted at varied levels on umbrella margin</td>
</tr>
<tr id="357234CFFF90005EFF750360FAFAE69E" box="[189,1398,1389,1415]" gridrow="4" pageId="3" pageNumber="4">
<th id="76A35DB3FF90005EFF750360FE2FE69E" box="[189,419,1389,1415]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF0D0360FE2FE69E" bold="true" box="[197,419,1389,1415]" pageId="3" pageNumber="4">Centripetal canals</emphasis>
</th>
<td id="76A35DB3FF90005EFE650360FD6EE69E" box="[429,738,1389,1415]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">Absent</td>
<td id="76A35DB3FF90005EFD3A0360FBABE69E" box="[754,1063,1389,1415]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">Present/absent</td>
<td id="76A35DB3FF90005EFBFF0360FAFAE69E" box="[1079,1398,1389,1415]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">Absent</td>
</tr>
<tr id="357234CFFF90005EFF750397FAFAE6E0" box="[189,1398,1434,1529]" gridrow="5" pageId="3" pageNumber="4">
<th id="76A35DB3FF90005EFF750397FE2FE6E0" box="[189,419,1434,1529]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF0D0397FEAFE6AD" bold="true" box="[197,291,1434,1460]" pageId="3" pageNumber="4">Gonads</emphasis>
</th>
<td id="76A35DB3FF90005EFE650397FD6EE6E0" box="[429,738,1434,1529]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">Ovoid; in pairs, on either side of the manubrial lobes</td>
<td id="76A35DB3FF90005EFD3A0397FBABE6E0" box="[754,1063,1434,1529]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">Ovoid; in pairs, on either side of the manubrial lobes</td>
<td id="76A35DB3FF90005EFBFF0397FAFAE6E0" box="[1079,1398,1434,1529]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">Elongated; simple; attached to the body of manubrium</td>
</tr>
<tr id="357234CFFF90005EFF750019FAFAE537" box="[189,1398,1556,1582]" gridrow="6" pageId="3" pageNumber="4">
<th id="76A35DB3FF90005EFF750019FE2FE537" box="[189,419,1556,1582]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF0D0019FE9FE537" bold="true" box="[197,275,1556,1582]" pageId="3" pageNumber="4">Velum</emphasis>
</th>
<td id="76A35DB3FF90005EFE650019FD6EE537" box="[429,738,1556,1582]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">Absent</td>
<td id="76A35DB3FF90005EFD3A0019FBABE537" box="[754,1063,1556,1582]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">Present</td>
<td id="76A35DB3FF90005EFBFF0019FAFAE537" box="[1079,1398,1556,1582]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">Present</td>
</tr>
<tr id="357234CFFF90005EFF75004DFAFAE542" box="[189,1398,1600,1627]" gridrow="7" pageId="3" pageNumber="4">
<th id="76A35DB3FF90005EFF75004DFE2FE542" box="[189,419,1600,1627]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF0D004DFEB3E543" bold="true" box="[197,319,1600,1626]" pageId="3" pageNumber="4">Statocysts</emphasis>
</th>
<td id="76A35DB3FF90005EFE65004DFD6EE542" box="[429,738,1600,1627]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">16</td>
<td id="76A35DB3FF90005EFD3A004DFBABE542" box="[754,1063,1600,1627]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">16</td>
<td id="76A35DB3FF90005EFBFF004DFAFAE542" box="[1079,1398,1600,1627]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">8</td>
</tr>
<tr id="357234CFFF90005EFF750063FAFAE590" box="[189,1398,1646,1673]" gridrow="8" pageId="3" pageNumber="4">
<th id="76A35DB3FF90005EFF750063FE2FE590" box="[189,419,1646,1673]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">
<emphasis id="B936EA9FFF90FFA2FF0D0063FEF2E591" bold="true" box="[197,382,1646,1672]" pageId="3" pageNumber="4">Ability to swim</emphasis>
</th>
<td id="76A35DB3FF90005EFE650063FD6EE590" box="[429,738,1646,1673]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">No</td>
<td id="76A35DB3FF90005EFD3A0063FBABE590" box="[754,1063,1646,1673]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">Yes</td>
<td id="76A35DB3FF90005EFBFF0063FAFAE590" box="[1079,1398,1646,1673]" gridcol="3" gridrow="8" pageId="3" pageNumber="4">Yes</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3586506FF90FFA2FF7500FFFAFBE4DB" pageId="3" pageNumber="4" type="key">
<paragraph id="8BFD368DFF90FFA2FF7500FFFE16E415" blockId="3.[189,1399,1778,1986]" box="[189,410,1778,1804]" pageId="3" pageNumber="4">
<heading id="D0B581E1FF90FFA2FF7500FFFE16E415" bold="true" box="[189,410,1778,1804]" fontSize="11" level="2" pageId="3" pageNumber="4" reason="6">
<emphasis id="B936EA9FFF90FFA2FF7500FFFE16E415" bold="true" box="[189,410,1778,1804]" pageId="3" pageNumber="4">Key to the genera</emphasis>
</heading>
</paragraph>
<paragraph id="8BFD368DFF90FFA2FF75012DFAFBE422" blockId="3.[189,1399,1778,1986]" box="[189,1399,1824,1851]" pageId="3" pageNumber="4">
1. Tentacles exclusively Fliform …………………………………………
<taxonomicName id="4C424D0EFF90FFA2FBE5012DFAFBE422" ID-CoL="7Q42C" authority="Beyer, 1959" authorityName="Beyer" authorityYear="1959" box="[1069,1399,1824,1851]" class="Hydrozoa" family="Ptychogastriidae" genus="Tesserogastria" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus">
<emphasis id="B936EA9FFF90FFA2FBE5012DFB56E423" bold="true" box="[1069,1242,1824,1850]" italics="true" pageId="3" pageNumber="4">Tesserogastria</emphasis>
<bibRefCitation id="EFD34B7CFF90FFA2FB2D012DFAFBE422" author="Beyer F." box="[1253,1399,1824,1851]" journalOrPublisher="Nytt Magasin for Zoologi" pageId="3" pageNumber="4" pagination="121 - 143" part="6" refString="Beyer F. 1959. A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps. Nytt Magasin for Zoologi 6: 121 - 143." title="A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps" type="journal article" year="1959">Beyer, 1959</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD368DFF90FFA2FF090149FAFAE444" blockId="3.[189,1399,1778,1986]" box="[193,1398,1859,1886]" pageId="3" pageNumber="4"> Tentacles of two types: Fliform and adhesive ………………………………………………………2</paragraph>
<paragraph id="8BFD368DFF90FFA2FF750188FAFBE486" blockId="3.[189,1399,1778,1986]" box="[189,1399,1924,1951]" pageId="3" pageNumber="4">
2. Umbrella dome-shaped, velum present ………………………………
<taxonomicName id="4C424D0EFF90FFA2FBEC0189FAFBE486" ID-CoL="7PWZ4" authority="Allman, 1878" authorityName="Allman" authorityYear="1878" box="[1060,1399,1924,1951]" class="Hydrozoa" family="Ptychogastriidae" genus="Ptychogastria" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus">
<emphasis id="B936EA9FFF90FFA2FBEC0189FB46E486" bold="true" box="[1060,1226,1924,1951]" italics="true" pageId="3" pageNumber="4">Ptychogastria</emphasis>
<bibRefCitation id="EFD34B7CFF90FFA2FB1A0189FAFBE486" author="Allman G. J." box="[1234,1399,1924,1951]" editor="Feilden H. W." journalOrPublisher="G. S. Nares" pageId="3" pageNumber="4" pagination="290 - 292" part="2" publicationUrl="http://dx.doi.org/10.5962/bhl.title.6828" refString="Allman G. J. 1878. Appendix No. XI. Hydrozoa. In: Feilden H. W. (ed.) Narrative of a voyage to the Polar Sea during 1875 - 6 in H. M. ships &quot; Alert &quot; and &quot; Discovery &quot; by Capt. G. S. Nares 2: 290 - 292. Sampson Low, Marston, Searle &amp; Rivington, London. http: // dx. doi. org / 10.5962 / bhl. title. 6828" title="Appendix No. XI. Hydrozoa" type="journal article" volumeTitle="Narrative of a voyage to the Polar Sea during 1875 - 6 in H. M. ships &quot; Alert &quot; and &quot; Discovery &quot; by Capt" year="1878">Allman, 1878</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD368DFF90FFA2FF0901A5FAFBE4DB" blockId="3.[189,1399,1778,1986]" box="[193,1399,1959,1986]" pageId="3" pageNumber="4">
Umbrella Fat, velum absent ………………………………………………
<taxonomicName id="4C424D0EFF90FFA2FB8601AAFA8DE4DB" ID-CoL="7PGWT" box="[1102,1281,1959,1986]" class="Hydrozoa" family="Ptychogastriidae" genus="Glaciambulata" kingdom="Animalia" order="Trachymedusae" pageId="3" pageNumber="4" phylum="Cnidaria" rank="genus" status="gen. nov.">
<emphasis id="B936EA9FFF90FFA2FB8601AAFA8DE4DB" bold="true" box="[1102,1281,1959,1986]" italics="true" pageId="3" pageNumber="4">Glaciambulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557E4FF90FFA2FAC201A5FAFBE4DB" box="[1290,1399,1960,1986]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>