501 lines
52 KiB
XML
501 lines
52 KiB
XML
<document id="C16CA1C3D4B003CD67AE7272725CBFD4" ID-DOI="10.11646/zootaxa.5497.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13618611" ID-ZooBank="C107D617-5F70-42C5-8FD4-E5C2DE0E48B1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725026357643" checkinUser="plazi" docAuthor="Khaustov, Alexander A., Frolov, Andrey V. & Akhmetova, Lilia A." docDate="2024" docId="03EB878FFFB6744781B43B0AFC07FE1A" docLanguage="en" docName="zootaxa.5497.3.2.pdf" docOrigin="Zootaxa 5497 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5497.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Spatulaphorus incisus Khaustov, Frolov & Akhmetova, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="357" masterDocId="FFD2FFF7FFA6745381233D58FFDEFFEF" masterDocTitle="Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)" masterLastPageNumber="368" masterPageNumber="337" pageNumber="353" updateTime="1725028125811" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="E3F08E9C9140AC162FCD5F6353E44BEC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8D5D589B7B0439B5B21117321DAFD516">
|
||
<mods:title id="643655EA2718DDDB89C2EEFFA132C78F">Seven new species of Spatulaphorus Rack (Acari: Pygmephoridae) phoretic on scarab beetles (Coleoptera: Scarabaeidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="0D0A9710C92655C22957C813A92E9AC6" type="personal">
|
||
<mods:role id="91D3898AE0F8CB78E929B0B6EC25DFE5">
|
||
<mods:roleTerm id="5C1197BD84A19B501CC3984746131BBC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="755134F9EF3552C62B85680B1F9BF48D">Khaustov, Alexander A.</mods:namePart>
|
||
<mods:affiliation id="C61E13983A613F04B1C4D55ACB0A1172">Tyumen State University, Tyumen, 6 Volodarskogo Str., 625003 Russia.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="577C3A92EF5ACA487FA96D1289C0465B" type="personal">
|
||
<mods:role id="2F724F0169EAB8E2E367AB7EDF014CD5">
|
||
<mods:roleTerm id="3D48AD60DE980F8B0CA36F19CF2B571D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="EE6CDD9833380E3D7F09B7D1ACD54034">Frolov, Andrey V.</mods:namePart>
|
||
<mods:affiliation id="713E8421BF5333E17A796FC41F5DF8EE">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="5F1A31B8783FB074F74FED8D20E1173A" type="personal">
|
||
<mods:role id="83715A8402D6046B559CFB9BCBA91A19">
|
||
<mods:roleTerm id="B9CC4CAAA315CD8A26657334F17E0A01">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="68F23597082CEA73862FE50EA17C9A9D">Akhmetova, Lilia A.</mods:namePart>
|
||
<mods:affiliation id="F5AC249481B72F8E587E5CADF353D839">Zoological Institute of Russian Academy of Sciences, Universitetskaya Embankment 1, 199034, Saint Petersburg, Russia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="A3C88067411FE127540206ECA907B837">text</mods:typeOfResource>
|
||
<mods:relatedItem id="C05C950EE87A07D7379867577A9D9961" type="host">
|
||
<mods:titleInfo id="9C71B5D9CE527D98B05ED34E2955DE04">
|
||
<mods:title id="D391EF760C20431571841D09CA4F4F12">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="95DF6CB1A3884619BE8DB3F712BFE4C1">
|
||
<mods:date id="31B87FF3C485433631340312D217E668">2024</mods:date>
|
||
<mods:detail id="E501BAD3850FD5EFD428812730C1108F" type="pubDate">
|
||
<mods:number id="12A09C3029031C1FC44ABA22BC4D956E">2024-08-26</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="54CC24B76F478A506A2C93E2C43B5955" type="volume">
|
||
<mods:number id="892310C5376CB03B05EF342C0CBEC1B2">5497</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="DFC78E093955B5BBB7CE38AD6713CAB7" type="issue">
|
||
<mods:number id="2AD8025876A849D567E8653C5162DCDA">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="2820E08B83F796A5F01887BA5A34089A" unit="page">
|
||
<mods:start id="42C39772C6038E56B2F61442A5F2E45B">337</mods:start>
|
||
<mods:end id="573C5E1D457B8E765B46E78F9EFF4540">368</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="580E0009CA11E845D774B0CA8B63713F">
|
||
<mods:url id="4DD8D30283DB55DAC4E7BF1A7D1235F9">http://dx.doi.org/10.11646/zootaxa.5497.3.2</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="11C8AFB9A83FC52C673C52B3D44684FB">journal article</mods:classification>
|
||
<mods:identifier id="4D9E1D590DFE2B6EF7C71C3F557CD36A" type="DOI">10.11646/zootaxa.5497.3.2</mods:identifier>
|
||
<mods:identifier id="C0A0DFB50204444291D348D2A7098060" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="854DBEC0451C39B2409853D5E452F25E" type="Zenodo-Dep">13618611</mods:identifier>
|
||
<mods:identifier id="17A9AD6E60B632F3BBD1DB8BD721F19D" type="ZooBank">C107D617-5F70-42C5-8FD4-E5C2DE0E48B1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03EB878FFFB6744781B43B0AFC07FE1A" LSID="urn:lsid:plazi:treatment:03EB878FFFB6744781B43B0AFC07FE1A" httpUri="http://treatment.plazi.org/id/03EB878FFFB6744781B43B0AFC07FE1A" lastPageId="20" lastPageNumber="357" pageId="16" pageNumber="353">
|
||
<subSubSection id="C3586512FFB6744381B43B0AFDD4F983" box="[151,522,1618,1645]" pageId="16" pageNumber="353" type="nomenclature">
|
||
<paragraph id="8BFD3699FFB6744381B43B0AFDD4F983" blockId="16.[151,522,1618,1680]" box="[151,522,1618,1645]" pageId="16" pageNumber="353">
|
||
<heading id="D0B581F5FFB6744381B43B0AFDD4F983" bold="true" box="[151,522,1618,1645]" fontSize="11" level="1" pageId="16" pageNumber="353" reason="1">
|
||
<emphasis id="B936EA8BFFB6744381B43B0AFDD4F983" bold="true" box="[151,522,1618,1645]" pageId="16" pageNumber="353">
|
||
<taxonomicName id="4C424D1AFFB6744381B43B0AFE7BF983" authorityName="Khaustov & Frolov & Akhmetova" authorityYear="2024" box="[151,421,1618,1645]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="353" phylum="Arthropoda" rank="species" species="incisus" status="sp. nov.">
|
||
<emphasis id="B936EA8BFFB6744381B43B0AFE7BF983" bold="true" box="[151,421,1618,1645]" italics="true" pageId="16" pageNumber="353">Spatulaphorus incisus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20557F0FFB67443808F3B0AFDD4F983" box="[428,522,1618,1644]" pageId="16" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3586512FFB6744081B43B2EFE4FF86B" lastPageId="19" lastPageNumber="356" pageId="16" pageNumber="353" type="description">
|
||
<paragraph id="8BFD3699FFB6744381B43B2EFEF9F97F" blockId="16.[151,522,1618,1680]" box="[151,295,1654,1680]" pageId="16" pageNumber="353">
|
||
(
|
||
<figureCitation id="13792A1CFFB6744381BC3B2EFEFEF97F" box="[159,288,1654,1680]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStart-3="FIGURE 20" captionStartId-0="17.[151,250,1496,1520]" captionStartId-1="18.[151,250,1430,1454]" captionStartId-2="19.[151,250,1827,1851]" captionStartId-3="20.[151,250,1555,1579]" captionTargetBox-0="[151,1435,187,1472]" captionTargetBox-1="[151,1436,276,1407]" captionTargetBox-2="[174,1413,189,1804]" captionTargetBox-3="[151,1436,525,1532]" captionTargetId-0="figure-17@17.[151,1435,181,1472]" captionTargetId-1="figure-87@18.[151,1436,273,1407]" captionTargetId-2="figure-17@19.[151,1436,181,1804]" captionTargetId-3="figure-181@20.[151,1436,525,1532]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="FIGURE 17. Spatulaphorus incisus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." captionText-1="FIGURE 18. Spatulaphorus incisus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." captionText-2="FIGURE 19. Spatulaphorus incisus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." captionText-3="FIGURE 20. DIC micrographs of Spatulaphorus incisus sp. nov., female: A—general view dorsally; B—general view ventrally." figureDoi-0="http://doi.org/10.5281/zenodo.13618651" figureDoi-1="http://doi.org/10.5281/zenodo.13618653" figureDoi-2="http://doi.org/10.5281/zenodo.13618655" figureDoi-3="http://doi.org/10.5281/zenodo.13618657" httpUri-0="https://zenodo.org/record/13618651/files/figure.png" httpUri-1="https://zenodo.org/record/13618653/files/figure.png" httpUri-2="https://zenodo.org/record/13618655/files/figure.png" httpUri-3="https://zenodo.org/record/13618657/files/figure.png" pageId="16" pageNumber="353">Figs 17–20</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BFD3699FFB6744381B43BE6FAE1F936" blockId="16.[151,1437,1726,2005]" box="[151,1343,1726,1753]" pageId="16" pageNumber="353">
|
||
<emphasis id="B936EA8BFFB6744381B43BE6FEFFF937" bold="true" box="[151,289,1726,1752]" pageId="16" pageNumber="353">Description</emphasis>
|
||
. FEMALE. Body well sclerotized. Length of idiosoma 180 (180–205), width 105 (105–125).
|
||
</paragraph>
|
||
<paragraph id="8BFD3699FFB6744381E43BBAFCC9F83B" blockId="16.[151,1437,1726,2005]" pageId="16" pageNumber="353">
|
||
<emphasis id="B936EA8BFFB6744381E43BBAFE88F913" box="[199,342,1762,1788]" italics="true" pageId="16" pageNumber="353">Gnathosoma</emphasis>
|
||
. Gnathosomal capsule, excluding palps, subrectangular, length 17 (17–20), width 18 (18–20). Dorsum with one pair of needle-like cheliceral setae (
|
||
<emphasis id="B936EA8BFFB67443823D3A5EFC98F8CF" box="[798,838,1798,1824]" italics="true" pageId="16" pageNumber="353">cha</emphasis>
|
||
) 3 (3–4). Dorsal median apodeme present, poorly developed. Postpalpal setae (
|
||
<emphasis id="B936EA8BFFB6744380C13A73FE20F8AB" box="[482,510,1835,1860]" italics="true" pageId="16" pageNumber="353">pp</emphasis>
|
||
) 2 (2) spiniform, situated laterad setae
|
||
<emphasis id="B936EA8BFFB6744382E03A72FC35F8AB" box="[963,1003,1834,1860]" italics="true" pageId="16" pageNumber="353">cha</emphasis>
|
||
. Ventral gnathosoma with one pair of smooth, pointed subcapitular setae
|
||
<emphasis id="B936EA8BFFB6744383003A17FDE9F887" box="[547,567,1871,1896]" italics="true" pageId="16" pageNumber="353">m</emphasis>
|
||
7 (6–7) and round pits
|
||
<emphasis id="B936EA8BFFB6744382633A17FC90F887" box="[832,846,1871,1896]" italics="true" pageId="16" pageNumber="353">n</emphasis>
|
||
situated posteriad
|
||
<emphasis id="B936EA8BFFB6744385013A17FBE8F887" box="[1058,1078,1871,1896]" italics="true" pageId="16" pageNumber="353">m</emphasis>
|
||
. Palps with smooth and weakly blunt-tipped setae
|
||
<emphasis id="B936EA8BFFB67443804D3A2AFE44F863" box="[366,410,1906,1932]" italics="true" pageId="16" pageNumber="353">dFe</emphasis>
|
||
3 (3) and pointed
|
||
<emphasis id="B936EA8BFFB67443835F3A2AFD75F863" box="[636,683,1906,1932]" italics="true" pageId="16" pageNumber="353">dGe</emphasis>
|
||
8 (8–9) dorsolaterally. Palps ventrally each with mushroom-like accessory setigenous structure (
|
||
<emphasis id="B936EA8BFFB6744380DA3ACFFDC3F85F" box="[505,541,1943,1968]" italics="true" pageId="16" pageNumber="353">ass</emphasis>
|
||
) and tiny solenidion
|
||
<emphasis id="B936EA8BFFB6744382293ACFFCC0F85F" box="[778,798,1943,1968]" italics="true" pageId="16" pageNumber="353">ω</emphasis>
|
||
. Palps terminated with small blunt-tipped tibial claw and tiny eupathid-like seta. Pharyngeal pumps poorly visible.
|
||
</paragraph>
|
||
<caption id="DF3D6611FFB7744281B43880FB34FA1F" ID-DOI="http://doi.org/10.5281/zenodo.13618651" ID-Zenodo-Dep="13618651" box="[151,1258,1496,1521]" httpUri="https://zenodo.org/record/13618651/files/figure.png" pageId="17" pageNumber="354" startId="17.[151,250,1496,1520]" targetBox="[151,1435,187,1472]" targetPageId="17" targetType="figure">
|
||
<paragraph id="8BFD3699FFB7744281B43880FB34FA1F" blockId="17.[151,1258,1496,1521]" box="[151,1258,1496,1521]" pageId="17" pageNumber="354">
|
||
<emphasis id="B936EA8BFFB7744281B43880FEC7FA1F" bold="true" box="[151,281,1496,1520]" pageId="17" pageNumber="354">FIGURE 17</emphasis>
|
||
.
|
||
<taxonomicName id="4C424D1AFFB7744280053880FDD8FA1F" authorityName="Khaustov & Frolov & Akhmetova" authorityYear="2024" box="[294,518,1496,1520]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="354" phylum="Arthropoda" rank="species" species="incisus" status="sp. nov.">
|
||
<emphasis id="B936EA8BFFB7744280053880FDD8FA1F" box="[294,518,1496,1520]" italics="true" pageId="17" pageNumber="354">Spatulaphorus incisus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B936EA8BFFB77442832E3880FD83FA1F" bold="true" box="[525,605,1496,1520]" pageId="17" pageNumber="354">
|
||
<taxonomicNameLabel id="A20557F0FFB77442832E3880FD83FA1F" box="[525,605,1496,1520]" pageId="17" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female: A—dorsum of body; B—venter of body. Legs omitted.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD3699FFB7744281E43B46FC86F8DA" blockId="17.[151,1437,1566,2025]" pageId="17" pageNumber="354">
|
||
<emphasis id="B936EA8BFFB7744281E43B46FE4BF9D7" box="[199,405,1566,1592]" italics="true" pageId="17" pageNumber="354">Idiosomal dorsum</emphasis>
|
||
(
|
||
<figureCitation id="13792A1CFFB7744280843B46FDCBF9D6" box="[423,533,1566,1593]" captionStart="FIGURE 17" captionStartId="17.[151,250,1496,1520]" captionTargetBox="[151,1435,187,1472]" captionTargetId="figure-17@17.[151,1435,181,1472]" captionTargetPageId="17" captionText="FIGURE 17. Spatulaphorus incisus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618651" httpUri="https://zenodo.org/record/13618651/files/figure.png" pageId="17" pageNumber="354">Figs 17A</figureCitation>
|
||
,
|
||
<figureCitation id="13792A1CFFB7744283073B47FD86F9D6" box="[548,600,1567,1593]" captionStart="FIGURE 20" captionStartId="20.[151,250,1555,1579]" captionTargetBox="[151,1436,525,1532]" captionTargetId="figure-181@20.[151,1436,525,1532]" captionTargetPageId="20" captionText="FIGURE 20. DIC micrographs of Spatulaphorus incisus sp. nov., female: A—general view dorsally; B—general view ventrally." figureDoi="http://doi.org/10.5281/zenodo.13618657" httpUri="https://zenodo.org/record/13618657/files/figure.png" pageId="17" pageNumber="354">20A</figureCitation>
|
||
). All dorsal shields with big puncta. Prodorsal shield with three pairs of setae (
|
||
<emphasis id="B936EA8BFFB7744281C33B1BFF24F9B2" box="[224,250,1603,1629]" italics="true" pageId="17" pageNumber="354">v1</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB77442802A3B1BFEFDF9B2" box="[265,291,1603,1629]" italics="true" pageId="17" pageNumber="354">v2</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB7744280123B1BFE89F9B2" box="[305,343,1603,1629]" italics="true" pageId="17" pageNumber="354">sc2</emphasis>
|
||
), one pair of weakly barbed capitate trichobothria (
|
||
<emphasis id="B936EA8BFFB7744282B83B1BFC1FF9B2" box="[923,961,1603,1629]" italics="true" pageId="17" pageNumber="354">sc1</emphasis>
|
||
) and one pair of round stigmata. Stigmata associated with well-developed tracheal trunks. All dorsal idiosomal setae blunt-tipped; setae
|
||
<emphasis id="B936EA8BFFB7744285FA3B3FFB2AF96E" box="[1241,1268,1639,1665]" italics="true" pageId="17" pageNumber="354">v2</emphasis>
|
||
smooth, other dorsal setae barbed. Alveoli of setae
|
||
<emphasis id="B936EA8BFFB7744283173BD3FD9EF94B" box="[564,576,1675,1700]" italics="true" pageId="17" pageNumber="354">e</emphasis>
|
||
and
|
||
<emphasis id="B936EA8BFFB77442835B3BD3FD5EF94A" box="[632,640,1675,1701]" italics="true" pageId="17" pageNumber="354">f</emphasis>
|
||
almost contiguous. Cupules
|
||
<emphasis id="B936EA8BFFB7744282E63BD3FC05F94B" box="[965,987,1675,1700]" italics="true" pageId="17" pageNumber="354">ia</emphasis>
|
||
on tergite D,
|
||
<emphasis id="B936EA8BFFB7744285553BD3FB4CF94B" box="[1142,1170,1675,1700]" italics="true" pageId="17" pageNumber="354">im</emphasis>
|
||
on tergite EF and
|
||
<emphasis id="B936EA8BFFB7744284403BD2FAA7F94B" box="[1379,1401,1674,1700]" italics="true" pageId="17" pageNumber="354">ih</emphasis>
|
||
on tergite H large, round, subequal. Lengths of dorsal setae:
|
||
<emphasis id="B936EA8BFFB7744282313BF7FCF2F926" box="[786,812,1711,1737]" italics="true" pageId="17" pageNumber="354">v1</emphasis>
|
||
15 (15–20),
|
||
<emphasis id="B936EA8BFFB7744282983BF7FC08F926" box="[955,982,1711,1737]" italics="true" pageId="17" pageNumber="354">v2</emphasis>
|
||
9 (9–10),
|
||
<emphasis id="B936EA8BFFB77442856B3BF7FBB0F926" box="[1096,1134,1711,1737]" italics="true" pageId="17" pageNumber="354">sc2</emphasis>
|
||
28 (26–32),
|
||
<emphasis id="B936EA8BFFB7744285DF3BF7FAC9F926" box="[1276,1303,1711,1737]" italics="true" pageId="17" pageNumber="354">c1</emphasis>
|
||
23 (23–26),
|
||
<emphasis id="B936EA8BFFB7744281B43B8BFF6CF902" box="[151,178,1747,1773]" italics="true" pageId="17" pageNumber="354">c2</emphasis>
|
||
28 (26–30),
|
||
<emphasis id="B936EA8BFFB7744280663B8AFE8DF903" box="[325,339,1746,1772]" italics="true" pageId="17" pageNumber="354">d</emphasis>
|
||
29 (28–32),
|
||
<emphasis id="B936EA8BFFB7744280C43B8BFE2DF903" box="[487,499,1747,1772]" italics="true" pageId="17" pageNumber="354">e</emphasis>
|
||
19 (19–24),
|
||
<emphasis id="B936EA8BFFB7744283A43B8BFD51F902" box="[647,655,1747,1773]" italics="true" pageId="17" pageNumber="354">f</emphasis>
|
||
33 (31–39),
|
||
<emphasis id="B936EA8BFFB7744282003B8AFCE0F903" box="[803,830,1746,1772]" italics="true" pageId="17" pageNumber="354">h1</emphasis>
|
||
37 (37–43),
|
||
<emphasis id="B936EA8BFFB7744282F03B8AFC31F903" box="[979,1007,1746,1772]" italics="true" pageId="17" pageNumber="354">h2</emphasis>
|
||
18 (18–24). Distances between setae:
|
||
<emphasis id="B936EA8BFFB7744281B43BAFFF04F8FE" box="[151,218,1783,1809]" italics="true" pageId="17" pageNumber="354">v1–v1</emphasis>
|
||
18 (18–20),
|
||
<emphasis id="B936EA8BFFB7744280533BAFFE6CF8FE" box="[368,434,1783,1809]" italics="true" pageId="17" pageNumber="354">v2–v2</emphasis>
|
||
33 (33–39),
|
||
<emphasis id="B936EA8BFFB77442836A3BAFFD7FF8FE" box="[585,673,1783,1809]" italics="true" pageId="17" pageNumber="354">sc2–sc2</emphasis>
|
||
38 (38–45),
|
||
<emphasis id="B936EA8BFFB7744282143BAFFCA4F8FE" box="[823,890,1783,1809]" italics="true" pageId="17" pageNumber="354">c1–c1</emphasis>
|
||
31 (31–37),
|
||
<emphasis id="B936EA8BFFB77442852D3BAFFB8FF8FE" box="[1038,1105,1783,1809]" italics="true" pageId="17" pageNumber="354">c1–c2</emphasis>
|
||
22 (22–25),
|
||
<emphasis id="B936EA8BFFB7744285C43BAEFACFF8FF" box="[1255,1297,1782,1809]" italics="true" pageId="17" pageNumber="354">d–d</emphasis>
|
||
51 (51–62),
|
||
<emphasis id="B936EA8BFFB7744281B43A43FF64F8DA" box="[151,186,1819,1845]" italics="true" pageId="17" pageNumber="354">e–f</emphasis>
|
||
5 (5–6),
|
||
<emphasis id="B936EA8BFFB7744280033A43FEE0F8DA" box="[288,318,1819,1845]" italics="true" pageId="17" pageNumber="354">f–f</emphasis>
|
||
43 (43–57),
|
||
<emphasis id="B936EA8BFFB7744280ED3A42FDCAF8DB" box="[462,532,1818,1845]" italics="true" pageId="17" pageNumber="354">h1–h1</emphasis>
|
||
30 (30–41),
|
||
<emphasis id="B936EA8BFFB7744283863A42FD35F8DB" box="[677,747,1818,1845]" italics="true" pageId="17" pageNumber="354">h1–h2</emphasis>
|
||
8 (8–11).
|
||
</paragraph>
|
||
<paragraph id="8BFD3699FFB7744181E43A66FA46FF16" blockId="17.[151,1437,1566,2025]" lastBlockId="18.[151,1437,151,249]" lastPageId="18" lastPageNumber="355" pageId="17" pageNumber="354">
|
||
<emphasis id="B936EA8BFFB7744281E43A66FE5DF8B7" box="[199,387,1854,1880]" italics="true" pageId="17" pageNumber="354">Idiosomal venter</emphasis>
|
||
(
|
||
<figureCitation id="13792A1CFFB7744280B33A66FE27F8B6" box="[400,505,1854,1881]" captionStart="FIGURE 17" captionStartId="17.[151,250,1496,1520]" captionTargetBox="[151,1435,187,1472]" captionTargetId="figure-17@17.[151,1435,181,1472]" captionTargetPageId="17" captionText="FIGURE 17. Spatulaphorus incisus sp. nov., female: A—dorsum of body; B—venter of body. Legs omitted." figureDoi="http://doi.org/10.5281/zenodo.13618651" httpUri="https://zenodo.org/record/13618651/files/figure.png" pageId="17" pageNumber="354">Figs 17B</figureCitation>
|
||
,
|
||
<figureCitation id="13792A1CFFB7744283203A67FDE8F8B6" box="[515,566,1855,1881]" captionStart="FIGURE 20" captionStartId="20.[151,250,1555,1579]" captionTargetBox="[151,1436,525,1532]" captionTargetId="figure-181@20.[151,1436,525,1532]" captionTargetPageId="20" captionText="FIGURE 20. DIC micrographs of Spatulaphorus incisus sp. nov., female: A—general view dorsally; B—general view ventrally." figureDoi="http://doi.org/10.5281/zenodo.13618657" httpUri="https://zenodo.org/record/13618657/files/figure.png" pageId="17" pageNumber="354">20B</figureCitation>
|
||
). Coxisternal fields I-II laterally with sparsely distributed big puncta. Setae
|
||
<emphasis id="B936EA8BFFB77442845A3A66FA48F8B7" box="[1401,1430,1854,1880]" italics="true" pageId="17" pageNumber="354">1b</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB7744281B43A3BFF6FF892" box="[151,177,1891,1917]" italics="true" pageId="17" pageNumber="354">2c</emphasis>
|
||
, and
|
||
<emphasis id="B936EA8BFFB7744281CD3A3AFED4F893" box="[238,266,1890,1916]" italics="true" pageId="17" pageNumber="354">4b</emphasis>
|
||
pointed; other ventral setae blunt-tipped; setae
|
||
<emphasis id="B936EA8BFFB7744282393A3AFCE8F893" box="[794,822,1890,1916]" italics="true" pageId="17" pageNumber="354">1b</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB7744282603A3BFC83F892" box="[835,861,1891,1917]" italics="true" pageId="17" pageNumber="354">2c</emphasis>
|
||
, and
|
||
<emphasis id="B936EA8BFFB7744282B93A3AFC68F893" box="[922,950,1890,1916]" italics="true" pageId="17" pageNumber="354">4b</emphasis>
|
||
with few weak barbs; setae
|
||
<emphasis id="B936EA8BFFB7744285CC3A3BFAF3F892" box="[1263,1325,1891,1917]" italics="true" pageId="17" pageNumber="354">ps1-3</emphasis>
|
||
distinctly barbed, other ventral setae smooth. Bases of setae
|
||
<emphasis id="B936EA8BFFB7744283FC3ADFFCD8F84E" box="[735,774,1927,1953]" italics="true" pageId="17" pageNumber="354">ps3</emphasis>
|
||
situated distinctly anteriad bases of setae
|
||
<emphasis id="B936EA8BFFB7744285CE3ADFFAEDF84E" box="[1261,1331,1927,1953]" italics="true" pageId="17" pageNumber="354">ps1-2.</emphasis>
|
||
Ap1 and ap2 well-developed and joined with thick appr; apsej developed only laterally; ap3 weak, not reaching appo; ap4 well developed, exceeding beyond bases of setae
|
||
<emphasis id="B936EA8BFFB7744283E03A96FD01F807" box="[707,735,1998,2024]" italics="true" pageId="17" pageNumber="354">3b</emphasis>
|
||
. Ap5 very short and thin. Ags small, cup-like; pgs small, oval. Posterior margin of posterior sternal plate deeply incised in middle part. Posterior margin of aggenital plate rounded. Lengths of ventral setae:
|
||
<emphasis id="B936EA8BFFB4744180863DE3FE1EFF3A" box="[421,448,187,213]" italics="true" pageId="18" pageNumber="355">1a</emphasis>
|
||
8 (8–11),
|
||
<emphasis id="B936EA8BFFB47441830D3DE3FD97FF3A" box="[558,585,187,213]" italics="true" pageId="18" pageNumber="355">1b</emphasis>
|
||
17 (17–19),
|
||
<emphasis id="B936EA8BFFB4744183F03DE3FD32FF3A" box="[723,748,187,213]" italics="true" pageId="18" pageNumber="355">1с</emphasis>
|
||
10 (10–12),
|
||
<emphasis id="B936EA8BFFB4744182553DE3FC4FFF3A" box="[886,913,187,213]" italics="true" pageId="18" pageNumber="355">2a</emphasis>
|
||
11 (11–13),
|
||
<emphasis id="B936EA8BFFB47441853B3DE3FBECFF3A" box="[1048,1074,187,213]" italics="true" pageId="18" pageNumber="355">2с</emphasis>
|
||
12 (12–17),
|
||
<emphasis id="B936EA8BFFB47441859F3DE3FB09FF3A" box="[1212,1239,187,213]" italics="true" pageId="18" pageNumber="355">3a</emphasis>
|
||
13 (13–15),
|
||
<emphasis id="B936EA8BFFB4744184433DE3FAA5FF3A" box="[1376,1403,187,213]" italics="true" pageId="18" pageNumber="355">3b</emphasis>
|
||
10 (10–14),
|
||
<emphasis id="B936EA8BFFB4744181D93D87FECDFF16" box="[250,275,223,249]" italics="true" pageId="18" pageNumber="355">3c</emphasis>
|
||
12 (12–14),
|
||
<emphasis id="B936EA8BFFB4744180BD3D87FE67FF16" box="[414,441,223,249]" italics="true" pageId="18" pageNumber="355">4a</emphasis>
|
||
12 (12–14),
|
||
<emphasis id="B936EA8BFFB4744183673D87FD81FF16" box="[580,607,223,249]" italics="true" pageId="18" pageNumber="355">4b</emphasis>
|
||
21 (21–29),
|
||
<emphasis id="B936EA8BFFB4744183CA3D87FCDCFF16" box="[745,770,223,249]" italics="true" pageId="18" pageNumber="355">4c</emphasis>
|
||
13 (13–15),
|
||
<emphasis id="B936EA8BFFB4744182AD3D87FC6DFF16" box="[910,947,223,249]" italics="true" pageId="18" pageNumber="355">ps1</emphasis>
|
||
18 (18–20),
|
||
<emphasis id="B936EA8BFFB47441851D3D87FBBDFF16" box="[1086,1123,223,249]" italics="true" pageId="18" pageNumber="355">ps2</emphasis>
|
||
18 (18–20),
|
||
<emphasis id="B936EA8BFFB4744185CD3D87FACDFF16" box="[1262,1299,223,249]" italics="true" pageId="18" pageNumber="355">ps3</emphasis>
|
||
19 (19–22).
|
||
</paragraph>
|
||
<caption id="DF3D6611FFB4744181B438CEFAC9FA41" ID-DOI="http://doi.org/10.5281/zenodo.13618653" ID-Zenodo-Dep="13618653" box="[151,1303,1430,1455]" httpUri="https://zenodo.org/record/13618653/files/figure.png" pageId="18" pageNumber="355" startId="18.[151,250,1430,1454]" targetBox="[151,1436,276,1407]" targetPageId="18" targetType="figure">
|
||
<paragraph id="8BFD3699FFB4744181B438CEFAC9FA41" blockId="18.[151,1303,1430,1455]" box="[151,1303,1430,1455]" pageId="18" pageNumber="355">
|
||
<emphasis id="B936EA8BFFB4744181B438CEFEC7FA41" bold="true" box="[151,281,1430,1454]" pageId="18" pageNumber="355">FIGURE 18</emphasis>
|
||
.
|
||
<taxonomicName id="4C424D1AFFB47441800538CFFDD8FA41" authorityName="Khaustov & Frolov & Akhmetova" authorityYear="2024" box="[294,518,1431,1455]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="355" phylum="Arthropoda" rank="species" species="incisus" status="sp. nov.">
|
||
<emphasis id="B936EA8BFFB47441800538CFFDD8FA41" box="[294,518,1431,1455]" italics="true" pageId="18" pageNumber="355">Spatulaphorus incisus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B936EA8BFFB47441832E38CFFD83FA40" bold="true" box="[525,605,1431,1455]" pageId="18" pageNumber="355">
|
||
<taxonomicNameLabel id="A20557F0FFB47441832E38CFFD83FA40" box="[525,605,1431,1455]" pageId="18" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD3699FFB4744181E43886FB8DF800" blockId="18.[151,1437,1501,2031]" pageId="18" pageNumber="355">
|
||
<emphasis id="B936EA8BFFB4744181E43886FF22FA18" box="[199,252,1502,1527]" italics="true" pageId="18" pageNumber="355">Legs</emphasis>
|
||
(
|
||
<figureCitation id="13792A1CFFB47441802F3885FEBAFA18" box="[268,356,1501,1527]" captionStart="FIGURE 18" captionStartId="18.[151,250,1430,1454]" captionTargetBox="[151,1436,276,1407]" captionTargetId="figure-87@18.[151,1436,273,1407]" captionTargetPageId="18" captionText="FIGURE 18. Spatulaphorus incisus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618653" httpUri="https://zenodo.org/record/13618653/files/figure.png" pageId="18" pageNumber="355">Figs 18</figureCitation>
|
||
,
|
||
<figureCitation id="13792A1CFFB4744180523885FE4EFA18" box="[369,400,1501,1527]" captionStart="FIGURE 19" captionStartId="19.[151,250,1827,1851]" captionTargetBox="[174,1413,189,1804]" captionTargetId="figure-17@19.[151,1436,181,1804]" captionTargetPageId="19" captionText="FIGURE 19. Spatulaphorus incisus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618655" httpUri="https://zenodo.org/record/13618655/files/figure.png" pageId="18" pageNumber="355">19</figureCitation>
|
||
). Setation of legs as in
|
||
<taxonomicName id="4C424D1AFFB4744183B93885FC9EFA18" authorityName="Khaustov & Frolov & Akhmetova" authorityYear="2024" box="[666,832,1501,1527]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="355" phylum="Arthropoda" rank="species" species="brevisetosus">
|
||
<emphasis id="B936EA8BFFB4744183B93885FC9EFA18" box="[666,832,1501,1527]" italics="true" pageId="18" pageNumber="355">S. brevisetosus</emphasis>
|
||
</taxonomicName>
|
||
. Leg I (
|
||
<figureCitation id="13792A1CFFB4744182BE3885FBD9FA18" box="[925,1031,1501,1527]" captionStart="FIGURE 18" captionStartId="18.[151,250,1430,1454]" captionTargetBox="[151,1436,276,1407]" captionTargetId="figure-87@18.[151,1436,273,1407]" captionTargetPageId="18" captionText="FIGURE 18. Spatulaphorus incisus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618653" httpUri="https://zenodo.org/record/13618653/files/figure.png" pageId="18" pageNumber="355">Fig. 18A</figureCitation>
|
||
). Tibiotarsus slightly enlarged, with large and long claw; tibiotarsus internally with oval strongly sclerotized structure near solenidion
|
||
<emphasis id="B936EA8BFFB4744184253B59FAFAF9F4" box="[1286,1316,1537,1563]" italics="true" pageId="18" pageNumber="355">φ2</emphasis>
|
||
. Setae (
|
||
<emphasis id="B936EA8BFFB4744184A63B5AFA4DF9F4" box="[1413,1427,1538,1563]" italics="true" pageId="18" pageNumber="355">u</emphasis>
|
||
) consolidated into a bifid horn-like structure opposing claw. Tarsal claw weakly blunt-tipped. Seta
|
||
<emphasis id="B936EA8BFFB47441843E3B7DFAF5F9D0" box="[1309,1323,1573,1599]" italics="true" pageId="18" pageNumber="355">d</emphasis>
|
||
of femur spatulate, with short subterminal projection. Setae (
|
||
<emphasis id="B936EA8BFFB4744183FD3B11FD38F98C" box="[734,742,1609,1635]" italics="true" pageId="18" pageNumber="355">l</emphasis>
|
||
) of genu and
|
||
<emphasis id="B936EA8BFFB4744182A63B11FC7EF98C" box="[901,928,1609,1635]" italics="true" pageId="18" pageNumber="355">v”</emphasis>
|
||
of tibiotarsus weakly barbed, other leg setae (except eupathidia
|
||
<emphasis id="B936EA8BFFB47441804D3B35FE5BF968" box="[366,389,1645,1671]" italics="true" pageId="18" pageNumber="355">p’</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB4744180B73B35FE6FF968" box="[404,433,1645,1671]" italics="true" pageId="18" pageNumber="355">p”</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB4744180E33B35FE00F968" box="[448,478,1645,1671]" italics="true" pageId="18" pageNumber="355">tc’</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB4744180CF3B35FDCEF968" box="[492,528,1645,1671]" italics="true" pageId="18" pageNumber="355">tc”</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB47441833C3B35FDE6F968" box="[543,568,1645,1671]" italics="true" pageId="18" pageNumber="355">ft’</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB4744183653B35FDB8F968" box="[582,614,1645,1671]" italics="true" pageId="18" pageNumber="355">ft”</emphasis>
|
||
) smooth. Setae
|
||
<emphasis id="B936EA8BFFB47441823F3B35FCF4F968" box="[796,810,1645,1671]" italics="true" pageId="18" pageNumber="355">l’</emphasis>
|
||
of femur, (
|
||
<emphasis id="B936EA8BFFB4744182883B35FC6DF968" box="[939,947,1645,1671]" italics="true" pageId="18" pageNumber="355">l</emphasis>
|
||
) of genu and
|
||
<emphasis id="B936EA8BFFB4744185723B35FB83F968" box="[1105,1117,1645,1671]" italics="true" pageId="18" pageNumber="355">k</emphasis>
|
||
of tibiotarsus weakly blunt-tipped; other leg setae (except eupathidia) pointed. Lengths of solenidia
|
||
<emphasis id="B936EA8BFFB4744182E23BC9FC3CF944" box="[961,994,1681,1707]" italics="true" pageId="18" pageNumber="355">ω1</emphasis>
|
||
6 (6–7),
|
||
<emphasis id="B936EA8BFFB47441856A3BC9FBB5F944" box="[1097,1131,1681,1707]" italics="true" pageId="18" pageNumber="355">ω2</emphasis>
|
||
4 (4),
|
||
<emphasis id="B936EA8BFFB4744185953BC9FB0DF944" box="[1206,1235,1681,1707]" italics="true" pageId="18" pageNumber="355">φ1</emphasis>
|
||
6 (6–7),
|
||
<emphasis id="B936EA8BFFB4744184193BC9FA86F944" box="[1338,1368,1681,1707]" italics="true" pageId="18" pageNumber="355">φ2</emphasis>
|
||
4 (4); solenidion
|
||
<emphasis id="B936EA8BFFB4744180313BEDFEEDF920" box="[274,307,1717,1743]" italics="true" pageId="18" pageNumber="355">ω1</emphasis>
|
||
digitiform; solenidion
|
||
<emphasis id="B936EA8BFFB4744183113BEDFD91F920" box="[562,591,1717,1743]" italics="true" pageId="18" pageNumber="355">φ1</emphasis>
|
||
thick, clavate; other solenidia weakly clavate. Leg II (
|
||
<figureCitation id="13792A1CFFB4744185863BEDFAD4F920" box="[1189,1290,1717,1743]" captionStart="FIGURE 18" captionStartId="18.[151,250,1430,1454]" captionTargetBox="[151,1436,276,1407]" captionTargetId="figure-87@18.[151,1436,273,1407]" captionTargetPageId="18" captionText="FIGURE 18. Spatulaphorus incisus sp. nov., female: A—right leg I, dorsal aspect; B—right leg II, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618653" httpUri="https://zenodo.org/record/13618653/files/figure.png" pageId="18" pageNumber="355">Fig. 18B</figureCitation>
|
||
). Tarsus with thickened basally claws and pad-like empodium. Solenidion
|
||
<emphasis id="B936EA8BFFB4744182643B82FC85F91C" box="[839,859,1754,1779]" italics="true" pageId="18" pageNumber="355">ω</emphasis>
|
||
5 (5–6) weakly clavate, solenidion
|
||
<emphasis id="B936EA8BFFB4744185D13B82FADFF91C" box="[1266,1281,1754,1779]" italics="true" pageId="18" pageNumber="355">φ</emphasis>
|
||
absent. Setae
|
||
<emphasis id="B936EA8BFFB4744181B43BA5FF76F8F8" box="[151,168,1789,1815]" italics="true" pageId="18" pageNumber="355">l’</emphasis>
|
||
of femur,
|
||
<emphasis id="B936EA8BFFB47441803D3BA5FEEBF8F8" box="[286,309,1789,1815]" italics="true" pageId="18" pageNumber="355">u’</emphasis>
|
||
, and
|
||
<emphasis id="B936EA8BFFB4744180573BA5FE46F8F8" box="[372,408,1789,1815]" italics="true" pageId="18" pageNumber="355">tc”</emphasis>
|
||
of tarsus smooth, other setae barbed; setae
|
||
<emphasis id="B936EA8BFFB4744182A13BA5FC4EF8F8" box="[898,912,1789,1815]" italics="true" pageId="18" pageNumber="355">d</emphasis>
|
||
and
|
||
<emphasis id="B936EA8BFFB4744182E43BA5FC06F8F8" box="[967,984,1789,1815]" italics="true" pageId="18" pageNumber="355">l’</emphasis>
|
||
of femur weakly blunt-tipped; other leg setae pointed. Leg III (
|
||
<figureCitation id="13792A1CFFB4744180BA3A79FDDCF8D4" box="[409,514,1825,1851]" captionStart="FIGURE 19" captionStartId="19.[151,250,1827,1851]" captionTargetBox="[174,1413,189,1804]" captionTargetId="figure-17@19.[151,1436,181,1804]" captionTargetPageId="19" captionText="FIGURE 19. Spatulaphorus incisus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618655" httpUri="https://zenodo.org/record/13618655/files/figure.png" pageId="18" pageNumber="355">Fig. 19A</figureCitation>
|
||
). Femur divided into basi- and telofemur. Claws and empodium of same shape as on tarsus II. Setae
|
||
<emphasis id="B936EA8BFFB4744180453A1DFEA2F8B0" box="[358,380,1861,1887]" italics="true" pageId="18" pageNumber="355">v’</emphasis>
|
||
of trochanter and
|
||
<emphasis id="B936EA8BFFB47441836B3A1DFD81F8B0" box="[584,607,1861,1887]" italics="true" pageId="18" pageNumber="355">u’</emphasis>
|
||
of tarsus smooth, other setae barbed; setae
|
||
<emphasis id="B936EA8BFFB4744185663A1DFB85F8B0" box="[1093,1115,1861,1887]" italics="true" pageId="18" pageNumber="355">v’</emphasis>
|
||
of trochanter,
|
||
<emphasis id="B936EA8BFFB4744185DE3A1DFAD5F8B0" box="[1277,1291,1861,1887]" italics="true" pageId="18" pageNumber="355">d</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB47441843A3A1DFAF1F8B0" box="[1305,1327,1861,1887]" italics="true" pageId="18" pageNumber="355">v’</emphasis>
|
||
of femur, and
|
||
<emphasis id="B936EA8BFFB4744181E53A31FF09F86C" box="[198,215,1897,1923]" italics="true" pageId="18" pageNumber="355">l’</emphasis>
|
||
of genu blunt-tipped, other leg setae pointed; solenidion
|
||
<emphasis id="B936EA8BFFB4744182723A32FCBEF86C" box="[849,864,1898,1923]" italics="true" pageId="18" pageNumber="355">φ</emphasis>
|
||
absent. Leg IV (
|
||
<figureCitation id="13792A1CFFB47441853B3A31FBA9F86C" box="[1048,1143,1897,1923]" captionStart="FIGURE 19" captionStartId="19.[151,250,1827,1851]" captionTargetBox="[174,1413,189,1804]" captionTargetId="figure-17@19.[151,1436,181,1804]" captionTargetPageId="19" captionText="FIGURE 19. Spatulaphorus incisus sp. nov., female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.13618655" httpUri="https://zenodo.org/record/13618655/files/figure.png" pageId="18" pageNumber="355">Fig 19B</figureCitation>
|
||
). Femur divided into basi- and telofemur. Claws simple, empodium as on tarsi II and III. Setae
|
||
<emphasis id="B936EA8BFFB4744182AC3AD5FC7BF848" box="[911,933,1933,1959]" italics="true" pageId="18" pageNumber="355">v’</emphasis>
|
||
of trochanter and
|
||
<emphasis id="B936EA8BFFB4744185533AD5FB48F848" box="[1136,1174,1933,1959]" italics="true" pageId="18" pageNumber="355">pl”</emphasis>
|
||
of tarsus smooth; other setae barbed; setae
|
||
<emphasis id="B936EA8BFFB4744180533AE9FE58F824" box="[368,390,1969,1995]" italics="true" pageId="18" pageNumber="355">v’</emphasis>
|
||
of trochanter,
|
||
<emphasis id="B936EA8BFFB4744183093AE9FDE6F824" box="[554,568,1969,1995]" italics="true" pageId="18" pageNumber="355">d</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB4744183653AE9FD82F824" box="[582,604,1969,1995]" italics="true" pageId="18" pageNumber="355">v’</emphasis>
|
||
of femur,
|
||
<emphasis id="B936EA8BFFB4744183F13AE9FD3DF824" box="[722,739,1969,1995]" italics="true" pageId="18" pageNumber="355">l’</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB4744183D23AE9FCD2F824" box="[753,780,1969,1995]" italics="true" pageId="18" pageNumber="355">v”</emphasis>
|
||
of tibia and
|
||
<emphasis id="B936EA8BFFB4744182B83AE9FC1FF824" box="[923,961,1969,1995]" italics="true" pageId="18" pageNumber="355">pl”</emphasis>
|
||
of tarsus blunt-tipped; tips of setae
|
||
<emphasis id="B936EA8BFFB4744184703AE9FAB4F824" box="[1363,1386,1969,1995]" italics="true" pageId="18" pageNumber="355">u’</emphasis>
|
||
,
|
||
<emphasis id="B936EA8BFFB47441845B3AE9FA42F824" box="[1400,1436,1969,1995]" italics="true" pageId="18" pageNumber="355">tc”</emphasis>
|
||
and
|
||
<emphasis id="B936EA8BFFB4744181E43A8DFF2EF800" box="[199,240,2005,2031]" italics="true" pageId="18" pageNumber="355">pv”</emphasis>
|
||
slightly thickened and flattened; other leg setae pointed; solenidion
|
||
<emphasis id="B936EA8BFFB4744182CD3A8EFC23F800" box="[1006,1021,2006,2031]" italics="true" pageId="18" pageNumber="355">φ</emphasis>
|
||
absent.
|
||
</paragraph>
|
||
<caption id="DF3D6611FFB5744081B43A7BFAF0F8D4" ID-DOI="http://doi.org/10.5281/zenodo.13618655" ID-Zenodo-Dep="13618655" box="[151,1326,1827,1852]" httpUri="https://zenodo.org/record/13618655/files/figure.png" pageId="19" pageNumber="356" startId="19.[151,250,1827,1851]" targetBox="[174,1413,189,1804]" targetPageId="19" targetType="figure">
|
||
<paragraph id="8BFD3699FFB5744081B43A7BFAF0F8D4" blockId="19.[151,1326,1827,1852]" box="[151,1326,1827,1852]" pageId="19" pageNumber="356">
|
||
<emphasis id="B936EA8BFFB5744081B43A7BFEC7F8D4" bold="true" box="[151,281,1827,1851]" pageId="19" pageNumber="356">FIGURE 19</emphasis>
|
||
.
|
||
<taxonomicName id="4C424D1AFFB5744080053A7BFDD8F8D4" authorityName="Khaustov & Frolov & Akhmetova" authorityYear="2024" box="[294,518,1827,1851]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="19" pageNumber="356" phylum="Arthropoda" rank="species" species="incisus" status="sp. nov.">
|
||
<emphasis id="B936EA8BFFB5744080053A7BFDD8F8D4" box="[294,518,1827,1851]" italics="true" pageId="19" pageNumber="356">Spatulaphorus incisus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B936EA8BFFB57440832E3A7CFD83F8D3" bold="true" box="[525,605,1828,1852]" pageId="19" pageNumber="356">
|
||
<taxonomicNameLabel id="A20557F0FFB57440832E3A7CFD83F8D3" box="[525,605,1828,1852]" pageId="19" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female: A—right leg III, dorsal aspect; B—right leg IV, dorsal aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD3699FFB5744081EB3A32FE4FF86B" blockId="19.[151,1436,1898,2032]" box="[200,401,1898,1924]" pageId="19" pageNumber="356">MALE unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3586512FFB5744781E43AD6FD9BFF3A" lastPageId="20" lastPageNumber="357" pageId="19" pageNumber="356" type="materials_examined">
|
||
<paragraph id="8BFD3699FFB5744081E43AD6FBF1F823" blockId="19.[151,1436,1898,2032]" pageId="19" pageNumber="356">
|
||
<emphasis id="B936EA8BFFB5744081E43AD6FEB0F848" bold="true" box="[199,366,1933,1960]" pageId="19" pageNumber="356">Type material</emphasis>
|
||
. Female
|
||
<materialsCitation id="3B2A3CC4FFB5744080F63AD6FD75F823" collectingDate="2017-02-26" collectionCode="ZISP, ESEC, CEMT" country="Brazil" location="Rio Ronuro" pageId="19" pageNumber="356" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
|
||
<typeStatus id="54F9883BFFB5744080F63AD6FDE4F847" box="[469,570,1934,1960]" pageId="19" pageNumber="356" type="holotype">holotype</typeStatus>
|
||
, slide
|
||
<collectionCode id="ED53AE5CFFB5744083A33AD6FD67F847" box="[640,697,1934,1960]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="19" pageNumber="356">ZISP</collectionCode>
|
||
T-Pygm-014,
|
||
<collectingCountry id="F3557609FFB5744082793AD6FC7FF847" box="[858,929,1934,1960]" name="Brazil" pageId="19" pageNumber="356">Brazil</collectingCountry>
|
||
,
|
||
<collectingRegion id="4986F87BFFB57440828F3AD6FB9EF847" box="[940,1088,1934,1960]" country="Brazil" name="Mato Grosso" pageId="19" pageNumber="356">Mato Grosso</collectingRegion>
|
||
,
|
||
<collectionCode id="ED53AE5CFFB5744085683AD6FB51F847" box="[1099,1167,1934,1960]" country="Egypt" httpUri="http://grbio.org/cool/4cts-0w01" name="Entomological Society of Egypt" pageId="19" pageNumber="356">ESEC</collectionCode>
|
||
<location id="8E9D6042FFB5744085B53AD6FAC3F847" LSID="urn:lsid:plazi:treatment:03EB878FFFB6744781B43B0AFC07FE1A:8E9D6042FFB5744085B53AD6FAC3F847" box="[1174,1309,1934,1960]" country="Brazil" name="Rio Ronuro" pageId="19" pageNumber="356" stateProvince="Mato Grosso">Rio Ronuro</location>
|
||
,
|
||
<date id="FFFC1059FFB57440840B3AD6FA42F847" box="[1320,1436,1934,1960]" pageId="19" pageNumber="356" value="2017-02-26">
|
||
<collectingDate id="EFB8E9B1FFB57440840B3AD6FA42F847" box="[1320,1436,1934,1960]" pageId="19" pageNumber="356" value="2017-02-26">26.II.2017</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED53AE5CFFB5744081BC3AEAFF2CF823" box="[159,242,1970,1996]" pageId="19" pageNumber="356">CEMT</collectionCode>
|
||
),
|
||
<collectedFrom id="385E7564FFB5744080253AEAFD75F823" box="[262,683,1970,1996]" pageId="19" pageNumber="356">
|
||
on
|
||
<taxonomicName id="4C424D1AFFB57440800A3AEAFD75F823" authority="(MacLeay)" baseAuthorityName="MacLeay" box="[297,683,1970,1996]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="356" phylum="Arthropoda" rank="species" species="faunus">
|
||
<emphasis id="B936EA8BFFB57440800A3AEAFDFAF823" box="[297,548,1970,1996]" italics="true" pageId="19" pageNumber="356">Sulcophanaeus faunus</emphasis>
|
||
(MacLeay)
|
||
</taxonomicName>
|
||
</collectedFrom>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B2A3CC4FFB5744083953AEAFBF5F823" box="[694,1067,1970,1996]" collectingDate="2017-02-26" collectionCode="Male" country="Brazil" location="Rio Ronuro" pageId="19" pageNumber="356" specimenCount="42" specimenCount-female="42" stateProvince="Mato Grosso" typeStatus="paratype">
|
||
<typeStatus id="54F9883BFFB5744083953AEAFCF8F823" box="[694,806,1970,1996]" pageId="19" pageNumber="356" type="paratype">paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D44FD10FFB5744082133AEAFC73F823" box="[816,941,1970,1996]" count="42" pageId="19" pageNumber="356" type="female">42 females</specimenCount>
|
||
, same data
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFD3699FFB5744781E43A8EFD9BFF3A" blockId="19.[151,1436,1898,2032]" lastBlockId="20.[151,1437,151,501]" lastPageId="20" lastPageNumber="357" pageId="19" pageNumber="356">
|
||
<emphasis id="B936EA8BFFB5744081E43A8EFE58F800" bold="true" box="[199,390,2005,2032]" pageId="19" pageNumber="356">Type deposition</emphasis>
|
||
.
|
||
<materialsCitation id="3B2A3CC4FFB5744780B43A8EFE25FF5E" country="Russia" lastPageId="20" lastPageNumber="357" location="Zoological Institute" pageId="19" pageNumber="356" specimenCount="4" stateProvince="Saint Petersburg" typeStatus="holotype">
|
||
The
|
||
<typeStatus id="54F9883BFFB5744080EF3A8EFDF0F81F" box="[460,558,2006,2032]" pageId="19" pageNumber="356" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D44FD10FFB5744083493A8EFCD3F81F" box="[618,781,2006,2032]" count="4" pageId="19" pageNumber="356" type="generic" typeStatus="paratype">four paratypes</specimenCount>
|
||
are deposited in the collection of
|
||
<location id="8E9D6042FFB5744085BF3A8EFAA5F81F" LSID="urn:lsid:plazi:treatment:03EB878FFFB6744781B43B0AFC07FE1A:8E9D6042FFB5744085BF3A8EFAA5F81F" box="[1180,1403,2006,2032]" country="Russia" name="Zoological Institute" pageId="19" pageNumber="356" stateProvince="Saint Petersburg">Zoological Institute</location>
|
||
of RAS,
|
||
<collectingRegion id="4986F87BFFB2744781FC3DCFFE40FF5E" box="[223,414,151,177]" country="Russia" name="Saint Petersburg" pageId="20" pageNumber="357">Saint Petersburg</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3557609FFB27447808E3DCFFE25FF5E" box="[429,507,151,177]" name="Russia" pageId="20" pageNumber="357">Russia</collectingCountry>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B2A3CC4FFB2744783293DCFFAC5FF5E" box="[522,1307,151,177]" country="Russia" location="Tyumen State" pageId="20" pageNumber="357" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
|
||
other
|
||
<typeStatus id="54F9883BFFB27447836D3DCFFD67FF5E" box="[590,697,151,177]" pageId="20" pageNumber="357" type="paratype">paratypes</typeStatus>
|
||
are deposited in the collection of the
|
||
<collectingRegion id="4986F87BFFB27447855E3DCFFAC5FF5E" box="[1149,1307,151,177]" country="Russia" name="Tyumen'" pageId="20" pageNumber="357">Tyumen State</collectingRegion>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B2A3CC4FFB2744784053DCFFD9EFF3A" country="Russia" location="University Museum of Zoology" pageId="20" pageNumber="357" specimenCount="1" stateProvince="Tyumen'" typeStatus="paratype">
|
||
<location id="8E9D6042FFB2744784053DCFFEA1FF3A" LSID="urn:lsid:plazi:treatment:03EB878FFFB6744781B43B0AFC07FE1A:8E9D6042FFB2744784053DCFFEA1FF3A" country="Russia" name="University Museum of Zoology" pageId="20" pageNumber="357" stateProvince="Tyumen'">University Museum of Zoology</location>
|
||
,
|
||
<collectingRegion id="4986F87BFFB2744780AA3DE3FE39FF3A" box="[393,487,187,213]" country="Russia" name="Tyumen'" pageId="20" pageNumber="357">Tyumen</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3557609FFB2744780D13DE3FD9EFF3A" box="[498,576,187,213]" name="Russia" pageId="20" pageNumber="357">Russia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3586512FFB2744781E43D86FD70FE42" pageId="20" pageNumber="357" type="diagnosis">
|
||
<paragraph id="8BFD3699FFB2744781E43D86FD70FE42" blockId="20.[151,1437,151,501]" pageId="20" pageNumber="357">
|
||
<emphasis id="B936EA8BFFB2744781E43D86FE1BFF17" bold="true" box="[199,453,222,248]" pageId="20" pageNumber="357">Differential diagnosis</emphasis>
|
||
. The new species is very similar to
|
||
<taxonomicName id="4C424D1AFFB2744782713D87FAF9FF16" authority="Khaustov and Frolov, 2017" authorityName="Khaustov and Frolov" authorityYear="2017" box="[850,1319,223,249]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B936EA8BFFB2744782713D87FCBEFF16" box="[850,864,223,249]" italics="true" pageId="20" pageNumber="357">S</emphasis>
|
||
.
|
||
<emphasis id="B936EA8BFFB27447824D3D87FC31FF16" box="[878,1007,223,249]" italics="true" pageId="20" pageNumber="357">brasiliensis</emphasis>
|
||
<bibRefCitation id="EFD34B68FFB2744782D63D87FAF9FF16" author="Khaustov, A. A. & Frolov, A. V." box="[1013,1319,223,249]" pageId="20" pageNumber="357" pagination="501 - 521" refId="ref16128" refString="Khaustov, A. A. & Frolov, A. V. (2017) New species of heterostigmatic mites (Acari: Heterostigmata: Athyreacaridae, Dolichocybidae, Pygmephoridae) associated with scarab beetles (Coleoptera: Geotrupidae, Scarabaeidae) from Brazil. Zootaxa, 4294 (5), 501 - 521. https: // doi. org / 10.11646 / zootaxa. 4294.5.1" type="journal article" year="2017">Khaustov and Frolov, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
, described from
|
||
<collectingCountry id="F3557609FFB2744781F73C5BFEC6FEF2" box="[212,280,259,285]" name="Brazil" pageId="20" pageNumber="357">Brazil</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFD34B68FFB2744780053C5BFD9DFEF2" author="Khaustov, A. A. & Frolov, A. V." box="[294,579,259,285]" pageId="20" pageNumber="357" pagination="501 - 521" refId="ref16128" refString="Khaustov, A. A. & Frolov, A. V. (2017) New species of heterostigmatic mites (Acari: Heterostigmata: Athyreacaridae, Dolichocybidae, Pygmephoridae) associated with scarab beetles (Coleoptera: Geotrupidae, Scarabaeidae) from Brazil. Zootaxa, 4294 (5), 501 - 521. https: // doi. org / 10.11646 / zootaxa. 4294.5.1" type="journal article" year="2017">Khaustov & Frolov 2017</bibRefCitation>
|
||
), in having unmodified setae on tarsi II and III, presence of big puncta on the dorsal idiosomal sclerites, and basally thickened claws on tarsi II and III. The new species differs from
|
||
<taxonomicName id="4C424D1AFFB2744784223C7FFA42FEAE" authorityName="Khaustov and Frolov" authorityYear="2017" box="[1281,1436,295,321]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B936EA8BFFB2744784223C7FFAD1FEAE" box="[1281,1295,295,321]" italics="true" pageId="20" pageNumber="357">S</emphasis>
|
||
.
|
||
<emphasis id="B936EA8BFFB2744784383C7FFA42FEAE" box="[1307,1436,295,321]" italics="true" pageId="20" pageNumber="357">brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
in having setae
|
||
<emphasis id="B936EA8BFFB2744780653C13FE5AFE8A" box="[326,388,331,357]" italics="true" pageId="20" pageNumber="357">ps1-2</emphasis>
|
||
barbed and weakly blunt-tipped (vs. setae
|
||
<emphasis id="B936EA8BFFB2744782473C13FC7CFE8A" box="[868,930,331,357]" italics="true" pageId="20" pageNumber="357">ps1-2</emphasis>
|
||
smooth and pointed in
|
||
<taxonomicName id="4C424D1AFFB2744785893C13FA99FE8A" authorityName="Khaustov and Frolov" authorityYear="2017" box="[1194,1351,331,357]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B936EA8BFFB2744785893C13FA99FE8A" box="[1194,1351,331,357]" italics="true" pageId="20" pageNumber="357">S. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
) and in having an unusually deeply incised posterior margin of the posterior sternal plate (vs. posterior margin of posterior sternal plate slightly concave in
|
||
<taxonomicName id="4C424D1AFFB2744783223CCBFD40FE42" authorityName="Khaustov and Frolov" authorityYear="2017" box="[513,670,403,429]" class="Insecta" family="Scarabaeidae" genus="Sulcophanaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B936EA8BFFB2744783223CCBFD40FE42" box="[513,670,403,429]" italics="true" pageId="20" pageNumber="357">S. brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3586512FFB2744781E43CEFFC07FE1A" pageId="20" pageNumber="357" type="etymology">
|
||
<paragraph id="8BFD3699FFB2744781E43CEFFC07FE1A" blockId="20.[151,1437,151,501]" pageId="20" pageNumber="357">
|
||
<emphasis id="B936EA8BFFB2744781E43CEFFE96FE3E" bold="true" box="[199,328,439,465]" pageId="20" pageNumber="357">Etymology</emphasis>
|
||
. The name of the new species is derived from Latin
|
||
<taxonomicName id="4C424D1AFFB2744782993CEFFBD6FE3F" authorityName="Khaustov & Frolov & Akhmetova" authorityYear="2024" box="[954,1032,439,464]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="357" phylum="Arthropoda" rank="species" species="incisus">
|
||
<emphasis id="B936EA8BFFB2744782993CEFFBD6FE3F" box="[954,1032,439,464]" italics="true" pageId="20" pageNumber="357">incisus</emphasis>
|
||
</taxonomicName>
|
||
meaning
|
||
<emphasis id="B936EA8BFFB27447855F3CEFFB15FE3E" box="[1148,1227,439,465]" italics="true" pageId="20" pageNumber="357">incised</emphasis>
|
||
, and refers to the unusual deep incision in the posterior margin of the posterior sternal plate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |