233 lines
34 KiB
XML
233 lines
34 KiB
XML
<document id="45A165E0ECA0B0547184853BA52C3F45" ID-DOI="10.1080/00222933.2018.1481235" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747141" ID-ZooBank="63A31AD2-F049-42CB-A45B-557014DC286E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620670765052" checkinUser="carolina" docAuthor="Taylor, Paul D., Martha, Silviu O. & Gordon, Dennis P." docDate="2018" docId="03EB8789FFDA4443A1D7771BFCEEFE8D" docLanguage="en" docName="JNATHIST.52.25-26.1657-1721.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222933.2018.1481235" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Tyloporella Voigt 1989" docType="treatment" docVersion="4" lastPageNumber="1708" masterDocId="FFD2FFF1FFE84477A00C7447FF92FFBE" masterDocTitle="Synopsis of ‘ onychocellid’ cheilostome bryozoan genera" masterLastPageNumber="1721" masterPageNumber="1657" pageNumber="1706" updateTime="1698914560039" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EE6706E7F2769895457F685609F05EB0">Synopsis of ‘ onychocellid’ cheilostome bryozoan genera</mods:title>
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<mods:namePart id="6B8D27260A9E9B4D084912E8B7D04873">Taylor, Paul D.</mods:namePart>
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<mods:namePart id="EE9725C3C9BD604359C804B181207D23">Martha, Silviu O.</mods:namePart>
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<mods:namePart id="F83F4F909A5B8616F5E51949B517FC95">Gordon, Dennis P.</mods:namePart>
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<mods:title id="A86C8A9D3DE75175EDBA63CD14F33FA3">Journal of Natural History</mods:title>
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<mods:title id="5532A17523D61F336540AC4042B47959">J. Nat. Hist.</mods:title>
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<subSubSection id="C3586514FFDA4445A1D7771BFCDAFCCB" box="[475,840,860,885]" pageId="50" pageNumber="1706" type="nomenclature">
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<paragraph id="8BFD369FFFDA4445A1D7771BFCDAFCCB" blockId="50.[475,840,860,920]" box="[475,840,860,885]" pageId="50" pageNumber="1706">
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<heading id="D0B581F3FFDA4445A1D7771BFCDAFCCB" box="[475,840,860,885]" centered="true" fontSize="11" level="2" pageId="50" pageNumber="1706" reason="2">
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Genus
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<taxonomicName id="4C424D1CFFDA4445A222771BFCDAFCCB" ID-CoL="85JQ7" authority="Voigt, 1989" authorityName="Voigt" authorityYear="1989" box="[558,840,860,885]" class="Gymnolaemata" family="Onychocellidae" genus="Tyloporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="50" pageNumber="1706" phylum="Bryozoa" rank="genus">
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<emphasis id="B936EA8DFFDA4445A222771BFD2AFCCB" bold="true" box="[558,696,860,885]" italics="true" pageId="50" pageNumber="1706">Tyloporella</emphasis>
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<bibRefCitation id="EFD34B6EFFDA4445A2CD771BFCDAFCCB" author="Voigt E" box="[705,840,860,885]" pageId="50" pageNumber="1706" pagination="8 - 87" refId="ref29174" refString="Voigt E. 1989. Beitrag zur Bryozoen-Fauna des sachsischen Cenomaniums. Revision von A. E. Reuss' " Die Bryozoen des unteren Quaders " in H. B. Geinitz' " Das Elbthalgebirge in Sachsen " (1872). I: Cheilostomata. Abh Staatl Mus Mineral Geol Dresden. 36: 8 - 87." type="journal article" year="1989">Voigt, 1989</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3586514FFDA4445A25D7738FD40FC26" box="[593,722,895,920]" pageId="50" pageNumber="1706" type="description">
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<paragraph id="8BFD369FFFDA4445A25D7738FD40FC26" blockId="50.[475,840,860,920]" box="[593,722,895,920]" pageId="50" pageNumber="1706">
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(
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||
<figureCitation id="13792A1AFFDA4445A2557738FD58FC26" box="[601,714,895,920]" captionStart="Figure 33" captionStartId="51.[160,225,946,969]" captionTargetBox="[163,1153,142,915]" captionTargetId="figure-188@51.[163,1153,142,915]" captionTargetPageId="51" captionText="Figure 33. Tyloporella reussi Voigt. (a,b) Upper Cenomanian (Actinocamax plenus zone), Hoher Stein Quarry in Dresden-Plauen, Saxony, Germany; (a) holotype, MMG SaK 1617; part of colony with two broken ovicells indicated by arrows. (b) MMG SaK 15777a; autozooids and avicularia at growing edge. (c) NMP, Novak Collection (figured Novak 1877, pl. 1, fig. 19, 20 as Membranipora irregularis (von Hagenow)); Late Cretaceous,?Turonian, Kamajka, Bohemia, Czech Republic; autozooids and avicularia; an autozooid with the opesia covered by a closure plate is arrowed. (d) NHMUK BZ 8221; Cretaceous. Late Cenomanian or early Turonian, Kaňk, Central Bohemian Region, Czech Republic; autozooids (some with closure plates) and avicularia. Scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747220" httpUri="https://zenodo.org/record/4747220/files/figure.png" pageId="50" pageNumber="1706">Figure 33</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3586514FFDA4445A0AC7780FE9EFBF6" pageId="50" pageNumber="1706" type="type_taxon">
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<paragraph id="8BFD369FFFDA4445A0AC7780FEA6FC5E" blockId="50.[160,1156,967,1096]" box="[160,308,967,992]" pageId="50" pageNumber="1706">
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<emphasis id="B936EA8DFFDA4445A0AC7780FEA6FC5E" bold="true" box="[160,308,967,992]" italics="true" pageId="50" pageNumber="1706">
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<typeStatus id="54F9883DFFDA4445A0AC7780FF4AFC5E" box="[160,216,967,992]" pageId="50" pageNumber="1706">Type</typeStatus>
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species
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</emphasis>
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</paragraph>
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<paragraph id="8BFD369FFFDA4445A0AC77ACFE9EFBF6" blockId="50.[160,1156,967,1096]" pageId="50" pageNumber="1706">
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<taxonomicName id="4C424D1CFFDA4445A0AC77ACFE64FBBD" ID-CoL="86V48" authority="Voigt, 1989" authorityName="Voigt" authorityYear="1989" box="[160,502,1003,1027]" class="Gymnolaemata" family="Onychocellidae" genus="Tyloporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="50" pageNumber="1706" phylum="Bryozoa" rank="species" species="reussi">
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<emphasis id="B936EA8DFFDA4445A0AC77ACFEF0FBBD" box="[160,354,1003,1027]" italics="true" pageId="50" pageNumber="1706">Tyloporella reussi</emphasis>
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<bibRefCitation id="EFD34B6EFFDA4445A17C77ACFE64FBBD" author="Voigt E" box="[368,502,1003,1027]" pageId="50" pageNumber="1706" pagination="8 - 87" refId="ref29174" refString="Voigt E. 1989. Beitrag zur Bryozoen-Fauna des sachsischen Cenomaniums. Revision von A. E. Reuss' " Die Bryozoen des unteren Quaders " in H. B. Geinitz' " Das Elbthalgebirge in Sachsen " (1872). I: Cheilostomata. Abh Staatl Mus Mineral Geol Dresden. 36: 8 - 87." type="journal article" year="1989">Voigt, 1989</bibRefCitation>
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</taxonomicName>
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, by original designation.
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<materialsCitation id="3B2A3CC2FFDA4445A32277ACFE9BFBF6" country="Germany" location="Saxony" pageId="50" pageNumber="1706" specimenCount="1" stateProvince="Sachsen" typeStatus="holotype">
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Cretaceous, late Cenomanian (
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<taxonomicName id="4C424D1CFFDA4445A0AB7049FE18FB98" ID-CoL="BT6H7" baseAuthorityName="de Blainville" baseAuthorityYear="1827" box="[167,394,1038,1062]" class="Cephalopoda" family="Belemnitellidae" genus="Actinocamax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Belemnitida" pageId="50" pageNumber="1706" phylum="Mollusca" rank="species" species="plenus">
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<emphasis id="B936EA8DFFDA4445A0AB7049FE18FB98" box="[167,394,1038,1062]" italics="true" pageId="50" pageNumber="1706">Actinocamax plenus</emphasis>
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</taxonomicName>
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ammonite Zone), Hoher Stein quarry, Dresden-Plauen,
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<collectingRegion id="4986F87DFFDA4445A4217049FB12FB98" box="[1069,1152,1038,1062]" country="Germany" name="Sachsen" pageId="50" pageNumber="1706">Saxony</collectingRegion>
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,
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<collectingCountry id="F355760FFFDA4445A0AC7077FE9BFBF6" box="[160,265,1072,1096]" name="Germany" pageId="50" pageNumber="1706">Germany</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3586514FFDA4445A0AC703FFEBAFA77" pageId="50" pageNumber="1706" type="diagnosis">
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<paragraph id="8BFD369FFFDA4445A0AC703FFE86FB2F" blockId="50.[160,1156,1144,1481]" box="[160,276,1144,1169]" pageId="50" pageNumber="1706">
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<emphasis id="B936EA8DFFDA4445A0AC703FFE86FB2F" bold="true" box="[160,276,1144,1169]" italics="true" pageId="50" pageNumber="1706">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BFD369FFFDA4445A0AC70DBFEBAFA77" blockId="50.[160,1156,1144,1481]" pageId="50" pageNumber="1706">
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Colony encrusting, unilamellar (
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<figureCitation id="13792A1AFFDA4445A21F70DBFD17FB0A" box="[531,645,1180,1204]" captionStart="Figure 33" captionStartId="51.[160,225,946,969]" captionTargetBox="[163,1153,142,915]" captionTargetId="figure-188@51.[163,1153,142,915]" captionTargetPageId="51" captionText="Figure 33. Tyloporella reussi Voigt. (a,b) Upper Cenomanian (Actinocamax plenus zone), Hoher Stein Quarry in Dresden-Plauen, Saxony, Germany; (a) holotype, MMG SaK 1617; part of colony with two broken ovicells indicated by arrows. (b) MMG SaK 15777a; autozooids and avicularia at growing edge. (c) NMP, Novak Collection (figured Novak 1877, pl. 1, fig. 19, 20 as Membranipora irregularis (von Hagenow)); Late Cretaceous,?Turonian, Kamajka, Bohemia, Czech Republic; autozooids and avicularia; an autozooid with the opesia covered by a closure plate is arrowed. (d) NHMUK BZ 8221; Cretaceous. Late Cenomanian or early Turonian, Kaňk, Central Bohemian Region, Czech Republic; autozooids (some with closure plates) and avicularia. Scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747220" httpUri="https://zenodo.org/record/4747220/files/figure.png" pageId="50" pageNumber="1706">Figure 33</figureCitation>
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(a,b)). Autozooids pyriform to subhexagonal (
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<figureCitation id="13792A1AFFDA4445A0AB70F9FE87FB68" box="[167,277,1214,1238]" captionStart="Figure 33" captionStartId="51.[160,225,946,969]" captionTargetBox="[163,1153,142,915]" captionTargetId="figure-188@51.[163,1153,142,915]" captionTargetPageId="51" captionText="Figure 33. Tyloporella reussi Voigt. (a,b) Upper Cenomanian (Actinocamax plenus zone), Hoher Stein Quarry in Dresden-Plauen, Saxony, Germany; (a) holotype, MMG SaK 1617; part of colony with two broken ovicells indicated by arrows. (b) MMG SaK 15777a; autozooids and avicularia at growing edge. (c) NMP, Novak Collection (figured Novak 1877, pl. 1, fig. 19, 20 as Membranipora irregularis (von Hagenow)); Late Cretaceous,?Turonian, Kamajka, Bohemia, Czech Republic; autozooids and avicularia; an autozooid with the opesia covered by a closure plate is arrowed. (d) NHMUK BZ 8221; Cretaceous. Late Cenomanian or early Turonian, Kaňk, Central Bohemian Region, Czech Republic; autozooids (some with closure plates) and avicularia. Scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747220" httpUri="https://zenodo.org/record/4747220/files/figure.png" pageId="50" pageNumber="1706">Figure 33</figureCitation>
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(c,d)); zooidal boundaries grooved. Cryptocyst extensive, granular. Gymnocyst, when visible, narrow, surrounding cryptocyst proximally and proximolaterally. Opesia terminal, occupying about one-third of the frontal surface, semielliptical; two or three pairs of minute oral spine bases in a few autozooids; closure plates often bearing impressions of opercular occlusor muscles present in some zooids (
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<figureCitation id="13792A1AFFDA4445A3DA710EFBD8FADF" box="[982,1098,1353,1377]" captionStart="Figure 33" captionStartId="51.[160,225,946,969]" captionTargetBox="[163,1153,142,915]" captionTargetId="figure-188@51.[163,1153,142,915]" captionTargetPageId="51" captionText="Figure 33. Tyloporella reussi Voigt. (a,b) Upper Cenomanian (Actinocamax plenus zone), Hoher Stein Quarry in Dresden-Plauen, Saxony, Germany; (a) holotype, MMG SaK 1617; part of colony with two broken ovicells indicated by arrows. (b) MMG SaK 15777a; autozooids and avicularia at growing edge. (c) NMP, Novak Collection (figured Novak 1877, pl. 1, fig. 19, 20 as Membranipora irregularis (von Hagenow)); Late Cretaceous,?Turonian, Kamajka, Bohemia, Czech Republic; autozooids and avicularia; an autozooid with the opesia covered by a closure plate is arrowed. (d) NHMUK BZ 8221; Cretaceous. Late Cenomanian or early Turonian, Kaňk, Central Bohemian Region, Czech Republic; autozooids (some with closure plates) and avicularia. Scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747220" httpUri="https://zenodo.org/record/4747220/files/figure.png" pageId="50" pageNumber="1706">Figure 33</figureCitation>
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(c,d)). Ovicells hyperstomial (
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<figureCitation id="13792A1AFFDA4445A1BD712BFDB6FA3A" box="[433,548,1388,1412]" captionStart="Figure 33" captionStartId="51.[160,225,946,969]" captionTargetBox="[163,1153,142,915]" captionTargetId="figure-188@51.[163,1153,142,915]" captionTargetPageId="51" captionText="Figure 33. Tyloporella reussi Voigt. (a,b) Upper Cenomanian (Actinocamax plenus zone), Hoher Stein Quarry in Dresden-Plauen, Saxony, Germany; (a) holotype, MMG SaK 1617; part of colony with two broken ovicells indicated by arrows. (b) MMG SaK 15777a; autozooids and avicularia at growing edge. (c) NMP, Novak Collection (figured Novak 1877, pl. 1, fig. 19, 20 as Membranipora irregularis (von Hagenow)); Late Cretaceous,?Turonian, Kamajka, Bohemia, Czech Republic; autozooids and avicularia; an autozooid with the opesia covered by a closure plate is arrowed. (d) NHMUK BZ 8221; Cretaceous. Late Cenomanian or early Turonian, Kaňk, Central Bohemian Region, Czech Republic; autozooids (some with closure plates) and avicularia. Scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747220" httpUri="https://zenodo.org/record/4747220/files/figure.png" pageId="50" pageNumber="1706">Figure 33</figureCitation>
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(a)). Avicularia interzooidal, small, irregularly interspersed among autozooids (
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<figureCitation id="13792A1AFFDA4445A1EF71C9FDC2FA18" box="[483,592,1422,1446]" captionStart="Figure 33" captionStartId="51.[160,225,946,969]" captionTargetBox="[163,1153,142,915]" captionTargetId="figure-188@51.[163,1153,142,915]" captionTargetPageId="51" captionText="Figure 33. Tyloporella reussi Voigt. (a,b) Upper Cenomanian (Actinocamax plenus zone), Hoher Stein Quarry in Dresden-Plauen, Saxony, Germany; (a) holotype, MMG SaK 1617; part of colony with two broken ovicells indicated by arrows. (b) MMG SaK 15777a; autozooids and avicularia at growing edge. (c) NMP, Novak Collection (figured Novak 1877, pl. 1, fig. 19, 20 as Membranipora irregularis (von Hagenow)); Late Cretaceous,?Turonian, Kamajka, Bohemia, Czech Republic; autozooids and avicularia; an autozooid with the opesia covered by a closure plate is arrowed. (d) NHMUK BZ 8221; Cretaceous. Late Cenomanian or early Turonian, Kaňk, Central Bohemian Region, Czech Republic; autozooids (some with closure plates) and avicularia. Scale bars = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4747220" httpUri="https://zenodo.org/record/4747220/files/figure.png" pageId="50" pageNumber="1706">Figure 33</figureCitation>
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(a,c)), located distally or laterodistally of autozooidal opesiae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3586514FFDA4443A0AC71BEFCEEFE8D" lastPageId="52" lastPageNumber="1708" pageId="50" pageNumber="1706" type="discussion">
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<paragraph id="8BFD369FFFDA4445A0AC71BEFE94F9AC" blockId="50.[160,1157,1529,1727]" box="[160,262,1529,1554]" pageId="50" pageNumber="1706">
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<emphasis id="B936EA8DFFDA4445A0AC71BEFE94F9AC" bold="true" box="[160,262,1529,1554]" italics="true" pageId="50" pageNumber="1706">Remarks</emphasis>
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</paragraph>
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<paragraph id="8BFD369FFFDA4444A0AC725AFBFCFA49" blockId="50.[160,1157,1529,1727]" lastBlockId="51.[160,1157,1261,1701]" lastPageId="51" lastPageNumber="1707" pageId="50" pageNumber="1706">
|
||
<bibRefCitation id="EFD34B6EFFDA4445A0AC725AFEB3F98B" author="Voigt E" box="[160,289,1565,1589]" pageId="50" pageNumber="1706" pagination="8 - 87" refId="ref29174" refString="Voigt E. 1989. Beitrag zur Bryozoen-Fauna des sachsischen Cenomaniums. Revision von A. E. Reuss' " Die Bryozoen des unteren Quaders " in H. B. Geinitz' " Das Elbthalgebirge in Sachsen " (1872). I: Cheilostomata. Abh Staatl Mus Mineral Geol Dresden. 36: 8 - 87." type="journal article" year="1989">Voigt (1989</bibRefCitation>
|
||
, p. 59) introduced the genus
|
||
<taxonomicName id="4C424D1CFFDA4445A261725AFD76F98B" authorityName="Voigt" authorityYear="1989" box="[621,740,1565,1589]" class="Gymnolaemata" family="Onychocellidae" genus="Tyloporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="50" pageNumber="1706" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B936EA8DFFDA4445A261725AFD76F98B" box="[621,740,1565,1589]" italics="true" pageId="50" pageNumber="1706">Tyloporella</emphasis>
|
||
</taxonomicName>
|
||
for a new species –
|
||
<taxonomicName id="4C424D1CFFDA4445A3C4725AFB16F98B" box="[968,1156,1565,1589]" class="Gymnolaemata" family="Onychocellidae" genus="Tyloporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="50" pageNumber="1706" phylum="Bryozoa" rank="species" species="reussi">
|
||
<emphasis id="B936EA8DFFDA4445A3C4725AFB16F98B" box="[968,1156,1565,1589]" italics="true" pageId="50" pageNumber="1706">Tyloporella reussi</emphasis>
|
||
</taxonomicName>
|
||
– that had been misidentified by
|
||
<bibRefCitation id="EFD34B6EFFDA4445A2307278FD5DF9E9" author="Reuss AE" box="[572,719,1599,1623]" pageId="50" pageNumber="1706" refId="ref27889" refString="Reuss AE. 1872. Die Bryozoen und Foraminiferen des unteren Quaders. In: Das Elbthalgebirge in Sachsen. I. Theil: Der untere Quader (Geinitz, HB editor); p. 95 - 144. Palaeontographica 20. Cassel: Theodor Fischer." type="book" year="1872">Reuss (1872)</bibRefCitation>
|
||
as ‘
|
||
<taxonomicName id="4C424D1CFFDA4445A30B7278FF72F9C4" authority="' (Reuss, 1846)" authorityName="' (Reuss" authorityYear="1846" baseAuthorityName="Reuss" baseAuthorityYear="1846" class="Gymnolaemata" family="Membraniporidae" genus="Membranipora" kingdom="Animalia" order="Cheilostomatida" pageId="50" pageNumber="1706" phylum="Bryozoa" rank="species" species="tenuisulca">
|
||
<emphasis id="B936EA8DFFDA4445A30B7278FBB6F9E9" box="[775,1060,1599,1623]" italics="true" pageId="50" pageNumber="1706">Membranipora tenuisulca</emphasis>
|
||
’ (
|
||
<bibRefCitation id="EFD34B6EFFDA4445A4307278FF45F9C4" author="Reuss AE" pageId="50" pageNumber="1706" refId="ref27818" refString="Reuss AE. 1846. Die Versteinerungen der bohmischen Kreideformation. Stuttgart: Schweizerbart' sche Verlagsbuchhandlung." type="book" year="1846">Reuss, 1846</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the latter being a possible candidate for assignment to
|
||
<taxonomicName id="4C424D1CFFDA4445A38B7225FB9EF9C4" authority="Taylor & Martha & Gordon, 2018" authorityName="Taylor & Martha & Gordon" authorityYear="2018" box="[903,1036,1634,1658]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="50" pageNumber="1706" phylum="Bryozoa" rank="genus" status="gen. nov.">
|
||
<emphasis id="B936EA8DFFDA4445A38B7225FB9EF9C4" box="[903,1036,1634,1658]" italics="true" pageId="50" pageNumber="1706">Aechmellina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20557F6FFDA4445A41B7225FB16F9C4" box="[1047,1156,1634,1658]" pageId="50" pageNumber="1706" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
The name alludes to similarities with
|
||
<taxonomicName id="4C424D1CFFDA4445A24872C2FCB7F923" authority="Lang, 1917" authorityName="Lang" authorityYear="1917" box="[580,805,1669,1693]" class="Gymnolaemata" family="Calloporidae" genus="Tylopora" higherTaxonomySource="CoL" kingdom="Animalia" order="Cheilostomatida" pageId="50" pageNumber="1706" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B936EA8DFFDA4445A24872C2FD31F923" box="[580,675,1669,1693]" italics="true" pageId="50" pageNumber="1706">Tylopora</emphasis>
|
||
<bibRefCitation id="EFD34B6EFFDA4445A2A672C2FCB7F923" author="Lang WD" box="[682,805,1669,1693]" pageId="50" pageNumber="1706" pagination="256 - 258" refId="ref26997" refString="Lang WD. 1917. IV. - on some new Cenomanian and Turonian Cheilostome Polyzoa. Geol Mag. 4: 256 - 258." type="journal article" year="1917">Lang, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
, which, however, has pauciserial, ribbon-like encrusting branches. The most recent redescription of
|
||
<taxonomicName id="4C424D1CFFDA4445A3ED72E0FBAFF901" box="[993,1085,1703,1727]" class="Gymnolaemata" family="Onychocellidae" genus="Tyloporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="50" pageNumber="1706" phylum="Bryozoa" rank="species" species="reussi">
|
||
<emphasis id="B936EA8DFFDA4445A3ED72E0FBAFF901" box="[993,1085,1703,1727]" italics="true" pageId="50" pageNumber="1706">T. reussi</emphasis>
|
||
</taxonomicName>
|
||
is by
|
||
<bibRefCitation id="EFD34B6EFFDB4444A0AC70AAFE16FABB" author="Martha SO & Niebuhr B & Scholz J" box="[160,388,1261,1285]" pageId="51" pageNumber="1707" pagination="11 - 52" refId="ref27354" refString="Martha SO, Niebuhr B, Scholz J. 2017. Cheilostome Bryozoen. Geol Saxonica. 62: 11 - 52." type="journal article" year="2017">Martha et al. (2017)</bibRefCitation>
|
||
. Apart from the frequent, small avicularia interspersed among the autozooids, a proximal, smooth gymnocyst is characteristic for this genus according to
|
||
<bibRefCitation id="EFD34B6EFFDB4444A0AC7175FEBEFAF4" author="Voigt E" box="[160,300,1330,1354]" pageId="51" pageNumber="1707" pagination="8 - 87" refId="ref29174" refString="Voigt E. 1989. Beitrag zur Bryozoen-Fauna des sachsischen Cenomaniums. Revision von A. E. Reuss' " Die Bryozoen des unteren Quaders " in H. B. Geinitz' " Das Elbthalgebirge in Sachsen " (1872). I: Cheilostomata. Abh Staatl Mus Mineral Geol Dresden. 36: 8 - 87." type="journal article" year="1989">Voigt (1989)</bibRefCitation>
|
||
and was taken as justification to assign the genus to
|
||
<taxonomicName id="4C424D1CFFDB4444A3A47175FBA4FAF4" box="[936,1078,1330,1354]" class="Gymnolaemata" family="Calloporidae" kingdom="Animalia" order="Cheilostomatida" pageId="51" pageNumber="1707" phylum="Bryozoa" rank="family">Calloporidae</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="4C424D1CFFDB4444A0D77112FE1AFAD3" authorityName="Jullien" authorityYear="1882" box="[219,392,1365,1389]" class="Gymnolaemata" family="Onychocellidae" kingdom="Animalia" order="Cheilostomatida" pageId="51" pageNumber="1707" phylum="Bryozoa" rank="family">Onychocellidae</taxonomicName>
|
||
by some subsequent taxonomists (e.g.
|
||
<bibRefCitation id="EFD34B6EFFDB4444A3477112FBE4FAD3" author="Taylor PD & McKinney FK" box="[843,1142,1365,1389]" pageId="51" pageNumber="1707" pagination="1 - 346" refId="ref28471" refString="Taylor PD, McKinney FK. 2006. Cretaceous Bryozoa from the Campanian and Maastrichtian of the Atlantic and Gulf Coastal Plains, United States. Scripta Geol. 132: 1 - 346." type="journal article" year="2006">Taylor and McKinney 2006</bibRefCitation>
|
||
). A gymnocyst is visible only in some better preserved specimens from the type locality, but not in the
|
||
<typeStatus id="54F9883DFFDB4444A15C71DDFE25FA0C" box="[336,439,1434,1458]" pageId="51" pageNumber="1707" type="holotype">holotype</typeStatus>
|
||
, which is poorly preserved. Possible minute spine bases were observed in few autozooids of the
|
||
<typeStatus id="54F9883DFFDB4444A22771FAFD1DFA6B" box="[555,655,1469,1493]" pageId="51" pageNumber="1707" type="holotype">holotype</typeStatus>
|
||
colony (
|
||
<bibRefCitation id="EFD34B6EFFDB4444A2FC71FAFCFEFA6B" author="Voigt E" box="[752,876,1469,1493]" pageId="51" pageNumber="1707" pagination="8 - 87" refId="ref29174" refString="Voigt E. 1989. Beitrag zur Bryozoen-Fauna des sachsischen Cenomaniums. Revision von A. E. Reuss' " Die Bryozoen des unteren Quaders " in H. B. Geinitz' " Das Elbthalgebirge in Sachsen " (1872). I: Cheilostomata. Abh Staatl Mus Mineral Geol Dresden. 36: 8 - 87." type="journal article" year="1989">Voigt 1989</bibRefCitation>
|
||
, p. 8, fig. 4), while other, better preserved, colonies show no evidence for spine bases (see
|
||
<bibRefCitation id="EFD34B6EFFDB4444A39E7198FBF0FA49" author="Martha SO & Niebuhr B & Scholz J" box="[914,1122,1503,1527]" pageId="51" pageNumber="1707" pagination="11 - 52" refId="ref27354" refString="Martha SO, Niebuhr B, Scholz J. 2017. Cheilostome Bryozoen. Geol Saxonica. 62: 11 - 52." type="journal article" year="2017">Martha et al. 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF3D6617FFDB4444A0AC77F5FC01FB29" ID-DOI="http://doi.org/10.5281/zenodo.4747220" ID-Zenodo-Dep="4747220" httpUri="https://zenodo.org/record/4747220/files/figure.png" pageId="51" pageNumber="1707" startId="51.[160,225,946,969]" targetBox="[163,1153,142,915]" targetPageId="51">
|
||
<paragraph id="8BFD369FFFDB4444A0AC77F5FC01FB29" blockId="51.[160,1156,946,1175]" pageId="51" pageNumber="1707">
|
||
<emphasis id="B936EA8DFFDB4444A0AC77F5FE9AFC77" bold="true" box="[160,264,946,969]" pageId="51" pageNumber="1707">Figure 33.</emphasis>
|
||
<taxonomicName id="4C424D1CFFDB4444A11D77F5FE6BFC77" authority="Voigt." authorityName="Voigt." box="[273,505,946,969]" class="Gymnolaemata" family="Onychocellidae" genus="Tyloporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="51" pageNumber="1707" phylum="Bryozoa" rank="species" species="reussi">
|
||
<emphasis id="B936EA8DFFDB4444A11D77F5FE24FC77" box="[273,438,946,969]" italics="true" pageId="51" pageNumber="1707">Tyloporella reussi</emphasis>
|
||
Voigt.
|
||
</taxonomicName>
|
||
(a,b) Upper Cenomanian (
|
||
<taxonomicName id="4C424D1CFFDB4444A30E77F5FC51FC77" baseAuthorityName="de Blainville" baseAuthorityYear="1827" box="[770,963,946,969]" class="Cephalopoda" family="Belemnitellidae" genus="Actinocamax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Belemnitida" pageId="51" pageNumber="1707" phylum="Mollusca" rank="species" species="plenus">
|
||
<emphasis id="B936EA8DFFDB4444A30E77F5FC51FC77" box="[770,963,946,969]" italics="true" pageId="51" pageNumber="1707">Actinocamax plenus</emphasis>
|
||
</taxonomicName>
|
||
zone), Hoher Stein Quarry in Dresden-Plauen, Saxony, Germany; (a) holotype, MMG SaK 1617; part of colony with two broken ovicells indicated by arrows. (b) MMG SaK 15777a; autozooids and avicularia at growing edge. (c) NMP, Novak Collection (figured Novak 1877, pl. 1, fig. 19, 20 as
|
||
<taxonomicName id="4C424D1CFFDB4444A382704CFED7FB81" authority="(von Hagenow))" baseAuthorityName="von Hagenow" class="Gymnolaemata" family="Membraniporidae" genus="Membranipora" kingdom="Animalia" order="Cheilostomatida" pageId="51" pageNumber="1707" phylum="Bryozoa" rank="species" species="irregularis">
|
||
<emphasis id="B936EA8DFFDB4444A382704CFB11FB9C" box="[910,1155,1035,1058]" italics="true" pageId="51" pageNumber="1707">Membranipora irregularis</emphasis>
|
||
(von Hagenow))
|
||
</taxonomicName>
|
||
; Late Cretaceous,?Turonian, Kamajka, Bohemia, Czech Republic; autozooids and avicularia; an autozooid with the opesia covered by a closure plate is arrowed. (d) NHMUK BZ 8221; Cretaceous. Late Cenomanian or early Turonian, Kaňk, Central Bohemian Region, Czech Republic; autozooids (some with closure plates) and avicularia. Scale bars = 500 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD369FFFDB4443A0CC7245FCE4FF77" blockId="51.[160,1157,1261,1701]" lastBlockId="52.[160,1155,142,201]" lastPageId="52" lastPageNumber="1708" pageId="51" pageNumber="1707">
|
||
The North American Maastrichtian species
|
||
<taxonomicName id="4C424D1CFFDB4444A2B27245FF4DF983" authority="(Canu and Bassler, 1926)" baseAuthorityName="Canu and Bassler" baseAuthorityYear="1926" class="Gymnolaemata" family="Onychocellidae" genus="Tyloporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="51" pageNumber="1707" phylum="Bryozoa" rank="species" species="cretacea">
|
||
<emphasis id="B936EA8DFFDB4444A2B27245FC0FF9A4" box="[702,925,1538,1562]" italics="true" pageId="51" pageNumber="1707">Tyloporella cretacea</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD34B6EFFDB4444A3BC7245FF45F983" author="Canu F & Bassler RS" pageId="51" pageNumber="1707" pagination="32 - 39" refId="ref24451" refString="Canu F, Bassler RS. 1926. Phylum Molluscoidea, Class Bryozoa. U. S. Geological Survey Professional Paper 137: 32 - 39. Washington (DC)." type="journal article" year="1926">Canu and Bassler, 1926</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
has opesiae that occupy about half of the zooidal frontal surface and ovicells with a smooth gymnocystal ooecium (
|
||
<bibRefCitation id="EFD34B6EFFDB4444A27C7200FC22F9E1" author="Taylor PD & McKinney FK" box="[624,944,1607,1631]" pageId="51" pageNumber="1707" pagination="1 - 346" refId="ref28471" refString="Taylor PD, McKinney FK. 2006. Cretaceous Bryozoa from the Campanian and Maastrichtian of the Atlantic and Gulf Coastal Plains, United States. Scripta Geol. 132: 1 - 346." type="journal article" year="2006">Taylor and McKinney 2006</bibRefCitation>
|
||
). A third species,
|
||
<taxonomicName id="4C424D1CFFDB4444A0AC722DFD38F93C" authority="Di Martino and Taylor, 2013" authorityName="Di Martino and Taylor" authorityYear="2013" box="[160,682,1642,1666]" class="Gymnolaemata" family="Onychocellidae" genus="Tyloporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="51" pageNumber="1707" phylum="Bryozoa" rank="species" species="smithi">
|
||
<emphasis id="B936EA8DFFDB4444A0AC722DFEF0F93C" box="[160,354,1642,1666]" italics="true" pageId="51" pageNumber="1707">Tyloporella smithi</emphasis>
|
||
<bibRefCitation id="EFD34B6EFFDB4444A167722DFD38F93C" author="Di Martino E & Taylor PD" box="[363,682,1642,1666]" pageId="51" pageNumber="1707" pagination="80 - 96" refId="ref25461" refString="Di Martino E, Taylor PD. 2013. First bryozoan fauna from a tropical Cretaceous carbonate: Simsima Formation, United Arab Emirates - Oman border region. Cret Res. 43: 80 - 96." type="journal article" year="2013">Di Martino and Taylor, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the Campanian of the
|
||
<collectingCountry id="F355760FFFDB4444A3F6722DFE91F91B" name="United Arab Emirates" pageId="51" pageNumber="1707">United Arab Emirates</collectingCountry>
|
||
, is bifoliate and can only be placed in this genus with some reservation, as is true for the specimen of
|
||
<taxonomicName id="4C424D1CFFDC4443A1CC74C9FC2CFF18" authority="(von Hagenow, 1851)" baseAuthorityName="von Hagenow" baseAuthorityYear="1851" box="[448,958,142,166]" class="Gymnolaemata" family="Calloporidae" genus="Amphiblestrum" kingdom="Animalia" order="Cheilostomatida" pageId="52" pageNumber="1708" phylum="Bryozoa" rank="species" species="elegans">
|
||
<emphasis id="B936EA8DFFDC4443A1CC74C9FD53FF18" box="[448,705,142,166]" italics="true" pageId="52" pageNumber="1708">Amphiblestrum elegans</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD34B6EFFDC4443A2DD74C9FC24FF18" author="von Hagenow F" box="[721,950,142,166]" pageId="52" pageNumber="1708" refId="ref29443" refString="von Hagenow F. 1851. Die Bryozoen der Maastrichter Kreidebildung. Cassel: Theodor Fischer." type="book" year="1851">von Hagenow, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
figured from the Danian of
|
||
<collectingCountry id="F355760FFFDC4443A11B74F6FEEFFF77" box="[279,381,177,201]" name="Denmark" pageId="52" pageNumber="1708">Denmark</collectingCountry>
|
||
by Berthelsen (
|
||
<bibRefCitation id="EFD34B6EFFDC4443A23C74F6FD7AFF77" author="Berthelsen O" box="[560,744,177,201]" pageId="52" pageNumber="1708" refId="ref23703" refString="Berthelsen O. 1962. Cheilostome Bryozoa in the Danian deposits of East Denmark. KObenhavn: C. A. Reitzel." type="book" year="1962">Berthelsen 1962</bibRefCitation>
|
||
, p. 9, fig. 3).
|
||
</paragraph>
|
||
<paragraph id="8BFD369FFFDC4443A0AC74BFFF79FEAF" blockId="52.[160,892,248,307]" box="[160,235,248,273]" pageId="52" pageNumber="1708">
|
||
<emphasis id="B936EA8DFFDC4443A0AC74BFFF79FEAF" bold="true" box="[160,235,248,273]" italics="true" pageId="52" pageNumber="1708">Range</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFD369FFFDC4443A0AC755CFCEEFE8D" blockId="52.[160,892,248,307]" box="[160,892,283,307]" pageId="52" pageNumber="1708">Cretaceous (Cenomanian to Maastrichtian),?Paleocene (Danian).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |