659 lines
86 KiB
XML
659 lines
86 KiB
XML
<document id="A241B9323E595D88D0F21F07F00B44D4" ID-DOI="10.11646/zootaxa.3986.2.2" ID-GBIF-Dataset="0679a97a-49b3-47c8-b520-46ed1a1536cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="244349" ID-ZooBank="986B0CFE-77D7-453F-8708-5F60F79624BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461261661897" checkinUser="plazi" docAuthor="Whiting, Martin J., Branch, William R., Pepper, Mitzy & Keogh, J. Scott" docDate="2015" docId="03E4BE09FF8A8B47FF69FF0DFBD62C31" docLanguage="en" docName="zt03986p192.pdf" docOrigin="Zootaxa 3986 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Platysaurus attenboroughi Whiting, Branch, Pepper & Keogh, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="190" masterDocId="FFDDC671FF868B56FFFEFF9AFFFF2F6C" masterDocTitle="A new species of spectacularly coloured flat lizard Platysaurus (Squamata: Cordylidae: Platysaurinae) from southern Africa" masterLastPageNumber="192" masterPageNumber="173" pageNumber="185" updateTime="1698609596299" updateUser="plazi">
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<mods:mods id="AA71ED0A8C0FD577DCBEB9C26FDC4E41" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FB294606A52ADFBCB96E39E02B3D59F6">
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<mods:title id="54AD80BC5552EB1BEFA70D61339ADE41">A new species of spectacularly coloured flat lizard Platysaurus (Squamata: Cordylidae: Platysaurinae) from southern Africa</mods:title>
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</mods:titleInfo>
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<mods:name id="4EC6D72D3D6F1866870A674A833ECD38" type="personal">
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<mods:roleTerm id="204B984F86417214244324C1F3264E11">Author</mods:roleTerm>
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<mods:namePart id="58EBAB54D08DDCD581D26D0908E29322">Whiting, Martin J.</mods:namePart>
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<mods:roleTerm id="C35B94AE251388D2B47FEC1AD26C8E05">Author</mods:roleTerm>
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<mods:namePart id="FDEEB09B6F031BFD2DCC134DA0117D71">Branch, William R.</mods:namePart>
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</mods:name>
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<mods:name id="BC42E5A2C31C8342DC66626444A05751" type="personal">
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<mods:roleTerm id="A47F53CDD74CA44054EA1E1DBA865AF4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="607A48E566B8C20A315C69D9CA6BFC00">Pepper, Mitzy</mods:namePart>
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</mods:name>
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<mods:name id="525510A7A51C31021829C5E168332EBF" type="personal">
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<mods:roleTerm id="0B43F103BB17AB8090FB45EBDF696EDE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5503E158C107A39B3DF0BAEDED77DB9E">Keogh, J. Scott</mods:namePart>
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<mods:typeOfResource id="C9F9F5CEFB229DF1CE8CC617028AE2F2">text</mods:typeOfResource>
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<mods:title id="DB1CFC09D514126B78AD31A16B0F0622">Zootaxa</mods:title>
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<mods:date id="654F066D238A99B7A28BC441DF707963">2015</mods:date>
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<mods:detail id="9AA57CC96EDACF4BFF10C90A59F0E9DE" type="volume">
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<mods:number id="8DDD4920874F8042477523251E802284">3986</mods:number>
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</mods:detail>
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<mods:detail id="4801DB95933959C4165B7BD432F23D03" type="issue">
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<mods:number id="93883472D7E1768EC4AC9226D6BD635D">2</mods:number>
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<mods:end id="057A13CAA055CA6A683B927C0193EA5A">192</mods:end>
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<mods:identifier id="6C8898E3EF12B7400B24DE3544E5EC6C" type="DOI">10.11646/zootaxa.3986.2.2</mods:identifier>
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<mods:identifier id="9103B70ED001DF264538222463CA570C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E4BE09FF8A8B47FF69FF0DFBD62C31" ID-DOI="http://doi.org/10.5281/zenodo.5629056" ID-GBIF-Taxon="119637116" ID-Zenodo-Dep="5629056" LSID="urn:lsid:plazi:treatment:03E4BE09FF8A8B47FF69FF0DFBD62C31" httpUri="http://treatment.plazi.org/id/03E4BE09FF8A8B47FF69FF0DFBD62C31" lastPageId="17" lastPageNumber="190" pageId="12" pageNumber="185">
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<subSubSection id="C3575C94FF8A8B5AFF69FF0DFB7A2E37" pageId="12" pageNumber="185" type="nomenclature">
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<paragraph id="8BF20F1FFF8A8B5AFF69FF0DFDC12FDE" blockId="12.[151,598,151,247]" box="[151,574,151,178]" pageId="12" pageNumber="185">
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<heading id="D0BAB873FF8A8B5AFF69FF0DFDC12FDE" bold="true" box="[151,574,151,178]" fontSize="11" level="1" pageId="12" pageNumber="185" reason="1">
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<emphasis id="B939D30DFF8A8B5AFF69FF0DFDC12FDE" bold="true" box="[151,574,151,178]" pageId="12" pageNumber="185">
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<taxonomicName id="4C4D749CFF8A8B5AFF69FF0DFE262FDD" box="[151,473,151,177]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
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<emphasis id="B939D30DFF8A8B5AFF69FF0DFE262FDD" bold="true" box="[151,473,151,177]" italics="true" pageId="12" pageNumber="185">Platysaurus attenboroughi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20A6E76FF8A8B5AFE1EFF02FDC12FDE" box="[480,574,152,178]" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF20F1FFF8A8B5AFF69FF20FDD32FBF" blockId="12.[151,598,151,247]" box="[151,556,186,211]" pageId="12" pageNumber="185">English: Attenborough’s Flat Lizard</paragraph>
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<paragraph id="8BF20F1FFF8A8B5AFF69FF44FDA92F9B" blockId="12.[151,598,151,247]" box="[151,598,222,247]" pageId="12" pageNumber="185">Afrikaans: Attenborough se Platakkedis</paragraph>
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<paragraph id="8BF20F1FFF8A8B5AFF69FEBCFB7A2E37" blockId="12.[151,1436,294,347]" pageId="12" pageNumber="185">
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<emphasis id="B939D30DFF8A8B5AFF69FEBCFEF22E51" bold="true" box="[151,269,294,317]" pageId="12" pageNumber="185">Synonymy.</emphasis>
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<taxonomicName id="4C4D749CFF8A8B5AFEEAFEBCFE142E51" box="[276,491,294,317]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="capensis">
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<emphasis id="B939D30DFF8A8B5AFEEAFEBCFE142E51" box="[276,491,294,317]" italics="true" pageId="12" pageNumber="185">Platysaurus capensis</emphasis>
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</taxonomicName>
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(part). FitzSimons, 1935: 535; 1943: 473;
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<bibRefCitation id="EFDC72EEFF8A8B5AFC61FEBCFBB52E50" author="Loveridge" box="[927,1098,294,316]" pageId="12" pageNumber="185" refString="Loveridge, A. (1944) Revision of the African lizards of the family Cordylidae. Bulletin of the Museum of Comparative Zoology Harvard, 95, 1 - 118." type="journal article" year="1944">Loveridge, 1944</bibRefCitation>
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: 97;
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<bibRefCitation id="EFDC72EEFF8A8B5AFB85FEBDFB0E2E50" author="Rose" box="[1147,1265,294,316]" pageId="12" pageNumber="185" refString="Rose, W. (1950) The Reptiles and Amphibians of Southern Africa. Maskew Miller, Cape Town, xxv + 378 pp., 213 figs., 7 pls." type="book" year="1950">Rose, 1950</bibRefCitation>
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: 155; 1962: 156;
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<bibRefCitation id="EFDC72EEFF8A8B5AFF39FEDFFE9A2E37" author="Broadley" box="[199,357,325,347]" pageId="12" pageNumber="185" refString="Broadley, D. G. (1978) A revision of the genus Platysaurus A. Smith (Sauria: Cordylidae). Occasional Papers of the National Museum of Rhodesia Series B Natural Sciences, 6, 129 - 185." type="journal article" year="1978">Broadley, 1978</bibRefCitation>
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: 157;
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<bibRefCitation id="EFDC72EEFF8A8B5AFE5CFEDFFDD12E37" author="Branch" box="[418,558,325,347]" pageId="12" pageNumber="185" refString="Branch, B. (1998) A field guide to snakes and other reptiles of southern Africa. 3 rd Edition. Struik Publishers, Cape Town, 399 pp." type="book" year="1998">Branch, 1998</bibRefCitation>
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: 165;
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<bibRefCitation id="EFDC72EEFF8A8B5AFD95FEDCFC842E37" author="Van" box="[619,891,325,347]" pageId="12" pageNumber="185" refString="Van Wyk, J. H. & Mouton, P. L. F. N. (1996) The reproductive cycles of the oviparous lizards Platysaurus capensis and P. minor: Evidence supporting a viviparity-oviparity reversal in the Cordylidae. Amphibia Reptilia, 17, 115 - 129. http: // dx. doi. org / 10.1163 / 156853896 X 00153" type="journal article" year="1996">Van Wyk & Mouton, 1996</bibRefCitation>
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: 117; Whiting, 2014: 214.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3575C94FF8A8B5AFF69FE1DFC7A2D14" pageId="12" pageNumber="185" type="materials_examined">
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||
<paragraph id="8BF20F1FFF8A8B5AFF69FE1DFC7A2D14" blockId="12.[151,1436,391,632]" pageId="12" pageNumber="185">
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||
<emphasis id="B939D30DFF8A8B5AFF69FE1DFEB62ECC" bold="true" box="[151,329,391,416]" pageId="12" pageNumber="185">
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<typeStatus id="54F6B1BDFF8A8B5AFF69FE1DFF2D2ECC" box="[151,210,391,416]" pageId="12" pageNumber="185">Type</typeStatus>
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material.
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</emphasis>
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||
<emphasis id="B939D30DFF8A8B5AFEAAFE1DFE442ECC" box="[340,443,391,416]" italics="true" pageId="12" pageNumber="185">
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<typeStatus id="54F6B1BDFF8A8B5AFEAAFE1DFE442ECC" box="[340,443,391,416]" pageId="12" pageNumber="185" type="holotype">Holotype</typeStatus>
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</emphasis>
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. TM 85806 (MJW 936), adult male collected by Bryan Maritz, Nick Tye and Chris Barichievy,
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<date id="FFF329DFFF8A8B5AFEDDFE37FE642EA8" box="[291,411,428,453]" pageId="12" pageNumber="185" value="2008-05">May 2008</date>
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||
(
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||
<tableCitation id="C6CF3AA4FF8A8B5AFE51FE36FDF42EA8" box="[431,523,428,453]" captionStart="TABLE 2" captionStartId="14.[151,239,620,643]" captionTargetBox="[159,1314,712,1508]" captionTargetPageId="14" captionText="TABLE 2. Measurements (mm) for the type series of Platysaurus attenboroughi sp. nov." httpUri="http://table.plazi.org/id/DF325F97FF888B58FF69FDF6FBB12DEF" pageId="12" pageNumber="185" tableUuid="DF325F97FF888B58FF69FDF6FBB12DEF">Table 2</tableCitation>
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||
,
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<figureCitation id="1376139AFF8A8B5AFDE3FE36FD5F2EA9" box="[541,672,428,453]" captionStart-0="FIGURE 4. a" captionStart-1="FIGURE 5. a" captionStartId-0="12.[151,250,1517,1539]" captionStartId-1="16.[151,250,1911,1933]" captionTargetBox-0="[154,1430,690,1492]" captionTargetBox-1="[343,1231,203,1871]" captionTargetId-0="figure@12.[151,1436,685,1495]" captionTargetId-1="figure@16.[298,1288,151,1890]" captionTargetPageId-0="12" captionTargetPageId-1="16" captionText-0="FIGURE 4. a.) Platysaurus attenboroughi sp. nov., male (paratype, TM 85805), Fish River Canyon; b.) P. attenboroughi sp. nov., female (allotype, TM 85807), Fish River Canyon; c.) P. attenboroughi sp. nov., male (holotype, TM 85806), venter, Fish River Canyon; d – e.) P. attenboroughi sp. nov., male (holotype, TM 85806), Fish River Canyon; f.) P. attenboroughi sp. nov., male, Richtersveld; g.) Platysaurus capensis, male, Kamiesberg near Kamieskroon; h.) P. capensis, female, Kamiesberg near Kamieskroon; and i.) P. attenboroughi sp. nov., habitat, Fish River Canyon (Photograph by N. Tye). All other photos by M. J. Whiting." captionText-1="FIGURE 5. a.) Spectral reflectance curves objectively measuring colour by body region in the holotype (TM 85806) in life. b.) Spectral reflectance of throat region for two males: (MJW 935 / TM 85805, MJW 936 / TM 85806 = holotype) and two females (MJW 934 / TM 85804, MJW 937 / TM 85807). Males have more distinct UV in their throats than females although it is still less conspicuous than what has been documented for the closely related P. broadleyi (Whiting et al., 2006)." httpUri-0="https://zenodo.org/record/244350/files/figure.png" httpUri-1="https://zenodo.org/record/244351/files/figure.png" pageId="12" pageNumber="185">Figure 4–5</figureCitation>
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).
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<emphasis id="B939D30DFF8A8B5AFD47FE37FCB12EA9" box="[697,846,428,453]" italics="true" pageId="12" pageNumber="185">
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<typeStatus id="54F6B1BDFF8A8B5AFD47FE37FD122EA8" box="[697,749,429,452]" pageId="12" pageNumber="185">Type</typeStatus>
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locality
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</emphasis>
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: Fish River Canyon, Karasburg District, southern
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<collectingCountry id="F35A4F8FFF8A8B5AFF69FE55FF072E84" box="[151,248,463,488]" name="Namibia" pageId="12" pageNumber="185">Namibia</collectingCountry>
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(27˚52'
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<geoCoordinate id="EE7969D8FF8A8B5AFEB0FE55FE6C2E84" box="[334,403,463,488]" direction="south" orientation="latitude" pageId="12" pageNumber="185" precision="5555" value="-21.7">21.7S</geoCoordinate>
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, 17˚31'
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<geoCoordinate id="EE7969D8FF8A8B5AFE1CFE55FDD42E84" box="[482,555,463,488]" direction="east" orientation="longitude" pageId="12" pageNumber="185" precision="5555" value="15.7">15.7E</geoCoordinate>
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; quarter-degree unit, 2717Dc).
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<emphasis id="B939D30DFF8A8B5AFC71FE55FC132E84" box="[911,1004,463,488]" italics="true" pageId="12" pageNumber="185">Allotype</emphasis>
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. TM 85807 (MJW 937), adult female collected by Bryan Maritz, Nick Tye and Chris Barichievy,
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<date id="FFF329DFFF8A8B5AFCD1FE6FFC5B2D60" box="[815,932,500,525]" pageId="12" pageNumber="185" value="2008-05">May 2008</date>
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; same locality data as
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<typeStatus id="54F6B1BDFF8A8B5AFB63FE6EFAFC2D61" box="[1181,1283,500,525]" pageId="12" pageNumber="185" type="holotype">holotype</typeStatus>
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.
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<emphasis id="B939D30DFF8A8B5AFAF2FE6FFA812D60" box="[1292,1406,501,524]" italics="true" pageId="12" pageNumber="185">
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||
<typeStatus id="54F6B1BDFF8A8B5AFAF2FE6FFA812D60" box="[1292,1406,501,524]" pageId="12" pageNumber="185" type="paratype">Paratypes</typeStatus>
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</emphasis>
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(
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<specimenCount id="9D4BC496FF8A8B5AFA73FE6EFEED2D5C" pageId="12" pageNumber="185" type="generic">2 specimens</specimenCount>
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): TM 85805 (MJW 935), adult male, TM 85804 (MJW 934), adult female, both collected by Bryan Maritz, Nick Tye and Chris Barichievy,
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<date id="FFF329DFFF8A8B5AFD8CFDA7FD132D38" box="[626,748,572,597]" pageId="12" pageNumber="185" value="2008-05">May 2008</date>
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, Fish River Canyon, Karasburg District, southern
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<collectingCountry id="F35A4F8FFF8A8B5AFAC2FDA6FA632D39" box="[1340,1436,572,597]" name="Namibia" pageId="12" pageNumber="185">Namibia</collectingCountry>
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(27˚50'
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<geoCoordinate id="EE7969D8FF8A8B5AFF1BFDC5FED52D14" box="[229,298,607,632]" direction="south" orientation="latitude" pageId="12" pageNumber="185" precision="5555" value="-3.7">03.7S</geoCoordinate>
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, 17˚32'
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<geoCoordinate id="EE7969D8FF8A8B5AFE86FDC5FE3E2D14" box="[376,449,607,632]" direction="east" orientation="longitude" pageId="12" pageNumber="185" precision="5555" value="47.5">47.5E</geoCoordinate>
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; quarter-degree unit, 2717Dc) (
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<tableCitation id="C6CF3AA4FF8A8B5AFCDEFDC5FC882D14" box="[800,887,607,632]" captionStart="TABLE 2" captionStartId="14.[151,239,620,643]" captionTargetBox="[159,1314,712,1508]" captionTargetPageId="14" captionText="TABLE 2. Measurements (mm) for the type series of Platysaurus attenboroughi sp. nov." httpUri="http://table.plazi.org/id/DF325F97FF888B58FF69FDF6FBB12DEF" pageId="12" pageNumber="185" tableUuid="DF325F97FF888B58FF69FDF6FBB12DEF">Table 2</tableCitation>
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).
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</paragraph>
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</subSubSection>
|
||
<caption id="DF325F97FF8A8B5AFF69FA77FF0E29F0" httpUri="https://zenodo.org/record/244350/files/figure.png" pageId="12" pageNumber="185" targetBox="[154,1430,690,1492]" targetPageId="12">
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<paragraph id="8BF20F1FFF8A8B5AFF69FA77FF0E29F0" blockId="12.[151,1436,1516,1692]" pageId="12" pageNumber="185">
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<emphasis id="B939D30DFF8A8B5AFF69FA77FEEB296E" bold="true" box="[151,276,1517,1539]" pageId="12" pageNumber="185">FIGURE 4.</emphasis>
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a.)
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<taxonomicName id="4C4D749CFF8A8B5AFEC2FA76FDB4296E" box="[316,587,1516,1538]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
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||
<emphasis id="B939D30DFF8A8B5AFEC2FA76FDB4296E" box="[316,587,1516,1538]" italics="true" pageId="12" pageNumber="185">Platysaurus attenboroughi</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B939D30DFF8A8B5AFDADFA77FD5B296E" bold="true" box="[595,676,1517,1538]" pageId="12" pageNumber="185">
|
||
<taxonomicNameLabel id="A20A6E76FF8A8B5AFDADFA77FD5B296E" box="[595,676,1517,1538]" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
, male (paratype, TM 85805), Fish River Canyon; b.)
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<taxonomicName id="4C4D749CFF8A8B5AFB30FA74FA89296E" box="[1230,1398,1516,1539]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8A8B5AFB30FA74FA89296E" box="[1230,1398,1516,1539]" italics="true" pageId="12" pageNumber="185">P. attenboroughi</emphasis>
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</taxonomicName>
|
||
<emphasis id="B939D30DFF8A8B5AFA83FA77FF3C294D" bold="true" pageId="12" pageNumber="185">
|
||
<taxonomicNameLabel id="A20A6E76FF8A8B5AFA83FA77FF3C294D" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female (allotype, TM 85807), Fish River Canyon; c.)
|
||
<taxonomicName id="4C4D749CFF8A8B5AFD0CF996FC66294D" box="[754,921,1547,1569]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8A8B5AFD0CF996FC66294D" box="[754,921,1547,1569]" italics="true" pageId="12" pageNumber="185">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8A8B5AFC5EF996FC0E294D" bold="true" box="[928,1009,1548,1569]" pageId="12" pageNumber="185">
|
||
<taxonomicNameLabel id="A20A6E76FF8A8B5AFC5EF996FC0E294D" box="[928,1009,1548,1569]" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (holotype, TM 85806), venter, Fish River Canyon; d–e.)
|
||
<taxonomicName id="4C4D749CFF8A8B5AFE90F9B1FDE9292C" box="[366,534,1578,1600]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8A8B5AFE90F9B1FDE9292C" box="[366,534,1578,1600]" italics="true" pageId="12" pageNumber="185">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8A8B5AFDE3F9B0FD902953" bold="true" box="[541,623,1578,1599]" pageId="12" pageNumber="185">
|
||
<taxonomicNameLabel id="A20A6E76FF8A8B5AFDE3F9B0FD902953" box="[541,623,1578,1599]" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (holotype, TM 85806), Fish River Canyon; f.)
|
||
<taxonomicName id="4C4D749CFF8A8B5AFB6AF9B1FAC3292C" box="[1172,1340,1578,1600]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8A8B5AFB6AF9B1FAC3292C" box="[1172,1340,1578,1600]" italics="true" pageId="12" pageNumber="185">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8A8B5AFABAF9B0FA6A2953" bold="true" box="[1348,1429,1578,1599]" pageId="12" pageNumber="185">
|
||
<taxonomicNameLabel id="A20A6E76FF8A8B5AFABAF9B0FA6A2953" box="[1348,1429,1578,1599]" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male, Richtersveld; g.)
|
||
<taxonomicName id="4C4D749CFF8A8B5AFE76F9D3FD9F2933" box="[392,608,1609,1631]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF8A8B5AFE76F9D3FD9F2933" box="[392,608,1609,1631]" italics="true" pageId="12" pageNumber="185">Platysaurus capensis</emphasis>
|
||
</taxonomicName>
|
||
, male, Kamiesberg near Kamieskroon; h.)
|
||
<taxonomicName id="4C4D749CFF8A8B5AFBE2F9D0FB722933" box="[1052,1165,1610,1631]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF8A8B5AFBE2F9D0FB722933" box="[1052,1165,1610,1631]" italics="true" pageId="12" pageNumber="185">P. capensis</emphasis>
|
||
</taxonomicName>
|
||
, female, Kamiesberg near Kamieskroon; and i.)
|
||
<taxonomicName id="4C4D749CFF8A8B5AFE88F9F3FDE02911" box="[374,543,1639,1662]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8A8B5AFE88F9F3FDE02911" box="[374,543,1639,1662]" italics="true" pageId="12" pageNumber="185">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8A8B5AFDD8F9F2FD882911" bold="true" box="[550,631,1640,1661]" pageId="12" pageNumber="185">
|
||
<taxonomicNameLabel id="A20A6E76FF8A8B5AFDD8F9F2FD882911" box="[550,631,1640,1661]" pageId="12" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitat, Fish River Canyon (Photograph by N. Tye). All other photos by M.J. Whiting.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3575C94FF8A8B5BFF39F952FC162D77" lastPageId="13" lastPageNumber="186" pageId="12" pageNumber="185" type="diagnosis">
|
||
<paragraph id="8BF20F1FFF8A8B5AFF39F952FA6528B1" blockId="12.[151,1437,1736,2014]" pageId="12" pageNumber="185">
|
||
<emphasis id="B939D30DFF8A8B5AFF39F952FEBE298D" bold="true" box="[199,321,1736,1761]" pageId="12" pageNumber="185">Diagnosis.</emphasis>
|
||
A medium-sized
|
||
<taxonomicName id="4C4D749CFF8A8B5AFDE5F952FD5E298D" box="[539,673,1736,1761]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="genus">
|
||
<emphasis id="B939D30DFF8A8B5AFDE5F952FD5E298D" box="[539,673,1736,1761]" italics="true" pageId="12" pageNumber="185">Platysaurus</emphasis>
|
||
</taxonomicName>
|
||
(Table 3,
|
||
<figureCitation id="1376139AFF8A8B5AFCDDF952FC77298D" box="[803,904,1736,1761]" captionStart="FIGURE 4. a" captionStartId="12.[151,250,1517,1539]" captionTargetBox="[154,1430,690,1492]" captionTargetId="figure@12.[151,1436,685,1495]" captionTargetPageId="12" captionText="FIGURE 4. a.) Platysaurus attenboroughi sp. nov., male (paratype, TM 85805), Fish River Canyon; b.) P. attenboroughi sp. nov., female (allotype, TM 85807), Fish River Canyon; c.) P. attenboroughi sp. nov., male (holotype, TM 85806), venter, Fish River Canyon; d – e.) P. attenboroughi sp. nov., male (holotype, TM 85806), Fish River Canyon; f.) P. attenboroughi sp. nov., male, Richtersveld; g.) Platysaurus capensis, male, Kamiesberg near Kamieskroon; h.) P. capensis, female, Kamiesberg near Kamieskroon; and i.) P. attenboroughi sp. nov., habitat, Fish River Canyon (Photograph by N. Tye). All other photos by M. J. Whiting." httpUri="https://zenodo.org/record/244350/files/figure.png" pageId="12" pageNumber="185">Figure 4</figureCitation>
|
||
) distinguished from all congeners, except
|
||
<taxonomicName id="4C4D749CFF8A8B5AFA79F950FF082869" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF8A8B5AFA79F950FF082869" italics="true" pageId="12" pageNumber="185">P. capensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B939D30DFF8A8B5AFED3F974FE452869" box="[301,442,1772,1797]" italics="true" pageId="12" pageNumber="185">
|
||
<taxonomicName id="4C4D749CFF8A8B5AFED3F974FE492869" box="[301,438,1772,1797]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="broadleyi">P. broadleyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
in that the scales on the side of the neck are indistinguishable from those on the dorsum. It can be distinguished from
|
||
<taxonomicName id="4C4D749CFF8A8B5AFE25F888FD9E2845" box="[475,609,1808,1833]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF8A8B5AFE25F888FD9E2845" box="[475,609,1808,1833]" italics="true" pageId="12" pageNumber="185">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
as follows: the breeding male has blue forelimbs (anterior surface) while that of
|
||
<taxonomicName id="4C4D749CFF8A8B5AFF19F8ACFE922822" box="[231,365,1845,1870]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF8A8B5AFF19F8ACFE922822" box="[231,365,1845,1870]" italics="true" pageId="12" pageNumber="185">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
may be orange, yellow, or a combination; the male has a light blue throat (
|
||
<figureCitation id="1376139AFF8A8B5AFB43F8AFFAFD2821" box="[1213,1282,1845,1870]" captionStart="FIGURE 4. a" captionStartId="12.[151,250,1517,1539]" captionTargetBox="[154,1430,690,1492]" captionTargetId="figure@12.[151,1436,685,1495]" captionTargetPageId="12" captionText="FIGURE 4. a.) Platysaurus attenboroughi sp. nov., male (paratype, TM 85805), Fish River Canyon; b.) P. attenboroughi sp. nov., female (allotype, TM 85807), Fish River Canyon; c.) P. attenboroughi sp. nov., male (holotype, TM 85806), venter, Fish River Canyon; d – e.) P. attenboroughi sp. nov., male (holotype, TM 85806), Fish River Canyon; f.) P. attenboroughi sp. nov., male, Richtersveld; g.) Platysaurus capensis, male, Kamiesberg near Kamieskroon; h.) P. capensis, female, Kamiesberg near Kamieskroon; and i.) P. attenboroughi sp. nov., habitat, Fish River Canyon (Photograph by N. Tye). All other photos by M. J. Whiting." httpUri="https://zenodo.org/record/244350/files/figure.png" pageId="12" pageNumber="185">Fig. 4</figureCitation>
|
||
,
|
||
<figureCitation id="1376139AFF8A8B5AFAF1F8AFFAE22822" box="[1295,1309,1845,1870]" captionStart="FIGURE 5. a" captionStartId="16.[151,250,1911,1933]" captionTargetBox="[343,1231,203,1871]" captionTargetId="figure@16.[298,1288,151,1890]" captionTargetPageId="16" captionText="FIGURE 5. a.) Spectral reflectance curves objectively measuring colour by body region in the holotype (TM 85806) in life. b.) Spectral reflectance of throat region for two males: (MJW 935 / TM 85805, MJW 936 / TM 85806 = holotype) and two females (MJW 934 / TM 85804, MJW 937 / TM 85807). Males have more distinct UV in their throats than females although it is still less conspicuous than what has been documented for the closely related P. broadleyi (Whiting et al., 2006)." httpUri="https://zenodo.org/record/244351/files/figure.png" pageId="12" pageNumber="185">5</figureCitation>
|
||
) compared to the dark blue of
|
||
<taxonomicName id="4C4D749CFF8A8B5AFE91F8C0FE0A281D" box="[367,501,1880,1905]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF8A8B5AFE91F8C0FE0A281D" box="[367,501,1880,1905]" italics="true" pageId="12" pageNumber="185">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
(although this may be highly variable and could be related to male fighting ability (Whiting
|
||
<emphasis id="B939D30DFF8A8B5AFEFAF8E4FEC128FA" box="[260,318,1917,1942]" italics="true" pageId="12" pageNumber="185">et al.</emphasis>
|
||
, 2006); and the male also has an extensive blue belly with a small orange lower abdominal patch and sometimes with an irregular black abdominal patch (centre), while
|
||
<taxonomicName id="4C4D749CFF8A8B5AFC44F838FBC028D5" box="[954,1087,1952,1977]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="185" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF8A8B5AFC44F838FBC028D5" box="[954,1087,1952,1977]" italics="true" pageId="12" pageNumber="185">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
has a darker (deep blue-black) abdomen with the lower abdomen usually orange (but may also be yellow or a mix). It also differs from
|
||
<emphasis id="B939D30DFF8A8B5AFA79F85CFA6528B1" box="[1415,1434,1990,2013]" italics="true" pageId="12" pageNumber="185">P.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF8B8B5BFF69FF0DFC162D77" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="186">
|
||
<taxonomicName id="4C4D749CFF8B8B5BFF69FF0DFEFE2FDC" box="[151,257,151,176]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF8B8B5BFF69FF0DFEFE2FDC" box="[151,257,151,176]" italics="true" pageId="13" pageNumber="186">broadleyi</emphasis>
|
||
</taxonomicName>
|
||
in some features of scalation, particularly the number of collar scales (mean
|
||
<taxonomicName id="4C4D749CFF8B8B5BFB82FF03FAC72FDC" box="[1148,1336,151,176]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8B8B5BFB82FF03FAC72FDC" box="[1148,1336,151,176]" italics="true" pageId="13" pageNumber="186">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8B8B5BFABEFF0DFA632FDC" bold="true" box="[1344,1436,151,176]" pageId="13" pageNumber="186">
|
||
<taxonomicNameLabel id="A20A6E76FF8B8B5BFABEFF0DFA632FDC" box="[1344,1436,151,176]" pageId="13" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
7.93 ± 1.06,
|
||
<taxonomicName id="4C4D749CFF8B8B5BFEDDFF27FE582FB9" box="[291,423,188,213]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF8B8B5BFEDDFF27FE582FB9" box="[291,423,188,213]" italics="true" pageId="13" pageNumber="186">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
9.22 ± 1.04), transverse dorsal scale rows (mean
|
||
<taxonomicName id="4C4D749CFF8B8B5BFC31FF27FB772FB9" box="[975,1160,188,213]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8B8B5BFC31FF27FB772FB9" box="[975,1160,188,213]" italics="true" pageId="13" pageNumber="186">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8B8B5BFB70FF26FB182FB9" bold="true" box="[1166,1255,188,213]" pageId="13" pageNumber="186">
|
||
<taxonomicNameLabel id="A20A6E76FF8B8B5BFB70FF26FB182FB9" box="[1166,1255,188,213]" pageId="13" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
85.23 ± 4.32,
|
||
<taxonomicName id="4C4D749CFF8B8B5BFA79FF27FEFE2F94" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF8B8B5BFA79FF27FEFE2F94" italics="true" pageId="13" pageNumber="186">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
104.96 ± 5.38) and the smaller upper forelimb scales (mean
|
||
<taxonomicName id="4C4D749CFF8B8B5BFC41FF7BFB852F94" box="[959,1146,223,248]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8B8B5BFC41FF7BFB852F94" box="[959,1146,223,248]" italics="true" pageId="13" pageNumber="186">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8B8B5BFB7DFF45FB212F94" bold="true" box="[1155,1246,223,248]" pageId="13" pageNumber="186">
|
||
<taxonomicNameLabel id="A20A6E76FF8B8B5BFB7DFF45FB212F94" box="[1155,1246,223,248]" pageId="13" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
17.17 ± 2.09,
|
||
<taxonomicName id="4C4D749CFF8B8B5BFA79FF7BFEFE2E71" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF8B8B5BFA79FF7BFEFE2E71" italics="true" pageId="13" pageNumber="186">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
25.82 ± 2.31; Table 3). Adult male coloration in
|
||
<taxonomicName id="4C4D749CFF8B8B5BFCDDFE9FFC242E71" box="[803,987,260,285]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8B8B5BFCDDFE9FFC242E71" box="[803,987,260,285]" italics="true" pageId="13" pageNumber="186">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8B8B5BFC1DFE9EFBC32E71" bold="true" box="[995,1084,260,285]" pageId="13" pageNumber="186">
|
||
<taxonomicNameLabel id="A20A6E76FF8B8B5BFC1DFE9EFBC32E71" box="[995,1084,260,285]" pageId="13" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is further distinguished from
|
||
<taxonomicName id="4C4D749CFF8B8B5BFA79FE9FFF082E2C" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF8B8B5BFA79FE9FFF082E2C" italics="true" pageId="13" pageNumber="186">P. capensis</emphasis>
|
||
</taxonomicName>
|
||
by having the dorsum more extensively covered with white spots (
|
||
<figureCitation id="1376139AFF8B8B5BFBF4FEBDFB902E2C" box="[1034,1135,295,320]" captionStart="FIGURE 4. a" captionStartId="12.[151,250,1517,1539]" captionTargetBox="[154,1430,690,1492]" captionTargetId="figure@12.[151,1436,685,1495]" captionTargetPageId="12" captionText="FIGURE 4. a.) Platysaurus attenboroughi sp. nov., male (paratype, TM 85805), Fish River Canyon; b.) P. attenboroughi sp. nov., female (allotype, TM 85807), Fish River Canyon; c.) P. attenboroughi sp. nov., male (holotype, TM 85806), venter, Fish River Canyon; d – e.) P. attenboroughi sp. nov., male (holotype, TM 85806), Fish River Canyon; f.) P. attenboroughi sp. nov., male, Richtersveld; g.) Platysaurus capensis, male, Kamiesberg near Kamieskroon; h.) P. capensis, female, Kamiesberg near Kamieskroon; and i.) P. attenboroughi sp. nov., habitat, Fish River Canyon (Photograph by N. Tye). All other photos by M. J. Whiting." httpUri="https://zenodo.org/record/244350/files/figure.png" pageId="13" pageNumber="186">Figure 4</figureCitation>
|
||
), and with a reduced fine vertebral stripe that only partially extends on to the hindbody, and with reduced (or absent) broad, dark paravertebral stripes; adult female coloration is more vaguely patterned than in either
|
||
<taxonomicName id="4C4D749CFF8B8B5BFB96FEEBFB192EE4" box="[1128,1254,369,392]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF8B8B5BFB96FEEBFB192EE4" box="[1128,1254,369,392]" italics="true" pageId="13" pageNumber="186">P. capensis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C4D749CFF8B8B5BFAF0FEEBFA6A2EE4" box="[1294,1429,367,392]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF8B8B5BFAF0FEEBFA6A2EE4" box="[1294,1429,367,392]" italics="true" pageId="13" pageNumber="186">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
, lacking the bold dark and pale paravertebral stripes and with scattered pale spots. It also differs from
|
||
<taxonomicName id="4C4D749CFF8B8B5BFAFDFE0FFA802EC0" box="[1283,1407,405,428]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF8B8B5BFAFDFE0FFA802EC0" box="[1283,1407,405,428]" italics="true" pageId="13" pageNumber="186">P. capensis</emphasis>
|
||
</taxonomicName>
|
||
in features of scalation (summarized in Table 3), particularly the greater number of upper forelimb scale rows (mean
|
||
<taxonomicName id="4C4D749CFF8B8B5BFF69FE7AFEAF2E9B" box="[151,336,478,503]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8B8B5BFF69FE7AFEAF2E9B" box="[151,336,478,503]" italics="true" pageId="13" pageNumber="186">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8B8B5BFEABFE44FE512E9B" bold="true" box="[341,430,478,503]" pageId="13" pageNumber="186">
|
||
<taxonomicNameLabel id="A20A6E76FF8B8B5BFEABFE44FE512E9B" box="[341,430,478,503]" pageId="13" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
17.17 ± 2.09,
|
||
<taxonomicName id="4C4D749CFF8B8B5BFDB0FE7AFD352E9B" box="[590,714,480,503]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF8B8B5BFDB0FE7AFD352E9B" box="[590,714,480,503]" italics="true" pageId="13" pageNumber="186">P. capensis</emphasis>
|
||
</taxonomicName>
|
||
14.62 ± 1.04) and number of subdigital lammelae beneath the 4th toe (mean
|
||
<taxonomicName id="4C4D749CFF8B8B5BFEF3FD9EFE392D77" box="[269,454,514,539]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF8B8B5BFEF3FD9EFE392D77" box="[269,454,514,539]" italics="true" pageId="13" pageNumber="186">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF8B8B5BFE33FD98FDD82D77" bold="true" box="[461,551,514,539]" pageId="13" pageNumber="186">
|
||
<taxonomicNameLabel id="A20A6E76FF8B8B5BFE33FD98FDD82D77" box="[461,551,514,539]" pageId="13" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
19.10 ± 1.11,
|
||
<taxonomicName id="4C4D749CFF8B8B5BFD36FD9EFCBB2D77" box="[712,836,516,539]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="186" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF8B8B5BFD36FD9EFCBB2D77" box="[712,836,516,539]" italics="true" pageId="13" pageNumber="186">P. capensis</emphasis>
|
||
</taxonomicName>
|
||
17.92 ± 0.64).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3575C94FF8B8B58FF39FDBCFAB3280E" lastPageId="14" lastPageNumber="187" pageId="13" pageNumber="186" type="description">
|
||
<paragraph id="8BF20F1FFF8B8B5BFF39FDBCFD7929AE" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="186">
|
||
<emphasis id="B939D30DFF8B8B5BFF39FDBCFEA62D53" bold="true" box="[199,345,550,575]" pageId="13" pageNumber="186">Description.</emphasis>
|
||
<emphasis id="B939D30DFF8B8B5BFE9CFDBCFE372D53" box="[354,456,550,575]" italics="true" pageId="13" pageNumber="186">
|
||
<typeStatus id="54F6B1BDFF8B8B5BFE9CFDBCFE372D53" box="[354,456,550,575]" pageId="13" pageNumber="186" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
. TM 85806, an adult male with original tail (lacking
|
||
<quantity id="4CB5A2FAFF8B8B5BFBC6FDBDFB672D53" box="[1080,1176,551,575]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="186" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
from tip, removed for DNA analysis). Head strongly depressed, much longer than broad (head length: tip of snout to anterior border of ear-opening, HL):
|
||
<quantity id="4CB5A2FAFF8B8B5BFE8FFDF4FE232DEB" box="[369,476,622,647]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="13" pageNumber="186" unit="mm" value="16.6">16.6 mm</quantity>
|
||
; head width (HW):
|
||
<quantity id="4CB5A2FAFF8B8B5BFD30FDF5FCC62DEB" box="[718,825,623,647]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="13" pageNumber="186" unit="mm" value="12.9">12.9 mm</quantity>
|
||
; HL/HW: 1.27). Large supranasals in broad contact behind rostral; nostril directed slightly backwards and piercing a very small nasal that contacts the rostral, first supralabial, a small postnasal and the large supranasal. Frontonasal hexagonal, as broad as long and in good contact on sides with loreal. Prefrontals in median contact. Frontal longer (
|
||
<quantity id="4CB5A2FAFF8B8B5BFC62FD41FBFA2D9F" box="[924,1029,731,755]" metricMagnitude="-3" metricUnit="m" metricValue="3.32" pageId="13" pageNumber="186" unit="mm" value="3.32">3.32 mm</quantity>
|
||
) than broad (max. width:
|
||
<quantity id="4CB5A2FAFF8B8B5BFACAFD41FA6F2D9F" box="[1332,1424,731,755]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="186" unit="mm" value="2.2">2.2 mm</quantity>
|
||
), much wider in front than behind. A pair of frontoparietals, each in contact laterally with middle pair of supraoculars. Interparietal small, diamond-shaped, set in the middle of the two pairs of parietals, the anterior pair the smallest, and with a conspicuous pineal pore. Occipital absent, two slightly enlarged triangular granules in its position. Four supraoculars, the first large and triangular and contacting the pentagonal preocular, the last the smallest. Four supraciliaries, anterior elongate, middle pair largest. Lower eyelid with a semi-transparent disc faintly divided into a number of vertical septa. Three elongate upper temporals bordering parietals on each side, middle one largest and longitudinally elongate, twice the length of posterior one. Two additional rows of enlarged scales present in dorsal temporal region, upper row (five right, four left) vertically elongate and more than twice the size of lower row. Ventral temporal region covered with five rows of irregular granules, slightly larger than those along backbone. A small postnasal; an elongate loreal and a preocular, the former much smaller than the latter. Four suboculars, the second elongate and extensively bordering the lip below. Rostral pentagonal, broader than deep. Six supralabials, five anterior to subocular. Mental subpentagonal. Seven infralabials, 5–7 very small and elongate and sandwiched between lip and very large fourth sublabial; five, all slightly larger than infralabials, the fourth by far the largest, the fifth the smallest. A longitudinal median series of four enlarged quadrangular or polygonal gular scales, bordered by scales that become smaller (subgranular and rounded) particularly laterally on the neck, and increase in size towards the collar, which consists of eight enlarged scales; in 25 rows between the angle of the jaws. Dorsal scales smooth, small, flat and rounded, without minute granules at junctions, and largest along backbone and on flanks, smallest dorso-laterally and minute on sides, in 89–91 transverse rows across midbody. Ventrals square or a little broader than long medially, arranged in 39 mainly regular transverse and 20 longitudinal series. Eight preanal plates, median pair largest and much larger than ventrals, reducing laterally towards the groin. Limbs long and slender, length of tibia subequal to head length. Upper forelimb and thighs with subequal granules above. Forearm and tibia with enlarged keeled scales; a row of nine large transverse plates on underside of tibia, largest at midcalf. Toes long, clawed and with smooth subdigital lamellae (17 on 4th toe on both sides). A series of 16 femoral pores on lower surface of right thigh, 13 on left, with 2–3 rows of modified generation gland scales (32 right thigh, 30 left) anterior to these pores. Tail depressed, tapering, with regular whorls of elongate, quadrangular scales, strongly keeled above laterally, smooth below. Measurements for the
|
||
<typeStatus id="54F6B1BDFF8B8B5BFAC4F91FFA6329F2" box="[1338,1436,1669,1694]" pageId="13" pageNumber="186" type="holotype">holotype</typeStatus>
|
||
and other
|
||
<typeStatus id="54F6B1BDFF8B8B5BFEF9F930FEC829AE" box="[263,311,1706,1730]" pageId="13" pageNumber="186">type</typeStatus>
|
||
material are given in
|
||
<tableCitation id="C6CF3AA4FF8B8B5BFDD2F933FD7D29AE" box="[556,642,1705,1730]" captionStart="TABLE 2" captionStartId="14.[151,239,620,643]" captionTargetBox="[159,1314,712,1508]" captionTargetPageId="14" captionText="TABLE 2. Measurements (mm) for the type series of Platysaurus attenboroughi sp. nov." httpUri="http://table.plazi.org/id/DF325F97FF888B58FF69FDF6FBB12DEF" pageId="13" pageNumber="186" tableUuid="DF325F97FF888B58FF69FDF6FBB12DEF">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF8B8B5BFF39F957FAC628AD" blockId="13.[151,1437,151,2021]" pageId="13" pageNumber="186">
|
||
<emphasis id="B939D30DFF8B8B5BFF39F957FEA6298A" bold="true" box="[199,345,1741,1766]" pageId="13" pageNumber="186">Description.</emphasis>
|
||
<emphasis id="B939D30DFF8B8B5BFE9FF957FE41298A" box="[353,446,1741,1766]" italics="true" pageId="13" pageNumber="186">Allotype</emphasis>
|
||
. TM 85807, an adult female with original tail (lacking
|
||
<quantity id="4CB5A2FAFF8B8B5BFBC2F954FB642989" box="[1084,1179,1742,1766]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="186" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
from tip, removed for DNA analysis). Scalation as for
|
||
<typeStatus id="54F6B1BDFF8B8B5BFDF6F96BFD952866" box="[520,618,1777,1802]" pageId="13" pageNumber="186" type="holotype">holotype</typeStatus>
|
||
except for: head length
|
||
<quantity id="4CB5A2FAFF8B8B5BFC7FF96BFC172865" box="[897,1000,1777,1802]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="13" pageNumber="186" unit="mm" value="16.4">16.4 mm</quantity>
|
||
, head width
|
||
<quantity id="4CB5A2FAFF8B8B5BFB85F968FB1D2865" box="[1147,1250,1778,1802]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="13" pageNumber="186" unit="mm" value="11.4">11.4 mm</quantity>
|
||
; HL/HW: 1.44); large supranasals in only narrow contact behind rostral; a small triangular occipital present at middle border of posterior parietals. Middle row of temporals comprised of six vertically elongate scales on each side of head; gulars in 26 rows between the angle of the jaws, and seven scales in collar; dorsal scales in 90 transverse rows across midbody; ventrals arranged in 38 mainly regular transverse and 18 longitudinal series; femoral pores minute, 17 on each thigh; thighs without modified scales with generation glands; subdigital lamellae under 4th toe 18/19.
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF8B8B58FF39F856FB442D51" blockId="13.[151,1437,151,2021]" lastBlockId="14.[151,1437,151,573]" lastPageId="14" lastPageNumber="187" pageId="13" pageNumber="186">
|
||
<emphasis id="B939D30DFF8B8B5BFF39F856FEB02889" bold="true" box="[199,335,1996,2021]" pageId="13" pageNumber="186">Coloration.</emphasis>
|
||
<typeStatus id="54F6B1BDFF8B8B5BFEA4F856FE382889" box="[346,455,1996,2021]" pageId="13" pageNumber="186" type="holotype">Holotype</typeStatus>
|
||
: In life: Above dull grey-brown dorsally, with a single narrow pale stripe along the backbone, running from the base of tail to crown of head, where it shallowly divides; scattered pale spots, 3–5 granules across, speckled body and flanks, with largest aligned in a longitudinal row on the upper flank which is confluent with a pair of pale lateral stripes on head that arise above each eye and travel backwards across upper temporal region; forebody flanks and front of forelimbsh bright blue, which extends through neck and onto upper lip; upper surface of tail and flanks anterior to groin orange-brown; proximal tail and upper surface of hindlimbs light brown at tail tip; upper hindbody and upper surface of tail grey-tan, darker along midline, with vague, paler spots laterally. Ventral surface of neck and first three-quarters of belly brilliant blue, paler under head, and with black ‘badge’ with irregular outline in centre of belly; lower surface of forelimbs ivory-pink; rear of belly, underside of tail and lower surface of hindlimbs light orange. In preservative (70% ethanol): faded, most orange and blue coloration lost; conspicuous central black belly ‘badge’ inconspicuous; thin dorsal stripe and pale spots only vaguely visible; upper and lower surface of head back to forelimb insertion dark grey, lighter towards hindlimbs; tail and lower surface of hindlimbs pinkish-ivory, darker on upper surface of limbs.
|
||
</paragraph>
|
||
<caption id="DF325F97FF888B58FF69FDF6FBB12DEF" ID-Table-UUID="DF325F97FF888B58FF69FDF6FBB12DEF" box="[151,1102,619,643]" httpUri="http://table.plazi.org/id/DF325F97FF888B58FF69FDF6FBB12DEF" pageId="14" pageNumber="187" targetBox="[159,1314,712,1508]" targetIsTable="true" targetPageId="14">
|
||
<paragraph id="8BF20F1FFF888B58FF69FDF6FBB12DEF" blockId="14.[151,1102,619,643]" box="[151,1102,619,643]" pageId="14" pageNumber="187">
|
||
<emphasis id="B939D30DFF888B58FF69FDF6FEF62DEF" bold="true" box="[151,265,620,643]" pageId="14" pageNumber="187">TABLE 2.</emphasis>
|
||
Measurements (mm) for the type series of
|
||
<taxonomicName id="4C4D749CFF888B58FD28FDF1FC0D2DEF" box="[726,1010,619,643]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="187" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF888B58FD28FDF1FC0D2DEF" box="[726,1010,619,643]" italics="true" pageId="14" pageNumber="187">Platysaurus attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A6E76FF888B58FC07FDF6FBB12DEF" box="[1017,1102,620,643]" pageId="14" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF20F1FFF888B58FE15FD05FB1A2962" blockId="14.[491,681,671,692]" lastBlockId="14.[159,1253,1527,1552]" pageId="14" pageNumber="187">
|
||
<emphasis id="B939D30DFF888B58FE15FD05FD562DD8" bold="true" box="[491,681,671,692]" pageId="14" pageNumber="187">Museum Number</emphasis>
|
||
Subdigital lamellae (4th toe) 17/17 18/19 18/16 19/18
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF888B58FE15FD52FB3B2A88" pageId="14" pageNumber="187">
|
||
<table id="F94DFDBFFF8874A9FF61FD52FADD2A88" box="[159,1314,712,1508]" gridcols="5" gridrows="7" pageId="14" pageNumber="187">
|
||
<tr id="357D0D5DFF8874A9FF61FD52FADD2D90" box="[159,1314,712,764]" gridrow="0" pageId="14" pageNumber="187" rowspan-0="1">
|
||
<th id="76AC6421FF8874A9FE15FD52FDA92D90" box="[491,598,712,764]" gridcol="1" gridrow="0" pageId="14" pageNumber="187">Holotype TM 85806</th>
|
||
<th id="76AC6421FF8874A9FD1CFD52FCB22D90" box="[738,845,712,764]" gridcol="2" gridrow="0" pageId="14" pageNumber="187">Allotype TM 85807</th>
|
||
<th id="76AC6421FF8874A9FC4FFD52FBD82D90" box="[945,1063,712,764]" gridcol="3" gridrow="0" pageId="14" pageNumber="187">Paratype TM 85805</th>
|
||
<th id="76AC6421FF8874A9FB55FD52FADD2D90" box="[1195,1314,712,764]" gridcol="4" gridrow="0" pageId="14" pageNumber="187">Paratype TM 85804</th>
|
||
</tr>
|
||
<tr id="357D0D5DFF8874A9FF61FC8AFADD2C14" box="[159,1314,784,888]" gridrow="1" pageId="14" pageNumber="187">
|
||
<th id="76AC6421FF8874A9FF61FC8AFEAC2C14" box="[159,339,784,888]" gridcol="0" gridrow="1" pageId="14" pageNumber="187">Sex Snout-vent length Tail length</th>
|
||
<td id="76AC6421FF8874A9FE15FC8AFDA92C14" box="[491,598,784,888]" gridcol="1" gridrow="1" pageId="14" pageNumber="187">M 86.6 121 (t) *</td>
|
||
<td id="76AC6421FF8874A9FD1CFC8AFCB22C14" box="[738,845,784,888]" gridcol="2" gridrow="1" pageId="14" pageNumber="187">F 78.5 124 (t) *</td>
|
||
<td id="76AC6421FF8874A9FC4FFC8AFBD82C14" box="[945,1063,784,888]" gridcol="3" gridrow="1" pageId="14" pageNumber="187">M 77.5 114 (r) **</td>
|
||
<td id="76AC6421FF8874A9FB55FC8AFADD2C14" box="[1195,1314,784,888]" gridcol="4" gridrow="1" pageId="14" pageNumber="187">F 74.0 107 (r) **</td>
|
||
</tr>
|
||
<tr id="357D0D5DFF8874A9FF61FC16FADD2C98" box="[159,1314,908,1012]" gridrow="2" pageId="14" pageNumber="187">
|
||
<th id="76AC6421FF8874A9FF61FC16FEAC2C98" box="[159,339,908,1012]" gridcol="0" gridrow="2" pageId="14" pageNumber="187">Total length Head length Head width</th>
|
||
<td id="76AC6421FF8874A9FE15FC16FDA92C98" box="[491,598,908,1012]" gridcol="1" gridrow="2" pageId="14" pageNumber="187">207.6 (t) * 16.6 12.2</td>
|
||
<td id="76AC6421FF8874A9FD1CFC16FCB22C98" box="[738,845,908,1012]" gridcol="2" gridrow="2" pageId="14" pageNumber="187">202.5 (t) * 16.4 11.4</td>
|
||
<td id="76AC6421FF8874A9FC4FFC16FBD82C98" box="[945,1063,908,1012]" gridcol="3" gridrow="2" pageId="14" pageNumber="187">191.5 (t) ** 16.8 11.9</td>
|
||
<td id="76AC6421FF8874A9FB55FC16FADD2C98" box="[1195,1314,908,1012]" gridcol="4" gridrow="2" pageId="14" pageNumber="187">181.0 (t) ** 16.5 11.2</td>
|
||
</tr>
|
||
<tr id="357D0D5DFF8874A9FF61FB92FADD2B1C" box="[159,1314,1032,1136]" gridrow="3" pageId="14" pageNumber="187">
|
||
<th id="76AC6421FF8874A9FF61FB92FEAC2B1C" box="[159,339,1032,1136]" gridcol="0" gridrow="3" pageId="14" pageNumber="187">Head height Snout length Eye diameter</th>
|
||
<td id="76AC6421FF8874A9FE15FB92FDA92B1C" box="[491,598,1032,1136]" gridcol="1" gridrow="3" pageId="14" pageNumber="187">6.0 6.7 4.8</td>
|
||
<td id="76AC6421FF8874A9FD1CFB92FCB22B1C" box="[738,845,1032,1136]" gridcol="2" gridrow="3" pageId="14" pageNumber="187">5.4 6.3 4.4</td>
|
||
<td id="76AC6421FF8874A9FC4FFB92FBD82B1C" box="[945,1063,1032,1136]" gridcol="3" gridrow="3" pageId="14" pageNumber="187">6.1 6.4 4.5</td>
|
||
<td id="76AC6421FF8874A9FB55FB92FADD2B1C" box="[1195,1314,1032,1136]" gridcol="4" gridrow="3" pageId="14" pageNumber="187">5.7 6.6 4.8</td>
|
||
</tr>
|
||
<tr id="357D0D5DFF8874A9FF61FB1EFADD2B80" box="[159,1314,1156,1260]" gridrow="4" pageId="14" pageNumber="187">
|
||
<th id="76AC6421FF8874A9FF61FB1EFEAC2B80" box="[159,339,1156,1260]" gridcol="0" gridrow="4" pageId="14" pageNumber="187">Inter-limb length Elbow to wrist Knee to heel</th>
|
||
<td id="76AC6421FF8874A9FE15FB1EFDA92B80" box="[491,598,1156,1260]" gridcol="1" gridrow="4" pageId="14" pageNumber="187">36.4 12.9 16.0</td>
|
||
<td id="76AC6421FF8874A9FD1CFB1EFCB22B80" box="[738,845,1156,1260]" gridcol="2" gridrow="4" pageId="14" pageNumber="187">40.1 11.3 15.0</td>
|
||
<td id="76AC6421FF8874A9FC4FFB1EFBD82B80" box="[945,1063,1156,1260]" gridcol="3" gridrow="4" pageId="14" pageNumber="187">35.0 12.5 16.2</td>
|
||
<td id="76AC6421FF8874A9FB55FB1EFADD2B80" box="[1195,1314,1156,1260]" gridcol="4" gridrow="4" pageId="14" pageNumber="187">35.8 10.2 16.3</td>
|
||
</tr>
|
||
<tr id="357D0D5DFF8874A9FF61FA9AFADD2A04" box="[159,1314,1280,1384]" gridrow="5" pageId="14" pageNumber="187">
|
||
<th id="76AC6421FF8874A9FF61FA9AFEAC2A04" box="[159,339,1280,1384]" gridcol="0" gridrow="5" pageId="14" pageNumber="187">Gulars Ventrals (long.) Ventrals (trans.)</th>
|
||
<td id="76AC6421FF8874A9FE15FA9AFDA92A04" box="[491,598,1280,1384]" gridcol="1" gridrow="5" pageId="14" pageNumber="187">25 39 20</td>
|
||
<td id="76AC6421FF8874A9FD1CFA9AFCB22A04" box="[738,845,1280,1384]" gridcol="2" gridrow="5" pageId="14" pageNumber="187">26 38 18</td>
|
||
<td id="76AC6421FF8874A9FC4FFA9AFBD82A04" box="[945,1063,1280,1384]" gridcol="3" gridrow="5" pageId="14" pageNumber="187">27 39 20</td>
|
||
<td id="76AC6421FF8874A9FB55FA9AFADD2A04" box="[1195,1314,1280,1384]" gridcol="4" gridrow="5" pageId="14" pageNumber="187">26 37 18</td>
|
||
</tr>
|
||
<tr id="357D0D5DFF8874A9FF61FAE6FADD2A88" box="[159,1314,1404,1508]" gridrow="6" pageId="14" pageNumber="187">
|
||
<th id="76AC6421FF8874A9FF61FAE6FEAC2A88" box="[159,339,1404,1508]" gridcol="0" gridrow="6" pageId="14" pageNumber="187">Femoral pores Dorsals (trans.) Collar scales</th>
|
||
<td id="76AC6421FF8874A9FE15FAE6FDA92A88" box="[491,598,1404,1508]" gridcol="1" gridrow="6" pageId="14" pageNumber="187">16/13 89-91 8</td>
|
||
<td id="76AC6421FF8874A9FD1CFAE6FCB22A88" box="[738,845,1404,1508]" gridcol="2" gridrow="6" pageId="14" pageNumber="187">17/17 90 7</td>
|
||
<td id="76AC6421FF8874A9FC4FFAE6FBD82A88" box="[945,1063,1404,1508]" gridcol="3" gridrow="6" pageId="14" pageNumber="187">17/18 88-91 8</td>
|
||
<td id="76AC6421FF8874A9FB55FAE6FADD2A88" box="[1195,1314,1404,1508]" gridcol="4" gridrow="6" pageId="14" pageNumber="187">16/16 89=90 10</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF888B58FF69F9ABFD2B2924" blockId="14.[151,807,1585,1640]" box="[151,724,1585,1608]" pageId="14" pageNumber="187">
|
||
* tail original, but
|
||
<quantity id="4CB5A2FAFF888B58FEA2F9ABFE512924" box="[348,430,1585,1608]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="14" pageNumber="187" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3mm</quantity>
|
||
removed for DNA analysis
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF888B58FF69F9CBFCD82904" blockId="14.[151,807,1585,1640]" box="[151,807,1617,1640]" pageId="14" pageNumber="187">
|
||
** tail tip regenerated and
|
||
<quantity id="4CB5A2FAFF888B58FE4AF9CBFDFF2904" box="[436,512,1617,1640]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="14" pageNumber="187" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3mm</quantity>
|
||
removed for DNA analysis
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF888B58FF39F90FFAB3280E" blockId="14.[151,1437,1685,2034]" pageId="14" pageNumber="187">Allotype: In life: above grey brown, with a thin vertebral stripe from top of head to tail base, bordered on each side from eye to groin by a pale dorso-lateral ‘stripe’ comprising a paler background underlying a longitudinal series of pale spots; back, flanks and upper surface of hindlimbs speckled with pale spots, 2–4 granules wide; top of head dark brown lighter with three pale stripes; upper lip and lower temporal region tan-cream; tail dirty gold, darker along midline, paler below; belly pale brown, lacking a central black ‘badge’. In preservative (70% ethanol): as above but much paler above and below; vertebral stripe almost inconspicuous and pale spots less visible.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3575C94FF888B59FF39F8F7FB1028B6" lastPageId="15" lastPageNumber="188" pageId="14" pageNumber="187" type="distribution">
|
||
<paragraph id="8BF20F1FFF888B58FF39F8F7FF4A28A2" blockId="14.[151,1437,1685,2034]" pageId="14" pageNumber="187">
|
||
<emphasis id="B939D30DFF888B58FF39F8F7FE9F28EA" bold="true" box="[199,352,1901,1926]" pageId="14" pageNumber="187">Distribution.</emphasis>
|
||
Along the lower Orange River from Goodhouse to the Richtersveld, extending north into
|
||
<collectingCountry id="F35A4F8FFF888B58FF69F80BFF0728C6" box="[151,248,1937,1962]" name="Namibia" pageId="14" pageNumber="187">Namibia</collectingCountry>
|
||
and recorded from the Hunsberg, Huamsib and Ploegberg mountains and the Fish River Canyon (Figure 1).
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF888B59FF39F843FCBE2E09" blockId="14.[151,1437,1685,2034]" lastBlockId="15.[151,1436,151,357]" lastPageId="15" lastPageNumber="188" pageId="14" pageNumber="187">
|
||
<emphasis id="B939D30DFF888B58FF39F843FE43289E" bold="true" box="[199,444,2009,2034]" pageId="14" pageNumber="187">Habitat and climate.</emphasis>
|
||
<taxonomicName id="4C4D749CFF888B58FE3CF843FD13289E" box="[450,748,2009,2034]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="187" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF888B58FE3CF843FD13289E" box="[450,748,2009,2034]" italics="true" pageId="14" pageNumber="187">Platysaurus attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF888B58FD0DF843FCB2289E" bold="true" box="[755,845,2009,2034]" pageId="14" pageNumber="187">
|
||
<taxonomicNameLabel id="A20A6E76FF888B58FD0DF843FCB2289E" box="[755,845,2009,2034]" pageId="14" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
occurs in the arid-subtropical region of the Northern Cape Province of
|
||
<collectingCountry id="F35A4F8FFF898B59FE9EFF0DFE0F2FDC" box="[352,496,151,176]" name="South Africa" pageId="15" pageNumber="188">South Africa</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="F35A4F8FFF898B59FD73FF0DFD122FDC" box="[653,749,151,176]" name="Namibia" pageId="15" pageNumber="188">Namibia</collectingCountry>
|
||
and specifically within the Gariep Desert Bioregion (
|
||
<bibRefCitation id="EFDC72EEFF898B59FABBFF0DFE852FB9" author="Mucina" pageId="15" pageNumber="188" refString="Mucina, L. & Rutherford, M. C. (2006) The Vegetation of South Africa, Lesotho and Swaziland. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford, 2006</bibRefCitation>
|
||
). This is an arid area characterized by low and erratic summer rainfall. Summers are typically hot and dry. Like all flat lizards, they are dependent on rock (mostly granite in this area) and take refuge in narrow rock fissures where they can escape suboptimal temperatures and predators. These areas are largely devoid of significant vegetation bar the occasional fig tree (
|
||
<taxonomicName id="4C4D749CFF898B59FD2FFEB3FCF02E2C" box="[721,783,297,320]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="188" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B939D30DFF898B59FD2FFEB3FCF02E2C" box="[721,783,297,320]" italics="true" pageId="15" pageNumber="188">Ficus</emphasis>
|
||
</taxonomicName>
|
||
) or succulent. For more detailed descriptions of climate, vegetation and topography see
|
||
<bibRefCitation id="EFDC72EEFF898B59FE0DFED6FCC32E09" author="Mucina" box="[499,828,332,357]" pageId="15" pageNumber="188" refString="Mucina, L. & Rutherford, M. C. (2006) The Vegetation of South Africa, Lesotho and Swaziland. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford (2006)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF69FE0EFB8A2D7C" blockId="15.[151,1436,404,528]" pageId="15" pageNumber="188">
|
||
<emphasis id="B939D30DFF898B59FF69FE0EFEF82EC7" bold="true" box="[151,263,404,427]" pageId="15" pageNumber="188">TABLE 3.</emphasis>
|
||
Summary statistics (n = sample size, mean ± 1 SD, range) for SVL (snout-vent-length) and meristic characters: gulars: scales transversely between posterior sublabials, UL: upper labials, Col: scales in collar, LRV: longitudinal rows of ventrals, TRV: transverse rows of ventrals, TD: transverse rows of dorsals, Forearm: upper forelimb scale rows, 4th toe: number of subdigital lamellae beneath 4th toe, and FP: number of femoral pores on left thigh.
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FDB6FB2C2D2E" blockId="15.[159,1262,556,1448]" box="[159,1235,556,578]" pageId="15" pageNumber="188">
|
||
Character
|
||
<taxonomicName id="4C4D749CFF898B59FE1EFDB7FD782D2E" box="[480,647,556,578]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF898B59FE1EFDB7FD782D2E" box="[480,647,556,578]" italics="true" pageId="15" pageNumber="188">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF898B59FD70FDB6FD212D2D" bold="true" box="[654,734,556,577]" pageId="15" pageNumber="188">
|
||
<taxonomicNameLabel id="A20A6E76FF898B59FD70FDB6FD212D2D" box="[654,734,556,577]" pageId="15" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<emphasis id="B939D30DFF898B59FCDCFDB7FB2C2D2E" box="[802,1235,556,578]" italics="true" pageId="15" pageNumber="188">
|
||
<taxonomicName id="4C4D749CFF898B59FCDCFDB7FC642D2E" box="[802,923,556,578]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="broadleyi">P. broadleyi</taxonomicName>
|
||
<taxonomicName id="4C4D749CFF898B59FB9DFDB7FB2C2D2E" box="[1123,1235,557,578]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="capensis">P. capensis</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FDCCFB822D07" blockId="15.[159,1262,556,1448]" box="[159,1149,597,619]" pageId="15" pageNumber="188">N 87 45 13</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FDE4FB112DF8" blockId="15.[159,1262,556,1448]" box="[159,1262,638,660]" pageId="15" pageNumber="188">SVL 71.75 ± 6.92 70.18 ± 7.21 68.92 ± 11.00</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FE1EFD32FB5D2DD2" blockId="15.[159,1262,556,1448]" box="[480,1186,680,702]" pageId="15" pageNumber="188">36-86 40-80 37–82</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FD4BFB352D8B" blockId="15.[159,1262,556,1448]" box="[159,1226,721,743]" pageId="15" pageNumber="188">Gulars 25.5 ± 0.3 27.5 ± 0.4 23.8 ± 1.4</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FE1EFD60FB5C2C7C" blockId="15.[159,1262,556,1448]" box="[480,1187,762,784]" pageId="15" pageNumber="188">22–30 24–32 21–27</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FCBFFB502C0F" blockId="15.[159,1262,556,1448]" pageId="15" pageNumber="188">UL 4.95 ± 0.24 4.9 ± 0.27 5 ± 0.20 4–5.5 4–5 4.5–5.5</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FCECFB292CE0" blockId="15.[159,1262,556,1448]" box="[159,1238,886,908]" pageId="15" pageNumber="188">Col 7.93 ± 1.06 9.22 ± 1.04 7.69 ± 0.63</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FE1EFC3AFB762CDA" blockId="15.[159,1262,556,1448]" box="[480,1161,928,950]" pageId="15" pageNumber="188">
|
||
<date id="FFF329DFFF898B59FE1EFC3AFCAB2CDA" box="[480,852,928,950]" pageId="15" pageNumber="188" value="2010-05-07" valueMax="2010-05-11" valueMin="2010-05-07">5–10 7–11</date>
|
||
6–8
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FC50FB1D2CB3" blockId="15.[159,1262,556,1448]" box="[159,1250,969,991]" pageId="15" pageNumber="188">LRV 19.79 ± 1.08 20.73 ± 1.0 19.69 ± 0.63</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FE1EFC68FB5D2B64" blockId="15.[159,1262,556,1448]" box="[480,1186,1010,1032]" pageId="15" pageNumber="188">18–22 20–22 18–20</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FB87FB1D2B5E" blockId="15.[159,1262,556,1448]" box="[159,1250,1052,1074]" pageId="15" pageNumber="188">TRV 41.84 ± 2.43 43.32 ± 1.86 40.42 ± 1.08</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FE1EFBDFFB5D2B37" blockId="15.[159,1262,556,1448]" box="[480,1186,1093,1115]" pageId="15" pageNumber="188">36–48 41–47 38–42</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FBF5FB1D2BE8" blockId="15.[159,1262,556,1448]" box="[159,1250,1134,1156]" pageId="15" pageNumber="188">TD 85.23 ± 4.32 104.96 ± 5.38 83.67 ± 3.45</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FE1EFB02FB5D2BC2" blockId="15.[159,1262,556,1448]" box="[480,1186,1176,1198]" pageId="15" pageNumber="188">78–94 92–117 77–86</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FB58FB1D2BBB" blockId="15.[159,1262,556,1448]" box="[159,1250,1217,1239]" pageId="15" pageNumber="188">Forearm 17.17 ± 2.09 25.82 ± 2.31 14.62 ± 1.04</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FE1EFB70FB5D2A6C" blockId="15.[159,1262,556,1448]" box="[480,1186,1258,1280]" pageId="15" pageNumber="188">13–23 21–30 13–16</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FA8CFB1D2A46" blockId="15.[159,1262,556,1448]" box="[159,1250,1299,1324]" pageId="15" pageNumber="188">4th toe 19.10 ± 1.11 20.6 ± 1.03 17.92 ± 0.64</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FE1EFAA5FB5C2A39" blockId="15.[159,1262,556,1448]" box="[480,1187,1343,1365]" pageId="15" pageNumber="188">17–22 18–22 17–19</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FF61FAF0FB1D2A13" blockId="15.[159,1262,556,1448]" box="[159,1250,1385,1407]" pageId="15" pageNumber="188">FP 16.73 ± 1.33 16.68 ± 0.83 16.73 ± 1.33</paragraph>
|
||
<paragraph id="8BF20F1FFF898B59FE1EFA08FB1028B6" blockId="15.[159,1262,556,1448]" lastBlockId="15.[151,1437,1517,2010]" pageId="15" pageNumber="188">
|
||
14–19 15–18 14–18.5
|
||
<emphasis id="B939D30DFF898B59FF39FA77FDC2296A" bold="true" box="[199,573,1517,1542]" pageId="15" pageNumber="188">Natural history and behaviour.</emphasis>
|
||
In
|
||
<collectingCountry id="F35A4F8FFF898B59FD9BFA77FD05296A" box="[613,762,1517,1542]" name="South Africa" pageId="15" pageNumber="188">South Africa</collectingCountry>
|
||
, it is mainly restricted to the Richtersveld region (
|
||
<bibRefCitation id="EFDC72EEFF898B59FAC7FA74FECA2945" author="Bauer" pageId="15" pageNumber="188" refString="Bauer, A. M. & Branch, W. R. (2001) The herpetofauna of the Richterveld National Park and the adjacent northern Richtersveld, Northern Cape Province, Republic of South Africa. Herpetological Natural History, 8, 111 - 160." type="journal article" year="2001">Bauer & Branch, 2001</bibRefCitation>
|
||
; Whiting, 2014). There it is widespread and common in boulder-strewn areas and on broad rock faces, often far from river courses (e.g. Tierhoek). It does not form the high density populations recorded for
|
||
<taxonomicName id="4C4D749CFF898B59FA79F9ACFEFE291E" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF898B59FA79F9ACFEFE291E" italics="true" pageId="15" pageNumber="188">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
(MJW unpubl. data). All
|
||
<taxonomicName id="4C4D749CFF898B59FDDCF9C3FD57291E" box="[546,680,1625,1650]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="genus">
|
||
<emphasis id="B939D30DFF898B59FDDCF9C3FD57291E" box="[546,680,1625,1650]" italics="true" pageId="15" pageNumber="188">Platysaurus</emphasis>
|
||
</taxonomicName>
|
||
have a fixed clutch of two eggs (
|
||
<bibRefCitation id="EFDC72EEFF898B59FBE7F9C3FB36291D" author="Broadley" box="[1049,1225,1625,1650]" pageId="15" pageNumber="188" refString="Broadley, D. G. (1978) A revision of the genus Platysaurus A. Smith (Sauria: Cordylidae). Occasional Papers of the National Museum of Rhodesia Series B Natural Sciences, 6, 129 - 185." type="journal article" year="1978">Broadley, 1978</bibRefCitation>
|
||
). The reproductive cycles of
|
||
<taxonomicName id="4C4D749CFF898B59FEF3F9E4FE7129F9" box="[269,398,1662,1685]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF898B59FEF3F9E4FE7129F9" box="[269,398,1662,1685]" italics="true" pageId="15" pageNumber="188">P. capensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4D749CFF898B59FE5CF9E4FDD129FA" box="[418,558,1661,1686]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF898B59FE5CF9E4FDD129FA" box="[418,558,1661,1686]" italics="true" pageId="15" pageNumber="188">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D749CFF898B59FD91F9E4FCD129FA" box="[623,814,1661,1686]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF898B59FD91F9E4FCD129FA" box="[623,814,1661,1686]" italics="true" pageId="15" pageNumber="188">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF898B59FCC5F9E7FC6529FA" bold="true" box="[827,922,1661,1686]" pageId="15" pageNumber="188">
|
||
<taxonomicNameLabel id="A20A6E76FF898B59FCC5F9E7FC6529FA" box="[827,922,1661,1686]" pageId="15" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were collectively studied when these were considered a single species (
|
||
<bibRefCitation id="EFDC72EEFF898B59FE26F93BFCF329D6" author="Van" box="[472,780,1697,1722]" pageId="15" pageNumber="188" refString="Van Wyk, J. H. & Mouton, P. L. F. N. (1996) The reproductive cycles of the oviparous lizards Platysaurus capensis and P. minor: Evidence supporting a viviparity-oviparity reversal in the Cordylidae. Amphibia Reptilia, 17, 115 - 129. http: // dx. doi. org / 10.1163 / 156853896 X 00153" type="journal article" year="1996">Van Wyk & Mouton, 1996</bibRefCitation>
|
||
). The minimum size at sexual maturity is
|
||
<quantity id="4CB5A2FAFF898B59FB14F93BFAC529D5" box="[1258,1338,1697,1722]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="15" pageNumber="188" unit="mm" value="64.0">64 mm</quantity>
|
||
(sex not specified); eggs are likely laid in summer (November–December) (
|
||
<bibRefCitation id="EFDC72EEFF898B59FC66F95CFB2E29B2" author="Van" box="[920,1233,1733,1758]" pageId="15" pageNumber="188" refString="Van Wyk, J. H. & Mouton, P. L. F. N. (1996) The reproductive cycles of the oviparous lizards Platysaurus capensis and P. minor: Evidence supporting a viviparity-oviparity reversal in the Cordylidae. Amphibia Reptilia, 17, 115 - 129. http: // dx. doi. org / 10.1163 / 156853896 X 00153" type="journal article" year="1996">Van Wyk & Mouton, 1996</bibRefCitation>
|
||
). While we know very little about the diet of
|
||
<taxonomicName id="4C4D749CFF898B59FE2FF970FD74286E" box="[465,651,1769,1794]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF898B59FE2FF970FD74286E" box="[465,651,1769,1794]" italics="true" pageId="15" pageNumber="188">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF898B59FD6AF973FD10286E" bold="true" box="[660,751,1769,1794]" pageId="15" pageNumber="188">
|
||
<taxonomicNameLabel id="A20A6E76FF898B59FD6AF973FD10286E" box="[660,751,1769,1794]" pageId="15" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the closely related
|
||
<taxonomicName id="4C4D749CFF898B59FC28F970FBA2286E" box="[982,1117,1769,1794]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF898B59FC28F970FBA2286E" box="[982,1117,1769,1794]" italics="true" pageId="15" pageNumber="188">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
is an omnivore and lives in similar habitat. The marked sexual dichromatism suggests a classic sexual selection system in which males compete heavily for females, as is the case in
|
||
<taxonomicName id="4C4D749CFF898B59FD58F8A8FCD22826" box="[678,813,1841,1866]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF898B59FD58F8A8FCD22826" box="[678,813,1841,1866]" italics="true" pageId="15" pageNumber="188">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDC72EEFF898B59FCC3F8ABFBD82825" author="Whiting" box="[829,1063,1841,1866]" pageId="15" pageNumber="188" refString="Whiting, M. J., Nagy, K. A. & Bateman, P. W. (2003) Evolution and maintenance of social status signalling badges: experimental manipulations in lizards. In: Fox, S. F. McCoy, J. K. & Baird, T. A. (Eds.), Lizard Social Behavior. Johns Hopkins University Press, Baltimore, pp. 47 - 82." type="book chapter" year="2003">
|
||
Whiting
|
||
<emphasis id="B939D30DFF898B59FC5DF8A8FC222826" box="[931,989,1841,1866]" italics="true" pageId="15" pageNumber="188">et al.</emphasis>
|
||
, 2003
|
||
</bibRefCitation>
|
||
; Whiting
|
||
<emphasis id="B939D30DFF898B59FB67F8A8FB2C2826" box="[1177,1235,1841,1866]" italics="true" pageId="15" pageNumber="188">et al.</emphasis>
|
||
, 2006). Males do have UV-reflective throats, which suggests a role of this colour signal in either settling contests (as in
|
||
<taxonomicName id="4C4D749CFF898B59FAF3F8CCFA6D2802" box="[1293,1426,1877,1902]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF898B59FAF3F8CCFA6D2802" box="[1293,1426,1877,1902]" italics="true" pageId="15" pageNumber="188">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
) or in mate choice, although this remains to be tested. In the two males we measured, their throats had violet and blue and less pure UV than we would typically see in adult male
|
||
<taxonomicName id="4C4D749CFF898B59FC6DF804FBE328DA" box="[915,1052,1949,1974]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="188" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF898B59FC6DF804FBE328DA" box="[915,1052,1949,1974]" italics="true" pageId="15" pageNumber="188">P. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1376139AFF898B59FBD3F807FB7128DA" box="[1069,1166,1949,1974]" captionStart="FIGURE 5. a" captionStartId="16.[151,250,1911,1933]" captionTargetBox="[343,1231,203,1871]" captionTargetId="figure@16.[298,1288,151,1890]" captionTargetPageId="16" captionText="FIGURE 5. a.) Spectral reflectance curves objectively measuring colour by body region in the holotype (TM 85806) in life. b.) Spectral reflectance of throat region for two males: (MJW 935 / TM 85805, MJW 936 / TM 85806 = holotype) and two females (MJW 934 / TM 85804, MJW 937 / TM 85807). Males have more distinct UV in their throats than females although it is still less conspicuous than what has been documented for the closely related P. broadleyi (Whiting et al., 2006)." httpUri="https://zenodo.org/record/244351/files/figure.png" pageId="15" pageNumber="188">Figure 5</figureCitation>
|
||
)(Whiting
|
||
<emphasis id="B939D30DFF898B59FAF6F804FABC28DA" box="[1288,1347,1949,1974]" italics="true" pageId="15" pageNumber="188">et al.</emphasis>
|
||
, 2006). Measurement of spectral reflectance in additional individuals is necessary for a proper comparison.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF325F97FF968B46FF69F8EDFB56289D" httpUri="https://zenodo.org/record/244351/files/figure.png" pageId="16" pageNumber="189" targetBox="[343,1231,203,1871]" targetPageId="16">
|
||
<paragraph id="8BF20F1FFF968B46FF69F8EDFB56289D" blockId="16.[151,1436,1911,2033]" pageId="16" pageNumber="189">
|
||
<emphasis id="B939D30DFF968B46FF69F8EDFEED28E1" bold="true" box="[151,274,1911,1933]" pageId="16" pageNumber="189">FIGURE 5.</emphasis>
|
||
a.) Spectral reflectance curves objectively measuring colour by body region in the holotype (TM 85806) in life. b.) Spectral reflectance of throat region for two males: (MJW935/TM 85805, MJW936/TM 85806=holotype) and two females (MJW934/TM 85804, MJW937/TM 85807). Males have more distinct UV in their throats than females although it is still less conspicuous than what has been documented for the closely related
|
||
<taxonomicName id="4C4D749CFF968B46FCB8F846FB5A289D" authority="Whiting et al., 2006" authorityName="Whiting et al." authorityYear="2006" box="[838,1189,2011,2033]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="189" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF968B46FCB8F846FC40289D" box="[838,959,2011,2033]" italics="true" pageId="16" pageNumber="189">P. broadleyi</emphasis>
|
||
(Whiting
|
||
<emphasis id="B939D30DFF968B46FBD9F846FBA4289D" box="[1063,1115,2011,2033]" italics="true" pageId="16" pageNumber="189">et al.</emphasis>
|
||
, 2006)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3575C94FF978B47FF39FF0DFBD62C31" pageId="17" pageNumber="190" type="etymology">
|
||
<paragraph id="8BF20F1FFF978B47FF39FF0DFD4A2E70" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="190">
|
||
<emphasis id="B939D30DFF978B47FF39FF0DFEB12FDC" bold="true" box="[199,334,151,176]" pageId="17" pageNumber="190">Etymology.</emphasis>
|
||
We name this new species in honour of Sir David F. Attenborough (Fig. 3), in recognition of his immense contribution to the public understanding and appreciation of animals, plants, ecosystems and nature in general. David Attenborough made flat lizards, specifically the closely related
|
||
<taxonomicName id="4C4D749CFF978B47FBDEFF45FAE52F94" box="[1056,1306,223,248]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="190" phylum="Chordata" rank="species" species="broadleyi">
|
||
<emphasis id="B939D30DFF978B47FBDEFF45FAE52F94" box="[1056,1306,223,248]" italics="true" pageId="17" pageNumber="190">Platysaurus broadleyi</emphasis>
|
||
</taxonomicName>
|
||
, famous in the BBC documentary series
|
||
<emphasis id="B939D30DFF978B47FE21FE9EFD502E71" box="[479,687,260,285]" italics="true" pageId="17" pageNumber="190">Life in Cold Blood</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF978B47FF39FEBDFB9C2DA0" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="190">
|
||
<emphasis id="B939D30DFF978B47FF39FEBDFE432E2C" bold="true" box="[199,444,295,320]" pageId="17" pageNumber="190">Conservation status.</emphasis>
|
||
The conservation status of the Cape Flat Lizard (
|
||
<emphasis id="B939D30DFF978B47FC16FEB3FB9F2E2C" box="[1000,1120,297,320]" italics="true" pageId="17" pageNumber="190">P. c a pe n s i s</emphasis>
|
||
sensu lato) was assessed as of Least Concern (Whiting 2014). Our division of this taxon into two species therefore requires a re-assessment of their conservation status. Both have relatively restricted distributions, with that of
|
||
<emphasis id="B939D30DFF978B47FBAFFEEBFB332EE4" box="[1105,1228,369,392]" italics="true" pageId="17" pageNumber="190">P. c a pe n s i s</emphasis>
|
||
now restricted to only seven quarter-degree grid squares (Figure 1). Although the Kamiesberg region is not formally protected, it does form part of an envisaged expansion of the Namaqua National Park (San Parks 2013). The species has also been recorded from Carolusberg, Springbok, within or in close proximity to the Goegap Nature Reserve.
|
||
<taxonomicName id="4C4D749CFF978B47FF69FE65FE3B2D74" box="[151,452,511,536]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="190" phylum="Chordata" rank="species" species="attenboroughi">
|
||
<emphasis id="B939D30DFF978B47FF69FE65FE3B2D74" box="[151,452,511,536]" italics="true" pageId="17" pageNumber="190">Platysaurus attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
is recorded from numerous localities within the Richtersveld National Park, and the Fish River Canyon forms part of the Ai-Ais Hot Springs Game Park in
|
||
<collectingCountry id="F35A4F8FFF978B47FC28FDBEFBC52D51" box="[982,1082,548,573]" name="Namibia" pageId="17" pageNumber="190">Namibia</collectingCountry>
|
||
. Together these two conserved areas form the /Ai/Ais-Richtersveld Transfrontier Park, and thus protect a large proportion of the species’ range. The extensive granite outcrops of both the Kamiesberg region and the /Ai/Ais-Richtersveld Transfrontier Park form the main habitat for both species, and are subject to little existing habitat threat. We conclude that currently both
|
||
<taxonomicName id="4C4D749CFF978B47FF31FD2FFEB82DA0" box="[207,327,693,716]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="190" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF978B47FF31FD2FFEB82DA0" box="[207,327,693,716]" italics="true" pageId="17" pageNumber="190">P capensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D749CFF978B47FE80FD2FFDC72DA1" box="[382,568,692,717]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="190" phylum="Chordata" rank="species" species="attenboroughi">
|
||
<emphasis id="B939D30DFF978B47FE80FD2FFDC72DA1" box="[382,568,692,717]" italics="true" pageId="17" pageNumber="190">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
are not of conservation concern (Least Concern).
|
||
</paragraph>
|
||
<paragraph id="8BF20F1FFF978B47FF39FD42FBD62C31" blockId="17.[151,1437,151,861]" pageId="17" pageNumber="190">
|
||
In summary, we describe a new species of flat lizard (
|
||
<taxonomicName id="4C4D749CFF978B47FCD5FD4DFBAA2D9C" box="[811,1109,727,752]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="190" phylum="Chordata" rank="species" species="attenboroughi">
|
||
<emphasis id="B939D30DFF978B47FCD5FD4DFBAA2D9C" box="[811,1109,727,752]" italics="true" pageId="17" pageNumber="190">Platysaurus attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
) that was formerly confused with
|
||
<taxonomicName id="4C4D749CFF978B47FF2FFD67FEB12C78" box="[209,334,765,788]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="190" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B939D30DFF978B47FF2FFD67FEB12C78" box="[209,334,765,788]" italics="true" pageId="17" pageNumber="190">P. capensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C4D749CFF978B47FEA3FD66FD762C79" box="[349,649,764,789]" class="Reptilia" family="Cordylidae" genus="Platysaurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="190" phylum="Chordata" rank="species" species="attenboroughi" status="sp. nov.">
|
||
<emphasis id="B939D30DFF978B47FEA3FD66FD762C79" box="[349,649,764,789]" italics="true" pageId="17" pageNumber="190">Platysaurus attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939D30DFF978B47FD6FFD66FD132C79" bold="true" box="[657,748,764,789]" pageId="17" pageNumber="190">
|
||
<taxonomicNameLabel id="A20A6E76FF978B47FD6FFD66FD132C79" box="[657,748,764,789]" pageId="17" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
occurs on rock from the region of the Orange River west of Goodhouse, including the Richtersveld of the Northern Cape Province of
|
||
<collectingCountry id="F35A4F8FFF978B47FC27FC85FB952C54" box="[985,1130,799,824]" name="South Africa" pageId="17" pageNumber="190">South Africa</collectingCountry>
|
||
and into
|
||
<collectingCountry id="F35A4F8FFF978B47FB28FC85FAC82C54" box="[1238,1335,799,824]" name="Namibia" pageId="17" pageNumber="190">Namibia</collectingCountry>
|
||
as far as the Fish River Canyon. This poorly known species warrants detailed future study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |