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<document id="21B0402AA54A0FD63677B971BFA7CF84" ID="10.11646/zootaxa.4181.1.1" ID-CLB-Dataset="37923" ID-DOI="10.11646/zootaxa.4181.1.1" ID-GBIF-Dataset="7f55813e-d9fb-47d1-ae70-4207ea726e64" ID-ISSN="1175-5326" ID-Zenodo-Dep="164243" ID-ZooBank="639B9F0E-4E0C-4859-9A32-093511BEEFB8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478071931831" checkinUser="plazi" docAuthor="Stonis, Jonas R., Diškus, Arūnas, Remeikis, Andrius, Gerulaitis, Virginijus &amp; Karsholt, Ole" docDate="2016" docId="03E487C7FF9CD24AFF462785F092FA89" docLanguage="en" docName="zootaxa.4181.1.1.pdf" docOrigin="Zootaxa 4181 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Stigmella robusta Remeikis &amp; Stonis, sp. nov." docType="treatment" docUuid="E6DE77D9-E6FE-4F82-9D6A-C81A03C932A9" docUuidSource="ZooBank" docVersion="15" lastPageNumber="41" masterDocId="FFDDFFBFFFB4D262FFD12712F06DFFFE" masterDocTitle="Leaf-mining Nepticulidae (Lepidoptera) from record high altitudes: documenting an entire new fauna in the Andean páramo and puna" masterLastPageNumber="94" masterPageNumber="1" pageNumber="41" updateTime="1698430786409" updateUser="ExternalLinkService">
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<mods:title id="3470020B4246309D397CFC89A2444161">Leaf-mining Nepticulidae (Lepidoptera) from record high altitudes: documenting an entire new fauna in the Andean páramo and puna</mods:title>
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<mods:namePart id="C38421F97A5CFAF2E0AA38C695AACCD1">Stonis, Jonas R.</mods:namePart>
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<mods:namePart id="405B417DC83C587B9F414D6C675E2299">Diškus, Arūnas</mods:namePart>
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<mods:namePart id="386A3B77D3BDDB43A0EB8C8A17E43E36">Remeikis, Andrius</mods:namePart>
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<mods:namePart id="A559BD42123BA087FEC3C2F305847E75">Gerulaitis, Virginijus</mods:namePart>
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<mods:namePart id="A43B6663315F44B67033610CF9B9979A">Karsholt, Ole</mods:namePart>
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<mods:date id="A809379B999DDE444109D7BC8AB68BAE">2016</mods:date>
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<treatment id="03E487C7FF9CD24AFF462785F092FA89" ID-DOI="http://doi.org/10.5281/zenodo.3509052" ID-GBIF-Taxon="124560979" ID-Zenodo-Dep="3509052" LSID="urn:lsid:zoobank.org:act:E6DE77D9-E6FE-4F82-9D6A-C81A03C932A9" httpUri="http://treatment.plazi.org/id/03E487C7FF9CD24AFF462785F092FA89" lastPageNumber="41" pageId="40" pageNumber="41">
<subSubSection id="C357655AFF9CD24AFF462785F1FAFF2D" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8BF236D1FF9CD24AFF462785F2AEFF4C" blockId="40.[151,707,151,211]" box="[151,707,151,178]" pageId="40" pageNumber="41">
<heading id="D0BA81BDFF9CD24AFF462785F2AEFF4C" bold="true" box="[151,707,151,178]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<emphasis id="B939EAC3FF9CD24AFF462785F2AEFF4C" bold="true" box="[151,707,151,178]" pageId="40" pageNumber="41">
<taxonomicName id="4C4D4D52FF9CD24AFF462785F234FF4C" authority="Remeikis &amp; Stonis" authorityName="Remeikis &amp; Stonis" box="[151,601,151,178]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="B939EAC3FF9CD24AFF462785F107FF4F" bold="true" box="[151,362,151,177]" italics="true" pageId="40" pageNumber="41">Stigmella robusta</emphasis>
Remeikis &amp; Stonis
</taxonomicName>
,
<taxonomicNameLabel id="A20A57B8FF9CD24AFDB4278AF2AEFF4C" box="[613,707,152,178]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF236D1FF9CD24AFF4627A8F1FAFF2D" blockId="40.[151,707,151,211]" box="[151,407,186,211]" pageId="40" pageNumber="41">
(
<figureCitation id="13762A54FF9CD24AFF4E27A8F098FF2D" box="[159,245,186,211]" captionStart="FIGURES 18 19" captionStartId="11.[151,264,1935,1958]" captionTargetBox="[151,1436,193,1912]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 18 19. Distribution of high altitude Nepticulidae and collecting in the Peruvian Andes. 18, distribution map; 19, collector and co-author Ole Karsholt in Polylepis forest during Polylepis Expedition organized by Prof. Dr. Jon Fjeldså (the leader of the Polylepis Expeditions, ZMUC) in 1987." httpUri="https://zenodo.org/record/164248/files/figure.png" pageId="40" pageNumber="41">Figs 18</figureCitation>
,
<figureCitation id="13762A54FF9CD24AFED027A9F14DFF2D" box="[257,288,187,211]" captionStart="FIGURE 27" captionStartId="16.[151,250,2000,2022]" captionTargetBox="[163,1429,193,1982]" captionTargetId="figure@16.[163,1430,193,1983]" captionTargetPageId="16" captionText="FIGURE 27. Distribution map of the high-Andean Nepticulidae with height records." httpUri="https://zenodo.org/record/266413/files/figure.png" pageId="40" pageNumber="41">27</figureCitation>
,
<figureCitation id="13762A54FF9CD24AFEFA27A9F1E2FF2D" box="[299,399,187,211]" captionStart="FIGURES 103 107" captionStartId="41.[151,264,1872,1895]" captionTargetBox="[162,1418,265,1844]" captionTargetId="figure@41.[162,1429,265,1845]" captionTargetPageId="41" captionText="FIGURES 103 107. Stigmella robusta Remeikis &amp; Stonis, sp. nov. 103, male adult, holotype; 104, male genitalia, holotype, genitalia slide no. RA 566, phallus; 105, same, capsule without phallus; 106, same, dorsal view of capsule; 107, female genitalia, paratype, genitalia slide no. RA 580 (ZMUC)." httpUri="https://zenodo.org/record/164268/files/figure.png" pageId="40" pageNumber="41">103107</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C357655AFF9CD24AFF462610F538FE9D" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8BF236D1FF9CD24AFF462610F538FE9D" blockId="40.[151,1437,258,1399]" pageId="40" pageNumber="41">
<materialsCitation id="3B253C8CFF9CD24AFF462610F5F4FEC1" ID-GBIF-Occurrence="1324487220" collectingDate="1987-02-15" collectingDateMax="1987-02-18" collectingDateMin="1987-02-15" collectionCode="ZMUC" collectorName="O. Karsholt" country="Peru" elevation="4100" latitude="-9.680556" location="Quabrada Pucavado" longLatPrecision="21" longitude="-77.225555" municipality="Cerro Cahuish" pageId="40" pageNumber="41" specimenCount="2" stateProvince="Ancash" typeStatus="holotype">
<emphasis id="B939EAC3FF9CD24AFF462610F127FEE5" bold="true" box="[151,330,258,283]" pageId="40" pageNumber="41">
<typeStatus id="54F68873FF9CD24AFF462610F0BFFEE5" box="[151,210,258,283]" pageId="40" pageNumber="41">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54F68873FF9CD24AFE862610F1A9FEE5" box="[343,452,258,283]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
: Ƌ,
<collectingCountry id="F35A7641FF9CD24AFE2D2611F22BFEE4" box="[508,582,259,282]" name="Peru" pageId="40" pageNumber="41">PERU</collectingCountry>
, Dept.
<collectingRegion id="4989F833FF9CD24AFD702610F295FEE5" box="[673,760,258,283]" country="Peru" name="Ancash" pageId="40" pageNumber="41">Ancash</collectingRegion>
,
<locationDeviation id="9A402FF3FF9CD24AFCD92610F38EFEE4" box="[776,995,258,283]" pageId="40" pageNumber="41">
<quantity id="4CB59B34FF9CD24AFCD92610F338FEE5" box="[776,853,258,283]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="40" pageNumber="41" unit="km" value="35.0">35 km</quantity>
SE Huaraz
</locationDeviation>
,
<collectingMunicipality id="6B96ACABFF9CD24AFC222611F4F0FEE5" box="[1011,1181,258,283]" pageId="40" pageNumber="41">Cerro Cahuish</collectingMunicipality>
,
<location id="8E92600AFF9CD24AFB7E2610F5F5FEE5" LSID="urn:lsid:plazi:treatment:03E487C7FF9CD24AFF462785F092FA89:8E92600AFF9CD24AFB7E2610F5F5FEE5" box="[1199,1432,258,283]" country="Peru" latitude="-9.680556" longLatPrecision="21" longitude="-77.225555" municipality="Cerro Cahuish" name="Quabrada Pucavado" pageId="40" pageNumber="41" stateProvince="Ancash">Quabrada Pucavado</location>
,
<geoCoordinate id="EE795016FF9CD24AFF462634F167FEC1" box="[151,266,294,319]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="15" value="-9.680556">9°40'50&quot;S</geoCoordinate>
,
<geoCoordinate id="EE795016FF9CD24AFEC92635F1CFFEC1" box="[280,418,295,319]" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="15" value="-77.225555">77°13'32&quot;W</geoCoordinate>
, elevation ca.
<quantity id="4CB59B34FF9CD24AFD992635F2CCFEC0" box="[584,673,295,319]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="40" pageNumber="41" unit="m" value="4100.0">
<elevation id="0060D1E2FF9CD24AFD992635F2CCFEC0" box="[584,673,295,319]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="40" pageNumber="41" unit="m" value="4100.0">4100 m</elevation>
</quantity>
,
<date id="FFF31011FF9CD24AFD7E2634F320FEC1" box="[687,845,294,319]" pageId="40" pageNumber="41" value="1987-02-15" valueMax="1987-02-18" valueMin="1987-02-15">
<collectingDate id="EFB7E9F9FF9CD24AFD7E2634F320FEC1" box="[687,845,294,319]" pageId="40" pageNumber="41" value="1987-02-15" valueMax="1987-02-18" valueMin="1987-02-15">1518.ii.1987</collectingDate>
</date>
,
<collectorName id="26B85307FF9CD24AFC8A2635F38FFEC1" box="[859,994,294,319]" pageId="40" pageNumber="41">O. Karsholt</collectorName>
, genitalia slide no. RA566Ƌ (
<collectionCode id="ED5CAE14FF9CD24AFAE82635F5FDFEC1" box="[1337,1424,295,319]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="40" pageNumber="41">ZMUC</collectionCode>
)
</materialsCitation>
. Paratypes: 3 Ƌ, 3 ♀, same label data as holotype, genitalia slide nos RA575Ƌ, RA576Ƌ, RA580♀ (ZMUC).
</paragraph>
</subSubSection>
<subSubSection id="C357655AFF9CD24AFF16267CF3B0FE31" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8BF236D1FF9CD24AFF16267CF3B0FE31" blockId="40.[151,1437,258,1399]" pageId="40" pageNumber="41">
<emphasis id="B939EAC3FF9CD24AFF16267CF12CFE79" bold="true" box="[199,321,366,391]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
The combination of a gnathos with closely juxtaposed processes, angular inner lobe of valva, large, distally four-lobed uncus, set of 23 large cornuti and cluster of small spine-like cornuti, and pale, speckled forewing distinguishes
<taxonomicName id="4C4D4D52FF9CD24AFE4D26A5F261FE30" box="[412,524,438,462]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="B939EAC3FF9CD24AFE4D26A5F261FE30" box="[412,524,438,462]" italics="true" pageId="40" pageNumber="41">S. robusta</emphasis>
</taxonomicName>
<emphasis id="B939EAC3FF9CD24AFDC526A4F200FE31" bold="true" box="[532,621,438,463]" pageId="40" pageNumber="41">
<taxonomicNameLabel id="A20A57B8FF9CD24AFDC526A4F200FE31" box="[532,621,438,463]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from all other
<taxonomicName id="4C4D4D52FF9CD24AFCC426A4F313FE30" box="[789,894,438,462]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B939EAC3FF9CD24AFCC426A4F313FE30" box="[789,894,438,462]" italics="true" pageId="40" pageNumber="41">Stigmella</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C357655AFF9CD24AFF1626C8F23CFB3D" pageId="40" pageNumber="41" type="description">
<paragraph id="8BF236D1FF9CD24AFF1626C8F22BFCEC" blockId="40.[151,1437,258,1399]" pageId="40" pageNumber="41">
<emphasis id="B939EAC3FF9CD24AFF1626C8F169FE0D" bold="true" box="[199,260,474,499]" pageId="40" pageNumber="41">Male</emphasis>
(
<figureCitation id="13762A54FF9CD24AFEC426C8F114FE0D" box="[277,377,474,499]" captionStart="FIGURES 103 107" captionStartId="41.[151,264,1872,1895]" captionTargetBox="[162,1418,265,1844]" captionTargetId="figure@41.[162,1429,265,1845]" captionTargetPageId="41" captionText="FIGURES 103 107. Stigmella robusta Remeikis &amp; Stonis, sp. nov. 103, male adult, holotype; 104, male genitalia, holotype, genitalia slide no. RA 566, phallus; 105, same, capsule without phallus; 106, same, dorsal view of capsule; 107, female genitalia, paratype, genitalia slide no. RA 580 (ZMUC)." httpUri="https://zenodo.org/record/164268/files/figure.png" pageId="40" pageNumber="41">Fig. 103</figureCitation>
). Forewing length
<quantity id="4CB59B34FF9CD24AFD8326C8F2B3FE0C" box="[594,734,474,499]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="40" pageNumber="41" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.54.5 mm</quantity>
; wingspan
<quantity id="4CB59B34FF9CD24AFCB126C8F381FE0C" box="[864,1004,474,499]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" metricValueMax="9.6" metricValueMin="7.6" pageId="40" pageNumber="41" unit="mm" value="8.6" valueMax="9.6" valueMin="7.6">7.69.6 mm</quantity>
. Head: palpi cream; frontal tuft ochre cream; collar and scape yellowish cream; antenna as half the length of forewing; flagellum with 42 segments, yellowish grey to grey. Thorax and tegula yellowish cream (concolorous with collar and scape). Forewing glossy yellowish cream speckled with ochre-brown to dark ochre-brown scales (sometime with little purple iridescence); fascia absent; fringe grey cream to yellowish grey; underside of forewing ochre-brown to dark brown, with cream basal spots but no androconia. Hindwing pale grey to grey on upper side and underside, with no spots or androconia; its fringe yellowish grey. Legs yellowish cream, darkened with dark metalic grey on upper side. Abdomen fuscous grey on upper side, grey cream to pale brownish on underside; tufts short, pale grey to grey; genital segments cream to grey cream.
</paragraph>
<paragraph id="8BF236D1FF9CD24AFF16240CF18CFCC9" blockId="40.[151,1437,258,1399]" box="[199,481,798,823]" pageId="40" pageNumber="41">
<emphasis id="B939EAC3FF9CD24AFF16240CF173FCC9" bold="true" box="[199,286,798,823]" pageId="40" pageNumber="41">Female</emphasis>
. Similar to male.
</paragraph>
<paragraph id="8BF236D1FF9CD24AFF162450F3DFFBF1" blockId="40.[151,1437,258,1399]" pageId="40" pageNumber="41">
<emphasis id="B939EAC3FF9CD24AFF162450F11FFCA5" bold="true" box="[199,370,834,859]" pageId="40" pageNumber="41">Male genitalia</emphasis>
(
<figureCitation id="13762A54FF9CD24AFE502450F270FCA5" box="[385,541,834,859]" captionStart="FIGURES 103 107" captionStartId="41.[151,264,1872,1895]" captionTargetBox="[162,1418,265,1844]" captionTargetId="figure@41.[162,1429,265,1845]" captionTargetPageId="41" captionText="FIGURES 103 107. Stigmella robusta Remeikis &amp; Stonis, sp. nov. 103, male adult, holotype; 104, male genitalia, holotype, genitalia slide no. RA 566, phallus; 105, same, capsule without phallus; 106, same, dorsal view of capsule; 107, female genitalia, paratype, genitalia slide no. RA 580 (ZMUC)." httpUri="https://zenodo.org/record/164268/files/figure.png" pageId="40" pageNumber="41">Figs 104106</figureCitation>
). Capsule longer (325 µm) than wide (175 µm). Vinculum with long triangular lateral lobes and very long ventral plate. Uncus wide, with four small lobes distally. Gnathos with two slender, closely juxtaposed caudal processes. Valva 175180 µm long, 5460 µm wide, with two distinctive apical processes and prominent (conspicuous) angular inner lobe; transtilla with triangular corners (instead sublateral processes). Juxta membranous, indistinctive. Phallus (
<figureCitation id="13762A54FF9CD24AFD2724C0F338FC15" box="[758,853,978,1003]" captionStart="FIGURES 103 107" captionStartId="41.[151,264,1872,1895]" captionTargetBox="[162,1418,265,1844]" captionTargetId="figure@41.[162,1429,265,1845]" captionTargetPageId="41" captionText="FIGURES 103 107. Stigmella robusta Remeikis &amp; Stonis, sp. nov. 103, male adult, holotype; 104, male genitalia, holotype, genitalia slide no. RA 566, phallus; 105, same, capsule without phallus; 106, same, dorsal view of capsule; 107, female genitalia, paratype, genitalia slide no. RA 580 (ZMUC)." httpUri="https://zenodo.org/record/164268/files/figure.png" pageId="40" pageNumber="41">Fig. 104</figureCitation>
) 280285 µm long, 70 µm wide; vesica with two to three large cornuti and basal cluster of numerous small slender cornuti.
</paragraph>
<paragraph id="8BF236D1FF9CD24AFF162308F489FB61" blockId="40.[151,1437,258,1399]" pageId="40" pageNumber="41">
<emphasis id="B939EAC3FF9CD24AFF162308F1FDFBCD" bold="true" box="[199,400,1050,1075]" pageId="40" pageNumber="41">Female genitalia</emphasis>
(
<figureCitation id="13762A54FF9CD24AFE722308F267FBCD" box="[419,522,1050,1075]" captionStart="FIGURES 103 107" captionStartId="41.[151,264,1872,1895]" captionTargetBox="[162,1418,265,1844]" captionTargetId="figure@41.[162,1429,265,1845]" captionTargetPageId="41" captionText="FIGURES 103 107. Stigmella robusta Remeikis &amp; Stonis, sp. nov. 103, male adult, holotype; 104, male genitalia, holotype, genitalia slide no. RA 566, phallus; 105, same, capsule without phallus; 106, same, dorsal view of capsule; 107, female genitalia, paratype, genitalia slide no. RA 580 (ZMUC)." httpUri="https://zenodo.org/record/164268/files/figure.png" pageId="40" pageNumber="41">Fig. 107</figureCitation>
). Total length
<date id="FFF31011FF9CD24AFD672308F35FFBCD" box="[694,818,1050,1075]" pageId="40" pageNumber="41" value="1140" valueMax="1155">11401155</date>
µm. Anterior processes slender; posterior apophyses slightly longer, very slender. Vestibulum narrow, without sclerites. Corpus bursae with heavily folded distal part and round, 500505 µm long, 500 µm wide basal part with comb-like pectinations; signa absent. Accessory sac wide but short; ductus spermathecae with 4.5 convolutions. Abdominal tip narrowed and rounded.
</paragraph>
<paragraph id="8BF236D1FF9CD24AFF1623B8F23CFB3D" blockId="40.[151,1437,258,1399]" box="[199,593,1194,1219]" pageId="40" pageNumber="41">
<emphasis id="B939EAC3FF9CD24AFF1623B8F127FB3D" bold="true" box="[199,330,1194,1219]" pageId="40" pageNumber="41">Bionomics.</emphasis>
Adults fly in February.
</paragraph>
</subSubSection>
<subSubSection id="C357655AFF9CD24AFF1623DCF1F3FAF4" pageId="40" pageNumber="41" type="distribution">
<paragraph id="8BF236D1FF9CD24AFF1623DCF1F3FAF4" blockId="40.[151,1437,258,1399]" pageId="40" pageNumber="41">
<emphasis id="B939EAC3FF9CD24AFF1623DCF134FB19" bold="true" box="[199,345,1230,1255]" pageId="40" pageNumber="41">Distribution</emphasis>
(
<figureCitation id="13762A54FF9CD24AFEBB23DCF1AFFB19" box="[362,450,1230,1255]" captionStart="FIGURES 18 19" captionStartId="11.[151,264,1935,1958]" captionTargetBox="[151,1436,193,1912]" captionTargetId="figure@11.[151,1436,193,1914]" captionTargetPageId="11" captionText="FIGURES 18 19. Distribution of high altitude Nepticulidae and collecting in the Peruvian Andes. 18, distribution map; 19, collector and co-author Ole Karsholt in Polylepis forest during Polylepis Expedition organized by Prof. Dr. Jon Fjeldså (the leader of the Polylepis Expeditions, ZMUC) in 1987." httpUri="https://zenodo.org/record/164248/files/figure.png" pageId="40" pageNumber="41">Figs 18</figureCitation>
,
<figureCitation id="13762A54FF9CD24AFE0123DDF19DFB19" box="[464,496,1231,1255]" captionStart="FIGURE 27" captionStartId="16.[151,250,2000,2022]" captionTargetBox="[163,1429,193,1982]" captionTargetId="figure@16.[163,1430,193,1983]" captionTargetPageId="16" captionText="FIGURE 27. Distribution map of the high-Andean Nepticulidae with height records." httpUri="https://zenodo.org/record/266413/files/figure.png" pageId="40" pageNumber="41">27</figureCitation>
). This species occurs high in the Peruvian Andes (
<collectingCountry id="F35A7641FF9CD24AFBE423DDF403FB19" box="[1077,1134,1231,1255]" name="Peru" pageId="40" pageNumber="41">Peru</collectingCountry>
:
<collectingRegion id="4989F833FF9CD24AFBAB23DCF4A3FB19" box="[1146,1230,1230,1255]" country="Peru" name="Ancash" pageId="40" pageNumber="41">Ancash</collectingRegion>
Departamento) at altitudes about
<quantity id="4CB59B34FF9CD24AFE9323E1F1F7FAF4" box="[322,410,1267,1291]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="40" pageNumber="41" unit="m" value="4100.0">4100 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C357655AFF9CD24AFF162204F092FA89" pageId="40" pageNumber="41" type="etymology">
<paragraph id="8BF236D1FF9CD24AFF162204F092FA89" blockId="40.[151,1437,258,1399]" pageId="40" pageNumber="41">
<emphasis id="B939EAC3FF9CD24AFF162204F123FAD1" bold="true" box="[199,334,1302,1327]" pageId="40" pageNumber="41">Etymology.</emphasis>
The species name is derived from Latin
<emphasis id="B939EAC3FF9CD24AFC952204F3CEFAD0" box="[836,931,1302,1326]" italics="true" pageId="40" pageNumber="41">robustus</emphasis>
(stout, strong) in reference to the heavily hardened (sclerotized) uncus, large angular inner lobe of valva and large, strongly sclerotized vinculum in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>